306 lines
41 KiB
XML
306 lines
41 KiB
XML
<document id="5341BF5C8D6F6D60951D052D8819869A" ID-CLB-Dataset="4287" ID-DOI="10.18590/euscorpius.2014.vol2014.iss180.1" ID-GBIF-Dataset="9a8832b9-a96c-47ca-aa19-74c6b9775f3e" ID-ISSN="1536-9307" ID-Zenodo-Dep="5507889" ID-ZooBank="BE227D9C-6BC8-42AA-B75F-8A67F8E0DC12" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617134285422" checkinUser="carolina" docAuthor="Teruel, Rolando, Rivera, Mel J. & Sánchez, Alejandro J." docDate="2014" docId="039287DBFF81FFAA88C7FA85BEC1FE58" docLanguage="en" docName="Euscorpius.180.1-11.pdf" docOrigin="Euscorpius 180" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Microtityus (Parvabsonus) borincanus Teruel, Rivera et Sanchez 2014, sp. n." docType="treatment" docUuid="19B3B413-132C-4D7C-AA43-0B6392175B6D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="5" masterDocId="FFABFFA3FF82FFAC8810FC51BD4FFFA9" masterDocTitle="First record of the genus Microtityus Kjellesvig-Waering, 1966, from Puerto Rico, with description of two new species (Scorpiones: Buthidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1698935380908" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="1440AFEA216E9DEF40762982A3753AB4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="416FA9585914711E168A9A4095BEE950">
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<mods:title id="C8946DC5BB37271965FE2CC6ECC33625">First record of the genus Microtityus Kjellesvig-Waering, 1966, from Puerto Rico, with description of two new species (Scorpiones: Buthidae)</mods:title>
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<mods:name id="1F5270D5319853FD6BE913728717BCFE" type="personal">
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<mods:roleTerm id="C3777457B780421C8F53856A1B8A24A5">Author</mods:roleTerm>
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<mods:namePart id="216B05ACECD1974BAA88FEC3710FABF6">Teruel, Rolando</mods:namePart>
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<mods:roleTerm id="97D4DE62DDA555EED896261A13227890">Author</mods:roleTerm>
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<mods:namePart id="43E2211C521713E0C0DD50BD993174E2">Rivera, Mel J.</mods:namePart>
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</mods:name>
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<mods:name id="55A799B1AB0F35173FF02BCCA93519A0" type="personal">
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<mods:roleTerm id="D6B0E9569E7AEA1FF182BB1F24FAC5E4">Author</mods:roleTerm>
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<mods:namePart id="13A66F26160747E3EC808DC83AA5983D">Sánchez, Alejandro J.</mods:namePart>
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<mods:typeOfResource id="707E2A78A77C21773892BD0BDBCBA92C">text</mods:typeOfResource>
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<mods:title id="95363662BC1CCECB243C9F1015275646">Euscorpius</mods:title>
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<mods:part id="1D25E41C67C958CB99B0C02A779EF25C">
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<mods:date id="D044AFF0C1FCF12867A911003ABEEEAF">2014</mods:date>
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<mods:number id="7E35407332ABE0D31D11CD7D70B40798">2014-12-31</mods:number>
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<mods:number id="3C194E17E379F601EDB98F1ABE2F5797">180</mods:number>
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<mods:start id="B2B0AE851A36904B121C1C54407D9DA2">1</mods:start>
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<mods:end id="F64E6CC72D88FB77EF5E84CFEDB4F76A">11</mods:end>
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<mods:classification id="2E4B99499B5155886581840C0875EAE4">journal article</mods:classification>
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<mods:identifier id="2F6536D4C1CE17E8C9839CDA4AE54411" type="DOI">10.18590/euscorpius.2014.vol2014.iss180.1</mods:identifier>
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<mods:identifier id="88E23D28BE0DBC54BE904849D11A8341" type="ISSN">1536-9307</mods:identifier>
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<treatment id="039287DBFF81FFAA88C7FA85BEC1FE58" ID-DOI="http://doi.org/10.5281/zenodo.5507887" ID-GBIF-Taxon="187975132" ID-Zenodo-Dep="5507887" LSID="urn:lsid:zoobank.org:act:19B3B413-132C-4D7C-AA43-0B6392175B6D" httpUri="http://treatment.plazi.org/id/039287DBFF81FFAA88C7FA85BEC1FE58" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2">
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<subSubSection id="C3216546FF81FFAF88C7FA85BFC5F8A6" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8B8436CDFF81FFAF88C7FA85BFC5F8A6" blockId="3.[196,797,1748,1900]" pageId="3" pageNumber="2">
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<heading id="D0CC81A1FF81FFAF88C7FA85BFC5F8A6" centered="true" fontSize="11" level="2" pageId="3" pageNumber="2" reason="2">
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<emphasis id="B94FEADFFF81FFAF88C7FA85BFC5F8A6" bold="true" pageId="3" pageNumber="2">
