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<mods:title id="CDD63A94E2008B93AE0E69806A0C7726">Notes on Indian wolf spiders: III. Genera Acantholycosa Dahl, 1908, Evippomma Roewer, 1959, Hippasosa Roewer, 1960 and Trochosa C. L. Koch, 1847 (Araneae: Lycosidae)</mods:title>
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<mods:namePart id="9AD6479A2507E67C08BE3758F3065FE5">Sankaran, Pradeep M.</mods:namePart>
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<mods:affiliation id="640338E718A630FD53F58FFF44787702">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India pradeepmspala @ rediffmail. com; https: // orcid. org / 0000 - 0003 - 0884 - 5625</mods:affiliation>
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<mods:namePart id="46A7729CBD56EA10329C6650F5E74FC3">Caleb, John T. D.</mods:namePart>
<mods:affiliation id="DEF1E5496E967F3688AB513F4B6AB3F2">Department of Anatomy, Saveetha Medical College &amp; Hospital, Saveetha Institute of Medical and Technical Sciences, Saveetha University, Chennai, Tamil Nadu 602 105, India</mods:affiliation>
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<heading id="D0CC81A4EC54854DFF4FF99AFD2BF9CE" bold="true" box="[151,690,1626,1653]" fontSize="11" level="1" pageId="4" pageNumber="537" reason="1">
<emphasis id="B94FEADAEC54854DFF4FF99AFD2BF9CE" bold="true" box="[151,690,1626,1653]" pageId="4" pageNumber="537">
<taxonomicName id="4C3B4D4BEC54854DFF4FF99AFDBCF9CF" authority="(Gajbe, 2004)" authorityName="Sankaran &amp; Caleb" authorityYear="2023" baseAuthorityName="Gajbe" baseAuthorityYear="2004" box="[151,549,1626,1653]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="537" phylum="Arthropoda" rank="species" species="kalpiensis" status="comb. nov.">
<emphasis id="B94FEADAEC54854DFF4FF99AFEEDF9CE" bold="true" box="[151,372,1626,1653]" italics="true" pageId="4" pageNumber="537">Arctosa kalpiensis</emphasis>
(
<bibRefCitation id="EFAA4B39EC54854DFE5BF99BFD85F9CF" author="Gajbe, U. A." box="[387,540,1626,1652]" pageId="4" pageNumber="537" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe, 2004</bibRefCitation>
)
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<taxonomicNameLabel id="A27C57A1EC54854DFDF4F99AFD2BF9CE" box="[556,690,1627,1653]" pageId="4" pageNumber="537" rank="species">comb. nov.</taxonomicNameLabel>
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<paragraph id="8B8436C8EC54854DFF4FF9BEFF40F922" blockId="4.[151,690,1626,1689]" box="[151,217,1663,1689]" pageId="4" pageNumber="537">
<figureCitation id="13002A4DEC54854DFF4FF9BEFF40F922" box="[151,217,1663,1689]" captionStart="FIGURE 3" captionStartId="5.[151,250,1589,1613]" captionTargetBox="[151,1436,181,1566]" captionTargetId="figure-17@5.[151,1436,181,1566]" captionTargetPageId="5" captionText="FIGURE 3. Arctosa kalpiensis (Gajbe, 2004) comb. nov., holotype of Ocyale kalpiensis Gajbe, 2004. A subadult female habitus, dorsal. B pre-epigyne of the same, ventral. C pre-vulva of the same, dorsal. D labels from type tube. Scale bars: A, 1 mm; BC, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10145899" httpUri="https://zenodo.org/record/10145899/files/figure.png" pageId="4" pageNumber="537">Fig. 3</figureCitation>
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<paragraph id="8B8436C8EC54854DFF4FF902FD60F960" blockId="4.[151,832,1731,1786]" box="[151,761,1731,1755]" pageId="4" pageNumber="537">
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<treatmentCitation id="0A9A10D9EC54854DFF4FF902FE6FF960" author="Gajbe, U. A." box="[151,502,1731,1755]" page="29" pageId="4" pageNumber="537" year="2004">
<taxonomicName id="4C3B4D4BEC54854DFF4FF902FE6FF960" ID-CoL="48L77" authority="Gajbe, 2004: 29" authorityName="Gajbe" authorityPageNumber="29" authorityYear="2004" box="[151,502,1731,1755]" class="Arachnida" family="Lycosidae" genus="Ocyale" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="537" phylum="Arthropoda" rank="species" species="kalpiensis">
<emphasis id="B94FEADAEC54854DFF4FF902FED3F960" box="[151,330,1731,1755]" italics="true" pageId="4" pageNumber="537">Ocyale kalpiensis</emphasis>
<bibRefCitation id="EFAA4B39EC54854DFE89F902FE6FF960" author="Gajbe, U. A." box="[337,502,1731,1755]" pageId="4" pageNumber="537" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe, 2004: 29</bibRefCitation>
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, figs 3739 (subadult
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).
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</paragraph>
<paragraph id="8B8436C8EC54854DFF4FF923FCD9F941" blockId="4.[151,832,1731,1786]" box="[151,832,1762,1786]" pageId="4" pageNumber="537">
<treatmentCitationGroup id="AB2B11E6EC54854DFF4FF923FCD9F941" box="[151,832,1762,1786]" pageId="4" pageNumber="537">
<taxonomicName id="4C3B4D4BEC54854DFF4FF923FEF6F941" baseAuthorityName="Gajbe" baseAuthorityYear="2004" box="[151,367,1762,1786]" class="Arachnida" family="Lycosidae" genus="Hippasosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="537" phylum="Arthropoda" rank="species" species="kalpiensis">
<emphasis id="B94FEADAEC54854DFF4FF923FEF6F941" box="[151,367,1762,1786]" italics="true" pageId="4" pageNumber="537">Hippasosa kalpiensis</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9A10D9EC54854DFEA5F923FDC9F941" author="Sherwood, D." box="[381,592,1762,1786]" page="583" pageId="4" pageNumber="537" year="2022">
<bibRefCitation id="EFAA4B39EC54854DFEA5F923FDC9F941" author="Sherwood, D." box="[381,592,1762,1786]" pageId="4" pageNumber="537" pagination="582 - 584" refId="ref10459" refString="Sherwood, D. (2022) On the taxonomic and nomenclatural status of Ocyale Audouin, 1826 (Araneae: Pisauridae) and Hippasosa Roewer, 1960 (Araneae: Lycosidae), with notes on some other taxa. Arachnology, 19 (2), 582 - 584. https: // doi. org / 10.13156 / arac. 2022.19.2.582" type="journal article" year="2022">Sherwood 2022: 583</bibRefCitation>
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(transfer from
<taxonomicName id="4C3B4D4BEC54854DFD32F923FCABF941" ID-CoL="66R7" authorityName="Audouin" authorityYear="1826" box="[746,818,1762,1786]" class="Arachnida" family="Lycosidae" genus="Ocyale" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADAEC54854DFD32F923FCABF941" box="[746,818,1762,1786]" italics="true" pageId="4" pageNumber="537">Ocyale</emphasis>
</taxonomicName>
).
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<emphasis id="B94FEADAEC54854DFF4FF8E9FED3F8F9" bold="true" box="[151,330,1832,1858]" pageId="4" pageNumber="537">Type material.</emphasis>
<materialsCitation id="3B533C95EC54854DFE8EF8E9FE95F831" collectingDate="1982-06-15" collectorName="U. A Gajbe &amp; Kolkata" country="India" county="Mandla" elevation="580" latitude="22.9" location="Kalpi" longLatPrecision="1258" longitude="80.2" municipality="Narayanganj" pageId="4" pageNumber="537" specimenCount="1" specimenCount-female="1" stateProvince="Madhya Pradesh" typeStatus="holotype">
<typeStatus id="5480886AEC54854DFE8EF8E9FE27F8F9" box="[342,446,1832,1858]" pageId="4" pageNumber="537" type="holotype">Holotype</typeStatus>
subadult
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from
<emphasis id="B94FEADAEC54854DFD41F8E9FD72F8F9" bold="true" box="[665,747,1832,1858]" pageId="4" pageNumber="537">
<collectingCountry id="F32C7658EC54854DFD41F8E9FD72F8F9" box="[665,747,1832,1858]" name="India" pageId="4" pageNumber="537">INDIA</collectingCountry>
</emphasis>
:
<emphasis id="B94FEADAEC54854DFCD8F8E9FC5DF8F9" box="[768,964,1832,1858]" italics="true" pageId="4" pageNumber="537">