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<taxonomicName id="4C3B4D4EFF81FFAF88C7FA85BF75F8A6" ID-CoL="438SR" authority="Teruel, Rivera et Sanchez" authorityName="Teruel, Rivera et Sanchez" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="borincanus" status="sp. nov." subGenus="Parvabsonus">
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<emphasis id="B94FEADFFF81FFAF88C7FA85BC2DF947" bold="true" box="[215,354,1748,1774]" italics="true" pageId="3" pageNumber="2">Microtityus</emphasis>
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(
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<emphasis id="B94FEADFFF81FFAF8963FA85BF40F947" bold="true" box="[371,527,1748,1774]" italics="true" pageId="3" pageNumber="2">Parvabsonus</emphasis>
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)
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<emphasis id="B94FEADFFF81FFAF8A30FA85BFE6F947" bold="true" box="[544,681,1748,1774]" italics="true" pageId="3" pageNumber="2">borincanus</emphasis>
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Teruel, Rivera et Sánchez
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27C57A4FF81FFAF8A58FAA4BFC5F8A6" box="[584,650,1781,1807]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3216546FF81FFAF8987FB47BF89F8C5" pageId="3" pageNumber="2" type="description">
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<paragraph id="8B8436CDFF81FFAF8987FB47BF04F884" blockId="3.[196,797,1748,1900]" box="[407,587,1814,1838]" pageId="3" pageNumber="2">
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<figureCitation id="13002A48FF81FFAF8987FB47BF58F887" box="[407,535,1814,1838]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[192,267,1207,1231]" captionStartId-1="4.[192,267,1259,1283]" captionStartId-2="5.[192,267,1304,1328]" captionTargetBox-0="[228,1429,179,1157]" captionTargetBox-1="[219,1439,202,1204]" captionTargetBox-2="[271,1393,224,1233]" captionTargetId-0="figure-326@3.[192,1439,168,1177]" captionTargetId-1="figure-308@4.[192,1439,192,1230]" captionTargetId-2="figure-317@5.[192,1440,192,1275]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n., entire dorsal and ventral views." captionText-1="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." captionText-2="Figure 3: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n., coloration in life." figureDoi-0="http://doi.org/10.5281/zenodo.5507891" figureDoi-1="http://doi.org/10.5281/zenodo.5507895" figureDoi-2="http://doi.org/10.5281/zenodo.5507897" httpUri-0="https://zenodo.org/record/5507891/files/figure.png" httpUri-1="https://zenodo.org/record/5507895/files/figure.png" httpUri-2="https://zenodo.org/record/5507897/files/figure.png" pageId="3" pageNumber="2">Figures 1–3</figureCitation>
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,
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<figureCitation id="13002A48FF81FFAF8A33FB47BF04F884" box="[547,587,1814,1838]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStartId-0="9.[192,267,1422,1446]" captionStartId-1="10.[192,267,1880,1904]" captionTargetBox-0="[192,1439,192,1392]" captionTargetBox-1="[198,1453,170,1849]" captionTargetId-0="figure-265@9.[192,1439,192,1392]" captionTargetId-1="figure-29@10.[199,1453,171,1848]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figure 6: Distribution of the genus Microtityus in the Greater Antilles (above), and the Puerto Rican Microtityus (Parvabsonus) borincanus, sp. n. (1) and Microtityus (Parvabsonus) santosi, sp. n. (2)." captionText-1="Figure 7: Two views of Susúa State Forest, habitat of Microtityus (Parvabsonus) borincanus." figureDoi-0="http://doi.org/10.5281/zenodo.5507903" figureDoi-1="http://doi.org/10.5281/zenodo.5507905" httpUri-0="https://zenodo.org/record/5507903/files/figure.png" httpUri-1="https://zenodo.org/record/5507905/files/figure.png" pageId="3" pageNumber="2">6–7</figureCitation>
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</paragraph>
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<paragraph id="8B8436CDFF81FFAF88D4FB64BF89F8C5" blockId="3.[196,797,1748,1900]" pageId="3" pageNumber="2">
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<emphasis id="B94FEADFFF81FFAF88D4FB64BF89F8C5" bold="true" pageId="3" pageNumber="2">http://zoobank.org/urn:lsid:zoobank.org:act:19B3B4 13-132C-4D7C-AA43-0B6392175B6D</emphasis>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3216546FF81FFAF8B2FF8A5B9DEFACE" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="8B8436CDFF81FFAF8B2FF8A5B9DEFACE" blockId="3.[831,1440,1267,1383]" pageId="3" pageNumber="2">
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<materialsCitation id="3B533C90FF81FFAF8B2FF8A5B8D2FAA2" ID-GBIF-Occurrence="3352132305" box="[831,1437,1267,1291]" collectionCode="TYPE, DATA, RTO" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<collectionCode id="ED2AAE08FF81FFAF8B2FF8A5BE8EFAA3" box="[831,961,1268,1291]" pageId="3" pageNumber="2">TYPE DATA</collectionCode>
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.