<collectingRegion id="49FFF82AEC54854DFCD8F8E9FC5DF8F9" box="[768,964,1832,1858]" country="India" name="Madhya Pradesh" pageId="4" pageNumber="537">Madhya Pradesh</collectingRegion>
</emphasis>
:
<collectingCounty id="62E54E44EC54854DFC01F8E9FBAAF8F9" box="[985,1075,1832,1858]" pageId="4" pageNumber="537">Mandla</collectingCounty>
:
<collectingMunicipality id="6BE0ACB2EC54854DFB9AF8E9FB4FF8F9" box="[1090,1238,1832,1858]" pageId="4" pageNumber="537">Narayanganj</collectingMunicipality>
:
<location id="8EE46013EC54854DFB3DF8E9FABAF8F9" LSID="urn:lsid:plazi:treatment:039287DEEC54854FFF4FF99AFC94FEDE:8EE46013EC54854DFB3DF8E9FABAF8F9" box="[1253,1315,1832,1858]" country="India" county="Mandla" latitude="22.9" longLatPrecision="1258" longitude="80.2" municipality="Narayanganj" name="Kalpi" pageId="4" pageNumber="537" stateProvince="Madhya Pradesh">Kalpi</location>
(
<geoCoordinate id="EE0F500FEC54854DFAE0F8E9FA01F8F8" box="[1336,1432,1832,1859]" degrees="22" direction="north" minutes="54" orientation="latitude" pageId="4" pageNumber="537" precision="925" value="22.9">22°54'N</geoCoordinate>
,
<geoCoordinate id="EE0F500FEC54854DFF4FF88DFF6DF8DD" box="[151,244,1868,1894]" degrees="80" direction="east" minutes="12" orientation="longitude" pageId="4" pageNumber="537" precision="925" value="80.2">80°12'E</geoCoordinate>
;
<quantity id="4CC39B2DEC54854DFF26F88DFEDAF8DC" box="[254,323,1868,1895]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="537" unit="m" value="580.0">
<elevation id="0016D1FBEC54854DFF26F88DFEDAF8DC" box="[254,323,1868,1895]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="537" unit="m" value="580.0">580 m</elevation>
</quantity>
alt.), leg.
<collectorName id="26CE531EEC54854DFE77F88DFDB1F8DD" box="[431,552,1868,1894]" pageId="4" pageNumber="537">U.A Gajbe</collectorName>
,
<date id="FF851008EC54854DFDEAF88DFD5EF8DD" box="[562,711,1868,1895]" pageId="4" pageNumber="537" value="1982-06-15">
<collectingDate id="EFC1E9E0EC54854DFDEAF88DFD5EF8DD" box="[562,711,1868,1895]" pageId="4" pageNumber="537" value="1982-06-15">15 June 1982</collectingDate>
</date>
, repository NZC-ZSI,
<collectorName id="26CE531EEC54854DFC66F88DFB8EF8DD" box="[958,1047,1868,1894]" pageId="4" pageNumber="537">Kolkata</collectorName>
(no registration number specified), examined.
</materialsCitation>
</paragraph>
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<subSubSection id="C3216543EC54854FFF1FF855FD38FEFA" lastPageId="6" lastPageNumber="539" pageId="4" pageNumber="537" type="description">
<paragraph id="8B8436C8EC54854DFF1FF855FC46F815" blockId="4.[151,1437,1832,2003]" box="[199,991,1940,1967]" pageId="4" pageNumber="537">
<emphasis id="B94FEADAEC54854DFF1FF855FEC0F815" bold="true" box="[199,345,1940,1966]" pageId="4" pageNumber="537">Description.</emphasis>
For description of the subadult female, see
<bibRefCitation id="EFAA4B39EC54854DFC9BF855FC42F815" author="Gajbe, U. A." box="[835,987,1940,1966]" pageId="4" pageNumber="537" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8436C8EC54854DFF1FF879FE1CF869" blockId="4.[151,1437,1832,2003]" box="[199,389,1976,2003]" pageId="4" pageNumber="537">
<emphasis id="B94FEADAEC54854DFF1FF879FE98F869" box="[199,257,1976,2002]" italics="true" pageId="4" pageNumber="537">Male</emphasis>
. Unknown.
</paragraph>
<caption id="DF446640EC55854CFF4FF9F4FEC5F92E" ID-DOI="http://doi.org/10.5281/zenodo.10145899" ID-Zenodo-Dep="10145899" httpUri="https://zenodo.org/record/10145899/files/figure.png" pageId="5" pageNumber="538" startId="5.[151,250,1589,1613]" targetBox="[151,1436,181,1566]" targetPageId="5" targetType="figure">
<paragraph id="8B8436C8EC55854CFF4FF9F4FEC5F92E" blockId="5.[151,1437,1589,1686]" pageId="5" pageNumber="538">
<emphasis id="B94FEADAEC55854CFF4FF9F4FE8FF9F5" bold="true" box="[151,278,1589,1614]" pageId="5" pageNumber="538">FIGURE 3.</emphasis>
<taxonomicName id="4C3B4D4BEC55854CFEF8F9F7FDE1F9F5" authority="(Gajbe, 2004)" authorityName="Sankaran &amp; Caleb" authorityYear="2023" baseAuthorityName="Gajbe" baseAuthorityYear="2004" box="[288,632,1590,1614]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="kalpiensis" status="comb. nov.">
<emphasis id="B94FEADAEC55854CFEF8F9F7FE44F9F5" box="[288,477,1590,1614]" italics="true" pageId="5" pageNumber="538">Arctosa kalpiensis</emphasis>
(
<bibRefCitation id="EFAA4B39EC55854CFE36F9F7FDE8F9F5" author="Gajbe, U. A." box="[494,625,1590,1614]" pageId="5" pageNumber="538" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe, 2004</bibRefCitation>
)
</taxonomicName>
<emphasis id="B94FEADAEC55854CFD5AF9F7FD60F9F5" bold="true" box="[642,761,1590,1614]" pageId="5" pageNumber="538">
<taxonomicNameLabel id="A27C57A1EC55854CFD5AF9F7FD60F9F5" box="[642,761,1590,1614]" pageId="5" pageNumber="538" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, holotype of
<taxonomicName id="4C3B4D4BEC55854CFC53F9F7FB56F9F5" authority="Gajbe, 2004" authorityName="Gajbe" authorityYear="2004" box="[907,1231,1590,1614]" class="Arachnida" family="Lycosidae" genus="Ocyale" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="kalpiensis">
<emphasis id="B94FEADAEC55854CFC53F9F7FBDBF9F5" box="[907,1090,1590,1614]" italics="true" pageId="5" pageNumber="538">Ocyale kalpiensis</emphasis>
<bibRefCitation id="EFAA4B39EC55854CFB94F9F7FB56F9F5" author="Gajbe, U. A." box="[1100,1231,1590,1614]" pageId="5" pageNumber="538" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe, 2004</bibRefCitation>
</taxonomicName>
.
<emphasis id="B94FEADAEC55854CFB03F9F4FB74F9F6" bold="true" box="[1243,1261,1589,1613]" pageId="5" pageNumber="538">A</emphasis>
subadult female habitus, dorsal.
<emphasis id="B94FEADAEC55854CFEE1F99BFED3F9C9" bold="true" box="[313,330,1626,1650]" pageId="5" pageNumber="538">B</emphasis>
pre-epigyne of the same, ventral.
<emphasis id="B94FEADAEC55854CFD71F998FD22F9CA" bold="true" box="[681,699,1625,1649]" pageId="5" pageNumber="538">C</emphasis>
pre-vulva of the same, dorsal.