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<specimenCount id="9D3DFD44FF81FFAF8BC7F8A5BEA4FAA2" box="[983,1003,1268,1291]" pageId="3" pageNumber="2" type="female">♀</specimenCount>
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<typeStatus id="5480886FFF81FFAF8BEBF8A2B917FAA2" box="[1019,1112,1267,1291]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
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(to be deposited in
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<collectionCode id="ED2AAE08FF81FFAF8D4AF8A2B8DAFAA2" box="[1370,1429,1267,1291]" pageId="3" pageNumber="2">RTO</collectionCode>
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)
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||
</materialsCitation>
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:
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<materialsCitation id="3B533C90FF81FFAF8B2FF943B9DEFACE" ID-GBIF-Occurrence="3352132301" collectingDate="2013-11-22" collectionCode="RTO" collectorName="M. J. Rivera." country="Puerto Rico" county="Municipality" latitude="18.09278" location="Susua State Forest" longLatPrecision="21" longitude="-66.90889" municipality="Barrio Susua" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Sabana Grande" typeStatus="holotype">
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||
<collectingCountry id="F32C765DFF81FFAF8B2FF943BE8EFA83" box="[831,961,1298,1322]" name="Puerto Rico" pageId="3" pageNumber="2">Puerto Rico</collectingCountry>
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,
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||
<collectingRegion id="49FFF82FFF81FFAF8BDCF943B920FA83" box="[972,1135,1298,1322]" country="Puerto Rico" name="Sabana Grande" pageId="3" pageNumber="2">Sabana Grande</collectingRegion>
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<collectingCounty id="62E54E41FF81FFAF8C66F943B84DFA83" box="[1142,1282,1298,1322]" pageId="3" pageNumber="2">Municipality</collectingCounty>
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,
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<collectingMunicipality id="6BE0ACB7FF81FFAF8D1EF943B8D4FA83" box="[1294,1435,1298,1322]" pageId="3" pageNumber="2">Barrio Susúa</collectingMunicipality>
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,
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||
<location id="8EE46016FF81FFAF8B2FF961B95AFAE1" LSID="urn:lsid:plazi:treatment:039287DBFF81FFAA88C7FA85BEC1FE58:8EE46016FF81FFAF8B2FF961B95AFAE1" box="[831,1045,1328,1352]" country="Puerto Rico" county="Municipality" latitude="18.09278" longLatPrecision="21" longitude="-66.90889" municipality="Barrio Susua" name="Susua State Forest" pageId="3" pageNumber="2" stateProvince="Sabana Grande">Susúa State Forest</location>
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||
,
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<geoCoordinate id="EE0F500AFF81FFAF8C38F961B9E9FAE1" box="[1064,1190,1328,1352]" degrees="18" direction="north" minutes="05" orientation="latitude" pageId="3" pageNumber="2" precision="15" seconds="34" value="18.09278">18°05'34"N</geoCoordinate>
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<geoCoordinate id="EE0F500AFF81FFAF8CA5F961B875FAE1" box="[1205,1338,1328,1352]" degrees="66" direction="west" minutes="54" orientation="longitude" pageId="3" pageNumber="2" precision="15" seconds="32" value="-66.90889">66°54'32"W</geoCoordinate>
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,
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<date id="FF85100DFF81FFAF8D5DF961BE82FACE" pageId="3" pageNumber="2" value="2013-11-22">
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<collectingDate id="EFC1E9E5FF81FFAF8D5DF961BE82FACE" pageId="3" pageNumber="2" value="2013-11-22">22 November 2013</collectingDate>
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</date>
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, leg.
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<collectorName id="26CE531BFF81FFAF8C16F91EB9DEFACE" box="[1030,1169,1359,1383]" pageId="3" pageNumber="2">M. J. Rivera.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3216546FF81FFA88B2FF9DBBF14FA2F" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="diagnosis">
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<paragraph id="8B8436CDFF81FFA88B2FF9DBBF14FA2F" blockId="3.[831,1441,1417,1901]" lastBlockId="4.[192,801,1359,1414]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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DIAGNOSIS. Adult size medium (
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<quantity id="4CC39B28FF81FFAF8CB8F9D8B9B7FA08" box="[1192,1272,1417,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="2" unit="mm" value="17.0">17 mm</quantity>
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) for the genus. Coloration yellowish brown, with a very dense pattern of blackish spots all over the body and appendages, metasomal segment V and telson slightly more reddish, tergites without clearly defined longitudinal dark bands, pedipalp fingers blackish with yellowish tips. Pedipalps orthobothriotaxic (femoral trichobothrium
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||
<emphasis id="B94FEADFFF81FFAF8D0FFA10B87AF9F0" box="[1311,1333,1601,1625]" italics="true" pageId="3" pageNumber="2">