<emphasis id="B94FEADAEC55854CFC23F99BFB94F9C9" bold="true" box="[1019,1037,1626,1650]" pageId="5" pageNumber="538">D</emphasis>
labels from type tube. Scale bars: A, 1 mm; BC, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8436C8EC55854FFF1FF929FF6DFF6E" blockId="5.[151,1437,1768,2010]" lastBlockId="6.[151,1437,150,357]" lastPageId="6" lastPageNumber="539" pageId="5" pageNumber="538">
<emphasis id="B94FEADAEC55854CFF1FF929FE88F8B9" bold="true" box="[199,273,1768,1794]" pageId="5" pageNumber="538">Notes.</emphasis>
This species was described based on a single female specimen collected from
<collectingRegion id="49FFF82AEC55854CFB54F929FAD0F8B9" box="[1164,1353,1768,1794]" country="India" name="Madhya Pradesh" pageId="5" pageNumber="538">Madhya Pradesh</collectingRegion>
(
<bibRefCitation id="EFAA4B39EC55854CFA8FF929FF4DF89D" author="Gajbe, U. A." pageId="5" pageNumber="538" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe 2004</bibRefCitation>
). It was recently transferred to
<taxonomicName id="4C3B4D4BEC55854CFDFFF8CCFD06F89D" authorityName="Roewer" authorityYear="1960" box="[551,671,1805,1830]" class="Arachnida" family="Lycosidae" genus="Hippasosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADAEC55854CFDFFF8CCFD06F89D" box="[551,671,1805,1830]" italics="true" pageId="5" pageNumber="538">Hippasosa</emphasis>
</taxonomicName>
as a consequence of the resurrection of this genus from its synonymy with
<taxonomicName id="4C3B4D4BEC55854CFF17F8F1FE73F8F1" authority="(Sherwood 2022)" baseAuthorityName="Sherwood" baseAuthorityYear="2022" box="[207,490,1840,1866]" class="Arachnida" family="Lycosidae" genus="Ocyale" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADAEC55854CFF17F8F1FE87F8F1" box="[207,286,1840,1866]" italics="true" pageId="5" pageNumber="538">Ocyale</emphasis>
(
<bibRefCitation id="EFAA4B39EC55854CFEF4F8F1FE7AF8F1" author="Sherwood, D." box="[300,483,1840,1866]" pageId="5" pageNumber="538" pagination="582 - 584" refId="ref10459" refString="Sherwood, D. (2022) On the taxonomic and nomenclatural status of Ocyale Audouin, 1826 (Araneae: Pisauridae) and Hippasosa Roewer, 1960 (Araneae: Lycosidae), with notes on some other taxa. Arachnology, 19 (2), 582 - 584. https: // doi. org / 10.13156 / arac. 2022.19.2.582" type="journal article" year="2022">Sherwood 2022</bibRefCitation>
)
</taxonomicName>
. Detailed examination of the type material of
<taxonomicName id="4C3B4D4BEC55854CFC3DF8F1FBEEF8F1" authorityName="Gajbe" authorityYear="2004" box="[997,1143,1840,1866]" class="Arachnida" family="Lycosidae" genus="Ocyale" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="kalpiensis">
<emphasis id="B94FEADAEC55854CFC3DF8F1FC60F8F1" box="[997,1017,1840,1866]" italics="true" pageId="5" pageNumber="538">O</emphasis>
.
<emphasis id="B94FEADAEC55854CFBDFF8F1FBEEF8F1" box="[1031,1143,1840,1866]" italics="true" pageId="5" pageNumber="538">kalpiensis</emphasis>
</taxonomicName>
revealed that the
<typeStatus id="5480886AEC55854CFAE2F8F1FA05F8F1" box="[1338,1436,1840,1866]" pageId="5" pageNumber="538" type="holotype">holotype</typeStatus>
female is a subadult (
<figureCitation id="13002A4DEC55854CFE5CF895FE44F8D5" box="[388,477,1876,1902]" captionStart="FIGURE 3" captionStartId="5.[151,250,1589,1613]" captionTargetBox="[151,1436,181,1566]" captionTargetId="figure-17@5.[151,1436,181,1566]" captionTargetPageId="5" captionText="FIGURE 3. Arctosa kalpiensis (Gajbe, 2004) comb. nov., holotype of Ocyale kalpiensis Gajbe, 2004. A subadult female habitus, dorsal. B pre-epigyne of the same, ventral. C pre-vulva of the same, dorsal. D labels from type tube. Scale bars: A, 1 mm; BC, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10145899" httpUri="https://zenodo.org/record/10145899/files/figure.png" pageId="5" pageNumber="538">Fig. 3A</figureCitation>
). The pre-epigyne of this species lacks the inverted T-shaped septum, which is present in the epigyne of