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||
d
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<subScript id="17BF3488FF81FFAF8D3CFA1AB87AF9F0" attach="left" box="[1324,1333,1611,1625]" fontSize="6" pageId="3" pageNumber="2">2</subScript>
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</emphasis>
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present), fixed and movable fingers with 10 principal rows of denticles, basal lobe/notch combination absent. Sternite V with the smooth patch small and teardrop-shaped. Metasoma moderately slender, with 10-10-8-8-5 complete carinae, most of which are finely serrate, segments II–IV dorsolateral carinae with terminal denticle enlarged, all intercarinal spaces coriaceous to finely granulose. Telson oval, vesicle coriaceous, subaculear tubercle moderate and irregular, aculeus shorter than vesicle. Pectines with 10/10 teeth, basal plate conspicuously longer than wide and spatulate.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF446645FF86FFA888D0F8BAB90DFAB5" ID-DOI="http://doi.org/10.5281/zenodo.5507895" ID-Zenodo-Dep="5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,267,1259,1283]" targetBox="[219,1439,202,1204]" targetPageId="4">
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||
<paragraph id="8B8436CDFF86FFA888D0F8BAB90DFAB5" blockId="4.[192,1440,1259,1308]" pageId="4" pageNumber="3">
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||
<emphasis id="B94FEADFFF86FFA888D0F8BABC65FAAA" bold="true" box="[192,298,1259,1283]" pageId="4" pageNumber="3">Figure 2:</emphasis>
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Female holotype of
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<taxonomicName id="4C3B4D4EFF86FFA889E6F8BFBE22FAAA" authority="Teruel & Rivera & Sánchez, 2014" authorityName="Teruel & Rivera & Sánchez" authorityYear="2014" box="[502,877,1261,1283]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="borincanus" status="sp. nov." subGenus="Parvabsonus">
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||
<emphasis id="B94FEADFFF86FFA889E6F8BFBF2AFAAA" box="[502,613,1262,1283]" italics="true" pageId="4" pageNumber="3">Microtityus</emphasis>
|
||
(
|
||
<emphasis id="B94FEADFFF86FFA88A64F8BCBFBEFAAA" box="[628,753,1261,1283]" italics="true" pageId="4" pageNumber="3">Parvabsonus</emphasis>
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||
)
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||
<emphasis id="B94FEADFFF86FFA88B11F8BCBE22FAAA" box="[769,877,1261,1283]" italics="true" pageId="4" pageNumber="3">borincanus</emphasis>
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</taxonomicName>
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||
,
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||
<emphasis id="B94FEADFFF86FFA88B6BF8BCBEFDFAAB" bold="true" box="[891,946,1261,1282]" pageId="4" pageNumber="3">
|
||
<taxonomicNameLabel id="A27C57A4FF86FFA88B6BF8BCBEFDFAAB" box="[891,946,1261,1282]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
:
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<emphasis id="B94FEADFFF86FFA88BD0F8BCBE9CFAAB" bold="true" box="[960,979,1261,1282]" pageId="4" pageNumber="3">a)</emphasis>
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pedipalps;
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<emphasis id="B94FEADFFF86FFA88C55F8BCB915FAAB" bold="true" box="[1093,1114,1261,1282]" pageId="4" pageNumber="3">b)</emphasis>
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carapace and anterior tergites;
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<emphasis id="B94FEADFFF86FFA88D9EF8BCB8EFFAAB" bold="true" box="[1422,1440,1261,1282]" pageId="4" pageNumber="3">c)</emphasis>
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sternopectinal region and anterior sternites;
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<emphasis id="B94FEADFFF86FFA88A76F957BF34FAB2" bold="true" box="[614,635,1286,1307]" pageId="4" pageNumber="3">d)</emphasis>
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metasomal segment V and telson, lateral view.
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</paragraph>
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</caption>
|
||
<subSubSection id="C3216546FF86FFA888D0F9E2BF01F92E" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8B8436CDFF86FFA888D0F9E2BF01F92E" blockId="4.[192,802,1459,1671]" pageId="4" pageNumber="3">
|
||
ETYMOLOGY. The selected name is a Latinized adjective taken from one of the two Spanish words used to name the people native to the island of
|
||
<collectingCountry id="F32C765DFF86FFA88A56F9A3BF82F9A0" box="[582,717,1521,1545]" name="Puerto Rico" pageId="4" pageNumber="3">Puerto Rico</collectingCountry>
|
||
, whose ancient Taino name was "Borikén", and which modern variant is "Borinquen". It highlights the occurrence of this species here, a long awaited discovery among scorpiologists but so far unaccomplished.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3216546FF86FFA888D0FAE6BF46F8E2" pageId="4" pageNumber="3" type="distribution">
|
||
<paragraph id="8B8436CDFF86FFA888D0FAE6BF46F8E2" blockId="4.[192,802,1718,1867]" pageId="4" pageNumber="3">
|
||
DISTRIBUTION (
|
||
<figureCitation id="13002A48FF86FFA88962FAE7BCF1F967" box="[370,446,1718,1742]" captionStart="Figure 6" captionStartId="9.[192,267,1422,1446]" captionTargetBox="[192,1439,192,1392]" captionTargetId="figure-265@9.[192,1439,192,1392]" captionTargetPageId="9" captionText="Figure 6: Distribution of the genus Microtityus in the Greater Antilles (above), and the Puerto Rican Microtityus (Parvabsonus) borincanus, sp. n. (1) and Microtityus (Parvabsonus) santosi, sp. n. (2)." figureDoi="http://doi.org/10.5281/zenodo.5507903" httpUri="https://zenodo.org/record/5507903/files/figure.png" pageId="4" pageNumber="3">Fig. 6</figureCitation>
|
||
). Known only from the
|
||
<typeStatus id="5480886FFF86FFA88AE3FAE6BE6EF966" box="[755,801,1719,1743]" pageId="4" pageNumber="3">type</typeStatus>
|
||
locality, in southwestern
|
||
<collectingCountry id="F32C765DFF86FFA889C7FA87BF12F944" box="[471,605,1749,1773]" name="Puerto Rico" pageId="4" pageNumber="3">Puerto Rico</collectingCountry>
|
||
. It represents one of the few serpentine outcrops of the island, characterized by high levels of plant endemism (Cedeño- Maldonado &
|
||
<bibRefCitation id="EFAA4B3CFF86FFA8894BFB62BCB5F8E2" author="BRECKON" box="[347,506,1843,1867]" pageId="4" pageNumber="3" pagination="348 - 356" refId="ref4768" refString="CEDENO-MALDONADO, J. A. & G. J. BRECKON. 1996. Serpentine endemism in the flora of Puerto Rico. Caribbean Journal of Science, 32 (4): 348 - 356." type="journal article" year="1996">Breckon, 1996</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3216546FF86FFAA8B2FF91EBEC1FE58" lastPageId="6" lastPageNumber="5" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="8B8436CDFF86FFAA8B2FF91EBC5FFAE4" blockId="4.[831,1440,1358,1864]" lastBlockId="6.[192,802,198,1357]" lastPageId="6" lastPageNumber="5" pageId="4" pageNumber="3">
|
||
DESCRIPTION (female
|
||
<typeStatus id="5480886FFF86FFA88C3FF91FB9DEFACF" box="[1071,1169,1358,1382]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
).
|
||
<emphasis id="B94FEADFFF86FFA88CB5F91EB86FFACE" bold="true" box="[1189,1312,1359,1383]" pageId="4" pageNumber="3">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF86FFA88D20F91FB834FACF" box="[1328,1403,1358,1382]" captionStart="Figure 1" captionStartId="3.[192,267,1207,1231]" captionTargetBox="[228,1429,179,1157]" captionTargetId="figure-326@3.[192,1439,168,1177]" captionTargetPageId="3" captionText="Figure 1: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n., entire dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.5507891" httpUri="https://zenodo.org/record/5507891/files/figure.png" pageId="4" pageNumber="3">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13002A48FF86FFA88D9AF91FB8D8FACF" box="[1418,1431,1358,1382]" captionStart="Figure 3" captionStartId="5.[192,267,1304,1328]" captionTargetBox="[271,1393,224,1233]" captionTargetId="figure-317@5.[192,1440,192,1275]" captionTargetPageId="5" captionText="Figure 3: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n., coloration in life." figureDoi="http://doi.org/10.5281/zenodo.5507897" httpUri="https://zenodo.org/record/5507897/files/figure.png" pageId="4" pageNumber="3">3</figureCitation>
|
||
) base yellowish brown, very densely spotted with blackish brown all over the body and appendages, including the ventral region. Chelicerae pale yellowish, with distal margin of manus and basal half of fingers deeply infuscate. Pedipalp femur very densely spotted with blackish brown on all surfaces except ventral, which it is essentially immaculate; patella very densely spotted with blackish brown on all surfaces except ventral, which possesses spots only in the distal portion; chela with hand essentially immaculate, only a few vestigial spots are present on external surface, fingers blackish with yellowish tips. Carapace symmetrically and densely spotted with blackish brown, but with frontal lobes almost immaculate; tergites densely spotted with blackish brown, but without clearly defined pattern of longitudinal bands; venter very densely spotted with blackish brown all over leg coxae, sternum, genital operculum, pectines and sternites, but with a thin yellowish strip along midline of sternites III–VI. Legs with trochanter, femur and patella very densely spotted with blackish brown externally, sparsely spotted both dorsally and ventrally, but immaculate internally, tarsi each conspicuously annulated: basal half blackish brown, distal half pale yellowish. Metasoma densely spotted with blackish brown on all surfaces, spotted pattern becoming somewhat denser and darker distally in each segment; segments IV–V and telson slightly more reddish than I– III; telson vesicle densely spotted with blackish brown, fainter on distal half; aculeus immaculate, with the distal half blackish brown.
|
||
<emphasis id="B94FEADFFF87FFA989BAFAA3BF57F8A3" bold="true" box="[426,536,1778,1802]" pageId="5" pageNumber="4">Carapace</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF87FFA98A39FAA3BF36F8A3" box="[553,633,1778,1802]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="5" pageNumber="4">Fig. 2b</figureCitation>
|
||
) slightly wider than long, subtriangular. Anterior margin deeply bilobed. Carinae: anterior median, lateral ocular, central median and posterior median granulose, superciliary subgranulose, other carinae obsolete to absent. Tegument coriaceous to finely granulose, with coarser granules scattered. Median eyes separated by more than one ocular diameter; three pairs of lateral eyes, which are all much smaller than median eyes.