<taxonomicName id="4C3B4D4BEC55854CFEBAF8B8FDADF829" box="[354,564,1912,1938]" pageId="5" pageNumber="538">
<emphasis id="B94FEADAEC55854CFEBAF8B8FE43F829" box="[354,474,1913,1938]" italics="true" pageId="5" pageNumber="538">Hippasosa</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EFAA4B39EC55854CFD9CF8B9FCBFF829" author="Alderweireldt, M." box="[580,806,1912,1938]" pageId="5" pageNumber="538" pagination="1349 - 1365" refId="ref8357" refString="Alderweireldt, M. (1996) A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History, 30 (9), 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261" type="journal article" year="1996">Alderweireldt 1996</bibRefCitation>
: figs 411, 2024), indicating the misplacement of this species in
<taxonomicName id="4C3B4D4BEC55854CFED3F85CFE1AF80D" authorityName="Roewer" authorityYear="1960" box="[267,387,1949,1974]" class="Arachnida" family="Lycosidae" genus="Hippasosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADAEC55854CFED3F85CFE1AF80D" box="[267,387,1949,1974]" italics="true" pageId="5" pageNumber="538">Hippasosa</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="EFAA4B39EC55854CFE29F85DFD58F80D" author="Jocque, M. &amp; Wellens, S. &amp; Andrianarivosoa, J. D. &amp; Rakotondraparany, F. &amp; The Seing &amp; Jocque, R." box="[497,705,1948,1974]" pageId="5" pageNumber="538" pagination="1 - 13" refId="ref9247" refString="Jocque, M., Wellens, S., Andrianarivosoa, J. D., Rakotondraparany, F., The Seing, S. &amp; Jocque, R. (2017) A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus. European Journal of Taxonomy, 355, 1 - 13. https: // doi. org / 10.5852 / ejt. 2017.355" type="journal article" year="2017">
Jocque
<emphasis id="B94FEADAEC55854CFD9DF85CFDE4F80D" box="[581,637,1948,1974]" italics="true" pageId="5" pageNumber="538">et al.</emphasis>
2017
</bibRefCitation>
). Instead, the pre-epigyne with a conspicuous atrium divided by a median septum as seen in
<taxonomicName id="4C3B4D4BEC55854CFE64F801FDB4F861" baseAuthorityName="Fabricius" baseAuthorityYear="1777" box="[444,557,1984,2010]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B94FEADAEC55854CFE64F801FDB4F861" box="[444,557,1984,2010]" italics="true" pageId="5" pageNumber="538">A. cinerea</emphasis>
</taxonomicName>
supports an interim inclusion of this species in
<taxonomicName id="4C3B4D4BEC55854CFB9AF801FB01F861" authorityName="C. L. Koch" authorityYear="1847" box="[1090,1176,1984,2010]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADAEC55854CFB9AF801FB01F861" box="[1090,1176,1984,2010]" italics="true" pageId="5" pageNumber="538">Arctosa</emphasis>
</taxonomicName>
(compare
<figureCitation id="13002A4DEC55854CFAC8F801FAFAF861" box="[1296,1379,1984,2010]" captionStart="FIGURE 3" captionStartId="5.[151,250,1589,1613]" captionTargetBox="[151,1436,181,1566]" captionTargetId="figure-17@5.[151,1436,181,1566]" captionTargetPageId="5" captionText="FIGURE 3. Arctosa kalpiensis (Gajbe, 2004) comb. nov., holotype of Ocyale kalpiensis Gajbe, 2004. A subadult female habitus, dorsal. B pre-epigyne of the same, ventral. C pre-vulva of the same, dorsal. D labels from type tube. Scale bars: A, 1 mm; BC, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10145899" httpUri="https://zenodo.org/record/10145899/files/figure.png" pageId="5" pageNumber="538">Fig. 3B</figureCitation>
with
<bibRefCitation id="EFAA4B39EC56854FFF4FFF56FEF4FF0A" author="Buchar, J. &amp; Knoflach, B. &amp; Thaler, K." box="[151,365,151,177]" pageId="6" pageNumber="539" pagination="329 - 336" refId="ref8561" refString="Buchar, J., Knoflach, B. &amp; Thaler, K. (2006) On the identity of Arctosa varians C. L. Koch and Arctosa similis Schenkel, with notes on related species. Bulletin of the British Arachnological Society, 13 (9), 329 - 336." type="journal article" year="2006">
Buchar
<emphasis id="B94FEADAEC56854FFF28FF56FEB0FF0A" box="[240,297,151,177]" italics="true" pageId="6" pageNumber="539">et al.</emphasis>
2006
</bibRefCitation>
: fig. 25;