|
||
<emphasis id="B94FEADFFF87FFA98D14F98FB82EFA5F" bold="true" box="[1284,1377,1502,1526]" pageId="5" pageNumber="4">Tergites</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF87FFA98D65F98CBE16F9BD" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="5" pageNumber="4">Fig. 2a</figureCitation>
|
||
) with the same granular sculpture as on carapace; I– VI with three longitudinal carinae, which are granulose and sharply projected beyond the posterior margin of every tergite, VII with five serrate longitudinal carinae.
|
||
<emphasis id="B94FEADFFF87FFA98B2FFA26BEF7F926" bold="true" box="[831,952,1655,1679]" pageId="5" pageNumber="4">Chelicerae</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF87FFA98BD7FA26B95CF926" box="[967,1043,1655,1679]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="5" pageNumber="4">Fig. 2a</figureCitation>
|
||
) with dentition typical for the genus, tegument smooth and shiny.
|
||
<emphasis id="B94FEADFFF87FFA98CA0FAC7B850F907" bold="true" box="[1200,1311,1686,1710]" pageId="5" pageNumber="4">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF87FFA98D2DFAC7B8D6F907" box="[1341,1433,1686,1710]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="5" pageNumber="4">Fig. 2a</figureCitation>
|
||
) orthobothriotaxic A-α (femur with
|
||
<emphasis id="B94FEADFFF87FFA98CD0FAE5B999F965" box="[1216,1238,1716,1740]" italics="true" pageId="5" pageNumber="4">
|
||
d
|
||
<subScript id="17BF3488FF87FFA98CDDFAEFB999F965" attach="left" box="[1229,1238,1726,1740]" fontSize="6" pageId="5" pageNumber="4">2</subScript>
|
||
</emphasis>
|
||
). Femur with five serrate carinae, intercarinal tegument coriaceous to finely granulose, with coarser granules scattered, internal surface with the four
|
||
<emphasis id="B94FEADFFF87FFA98C71FB40B927F881" box="[1121,1128,1809,1832]" italics="true" pageId="5" pageNumber="4">i</emphasis>
|
||
trichobothria surrounding a spur of moderate size (smaller in left pedipalp). Patella with seven serrate carinae, intercarinal tegument with the same granular sculpture as on femur, internal surface with 2–3 slightly larger granules. Chela oval and narrower than patella; hand with nine finely serrate carinae, intercarinal tegument coriaceous to finely granulose; fingers without basal lobe/notch combination, fixed finger with 10/10 principal rows of denticles, movable finger with 10/10 plus apical subrow composed by four granules aligned similar to principal rows (missing in right finger).
|
||
<emphasis id="B94FEADFFF84FFAA89C2FDEDBF48FE7D" bold="true" box="[466,519,444,468]" pageId="6" pageNumber="5">Legs</emphasis>
|
||
with all carinae serrate to granulose; intercarinal tegument coriaceous to finely granulose, with coarser granules scattered.
|
||
<emphasis id="B94FEADFFF84FFAA8AAEFDA8BE6EFDB8" bold="true" box="[702,801,505,529]" pageId="6" pageNumber="5">Sternum</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF84FFAA88D8FE46BC59FD86" box="[200,278,535,559]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="6" pageNumber="5">Fig. 2c</figureCitation>
|
||
) type 1, subtriangular.
|
||
<emphasis id="B94FEADFFF84FFAA8A09FE46BF39FD86" bold="true" box="[537,630,535,559]" pageId="6" pageNumber="5">Pectines</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF84FFAA8A97FE46BF9AFD86" box="[647,725,535,559]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="6" pageNumber="5">Fig. 2c</figureCitation>
|
||
) about half the length of leg IV coxa. Tooth count 10/10, fulcra very well developed. Basal middle lamella slightly enlarged, oval. Basal plate longer than wide, not spatulate, with posterior margin narrowly convex.
|
||
<emphasis id="B94FEADFFF84FFAA8AADFEC3BE6DFD03" bold="true" box="[701,802,658,682]" pageId="6" pageNumber="5">Sternites</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF84FFAA88D8FEE1BC58FD61" box="[200,279,688,712]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="6" pageNumber="5">Fig. 2c</figureCitation>
|
||
) with short-oval spiracles. Sternite III with the lateral areas shallowly depressed and essentially smooth; IV–VI coriaceous to finely granulose, with coarser granules scattered; VII finely and densely granulose. Posterior margin of V essentially straight, with the smooth patch small, teardrop-shaped, slightly longer than wide, translucent, and not bulky nor protruding from posterior margin of sternite; VI–VII with two pairs of granulose lateral carinae.