<bibRefCitation id="EFAA4B39EC56854FFE0AFF56FD01FF0A" author="Wang, L. Y. &amp; Marusik, Y. M. &amp; Zhang, Z. S." box="[466,664,151,177]" pageId="6" pageNumber="539" pagination="53 - 68" refId="ref11069" refString="Wang, L. Y., Marusik, Y. M. &amp; Zhang, Z. S. (2012) Notes on three poorly known Arctosa species from China (Araneae: Lycosidae). Zootaxa, 3404 (1), 53 - 68. https: // doi. org / 10.11646 / zootaxa. 3404.1.2" type="journal article" year="2012">
Wang
<emphasis id="B94FEADAEC56854FFDC3FF56FDCDFF0A" box="[539,596,151,177]" italics="true" pageId="6" pageNumber="539">et al.</emphasis>
2012
</bibRefCitation>
: fig. 12E). Based on these observations, we transfer
<taxonomicName id="4C3B4D4BEC56854FFB33FF56FAE7FF0A" authorityName="Gajbe" authorityYear="2004" box="[1259,1406,151,177]" class="Arachnida" family="Lycosidae" genus="Ocyale" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="539" phylum="Arthropoda" rank="species" species="kalpiensis">
<emphasis id="B94FEADAEC56854FFB33FF56FAE7FF0A" box="[1259,1406,151,177]" italics="true" pageId="6" pageNumber="539">H. kalpiensis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C3B4D4BEC56854FFF4FFF7AFF74FF6E" authorityName="C. L. Koch" authorityYear="1847" box="[151,237,187,213]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="539" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADAEC56854FFF4FFF7AFF74FF6E" box="[151,237,187,213]" italics="true" pageId="6" pageNumber="539">Arctosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8436C8EC56854FFF1FFF1EFD38FEFA" blockId="6.[151,1437,150,357]" pageId="6" pageNumber="539">
The NZC-ZSI collection has one glass tube for this species, labelled as holotype (no registration number specified) containing
<specimenCount id="9D3DFD41EC56854FFE53FEC2FD83FEA6" box="[395,538,259,285]" count="1" pageId="6" pageNumber="539" type="subadult">one subadult</specimenCount>
female specimen in fairly good condition. The same tube has a small glass vial containing the dissected genitalia (
<figureCitation id="13002A4DEC56854FFDC3FEE6FD0DFEFA" box="[539,660,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1589,1613]" captionTargetBox="[151,1436,181,1566]" captionTargetId="figure-17@5.[151,1436,181,1566]" captionTargetPageId="5" captionText="FIGURE 3. Arctosa kalpiensis (Gajbe, 2004) comb. nov., holotype of Ocyale kalpiensis Gajbe, 2004. A subadult female habitus, dorsal. B pre-epigyne of the same, ventral. C pre-vulva of the same, dorsal. D labels from type tube. Scale bars: A, 1 mm; BC, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10145899" httpUri="https://zenodo.org/record/10145899/files/figure.png" pageId="6" pageNumber="539">Fig. 3BC</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3216543EC56854FFF1FFE8BFC94FEDE" box="[199,781,330,357]" pageId="6" pageNumber="539" type="distribution">
<paragraph id="8B8436C8EC56854FFF1FFE8BFC94FEDE" blockId="6.[151,1437,150,357]" box="[199,781,330,357]" pageId="6" pageNumber="539">
<emphasis id="B94FEADAEC56854FFF1FFE8BFEF8FEDF" bold="true" box="[199,353,330,356]" pageId="6" pageNumber="539">Distribution.</emphasis>
<collectingCountry id="F32C7658EC56854FFEBFFE8AFE38FEDE" box="[359,417,331,357]" name="India" pageId="6" pageNumber="539">India</collectingCountry>
(
<collectingRegion id="49FFF82AEC56854FFE77FE8AFDE9FEDE" box="[431,624,331,357]" country="India" name="Madhya Pradesh" pageId="6" pageNumber="539">Madhya Pradesh</collectingRegion>
,
<bibRefCitation id="EFAA4B39EC56854FFDA3FE8AFC9BFEDE" author="Gajbe, U. A." box="[635,770,331,357]" pageId="6" pageNumber="539" pagination="1 - 40" refId="ref9047" refString="Gajbe, U. A. (2004) Studies on some spiders of the family Lycosidae (Araneae: Arachnida) from Madhya Pradesh, India. Records of the Zoological Survey of India, Occasional Paper, 221, 1 - 40. https: // doi. org / 10.26515 / rzsi / v 105 / i 1 - 2 / 2005 / 159247" type="journal article" year="2004">Gajbe 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>