|
||
<emphasis id="B94FEADFFF84FFAA8A1CFFF7BFCDFC17" bold="true" box="[524,642,934,958]" pageId="6" pageNumber="5">Metasoma</emphasis>
|
||
(
|
||
<figureCitation id="13002A48FF84FFAA8A88FFF7BFA6FC17" box="[664,745,934,958]" captionStart="Figure 1" captionStartId="3.[192,267,1207,1231]" captionTargetBox="[228,1429,179,1157]" captionTargetId="figure-326@3.[192,1439,168,1177]" captionTargetPageId="3" captionText="Figure 1: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n., entire dorsal and ventral views." figureDoi="http://doi.org/10.5281/zenodo.5507891" httpUri="https://zenodo.org/record/5507891/files/figure.png" pageId="6" pageNumber="5">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13002A48FF84FFAA8AEEFFF7BE55FC17" box="[766,794,934,958]" captionStart="Figure 2" captionStartId="4.[192,267,1259,1283]" captionTargetBox="[219,1439,202,1204]" captionTargetId="figure-308@4.[192,1439,192,1230]" captionTargetPageId="4" captionText="Figure 2: Female holotype of Microtityus (Parvabsonus) borincanus, sp. n.: a) pedipalps; b) carapace and anterior tergites; c) sternopectinal region and anterior sternites; d) metasomal segment V and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5507895" httpUri="https://zenodo.org/record/5507895/files/figure.png" pageId="6" pageNumber="5">2d</figureCitation>
|
||
) with intercarinal tegument coriaceous to very finely and densely granulose, with coarser granules scattered; segments I–II with ten complete carinae, II–IV with eight, V with five, all sharply serrate, dorsolateral carinae on I– IV with terminal denticle conspicuously enlarged and very sharp. Telson vesicle oval, smooth and shiny, with a weakly subgranulose ventromedian carina that abruptly rises into the subaculear tubercle, which points towards the tip of the aculeus or slightly beyond, and is moderately-sized, irregularly conical, essentially smooth, and possesses two vestigial dorsal granules; aculeus shorter than vesicle, very sharp, and evenly curved.
|
||
</paragraph>
|
||
<caption id="DF446645FF87FFA988D0F949B917FA86" ID-DOI="http://doi.org/10.5281/zenodo.5507897" ID-Zenodo-Dep="5507897" box="[192,1112,1304,1328]" httpUri="https://zenodo.org/record/5507897/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,267,1304,1328]" targetBox="[271,1393,224,1233]" targetPageId="5">
|
||
<paragraph id="8B8436CDFF87FFA988D0F949B917FA86" blockId="5.[192,1112,1304,1328]" box="[192,1112,1304,1328]" pageId="5" pageNumber="4">
|
||
<emphasis id="B94FEADFFF87FFA988D0F949BC67FA99" bold="true" box="[192,296,1304,1328]" pageId="5" pageNumber="4">Figure 3:</emphasis>
|
||
Female holotype of
|
||
<taxonomicName id="4C3B4D4EFF87FFA989FFF94BBE2CFA86" authority="Teruel & Rivera & Sánchez, 2014" authorityName="Teruel & Rivera & Sánchez" authorityYear="2014" box="[495,867,1305,1327]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="borincanus" status="sp. nov." subGenus="Parvabsonus">
|
||
<emphasis id="B94FEADFFF87FFA989FFF94BBF11FA86" box="[495,606,1306,1327]" italics="true" pageId="5" pageNumber="4">Microtityus</emphasis>
|
||
(
|
||
<emphasis id="B94FEADFFF87FFA98A7CF948BFA6FA86" box="[620,745,1305,1327]" italics="true" pageId="5" pageNumber="4">Parvabsonus</emphasis>
|
||
)
|
||
<emphasis id="B94FEADFFF87FFA98AE7F948BE2CFA86" box="[759,867,1305,1327]" italics="true" pageId="5" pageNumber="4">borincanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B94FEADFFF87FFA98B60F94BBEE9FA86" bold="true" box="[880,934,1306,1327]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A27C57A4FF87FFA98B60F94BBEE9FA86" box="[880,934,1306,1327]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, coloration in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8436CDFF84FFAA88D0F929BFBCF8F5" blockId="6.[192,802,1400,1884]" pageId="6" pageNumber="5">
|
||
COMPARISONS (adult female only). This species closely resembles
|
||
<taxonomicName id="4C3B4D4EFF84FFAA892DF9C6BF9CFA07" authority="Armas, 1999" authorityName="Armas" authorityYear="1999" box="[317,723,1430,1454]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="virginiae">
|
||
<emphasis id="B94FEADFFF84FFAA892DF9C6BF64FA07" box="[317,555,1431,1454]" italics="true" pageId="6" pageNumber="5">Microtityus virginiae</emphasis>
|
||
Armas, 1999
|
||
</taxonomicName>
|
||
(from Sierra de Neiba in southern Hispaniola) as both share a unique combination of characters that unequivocally distinguish them both from the remaining members of the subgenus
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8947FA40BCADF980" authorityName="Armas" authorityYear="1974" box="[343,482,1553,1577]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="subGenus" subGenus="Parvabsonus">
|
||
<emphasis id="B94FEADFFF84FFAA8947FA40BCADF980" box="[343,482,1553,1577]" italics="true" pageId="6" pageNumber="5">Parvabsonus</emphasis>
|
||
</taxonomicName>
|
||
: medium size, overall coloration very dark, metasoma slender, subaculear tubercle ventrally not serrate, and low pectinal tooth counts. But the latter can be easily distinguished from
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8A96FA3FBE6EF92C" authority="Teruel & Rivera & Sánchez, 2014" authorityName="Teruel & Rivera & Sánchez" authorityYear="2014" box="[646,801,1645,1669]" class="Arachnida" family="Buthidae" genus="Microtityus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="borincanus" status="sp. nov.">
|
||
<emphasis id="B94FEADFFF84FFAA8A96FA3FBE6EF92C" box="[646,801,1645,1669]" italics="true" pageId="6" pageNumber="5">M. borincanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94FEADFFF84FFAA88D0FADDBC4CF90D" bold="true" box="[192,259,1676,1700]" pageId="6" pageNumber="5">
|
||
<taxonomicNameLabel id="A27C57A4FF84FFAA88D0FADDBC4CF90D" box="[192,259,1676,1700]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by:
|
||
<emphasis id="B94FEADFFF84FFAA8950FADDBC18F90D" bold="true" box="[320,343,1676,1700]" pageId="6" pageNumber="5">1)</emphasis>
|
||
pedipalp chela with hand very densely spotted and fingers not uniformly blackish, but also irregularly spotted;
|
||
<emphasis id="B94FEADFFF84FFAA89A0FA98BC88F948" bold="true" box="[432,455,1737,1761]" pageId="6" pageNumber="5">2)</emphasis>
|
||
pedipalp fingers with nine principal rows of denticles;
|
||
<emphasis id="B94FEADFFF84FFAA8A11FAB9BF58F8A9" bold="true" box="[513,535,1768,1792]" pageId="6" pageNumber="5">3)</emphasis>
|
||
dorsolateral carinae on metasomal segments I–IV with terminal denticle only vestigially enlarged;
|
||
<emphasis id="B94FEADFFF84FFAA89ADFB77BC9CF897" bold="true" box="[445,467,1830,1854]" pageId="6" pageNumber="5">4)</emphasis>
|
||
telson conspicuously more slender, with subaculear tubercle longer and sharper.
|
||
</paragraph>
|
||
<paragraph id="8B8436CDFF84FFAA8B2FFC97BEC1FE58" blockId="6.[831,1441,198,497]" pageId="6" pageNumber="5">
|
||
ECOLOGICAL NOTES. The
|
||
<typeStatus id="5480886FFF84FFAA8C79FC97B989FF77" box="[1129,1222,198,222]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
and single known specimen was found inside a rotten log lying on the ground. Other scorpions that occur at Susúa State Forest are the scorpionid
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8C0CFD70BE38FEF1" authority="Francke, 1978" authorityName="Francke" authorityYear="1978" class="Arachnida" family="Diplocentridae" genus="Heteronebo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="portoricensis">
|
||
<emphasis id="B94FEADFFF84FFAA8C0CFD70B87AFE90" box="[1052,1333,289,313]" italics="true" pageId="6" pageNumber="5">Heteronebo portoricensis</emphasis>
|
||
Francke, 1978
|
||
</taxonomicName>
|
||
, and the buthids
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8C49FD11BE30FEDE" authority="(Koch, 1845)" baseAuthorityName="Koch" baseAuthorityYear="1845" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="griseus">
|
||
<emphasis id="B94FEADFFF84FFAA8C49FD11B80AFEF1" box="[1113,1349,320,344]" italics="true" pageId="6" pageNumber="5">Centruroides griseus</emphasis>
|
||
(Koch, 1845)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8B9BFD31B86AFEDE" authority="Santiago-Blay, 2009" authorityName="Santiago-Blay" authorityYear="2009" box="[907,1317,351,375]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="angelesae">
|
||
<emphasis id="B94FEADFFF84FFAA8B9BFD31B970FEDE" box="[907,1087,351,375]" italics="true" pageId="6" pageNumber="5">Tityus angelesae</emphasis>
|
||
Santiago-Blay, 2009
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8D4FFD31B974FE3F" authority="(Karsch, 1879)" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="obtusus">
|
||
<emphasis id="B94FEADFFF84FFAA8D4FFD31BEDEFE3F" italics="true" pageId="6" pageNumber="5">Tityus obtusus</emphasis>
|
||
(Karsch, 1879)
|
||
</taxonomicName>
|
||
; of them,
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8CB8FD2EB819FE3F" authorityName="Francke" authorityYear="1978" box="[1192,1366,383,406]" class="Arachnida" family="Diplocentridae" genus="Heteronebo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="portoricensis">
|
||
<emphasis id="B94FEADFFF84FFAA8CB8FD2EB819FE3F" box="[1192,1366,383,406]" italics="true" pageId="6" pageNumber="5">H. portoricensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D4EFF84FFAA8D9BFD2EBEE5FE1D" authorityName="Santiago-Blay" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="angelesae">
|
||
<emphasis id="B94FEADFFF84FFAA8D9BFD2EBEE5FE1D" italics="true" pageId="6" pageNumber="5">T. angelesae</emphasis>
|
||
</taxonomicName>
|
||
are soil-dwelling species, while the other two are more habitat-generalist and also occur on the vegetation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |