treatments-xml/data/03/92/87/039287B73135FFB8FF5DB1A5D195FB43.xml

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<mods:title id="F453AA77DBEF0062A5B0D6996FCCB0C7">Description of four new species of Burrowing Frogs in the Fejervarya rufescens complex (Dicroglossidae) with notes on morphological affinities of Fejervarya species in the Western Ghats</mods:title>
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<mods:namePart id="B5DCA2E48A769735DC51F1ABAE6659AB">Garg, Sonali</mods:namePart>
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<taxonomicName id="4C3B4D223135FF85FF5DB1A5D4B4FF1E" ID-CoL="6HR34" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[151,386,151,177]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FF5DB1A5D4B4FF1E" bold="true" box="[151,386,151,177]" italics="true" pageId="31" pageNumber="482">Fejervarya neilcoxi</emphasis>
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<paragraph id="8B8436A13135FF85FF5DB188D4FDFF58" blockId="31.[151,1158,151,283]" pageId="31" pageNumber="482">http://zoobank.org/urn:lsid:zoobank.org:act:E3B863F6-7443-495C-A698-8543FF6C4197 Neil Cox’s Burrowing Frog</paragraph>
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<subSubSection id="C321652A3135FF85FF5DB030D4F3FEB4" box="[151,453,258,283]" pageId="31" pageNumber="482" type="description">
<paragraph id="8B8436A13135FF85FF5DB030D4F3FEB4" blockId="31.[151,1158,151,283]" box="[151,453,258,283]" pageId="31" pageNumber="482">
(
<tableCitation id="C6B9031A3135FF85FF54B030D42BFEB4" box="[158,285,258,283]" captionStart-0="TABLE 1" captionStart-1="TABLE 4" captionStart-2="TABLE 5" captionStart-3="TABLE 6" captionStart-4="TABLE 7" captionStartId-0="11.[125,217,515,539]" captionStartId-1="14.[151,239,940,963]" captionStartId-2="15.[151,239,581,604]" captionStartId-3="15.[151,239,1081,1104]" captionStartId-4="18.[119,204,562,584]" captionText-0="TABLE 1. List of DNA sequences used in the study. Taxa are arranged in the order of their phylogenetic position in Figure 1 (Top to bottom)." captionText-1="TABLE 4. Factor loadings and percent variance explained by Principal Component (PC) 1 – 5 for 17 size-corrected morphometric variables from males of five species in the Fejervarya rufescens group. Values in bold indicate variables with the highest loadings." captionText-2="TABLE 5. Scores for discriminant function analysis of the principal components (PC 1 – 5). Values in bold indicate variables with the highest loadings for discriminant function roots of eigenvalue&gt; 1.0." captionText-3="TABLE 6. Classification matrix for adult male specimens of the Fejervarya rufescens group from the discriminant function analysis." captionText-4="TABLE 7. Morphometric measurements for five species in the Fejervarya rufescens group. All measurements are in millimeters (mm). The status of each specimen is given the museum number: Holotype (HT), Paratype (PT), Neotype (NT), Referred specimen (RS)." pageId="31" pageNumber="482">Tables 1–7</tableCitation>
; 
<figureCitation id="13002A243135FF85FEE3B030D4A5FEB4" box="[297,403,258,283]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[151,250,1885,1907]" captionStartId-1="5.[151,250,1618,1640]" captionStartId-2="6.[151,250,1765,1787]" captionStartId-3="8.[151,250,1820,1842]" captionStartId-4="9.[151,250,1518,1540]" captionStartId-5="10.[151,250,1726,1748]" captionTargetBox-0="[161,1435,204,1854]" captionTargetBox-1="[155,1424,193,1582]" captionTargetBox-2="[151,1436,197,1739]" captionTargetBox-3="[151,1436,180,1813]" captionTargetBox-4="[151,1436,193,1495]" captionTargetBox-5="[158,1434,882,1697]" captionTargetId-0="figure@4.[151,1436,193,1864]" captionTargetId-1="figure@5.[151,1436,193,1597]" captionTargetId-2="figure@6.[151,1436,193,1744]" captionTargetId-3="figure@8.[151,1436,180,1818]" captionTargetId-4="figure@9.[151,1436,193,1497]" captionTargetId-5="figure@10.[151,1437,879,1704]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of&gt; 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." captionText-1="FIGURE 2. Phylogenetic analyses. A. Maximum Likelihood phylogram based on mitochondrial (16 S, COI and Cytb) and nuclear (Rag 1) DNA of 2314 basepairs, showing phylogenetic relationships among members of the Fejervarya rufescens group. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of&gt; 50 % are indicated above and below the branches, respectively. Fejervarya granosa (2) was used as the outgroup taxon. Voucher number and locality details of the samples are referenced in Table 1; B. Median Joining haplotype network inferred from 554 basepair fragment of the nuclear Rag 1 gene. Circle sizes are proportional to the number of haplotypes, branch lengths are proportional to the number of mutational steps, and black bars represent mutational steps. Number of individuals used for each species: Fejervarya cepfi (3), Fejervarya kadar (1), Fejervarya manoharani (3), Fejervarya neilcoxi (2), and Fejervarya rufescens (5)." captionText-2="FIGURE 3. Geographical distribution of five species in Fejervarya rufescens group of the Western Ghats, Peninsular India. The Western Ghats biodiversity hotspot region is shown in orange colour. Locality details are referenced in Table 3." captionText-3="FIGURE 4. Four morphologically recognized Fejervarya groups in the Western Ghats. A – B. Fejervarya nilagirica group: A. Fejervarya nilagirica in life (male, SDBDU 2014.2772); B. Fejervarya keralensis in life (male, SDBDU 2015.3139); C. Fejervarya rufescens group, Fejervarya rufescens in life (male, SDBDU 2015.3070); D. Fejervarya sahyadris group, Fejervarya sahyadris in life (male, SDBDU 2015.3046); E – F. Fejervarya syhadrensis group: E. Fejervarya syhadrensis in life (male, SDBDU 2015.2934); F. Fejervarya granosa in life (male, SDBDU 2014.2541). Numbers on the panels represents: 1. dorsal view, 2. ventral view of foot showing the inner metatarsal tubercle, 3. schematic illustration of foot webbing, 4. thighs showing the marking and colour pattern, 5. lateral view showing the groin markings." captionText-4="FIGURE 5. Morphological comparison between members of the genus Sphaerotheca and Fejervarya rufescens group of the Western Ghats. A – H. Genus Sphaerotheca: A – E. Sphaerotheca cf. breviceps, in life (SDBDU 2014.2822): A. dorsolateral view, B. ventral view, C. ventral skin texture, D. lateral view showing the groin markings, E. thighs showing the marking and colour pattern; F – H. Inner metatarsal tubercle: F. Sphaerotheca breviceps (syntype ZMB 3351), G. S. dobsonii (Holotype BMNH 1947.2.28.45), H. S. rolandae (Holotype BMNH 1973.3024). I – R. Fejervarya rufescens group: I – K. F. rufescens in life (SDBDU 2015.3070): I. dorsolateral view, J. ventral view, K. ventral skin texture; L – M. F. kadar sp. nov. (Holotype ZSI / WGRC / V / A / 940): L. lateral view showing the groin markings, M. thighs showing the marking and colour pattern; N – R. Inner metatarsal tubercle: N. F. cepfi sp. nov. (Holotype ZSI / WGRC / V / A / 937), O. F. kadar sp. nov. (Holotype ZSI / WGRC / V / A / 940), P. F. manoharani sp. nov. (Holotype ZSI / WGRC / V / A / 945), Q. F. neilcoxi sp. nov. (Holotype ZSI / WGRC / V / A / 951), R. F. rufescens (SDBDU 2015.3070)." captionText-5="FIGURE 6. PCA plot for the first two principal component axes showing morphometric differentiation among adult males of five species in the Fejervarya rufescens groups of the Western Ghats. Factor loadings for each component are provided in Table 4." httpUri-0="https://zenodo.org/record/829835/files/figure.png" httpUri-1="https://zenodo.org/record/829837/files/figure.png" httpUri-2="https://zenodo.org/record/829839/files/figure.png" httpUri-3="https://zenodo.org/record/829841/files/figure.png" httpUri-4="https://zenodo.org/record/829843/files/figure.png" httpUri-5="https://zenodo.org/record/829845/files/figure.png" pageId="31" pageNumber="482">Figs. 1–6</figureCitation>
, 
<figureCitation id="13002A243135FF85FE55B031D48AFEB4" box="[415,444,259,283]" captionStart="FIGURE 12" captionStartId="33.[151,250,1954,1976]" captionTargetBox="[151,1436,193,1933]" captionTargetId="figure@33.[151,1437,193,1933]" captionTargetPageId="33" captionText="FIGURE 12. Holotype of Fejervarya neilcoxi sp. nov. (ZSI / WGRC / V / A / 951): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829857/files/figure.png" pageId="31" pageNumber="482">12</figureCitation>
)
</paragraph>
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<subSubSection id="C321652A3135FF85FF5DB078D012FE61" pageId="31" pageNumber="482" type="etymology">
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<emphasis id="B94FEAB33135FF85FF5DB078D428FECC" bold="true" box="[151,286,330,355]" pageId="31" pageNumber="482">Etymology.</emphasis>
This species is named for Dr Neil Cox, Manager of the IUCN-Conservation International Biodiversity Assessment Unit. Neil has been associated with the IUCN Red List in a variety of capacities including species assessment and management, and the new species is named particularly in appreciation of his contribution towards the Global Amphibian Assessment. The species epithet 
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<emphasis id="B94FEAB33135FF85FCC2B084D657FE60" box="[776,865,438,463]" italics="true" pageId="31" pageNumber="482">neilcoxi</emphasis>
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is treated as a noun in the genitive case.
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.
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ZSI/ 
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/V/A/951, an adult male, from 
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(
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asl
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), 
<collectingMunicipality id="6BE0ACDB3135FF85FF07B0CCD4BAFDB8" box="[205,396,510,535]" pageId="31" pageNumber="482">Palakkad district</collectingMunicipality>
, 
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state, 
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, collected by SDB and SG on 0 
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<date id="FF8510613135FF85FC00B0CDD166FDB8" box="[970,1104,510,535]" pageId="31" pageNumber="482" value="2015-07-08">8 July 2015</date>
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.
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<paragraph id="8B8436A13135FF85FF0DB310D729FD2C" blockId="31.[151,1437,330,2020]" pageId="31" pageNumber="482">
<materialsCitation id="3B533CFC3135FF85FF0DB310D729FD2C" ID-GBIF-Occurrence="1572402630" collectingDate="2012-08-12" collectionCode="WGRC" country="India" elevation="650" latitude="10.410138" location="Parambikulam" longLatPrecision="1" longitude="76.767944" municipality="Palakkad district" pageId="31" pageNumber="482" specimenCount="2" stateProvince="Kerala" typeStatus="paratype">
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<typeStatus id="548088033135FF85FF0DB310D477FD94" box="[199,321,546,571]" pageId="31" pageNumber="482" type="paratype">Paratypes</typeStatus>
.
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ZSI/ 
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/V/A/952–953, two adult males, and ZSI/ 
<collectionCode id="ED2AAE643135FF85FC7AB310D130FD94" box="[944,1030,546,571]" pageId="31" pageNumber="482">WGRC</collectionCode>
/V/A/954, an adult female collected along with the 
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; ZSI/ 
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/V/A/955, an adult female, from the same locality as holotype, collected by SDB and team on 
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<date id="FF8510613135FF85FEAEB359D72DFD2C" box="[356,539,619,643]" pageId="31" pageNumber="482" value="2012-08-12">12 August 2012</date>
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.
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<emphasis id="B94FEAB33135FF85FF0DB3BCD4F5FD08" bold="true" box="[199,451,654,679]" pageId="31" pageNumber="482">Genetic relationship.</emphasis>
Phylogenetically, 
<taxonomicName id="4C3B4D223135FF85FD50B3BDD64CFD08" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[666,890,654,679]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FD50B3BDD64CFD08" box="[666,890,654,679]" italics="true" pageId="31" pageNumber="482">Fejervarya neilcoxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FC4FB3BCD6D4FD08" box="[901,994,654,679]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FC4FB3BCD6D4FD08" bold="true" box="[901,994,654,679]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
is nested in the 
<taxonomicName id="4C3B4D223135FF85FB61B3BDD0AAFD08" authorityName="Jerdon" authorityYear="1853" box="[1195,1436,655,679]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="B94FEAB33135FF85FB61B3BDD0AAFD08" box="[1195,1436,655,679]" italics="true" pageId="31" pageNumber="482">Fejervarya rufescens</emphasis>
</taxonomicName>
group (
<figureCitation id="13002A243135FF85FF22B380D465FD64" box="[232,339,690,715]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1885,1907]" captionStartId-1="5.[151,250,1618,1640]" captionTargetBox-0="[161,1435,204,1854]" captionTargetBox-1="[155,1424,193,1582]" captionTargetId-0="figure@4.[151,1436,193,1864]" captionTargetId-1="figure@5.[151,1436,193,1597]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of&gt; 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." captionText-1="FIGURE 2. Phylogenetic analyses. A. Maximum Likelihood phylogram based on mitochondrial (16 S, COI and Cytb) and nuclear (Rag 1) DNA of 2314 basepairs, showing phylogenetic relationships among members of the Fejervarya rufescens group. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of&gt; 50 % are indicated above and below the branches, respectively. Fejervarya granosa (2) was used as the outgroup taxon. Voucher number and locality details of the samples are referenced in Table 1; B. Median Joining haplotype network inferred from 554 basepair fragment of the nuclear Rag 1 gene. Circle sizes are proportional to the number of haplotypes, branch lengths are proportional to the number of mutational steps, and black bars represent mutational steps. Number of individuals used for each species: Fejervarya cepfi (3), Fejervarya kadar (1), Fejervarya manoharani (3), Fejervarya neilcoxi (2), and Fejervarya rufescens (5)." httpUri-0="https://zenodo.org/record/829835/files/figure.png" httpUri-1="https://zenodo.org/record/829837/files/figure.png" pageId="31" pageNumber="482">Figs. 1–2</figureCitation>
) of the 
<collectingRegion id="49FFF8433135FF85FE67B381D73EFD65" box="[429,520,691,714]" country="Sri Lanka" name="Western" pageId="31" pageNumber="482">Western</collectingRegion>
Ghats. The average uncorrected pairwise genetic divergence with 
<taxonomicName id="4C3B4D223135FF85FB31B386D0B7FD64" authorityName="Jerdon" authorityYear="1853" box="[1275,1409,691,715]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="B94FEAB33135FF85FB31B386D0B7FD64" box="[1275,1409,691,715]" italics="true" pageId="31" pageNumber="482">F. rufescens</emphasis>
</taxonomicName>
is 3.9% (range 3.7–4.1%, 
<emphasis id="B94FEAB33135FF85FE68B3E5D483FD41" box="[418,437,727,750]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 14) for 16S, 11.2% (range 10.7–11.5%, 
<emphasis id="B94FEAB33135FF85FC5FB3E5D69EFD41" box="[917,936,727,750]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 10) for COI, and 9.7% (range 8.7–10.8%, 
<emphasis id="B94FEAB33135FF85FF5DB3CED59CFCBC" box="[151,170,764,787]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 10) for Cytb; 4.9% (range 4.9–5.0%, 
<emphasis id="B94FEAB33135FF85FDBDB3CED7BCFCBC" box="[631,650,764,787]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 8) for 16S, 11.0% (range 10.9–11.1%, 
<emphasis id="B94FEAB33135FF85FBAFB3CED14EFCBC" box="[1125,1144,764,787]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6) for COI, and 12.8% (range 12.7–12.9%, 
<emphasis id="B94FEAB33135FF85FEB0B22DD4BBFC99" box="[378,397,799,822]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6) for Cytb with 
<taxonomicName id="4C3B4D223135FF85FDAEB22DD782FC98" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[612,692,799,823]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FDAEB22DD782FC98" box="[612,692,799,823]" italics="true" pageId="31" pageNumber="482">F. cepfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FD76B22CD623FC98" box="[700,789,798,823]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FD76B22CD623FC98" bold="true" box="[700,789,798,823]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
; 4.7% (
<emphasis id="B94FEAB33135FF85FCA7B22DD6B6FC99" box="[877,896,799,822]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 2) for 16S, 11.1% (
<emphasis id="B94FEAB33135FF85FBB8B22DD1B3FC99" box="[1138,1157,799,822]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 2) for COI, and 10.8% (
<emphasis id="B94FEAB33135FF85FF6AB276D585FCF4" box="[160,179,836,859]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 2) for Cytb with 
<taxonomicName id="4C3B4D223135FF85FE6BB276D736FCF4" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[417,512,834,859]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FE6BB276D736FCF4" box="[417,512,834,859]" italics="true" pageId="31" pageNumber="482">F. kadar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FDC1B270D75EFCF4" box="[523,616,834,859]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FDC1B270D75EFCF4" bold="true" box="[523,616,834,859]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
; and 3.5% (
<emphasis id="B94FEAB33135FF85FD31B276D638FCF4" box="[763,782,836,859]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 8) for 16S, 8.9% (
<emphasis id="B94FEAB33135FF85FBC0B276D12BFCF4" box="[1034,1053,836,859]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6) for COI, and 10.4% (range 10.3–10.4%, 
<emphasis id="B94FEAB33135FF85FEE6B255D409FCD1" box="[300,319,871,894]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6) for Cytb with 
<taxonomicName id="4C3B4D223135FF85FDD3B255D78BFCD0" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[537,701,870,895]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="manoharani" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FDD3B255D78BFCD0" box="[537,701,870,895]" italics="true" pageId="31" pageNumber="482">F. manoharani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FD0EB254D628FCD0" box="[708,798,870,895]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FD0EB254D628FCD0" bold="true" box="[708,798,870,895]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
(Table 2).
</paragraph>
</subSubSection>
<subSubSection id="C321652A3135FFBAFF0DB2B8D4C6FE8A" lastPageId="32" lastPageNumber="483" pageId="31" pageNumber="482" type="diagnosis">
<paragraph id="8B8436A13135FF85FF0DB2B8D0A4FB30" blockId="31.[151,1437,330,2020]" pageId="31" pageNumber="482">
<emphasis id="B94FEAB33135FF85FF0DB2B8D477FC0C" bold="true" box="[199,321,906,931]" pageId="31" pageNumber="482">Diagnosis.</emphasis>
<taxonomicName id="4C3B4D223135FF85FE85B2BED705FC0C" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[335,563,906,931]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FE85B2BED705FC0C" box="[335,563,906,931]" italics="true" pageId="31" pageNumber="482">Fejervarya neilcoxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FD88B2B8D795FC0C" box="[578,675,906,931]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FD88B2B8D795FC0C" bold="true" box="[578,675,906,931]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from known congeners by the following combination of morphological characters: (1) medium male adult size (SVL 32.0– 
<quantity id="4CC39B443135FF85FBFBB29DD1A0FC69" box="[1073,1174,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="31" pageNumber="482" unit="mm" value="33.3">33.3 mm</quantity>
, 
<emphasis id="B94FEAB33135FF85FB6BB29DD182FC69" box="[1185,1204,943,966]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3); (2) stout body; (3) dorsal skin prominently granular with spinular projections; (4) snout subovoid in dorsal view and obtuse in lateral view; (5) presence of rictal gland at labial commissure of the mouth; (6) eye length nearly equal to snout length (male EL/SL ratio 97.6–97.7%, 
<emphasis id="B94FEAB33135FF85FD87B52ED756FB9C" box="[589,608,1052,1075]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3); (7) tympanum diameter less than half of eye length, (male TYD/EL ratio 38.1–39.0%, 
<emphasis id="B94FEAB33135FF85FEA3B50DD44AFBF9" box="[361,380,1087,1110]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3); (8) inter upper eyelid width nearly three-fourth of upper eyelid width (male IUE/UEW ratio 72.0–75.0%, 
<emphasis id="B94FEAB33135FF85FEA0B556D44BFBD4" box="[362,381,1124,1147]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3) and internarial distance (male IUE/IN ratio 70.0–75.0%, 
<emphasis id="B94FEAB33135FF85FB82B556D16DFBD4" box="[1096,1115,1124,1147]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3); (9) prominent shovelshaped inner metatarsal tubercle prominent and small outer metatarsal tubercle; (10) webbing between toes small.
</paragraph>
<paragraph id="8B8436A13135FF85FF0DB598D71AFAA4" blockId="31.[151,1437,330,2020]" pageId="31" pageNumber="482">
<emphasis id="B94FEAB33135FF85FF0DB598D726FB6C" bold="true" box="[199,528,1194,1219]" pageId="31" pageNumber="482">Morphological comparison.</emphasis>
Based on the overall morphology and comparable body size, 
<taxonomicName id="4C3B4D223135FF85FB0AB59ED0AAFB6C" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[1216,1436,1194,1219]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FB0AB59ED0AAFB6C" box="[1216,1436,1194,1219]" italics="true" pageId="31" pageNumber="482">Fejervarya neilcoxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FF5DB5FCD5C4FB48" box="[151,242,1230,1255]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FF5DB5FCD5C4FB48" bold="true" box="[151,242,1230,1255]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
could be confused with the known species 
<taxonomicName id="4C3B4D223135FF85FD28B5FDD65FFB48" authorityName="Jerdon" authorityYear="1853" box="[738,873,1231,1255]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="B94FEAB33135FF85FD28B5FDD65FFB48" box="[738,873,1231,1255]" italics="true" pageId="31" pageNumber="482">F. rufescens</emphasis>
</taxonomicName>
and three new species, 
<taxonomicName id="4C3B4D223135FF85FBB3B5FDD1FAFB48" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[1145,1228,1231,1255]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FBB3B5FDD1FAFB48" box="[1145,1228,1231,1255]" italics="true" pageId="31" pageNumber="482">F. cepfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FB1EB5FCD019FB48" box="[1236,1327,1230,1255]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FB1EB5FCD019FB48" bold="true" box="[1236,1327,1230,1255]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C3B4D223135FF85FAF4B5FDD0AFFB48" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[1342,1433,1230,1255]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FAF4B5FDD0AFFB48" box="[1342,1433,1230,1255]" italics="true" pageId="31" pageNumber="482">F. kadar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FF5DB5C0D5C6FAA4" box="[151,240,1266,1291]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FF5DB5C0D5C6FAA4" bold="true" box="[151,240,1266,1291]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C3B4D223135FF85FEEDB5C6D4FDFAA4" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[295,459,1266,1291]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="manoharani" status="sp. nov.">
<emphasis id="B94FEAB33135FF85FEEDB5C6D4FDFAA4" box="[295,459,1266,1291]" italics="true" pageId="31" pageNumber="482">F. manoharani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83135FF85FE18B5C0D71AFAA4" box="[466,556,1266,1291]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B94FEAB33135FF85FE18B5C0D71AFAA4" bold="true" box="[466,556,1266,1291]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8B8436A13135FF85FF0DB425D126F948" blockId="31.[151,1437,330,2020]" pageId="31" pageNumber="482">
However, 
<taxonomicName id="4C3B4D223135FF85FEF6B425D72FFA80" authorityName="Garg &amp; Biju" authorityYear="2017" box="[316,537,1302,1327]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="neilcoxi">
<emphasis id="B94FEAB33135FF85FEF6B425D72FFA80" box="[316,537,1302,1327]" italics="true" pageId="31" pageNumber="482">Fejervarya neilcoxi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B4D223135FF85FD79B425D60CFA80" authorityName="Jerdon" authorityYear="1853" box="[691,826,1303,1327]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="B94FEAB33135FF85FD79B425D60CFA80" box="[691,826,1303,1327]" italics="true" pageId="31" pageNumber="482">F. rufescens</emphasis>
</taxonomicName>
by its dorsal skin prominently granular with spinular projections (vs. shagreened to granular without prominent warts); snout obtuse in lateral view (vs. rounded); tympanum to eye distance relatively longer or nearly equal to tympanum diameter, male TYE 
<quantity id="4CC39B443135FF85FB01B46CD060FAD9" box="[1227,1366,1374,1399]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="31" pageNumber="482" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
, TYD 
<quantity id="4CC39B443135FF85FF5DB4B0D5C4FA35" box="[151,242,1410,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="31" pageNumber="482" unit="mm" value="1.6">1.6 mm</quantity>
, TYE/TYD ratio 87.5–100%, 
<emphasis id="B94FEAB33135FF85FD9AB4B6D755FA34" box="[592,611,1412,1435]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. shorter, TYE 
<quantity id="4CC39B443135FF85FCA5B4B1D6CDFA35" box="[879,1019,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="31" pageNumber="482" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
, TYD 
<quantity id="4CC39B443135FF85FB87B4B0D1EFFA35" box="[1101,1241,1410,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="31" pageNumber="482" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6–2.2 mm</quantity>
, TYE/TYD ratio 50.0–76.5%, 
<emphasis id="B94FEAB33135FF85FEE5B495D474FA11" box="[303,322,1447,1470]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6); inter upper eyelid width relatively wider than upper eyelid width, male IUE 
<quantity id="4CC39B443135FF85FAC1B495D0AEFA11" box="[1291,1432,1447,1471]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="31" pageNumber="482" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
, UEW 
<quantity id="4CC39B443135FF85FF2BB4F8D458FA4D" box="[225,366,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="31" pageNumber="482" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
, IUE/UEW ratio 72.0–75.0%, 
<emphasis id="B94FEAB33135FF85FD1CB4FED7DFFA4C" box="[726,745,1484,1507]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. relatively narrower, male IUE 
<quantity id="4CC39B443135FF85FB0AB4F9D07AFA4D" box="[1216,1356,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="31" pageNumber="482" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
, UEW 
<quantity id="4CC39B443135FF85FF5DB4DCD417F9A9" box="[151,289,1518,1543]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="31" pageNumber="482" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
, IUE/UEW ratio 42.9–58.6%, 
<emphasis id="B94FEAB33135FF85FDB5B4DDD7A4F9A9" box="[639,658,1519,1542]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6); thigh shorter than shank length, male TL 14.0– 
<quantity id="4CC39B443135FF85FB3EB4DCD06CF9A9" box="[1268,1370,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="31" pageNumber="482" unit="mm" value="14.5">14.5 mm</quantity>
, SHL 15.3–15.8, TL/SHL ratio 91.5–92.2%, 
<emphasis id="B94FEAB33135FF85FD9EB726D751F984" box="[596,615,1556,1579]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. equal or longer, male TL 14.7–15.8, SHL 14.5–15.8, TL/SHL ratio 100–102.1%, 
<emphasis id="B94FEAB33135FF85FEA4B705D4B7F9E1" box="[366,385,1591,1614]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6); thigh length shorter than foot length, male TL 14.0– 
<quantity id="4CC39B443135FF85FBD9B704D14EF9E1" box="[1043,1144,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="31" pageNumber="482" unit="mm" value="14.5">14.5 mm</quantity>
, FOL 
<quantity id="4CC39B443135FF85FB0AB704D050F9E1" box="[1216,1382,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="1.69" metricValueMin="1.63" pageId="31" pageNumber="482" unit="mm" value="16.6" valueMax="16.9" valueMin="16.3">16.3–16.9 mm</quantity>
, TL/ FOL ratio 85.4–87.1%, 
<emphasis id="B94FEAB33135FF85FE7AB76ED4F5F9DC" box="[432,451,1628,1651]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. nearly equal, male TL 
<quantity id="4CC39B443135FF85FC80B768D6C2F9DD" box="[842,1012,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.58" metricValueMin="1.47" pageId="31" pageNumber="482" unit="mm" value="15.25" valueMax="15.8" valueMin="14.7">14.7–15.8 mm</quantity>
, FOL 
<quantity id="4CC39B443135FF85FB8EB768D1D8F9DD" box="[1092,1262,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.59" metricValueMin="1.48" pageId="31" pageNumber="482" unit="mm" value="15.35" valueMax="15.9" valueMin="14.8">14.8–15.9 mm</quantity>
, TL/FOL ratio 98.0–99.4%, 
<emphasis id="B94FEAB33135FF85FEE7B7B0D476F936" box="[301,320,1666,1689]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 6); and relatively less webbing between toes, male 
<date id="FF8510613135FF85FC6BB7B0D68FF935" box="[929,953,1666,1690]" pageId="31" pageNumber="482">I2</date>
– –2 
<date id="FF8510613135FF85FC17B7B0D6CAF935" box="[989,1020,1666,1690]" pageId="31" pageNumber="482">II2</date>
– –3– 
<date id="FF8510613135FF85FBE2B7B0D164F935" box="[1064,1106,1666,1690]" pageId="31" pageNumber="482">III2</date>
1/2–3 1/2 
<date id="FF8510613135FF85FB56B7B0D1FEF935" box="[1180,1224,1666,1690]" pageId="31" pageNumber="482">IV3</date>
1/2– 
<date id="FF8510613135FF85FB24B7B0D039F935" box="[1262,1295,1666,1690]" pageId="31" pageNumber="482">2V</date>
, specifically the third toe webbing below the first subarticular tubercle on the outside and fourth toe webbing below the second subarticular tubercle on either side (vs. more, male 
<date id="FF8510613135FF85FD1CB7FDD7DBF948" box="[726,749,1743,1767]" pageId="31" pageNumber="482">I2</date>
– –2 
<date id="FF8510613135FF85FCD8B7FDD607F948" box="[786,817,1743,1767]" pageId="31" pageNumber="482">II2</date>
– – 
<date id="FF8510613135FF85FC83B7FDD64DF948" box="[841,891,1740,1767]" bridgedPair="-" pageId="31" pageNumber="482">3–III</date>
2– 
<date id="FF8510613135FF85FC53B7FDD6F3F948" box="[921,965,1743,1767]" pageId="31" pageNumber="482">3IV</date>
3– 
<date id="FF8510613135FF85FC2AB7FDD135F948" box="[992,1027,1743,1767]" pageId="31" pageNumber="482">2V</date>
).
</paragraph>
<paragraph id="8B8436A13135FFBAFF0DB7C6D4C6FE8A" blockId="31.[151,1437,330,2020]" lastBlockId="32.[151,1437,151,2010]" lastPageId="32" lastPageNumber="483" pageId="31" pageNumber="482">
<taxonomicName id="4C3B4D223135FF85FF0DB7C6D495F8A3" authorityName="Garg &amp; Biju" authorityYear="2017" box="[199,419,1779,1804]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="neilcoxi">
<emphasis id="B94FEAB33135FF85FF0DB7C6D495F8A3" box="[199,419,1779,1804]" italics="true" pageId="31" pageNumber="482">Fejervarya neilcoxi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B4D223135FF85FDF2B7C6D7BCF8A4" authorityName="Garg &amp; Biju" authorityYear="2017" box="[568,650,1779,1803]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="cepfi">
<emphasis id="B94FEAB33135FF85FDF2B7C6D7BCF8A4" box="[568,650,1779,1803]" italics="true" pageId="31" pageNumber="482">F. cepfi</emphasis>
</taxonomicName>
by its dorsal skin with relatively more prominent glandular warts (vs. less); horizontal diameter of eye nearly equal to snout length, EL 
<quantity id="4CC39B443135FF85FC63B625D101F880" box="[937,1079,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="31" pageNumber="482" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1–4.2 mm</quantity>
, SL 
<quantity id="4CC39B443135FF85FBBEB625D034F880" box="[1140,1282,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="31" pageNumber="482" unit="mm" value="4.25" valueMax="4.3" valueMin="4.2">4.2–4.3 mm</quantity>
, EL/SL ratio 97.6–97.7%, 
<emphasis id="B94FEAB33135FF85FEE6B60ED409F8FC" box="[300,319,1852,1875]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. smaller, male EL 
<quantity id="4CC39B443135FF85FDA9B609D7DDF8FC" box="[611,747,1851,1876]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="31" pageNumber="482" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
, SL 
<quantity id="4CC39B443135FF85FCD7B609D693F8FC" box="[797,933,1851,1876]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="31" pageNumber="482" unit="mm" value="4.699999999999999" valueMax="4.8" valueMin="4.6">4.6–4.8 mm</quantity>
, EL/SL ratio 72.9–73.9%, 
<emphasis id="B94FEAB33135FF85FB05B60ED1D4F8FC" box="[1231,1250,1852,1875]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 2); tympanum diameter less than half of horizontal diameter of eye, male TYD 
<quantity id="4CC39B443135FF85FC7FB66DD122F8D8" box="[949,1044,1887,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="31" pageNumber="482" unit="mm" value="1.6">1.6 mm</quantity>
, EL 
<quantity id="4CC39B443135FF85FB9CB66DD1D3F8D8" box="[1110,1253,1887,1911]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="31" pageNumber="482" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1–4.2 mm</quantity>
, TYD/EL ratio 38.1–39.0%, 
<emphasis id="B94FEAB33135FF85FEE7B6B6D476F834" box="[301,320,1924,1947]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. nearly half, male TYD 
<quantity id="4CC39B443135FF85FD64B6B1D630F834" box="[686,774,1923,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="482" unit="mm" value="1.8">1.8 mm</quantity>
, EL 
<quantity id="4CC39B443135FF85FCF7B6B1D6F1F834" box="[829,967,1923,1948]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="31" pageNumber="482" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
, TYD/EL ratio 51.4–52.9%, 
<emphasis id="B94FEAB33135FF85FAC5B6B6D014F834" box="[1295,1314,1924,1947]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 2); inter upper eyelid width narrower than upper eyelid width, male IUE 
<quantity id="4CC39B443135FF85FCBEB695D6C8F810" box="[884,1022,1959,1983]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="31" pageNumber="482" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
, UEW 
<quantity id="4CC39B443135FF85FB99B695D1EBF810" box="[1107,1245,1959,1984]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="31" pageNumber="482" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
, IUE/UEW ratio 72.0–75.0%, 
<emphasis id="B94FEAB33135FF85FEE6B6FED409F84C" box="[300,319,1996,2019]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 3 (vs. nearly equal, male IUE 
<quantity id="4CC39B443135FF85FD7BB6F9D60DF84C" box="[689,827,1995,2020]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="31" pageNumber="482" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
, UEW 
<quantity id="4CC39B443135FF85FC44B6F9D12EF84C" box="[910,1048,1995,2020]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="31" pageNumber="482" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
, IUE/UEW ratio 96.0–96.2%, 
<emphasis id="B94FEAB33135FF85FAB8B6FED0B3F84C" box="[1394,1413,1996,2019]" italics="true" pageId="31" pageNumber="482">N</emphasis>
= 2); inter upper eyelid width narrower than internarial distance, male IUE 
<quantity id="4CC39B44310AFFBAFC0EB1AAD17AFF00" box="[964,1100,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="32" pageNumber="483" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
, IN 2.5–3.0 mm, IUE/IN ratio 70.0–75.0%, 
<emphasis id="B94FEAB3310AFFBAFEE5B18FD474FF7B" box="[303,322,189,212]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. nearly equal, male IUE 
<quantity id="4CC39B44310AFFBAFD06B18ED66EFF7B" box="[716,856,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="32" pageNumber="483" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
, IN 
<quantity id="4CC39B44310AFFBAFC45B18ED6DDFF7B" box="[911,1003,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="483" unit="mm" value="2.5">2.5 mm</quantity>
, IUE/IN ratio 96–100%, 
<emphasis id="B94FEAB3310AFFBAFAD9B18FD010FF7B" box="[1299,1318,189,212]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 2); and relatively less webbing between toes, male 
<date id="FF851061310AFFBAFCEFB1D1D60AFF54" box="[805,828,227,251]" pageId="32" pageNumber="483">I2</date>
– –2 
<date id="FF851061310AFFBAFCAAB1D1D6B6FF54" box="[864,896,227,251]" pageId="32" pageNumber="483">II2</date>
– –3– 
<date id="FF851061310AFFBAFC66B1D1D6E3FF54" box="[940,981,227,251]" pageId="32" pageNumber="483">III2</date>
1/2–3 1/2 
<date id="FF851061310AFFBAFBEAB1D1D17DFF54" box="[1056,1099,227,251]" pageId="32" pageNumber="483">IV3</date>
1/2– 
<date id="FF851061310AFFBAFBBBB1D1D1A5FF54" box="[1137,1171,227,251]" pageId="32" pageNumber="483">2V</date>
(vs. more, male 
<date id="FF851061310AFFBAFF5DB03FD598FE8A" box="[151,174,269,293]" pageId="32" pageNumber="483">I1</date>
+–2– 
<date id="FF851061310AFFBAFF11B03FD5CDFE8A" box="[219,251,269,293]" pageId="32" pageNumber="483">II1</date>
+– 
<date id="FF851061310AFFBAFEDEB03FD473FE8A" box="[276,325,265,293]" bridgedPair="-" pageId="32" pageNumber="483">3–III</date>
2– 
<date id="FF851061310AFFBAFEA9B03FD4A6FE8A" box="[355,400,269,293]" pageId="32" pageNumber="483">3IV</date>
3–1 1/ 
<date id="FF851061310AFFBAFE0EB029D4D2FE8A" box="[452,484,269,297]" pageId="32" pageNumber="483">2V</date>
).
</paragraph>
</subSubSection>
<subSubSection id="C321652A310AFFBAFF0DB006D71DFBE4" pageId="32" pageNumber="483" type="description">
<paragraph id="8B8436A1310AFFBAFF0DB006D117FDC1" blockId="32.[151,1437,151,2010]" pageId="32" pageNumber="483">
<taxonomicName id="4C3B4D22310AFFBAFF0DB006D491FEE3" authorityName="Garg &amp; Biju" authorityYear="2017" box="[199,423,307,332]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="neilcoxi">
<emphasis id="B94FEAB3310AFFBAFF0DB006D491FEE3" box="[199,423,307,332]" italics="true" pageId="32" pageNumber="483">Fejervarya neilcoxi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B4D22310AFFBAFD82B006D793FEE3" authorityName="Garg &amp; Biju" authorityYear="2017" box="[584,677,307,332]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="kadar">
<emphasis id="B94FEAB3310AFFBAFD82B006D793FEE3" box="[584,677,307,332]" italics="true" pageId="32" pageNumber="483">F. kadar</emphasis>
</taxonomicName>
by its dorsal skin with prominent granular projections (vs. with scattered glandular warts); snout subovoid in dorsal view (vs. rounded); eye length nearly equal to snout length, male EL 
<quantity id="4CC39B44310AFFBAFF37B049D4B3FE3C" box="[253,389,379,403]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="32" pageNumber="483" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1–4.2 mm</quantity>
, SL 
<quantity id="4CC39B44310AFFBAFE7DB049D776FE3C" box="[439,576,379,403]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="32" pageNumber="483" unit="mm" value="4.25" valueMax="4.3" valueMin="4.2">4.2–4.3 mm</quantity>
, EL/SL ratio 97.6–97.7%, 
<emphasis id="B94FEAB3310AFFBAFCA1B04ED648FE3C" box="[875,894,380,403]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. relatively shorter, male EL 4.0– 
<quantity id="4CC39B44310AFFBAFA8AB049D0AEFE3C" box="[1344,1432,379,403]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="32" pageNumber="483" unit="mm" value="4.2">4.2 mm</quantity>
, SL 
<quantity id="4CC39B44310AFFBAFF74B0ADD470FE18" box="[190,326,415,440]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="32" pageNumber="483" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
, EL/SL ratio 88.9–93.3%, 
<emphasis id="B94FEAB3310AFFBAFDBBB092D7B2FE18" box="[625,644,416,439]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3); inter upper eyelid width relatively wider than upper eyelid width, male IUE 
<quantity id="4CC39B44310AFFBAFEC4B0F1D4ACFE74" box="[270,410,451,475]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="32" pageNumber="483" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
, UEW 
<quantity id="4CC39B44310AFFBAFE3BB0F1D74AFE74" box="[497,636,451,476]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="32" pageNumber="483" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
, IUE/UEW ratio 72.0–75.0%, 
<emphasis id="B94FEAB3310AFFBAFC17B0F6D6C6FE74" box="[989,1008,452,475]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. narrower, male IUE 
<quantity id="4CC39B44310AFFBAFA82B0F1D5F3FE50" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="32" pageNumber="483" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
, UEW 3.0– 
<quantity id="4CC39B44310AFFBAFE9AB0D5D49CFE50" box="[336,426,487,511]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="32" pageNumber="483" unit="mm" value="3.3">3.3 mm</quantity>
, IUE/UEW ratio 53.1–60.0%, 
<emphasis id="B94FEAB3310AFFBAFCC5B0DAD614FE50" box="[783,802,488,511]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3); thigh length shorter than shank length, male TL 14.0– 
<quantity id="4CC39B44310AFFBAFF1DB339D40BFD8C" box="[215,317,523,548]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="32" pageNumber="483" unit="mm" value="14.5">14.5 mm</quantity>
, SHL 
<quantity id="4CC39B44310AFFBAFE4DB339D71BFD8C" box="[391,557,523,548]" metricMagnitude="-2" metricUnit="m" metricValue="1.555" metricValueMax="1.58" metricValueMin="1.53" pageId="32" pageNumber="483" unit="mm" value="15.55" valueMax="15.8" valueMin="15.3">15.3–15.8 mm</quantity>
, TL/SHL ratio 91.5–92.2%, 
<emphasis id="B94FEAB3310AFFBAFCB9B33ED6B0FD8C" box="[883,902,524,547]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. nearly equal, male TL 
<quantity id="4CC39B44310AFFBAFB3BB339D0A1FD8C" box="[1265,1431,523,548]" metricMagnitude="-2" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.54" metricValueMin="1.45" pageId="32" pageNumber="483" unit="mm" value="14.95" valueMax="15.4" valueMin="14.5">14.5–15.4 mm</quantity>
, SHL 
<quantity id="4CC39B44310AFFBAFF10B31DD4B0FDE8" box="[218,390,559,584]" metricMagnitude="-2" metricUnit="m" metricValue="1.505" metricValueMax="1.55" metricValueMin="1.46" pageId="32" pageNumber="483" unit="mm" value="15.05" valueMax="15.5" valueMin="14.6">14.6–15.5 mm</quantity>
, TL/SHL ratio 99.3–99.4%, 
<emphasis id="B94FEAB3310AFFBAFD2FB302D7CEFDE8" box="[741,760,560,583]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3); and relatively more webbing between toes, male 
<date id="FF851061310AFFBAFF5DB364D598FDC1" box="[151,174,598,622]" pageId="32" pageNumber="483">I2</date>
– –2 
<date id="FF851061310AFFBAFF18B364D5C4FDC1" box="[210,242,598,622]" pageId="32" pageNumber="483">II2</date>
– –3– 
<date id="FF851061310AFFBAFED4B364D471FDC1" box="[286,327,598,622]" pageId="32" pageNumber="483">III2</date>
1/2–3 1/2 
<date id="FF851061310AFFBAFE58B364D48BFDC1" box="[402,445,598,622]" pageId="32" pageNumber="483">IV3</date>
1/2– 
<date id="FF851061310AFFBAFE2EB364D733FDC1" box="[484,517,598,622]" pageId="32" pageNumber="483">2V</date>
(vs. less, male 
<date id="FF851061310AFFBAFD78B364D7FFFDC1" box="[690,713,598,622]" pageId="32" pageNumber="483">I2</date>
– –2– 
<date id="FF851061310AFFBAFD3EB364D623FDC1" box="[756,789,598,622]" pageId="32" pageNumber="483">II2</date>
– – 
<date id="FF851061310AFFBAFCE6B364D66BFDC1" box="[812,861,594,622]" bridgedPair="-" pageId="32" pageNumber="483">3–III</date>
2–3 2/3 
<date id="FF851061310AFFBAFC57B364D6FFFDC1" box="[925,969,598,622]" pageId="32" pageNumber="483">IV3</date>
2/3– 
<date id="FF851061310AFFBAFC3AB364D122FDC1" box="[1008,1044,598,622]" pageId="32" pageNumber="483">2V</date>
).
</paragraph>
<paragraph id="8B8436A1310AFFBAFF0DB34FD71DFBE4" blockId="32.[151,1437,151,2010]" pageId="32" pageNumber="483">
<taxonomicName id="4C3B4D22310AFFBAFF0DB34FD49AFD3A" authorityName="Garg &amp; Biju" authorityYear="2017" box="[199,428,636,661]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="neilcoxi">
<emphasis id="B94FEAB3310AFFBAFF0DB34FD49AFD3A" box="[199,428,636,661]" italics="true" pageId="32" pageNumber="483">Fejervarya neilcoxi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B4D22310AFFBAFD92B34CD633FD3A" authorityName="Garg &amp; Biju" authorityYear="2017" box="[600,773,636,661]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="manoharani">
<emphasis id="B94FEAB3310AFFBAFD92B34CD633FD3A" box="[600,773,636,661]" italics="true" pageId="32" pageNumber="483">F. manoharani</emphasis>
</taxonomicName>
by its prominently granular dorsal skin with spinular projections (vs. glandular with interrupted linear warts); snout subovoid in dorsal view (vs. rounded) and obtuse in lateral view (vs. vertical); relatively larger snout-vent size, male SVL 32.0– 
<quantity id="4CC39B44310AFFBAFC2FB3F7D17CFD73" box="[997,1098,709,733]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="32" pageNumber="483" unit="mm" value="33.3">33.3 mm</quantity>
, 
<emphasis id="B94FEAB3310AFFBAFB9FB3F4D151FD72" box="[1109,1127,710,733]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. smaller, male SVL 28.1–30.0 mm, 
<emphasis id="B94FEAB3310AFFBAFE80B3D8D46BFCAE" box="[330,349,746,769]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 4); inter upper eyelid width relatively wider than upper eyelid width, male IUE 
<quantity id="4CC39B44310AFFBAFAC7B3DBD0AEFCAF" box="[1293,1432,745,769]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="32" pageNumber="483" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
, UEW 
<quantity id="4CC39B44310AFFBAFF15B23ED45FFC8B" box="[223,361,780,805]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="32" pageNumber="483" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
, IUE/UEW ratio 72.0–75.0%, 
<emphasis id="B94FEAB3310AFFBAFD0CB23CD7EFFC8A" box="[710,729,782,805]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. relatively narrower, male IUE 1.7–2.0 mm, UEW 3.0 mm, IUE/UEW ratio 56.7–66.7%, 
<emphasis id="B94FEAB3310AFFBAFDEDB200D70CFCE6" box="[551,570,818,841]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 4); forearm length relatively longer than hand length, male FAL 
<quantity id="4CC39B44310AFFBAFA82B202D5F3FCC3" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.2" metricValueMin="6.1" pageId="32" pageNumber="483" unit="mm" value="6.15" valueMax="6.2" valueMin="6.1">6.1–6.2 mm</quantity>
, HAL 
<quantity id="4CC39B44310AFFBAFEDFB267D497FCC3" box="[277,417,853,877]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.4" metricValueMin="7.1" pageId="32" pageNumber="483" unit="mm" value="7.25" valueMax="7.4" valueMin="7.1">7.1–7.4 mm</quantity>
, FAL/HAL ratio 83.8–86.1%, 
<emphasis id="B94FEAB3310AFFBAFD35B264D624FCC2" box="[767,786,854,877]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. relatively shorter, male FAL 
<quantity id="4CC39B44310AFFBAFB03B266D063FCC3" box="[1225,1365,852,877]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="32" pageNumber="483" unit="mm" value="5.25" valueMax="5.4" valueMin="5.1">5.1–5.4 mm</quantity>
, HAL 
<quantity id="4CC39B44310AFFBAFF5DB24AD412FC3F" box="[151,292,888,913]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.9" metricValueMin="6.4" pageId="32" pageNumber="483" unit="mm" value="6.65" valueMax="6.9" valueMin="6.4">6.4–6.9 mm</quantity>
, FAL/HAL ratio 76.8–79.7%, 
<emphasis id="B94FEAB3310AFFBAFD47B248D796FC3E" box="[653,672,890,913]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 4); thigh length relatively shorter than shank length, male TL 14.0– 
<quantity id="4CC39B44310AFFBAFF1DB2AED40BFC1B" box="[215,317,924,949]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="32" pageNumber="483" unit="mm" value="14.5">14.5 mm</quantity>
, SHL 
<quantity id="4CC39B44310AFFBAFE4CB2AED71AFC1B" box="[390,556,924,949]" metricMagnitude="-2" metricUnit="m" metricValue="1.555" metricValueMax="1.58" metricValueMin="1.53" pageId="32" pageNumber="483" unit="mm" value="15.55" valueMax="15.8" valueMin="15.3">15.3–15.8 mm</quantity>
, TL/SHL ratio 91.5–92.2%, 
<emphasis id="B94FEAB3310AFFBAFCBBB2ACD6B2FC1A" box="[881,900,926,949]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. relatively longer or nearly equal, male TL 
<quantity id="4CC39B44310AFFBAFF0BB2F2D451FC77" box="[193,359,960,985]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.35" metricValueMin="1.29" pageId="32" pageNumber="483" unit="mm" value="13.2" valueMax="13.5" valueMin="12.9">12.9–13.5 mm</quantity>
, SHL 
<quantity id="4CC39B44310AFFBAFE7AB2F3D760FC77" box="[432,598,961,985]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.38" metricValueMin="1.3199999999999998" pageId="32" pageNumber="483" unit="mm" value="13.5" valueMax="13.8" valueMin="13.2">13.2–13.8 mm</quantity>
, TL/SHL ratio 96.3 – 99.2%, 
<emphasis id="B94FEAB3310AFFBAFC60B2F0D68BFC76" box="[938,957,962,985]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 4); and relatively less webbing between toes, male 
<date id="FF851061310AFFBAFEC4B2DAD413FBAF" box="[270,293,1000,1024]" pageId="32" pageNumber="483">I2</date>
– –2 
<date id="FF851061310AFFBAFE83B2DAD45FFBAF" box="[329,361,1000,1024]" pageId="32" pageNumber="483">II2</date>
– –3– 
<date id="FF851061310AFFBAFE5FB2DAD488FBAF" box="[405,446,1000,1024]" pageId="32" pageNumber="483">III2</date>
1/2–3 1/2 
<date id="FF851061310AFFBAFDC3B2DAD703FBAF" box="[521,565,1000,1024]" pageId="32" pageNumber="483">IV3</date>
1/2– 
<date id="FF851061310AFFBAFD91B2DAD74AFBAF" box="[603,636,1000,1024]" pageId="32" pageNumber="483">2V</date>
(vs. more, male I2–2 
<date id="FF851061310AFFBAFCA3B2DAD6BCFBAF" box="[873,906,1000,1024]" pageId="32" pageNumber="483">II2</date>
– – 
<date id="FF851061310AFFBAFC6BB2DAD6E4FBAF" box="[929,978,996,1024]" bridgedPair="-" pageId="32" pageNumber="483">3–III</date>
2–3 1/3 
<date id="FF851061310AFFBAFBD8B2DAD10BFBAF" box="[1042,1085,1000,1024]" pageId="32" pageNumber="483">IV3</date>
1/3– 
<date id="FF851061310AFFBAFBA9B2DAD1B2FBAF" box="[1123,1156,1000,1024]" pageId="32" pageNumber="483">2V</date>
, specifically the third toe webbing upto the first subarticular tubercle on the outside and fourth toe webbing extending closer to the second subarticular tubercle on either side).
</paragraph>
</subSubSection>
<subSubSection id="C321652A310AFFBAFF0DB564D14BF987" pageId="32" pageNumber="483" type="reference_group">
<paragraph id="8B8436A1310AFFBAFF0DB564D14BF987" blockId="32.[151,1437,151,2010]" pageId="32" pageNumber="483">
Since 
<taxonomicName id="4C3B4D22310AFFBAFEDAB565D4C6FBC0" ID-CoL="6HR34" authorityName="Garg &amp; Biju" authorityYear="2017" box="[272,496,1110,1135]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="neilcoxi">
<emphasis id="B94FEAB3310AFFBAFEDAB565D4C6FBC0" box="[272,496,1110,1135]" italics="true" pageId="32" pageNumber="483">Fejervarya neilcoxi</emphasis>
</taxonomicName>
is found in the same geographical region as 
<taxonomicName id="4C3B4D22310AFFBAFBCCB565D5ECFB3C" ID-CoL="6HR2Y" authority="Rao 1937" authorityName="Rao" authorityYear="1937" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="parambikulamana">
<emphasis id="B94FEAB3310AFFBAFBCCB565D06CFBC0" box="[1030,1370,1110,1135]" italics="true" pageId="32" pageNumber="483">Fejervarya parambikulamana</emphasis>
(
<bibRefCitation id="EFAA4B50310AFFBAFAA7B565D5E4FB3C" author="Rao" pageId="32" pageNumber="483" refString="Rao, C. R. N. (1937) On some new forms of Batrachia from S. India. Proceedings of the Indian Academy of Sciences: Section B, 6, 387 - 427." type="journal article" year="1937">Rao 1937</bibRefCitation>
)
</taxonomicName>
, for which the 
<typeStatus id="54808803310AFFBAFE4BB549D487FB3C" box="[385,433,1147,1171]" pageId="32" pageNumber="483">type</typeStatus>
specimen is now lost (
<bibRefCitation id="EFAA4B50310AFFBAFD65B548D672FB3C" author="Dubois" box="[687,836,1146,1171]" pageId="32" pageNumber="483" refString="Dubois, A. (1984) Note preliminaire sur le groupe de Rana limnocharis Gravenhorst, 1829 (Amphibiens, Anoures). Alytes, 3 (4), 143 - 159." type="journal article" year="1984">Dubois 1984</bibRefCitation>
), we also compared this new species with the original description of 
<taxonomicName id="4C3B4D22310AFFBAFE92B5ADD68BFB18" authority="Rao 1937" authorityName="Rao" authorityYear="1937" box="[344,957,1182,1207]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="parambikulamana" subGenus="Tomopterna">
<emphasis id="B94FEAB3310AFFBAFE92B5ADD4A5FB19" box="[344,403,1183,1206]" italics="true" pageId="32" pageNumber="483">Rana</emphasis>
(
<emphasis id="B94FEAB3310AFFBAFE7EB5ADD70CFB19" box="[436,570,1183,1206]" italics="true" pageId="32" pageNumber="483">Tomopterna</emphasis>
) 
<emphasis id="B94FEAB3310AFFBAFD90B5ACD611FB18" box="[602,807,1182,1207]" italics="true" pageId="32" pageNumber="483">parambikulamana</emphasis>
<bibRefCitation id="EFAA4B50310AFFBAFCF4B5ADD68BFB18" author="Rao" box="[830,957,1183,1207]" pageId="32" pageNumber="483" refString="Rao, C. R. N. (1937) On some new forms of Batrachia from S. India. Proceedings of the Indian Academy of Sciences: Section B, 6, 387 - 427." type="journal article" year="1937">Rao 1937</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="4C3B4D22310AFFBAFC12B5ADD1F2FB18" ID-CoL="6HR34" authorityName="Garg &amp; Biju" authorityYear="2017" box="[984,1220,1182,1207]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="neilcoxi">
<emphasis id="B94FEAB3310AFFBAFC12B5ADD1F2FB18" box="[984,1220,1182,1207]" italics="true" pageId="32" pageNumber="483">Fejervarya neilcoxi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3B4D22310AFFBAFA4DB5ADD452FB74" authorityName="Rao" authorityYear="1937" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="parambikulamana">
<emphasis id="B94FEAB3310AFFBAFA4DB5ADD452FB74" italics="true" pageId="32" pageNumber="483">F. parambikulamana</emphasis>
</taxonomicName>
by its relatively smaller snout-vent size, male SVL 32.0– 
<quantity id="4CC39B44310AFFBAFC2CB5F1D17CFB75" box="[998,1098,1219,1243]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="32" pageNumber="483" unit="mm" value="33.3">33.3 mm</quantity>
, 
<emphasis id="B94FEAB3310AFFBAFB9FB5F1D151FB75" box="[1109,1127,1219,1242]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. larger, male “From snout to vent 39.00 mm”, 
<emphasis id="B94FEAB3310AFFBAFE00B5D5D4EBFB51" box="[458,477,1255,1278]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1); prominently granular dorsal skin with spinular projections (vs. “skin smooth above” and “no granulation on any part of the body”); head wider than long, male HW 
<quantity id="4CC39B44310AFFBAFB77B439D050FA8D" box="[1213,1382,1291,1315]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.21" metricValueMin="1.19" pageId="32" pageNumber="483" unit="mm" value="12.0" valueMax="12.1" valueMin="11.9">11.9–12.1 mm</quantity>
, HL 
<quantity id="4CC39B44310AFFBAFF5DB41DD40DFAE9" box="[151,315,1327,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.155" metricValueMax="1.17" metricValueMin="1.14" pageId="32" pageNumber="483" unit="mm" value="11.55" valueMax="11.7" valueMin="11.4">11.4–11.7 mm</quantity>
, 
<emphasis id="B94FEAB3310AFFBAFE83B41DD46AFAE9" box="[329,348,1327,1350]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. head “distinctly longer than broad”, HW “11.50” mm, HL “15.00” mm, 
<emphasis id="B94FEAB3310AFFBAFAC9B41DD020FAE9" box="[1283,1302,1327,1350]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1); snout nearly equal to the diameter of the eye, male SL 
<quantity id="4CC39B44310AFFBAFD72B461D676FAC5" box="[696,832,1363,1387]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="32" pageNumber="483" unit="mm" value="4.25" valueMax="4.3" valueMin="4.2">4.2–4.3 mm</quantity>
, EL 
<quantity id="4CC39B44310AFFBAFCBFB461D6C8FAC5" box="[885,1022,1363,1387]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="32" pageNumber="483" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1–4.2 mm</quantity>
, 
<emphasis id="B94FEAB3310AFFBAFBC3B461D12AFAC5" box="[1033,1052,1363,1386]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 3 (vs. “longer than the eye”, SL “6.50” mm, EL “5.00” mm, 
<emphasis id="B94FEAB3310AFFBAFE13B445D4DAFA21" box="[473,492,1399,1422]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1); first finger longer than the second, FL I 
<quantity id="4CC39B44310AFFBAFC26B444D173FA21" box="[1004,1093,1398,1423]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="32" pageNumber="483" unit="mm" value="3.6">3.6 mm</quantity>
, FL II 
<quantity id="4CC39B44310AFFBAFB4EB445D1EBFA21" box="[1156,1245,1399,1423]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="32" pageNumber="483" unit="mm" value="2.9">2.9 mm</quantity>
, 
<emphasis id="B94FEAB3310AFFBAFB20B445D1CBFA21" box="[1258,1277,1399,1422]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1 (vs. “first nearly equal to the 2nd”, FL I “6.00 mm”, FL II “ 
<quantity id="4CC39B44310AFFBAFD6AB4AFD63FFA19" box="[672,777,1437,1462]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" pageId="32" pageNumber="483" unit="mm" value="5.75">5.75 mm</quantity>
”, 
<emphasis id="B94FEAB3310AFFBAFCD4B4ACD607FA1A" box="[798,817,1438,1461]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1); and fourth finger length considerably shorter than the diameter of the eye, FL IV 
<quantity id="4CC39B44310AFFBAFE14B4F6D700FA73" box="[478,566,1476,1501]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="32" pageNumber="483" unit="mm" value="2.6">2.6 mm</quantity>
, EL 
<quantity id="4CC39B44310AFFBAFDA1B4F6D7F5FA73" box="[619,707,1476,1500]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="32" pageNumber="483" unit="mm" value="4.2">4.2 mm</quantity>
, 
<emphasis id="B94FEAB3310AFFBAFD04B4F7D7D7FA73" box="[718,737,1477,1500]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1 (vs. “fourth digit equals the diameter of the eye”, FL IV 5.0 mm, EL 5.0 mm, 
<emphasis id="B94FEAB3310AFFBAFE95B4DED444F9AC" box="[351,370,1516,1539]" italics="true" pageId="32" pageNumber="483">N</emphasis>
= 1). Based on the original description and the accompanying illustration (
<bibRefCitation id="EFAA4B50310AFFBAFB0BB4D9D5E1F988" author="Rao" pageId="32" pageNumber="483" refString="Rao, C. R. N. (1937) On some new forms of Batrachia from S. India. Proceedings of the Indian Academy of Sciences: Section B, 6, 387 - 427." type="journal article" year="1937">
Rao 1937, Pl. XXI, 
<figureCitation id="13002A24310AFFBAFF5DB73CD5E1F988" box="[151,215,1550,1575]" captionStart="FIGURE 1" captionStartId="4.[151,250,1885,1907]" captionTargetBox="[161,1435,204,1854]" captionTargetId="figure@4.[151,1436,193,1864]" captionTargetPageId="4" captionText="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of&gt; 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." httpUri="https://zenodo.org/record/829835/files/figure.png" pageId="32" pageNumber="483">Fig. 1</figureCitation>
</bibRefCitation>
), 
<taxonomicName id="4C3B4D22310AFFBAFF25B722D4EFF988" authorityName="Rao" authorityYear="1937" box="[239,473,1550,1575]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="parambikulamana">
<emphasis id="B94FEAB3310AFFBAFF25B722D4EFF988" box="[239,473,1550,1575]" italics="true" pageId="32" pageNumber="483">F. parambikulamana</emphasis>
</taxonomicName>
is likely to be a member of the 
<taxonomicName id="4C3B4D22310AFFBAFCF6B722D11BF988" ID-CoL="6HR2X" box="[828,1069,1550,1575]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="nilagirica">
<emphasis id="B94FEAB3310AFFBAFCF6B722D11BF988" box="[828,1069,1550,1575]" italics="true" pageId="32" pageNumber="483">Fejervarya nilagirica</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C321652A310AFFB8FF0DB700D66DFB98" lastPageId="34" lastPageNumber="485" pageId="32" pageNumber="483" type="description">
<paragraph id="8B8436A1310AFFB8FF0DB700D464FE23" blockId="32.[151,1437,151,2010]" lastBlockId="34.[151,1437,182,1260]" lastPageId="34" lastPageNumber="485" pageId="32" pageNumber="483">
<emphasis id="B94FEAB3310AFFBAFF0DB700D7CAF9E4" bold="true" box="[199,764,1586,1611]" pageId="32" pageNumber="483">
Description of 
<typeStatus id="54808803310AFFBAFEB3B700D4D6F9E4" box="[377,480,1586,1611]" pageId="32" pageNumber="483" type="holotype">holotype</typeStatus>
<emphasis id="B94FEAB3310AFFBAFE22B700D7C3F9E4" bold="true" box="[488,757,1586,1611]" italics="true" pageId="32" pageNumber="483">(measurements in mm)</emphasis>
.
</emphasis>
Adult male (SVL 33.3), rather stout; head wider than long (HW 12.1, HL 11.7); snout subovoid in dorsal view and obtuse in lateral view, its length (SL 4.3) nearly equal to horizontal diameter of eye (EL 4.2); loreal region acute with rounded canthus rostralis; interorbital space flat, narrower (IUE 2.1) than upper eyelid (UEW 2.9) and internarial distance (IN 3.0); nostril nearly as close to eye (EN 2.1) as to tip of snout (NS 2.0); tympanum (TYD 1.6) 38.1% of eye diameter (EL 4.2); tympanum-eye distance (TYE 1.4), 87.5% of the tympanum diameter (TYD 1.6); supratympanic fold well developed, extends from posterior corner of eye to near the shoulder; vomerine ridge present, bearing small teeth, at an angle of 45° to the body axis, as close to choanae as to each other; tongue moderately large, emarginated, bearing no median lingual process; rictal gland present at labial commissure of the mouth. Arms short, forearm length (FAL 6.2) shorter then hand length (HAL 7.4); relative length of fingers IV&lt;II&lt;I&lt;III (FL I 3.6, FL II 2.9, FL III 4.5, FL IV 2.6); finger tips rounded, slightly enlarged without discs, fingers without fringes, webbing between fingers absent; subarticular tubercles prominent, circular; one distinct palmar tubercle, oval, bifid; supernumerary tubercles absent. Hind limbs short, thigh (TL 14.5) shorter than shank (SHL 15.8) and foot (FOL 16.9), distance from the base of tarsus to the tip of toe IV (TFOL 22.7); toes long, relative length of toes I&lt;II&lt;V&lt;III&lt;IV; toe tips rounded, slightly enlarged without discs, toes without fringes, webbing between toes small: 
<date id="FF8510613108FFB8FBACB030D14BFEB5" box="[1126,1149,258,282]" pageId="34" pageNumber="485">I2</date>
– –2 
<date id="FF8510613108FFB8FB6BB030D1F7FEB5" box="[1185,1217,258,282]" pageId="34" pageNumber="485">II2</date>
– –3– 
<date id="FF8510613108FFB8FB26B030D020FEB5" box="[1260,1302,258,282]" pageId="34" pageNumber="485">III2</date>
1/2–3 1/2 
<date id="FF8510613108FFB8FAABB030D0BAFEB5" box="[1377,1420,258,282]" pageId="34" pageNumber="485">IV3</date>
1/ 2– 
<date id="FF8510613108FFB8FF64B01AD5E6FEEF" box="[174,208,296,320]" pageId="34" pageNumber="485">2V</date>
; inner toe length (ITL 2.8); subarticular tubercles prominent, all present, circular; inner metatarsal tubercle prominent, shovel-shaped (IMT 1.9); outer metatarsal tubercle small (OMT 0.7), rounded; supernumerary tubercles absent.
</paragraph>
<caption id="DF446629310BFFBBFF5DB690D16CF859" httpUri="https://zenodo.org/record/829857/files/figure.png" pageId="33" pageNumber="484" targetBox="[151,1436,193,1933]" targetPageId="33">
<paragraph id="8B8436A1310BFFBBFF5DB690D16CF859" blockId="33.[151,1436,1954,2038]" pageId="33" pageNumber="484">
<emphasis id="B94FEAB3310BFFBBFF5DB690D416F818" bold="true" box="[151,288,1954,1976]" pageId="33" pageNumber="484">FIGURE 12.</emphasis>
Holotype of 
<taxonomicName id="4C3B4D22310BFFBBFE0BB691D7BEF817" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[449,648,1954,1976]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
<emphasis id="B94FEAB3310BFFBBFE0BB691D7BEF817" box="[449,648,1954,1976]" italics="true" pageId="33" pageNumber="484">Fejervarya neilcoxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C8310BFFBBFD45B691D7D6F817" box="[655,736,1955,1976]" pageId="33" pageNumber="484" rank="species">
<emphasis id="B94FEAB3310BFFBBFD45B691D7D6F817" bold="true" box="[655,736,1955,1976]" pageId="33" pageNumber="484">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZSI/WGRC/V/A/951): 
<emphasis id="B94FEAB3310BFFBBFC15B690D6C1F818" bold="true" box="[991,1015,1954,1975]" pageId="33" pageNumber="484">A.</emphasis>
dorsolateral view, in life; 
<emphasis id="B94FEAB3310BFFBBFAC9B691D02CF817" bold="true" box="[1283,1306,1955,1976]" pageId="33" pageNumber="484">B.</emphasis>
dorsal view, in preservation; 
<emphasis id="B94FEAB3310BFFBBFE8DB6F3D469F878" bold="true" box="[327,351,1985,2007]" pageId="33" pageNumber="484">C.</emphasis>
ventral view, in preservation; 
<emphasis id="B94FEAB3310BFFBBFD7AB6F3D7FEF879" bold="true" box="[688,712,1985,2006]" pageId="33" pageNumber="484">D.</emphasis>
lateral view of head, in preservation; 
<emphasis id="B94FEAB3310BFFBBFBBAB6F3D1BEF879" bold="true" box="[1136,1160,1985,2006]" pageId="33" pageNumber="484">E.</emphasis>
ventral view of hand, in preservation; 
<emphasis id="B94FEAB3310BFFBBFEEBB6D2D405F85A" bold="true" box="[289,307,2016,2037]" pageId="33" pageNumber="484">F.</emphasis>
ventral view of foot, in preservation; 
<emphasis id="B94FEAB3310BFFBBFD7EB6D2D7FFF859" bold="true" box="[692,713,2016,2038]" pageId="33" pageNumber="484">G.</emphasis>
schematic illustration of foot webbing.
</paragraph>
</caption>
<paragraph id="8B8436A13108FFB8FF0DB0A4D1D9FD90" blockId="34.[151,1437,182,1260]" pageId="34" pageNumber="485">
Skin of snout shagreened to prominently granular, upper eyelids prominently tuberculate, anterior and posterior parts of back, and upper and lower parts of flank shagreened to prominently granular with spinular projections; interrupted inverse V-shaped ridge on center of dorsum; dorsal surfaces of forelimb, thigh and shank shagreened with scattered spinular projections. Ventral surface of throat, chest, belly and limbs shagreened, anterior part of thigh sparsely granular; fejervaryan line present on both sides of the belly (
<figureCitation id="13002A243108FFB8FB44B314D1D7FD90" box="[1166,1249,550,575]" captionStart="FIGURE 12" captionStartId="33.[151,250,1954,1976]" captionTargetBox="[151,1436,193,1933]" captionTargetId="figure@33.[151,1437,193,1933]" captionTargetPageId="33" captionText="FIGURE 12. Holotype of Fejervarya neilcoxi sp. nov. (ZSI / WGRC / V / A / 951): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829857/files/figure.png" pageId="34" pageNumber="485">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8B8436A13108FFB8FF0DB379D6F5FC2C" blockId="34.[151,1437,182,1260]" pageId="34" pageNumber="485">
<emphasis id="B94FEAB33108FFB8FF0DB379D486FDCB" bold="true" box="[199,432,587,612]" pageId="34" pageNumber="485">
Colour of 
<typeStatus id="548088033108FFB8FE88B379D49DFDCB" box="[322,427,587,612]" pageId="34" pageNumber="485" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="B94FEAB33108FFB8FE72B37ED735FDCB" box="[440,515,587,612]" italics="true" pageId="34" pageNumber="485">In life.</emphasis>
Snout, upper eyelids and anterior and posterior parts of back greyish-brown with prominent blackish-brown blotches (
<figureCitation id="13002A243108FFB8FDFEB35CD7B0FD28" box="[564,646,622,647]" captionStart="FIGURE 12" captionStartId="33.[151,250,1954,1976]" captionTargetBox="[151,1436,193,1933]" captionTargetId="figure@33.[151,1437,193,1933]" captionTargetPageId="33" captionText="FIGURE 12. Holotype of Fejervarya neilcoxi sp. nov. (ZSI / WGRC / V / A / 951): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829857/files/figure.png" pageId="34" pageNumber="485">Fig. 12</figureCitation>
A), upper and lower lip with faint alternate brown and light grey cross bands; tympanum light grey; flanks light grey with scattered dark grey mottling; forelimbs and hind limbs light brown with dark brown transverse bands; groin off-white with faint reticulations; anterior part of thigh light grey with greyish-brown reticulations; webbing light brown. Ventral surface of throat light flesh red with two lateral black calling patches; belly white; forearm and foreleg light flesh red. 
<emphasis id="B94FEAB33108FFB8FC04B232D14AFCB8" box="[974,1148,768,791]" italics="true" pageId="34" pageNumber="485">In preservation</emphasis>
. Dorsum dark grey with blackish-grey blotches, forelimbs and hind limbs light greyish-brown dark greyish-brown transverse bands, posterior part of thigh greyish-brown with faint dark brown reticulations. Ventral surface of throat light grey with two lateral black calling patches on either side; belly off-white (
<figureCitation id="13002A243108FFB8FCA8B259D680FC2C" box="[866,950,875,900]" captionStart="FIGURE 12" captionStartId="33.[151,250,1954,1976]" captionTargetBox="[151,1436,193,1933]" captionTargetId="figure@33.[151,1437,193,1933]" captionTargetPageId="33" captionText="FIGURE 12. Holotype of Fejervarya neilcoxi sp. nov. (ZSI / WGRC / V / A / 951): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829857/files/figure.png" pageId="34" pageNumber="485">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8B8436A13108FFB8FF0DB2BCD12FFC40" blockId="34.[151,1437,182,1260]" pageId="34" pageNumber="485">
<emphasis id="B94FEAB33108FFB8FF0DB2BCD47CFC08" bold="true" box="[199,330,910,935]" pageId="34" pageNumber="485">Variations.</emphasis>
Morphometric data from three adult males and two adult females, including the holotype, is given in 
<tableCitation id="C6B9031A3108FFB8FF7EB281D43DFC64" box="[180,267,947,972]" captionStart="TABLE 7" captionStartId="18.[119,204,562,584]" captionText="TABLE 7. Morphometric measurements for five species in the Fejervarya rufescens group. All measurements are in millimeters (mm). The status of each specimen is given the museum number: Holotype (HT), Paratype (PT), Neotype (NT), Referred specimen (RS)." pageId="34" pageNumber="485">Table 7</tableCitation>
. 
<emphasis id="B94FEAB33108FFB8FEDCB281D721FC64" box="[278,535,947,972]" italics="true" pageId="34" pageNumber="485">Colour in preservation</emphasis>
. ZSI/WGRC/V/A/953: dorsum light grey with dark grey blotches; ZSI/WGRC/ V/A/954 and ZSI/WGRC/V/A/955: ventral surface of throat with grey speckles.
</paragraph>
<paragraph id="8B8436A13108FFB8FF0DB2C9D66DFB98" blockId="34.[151,1437,182,1260]" pageId="34" pageNumber="485">
<emphasis id="B94FEAB33108FFB8FF0DB2C9D711FBBB" bold="true" box="[199,551,1019,1044]" pageId="34" pageNumber="485">Secondary sexual characters.</emphasis>
<emphasis id="B94FEAB33108FFB8FDFAB2C9D75FFBBB" box="[560,617,1019,1044]" italics="true" pageId="34" pageNumber="485">Male</emphasis>
: calling patches on either side of the throat. 
<emphasis id="B94FEAB33108FFB8FBBAB2C9D1F2FBBB" box="[1136,1220,1019,1044]" italics="true" pageId="34" pageNumber="485">Female</emphasis>
(ZSI/WGRC/V/A/ 955): pigmented eggs present (diameter 1.5 ± 
<quantity id="4CC39B443108FFB8FD51B52DD7C5FB98" box="[667,755,1055,1079]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="485" unit="mm" value="0.3">0.3 mm</quantity>
, 
<emphasis id="B94FEAB33108FFB8FD34B512D627FB98" box="[766,785,1056,1079]" italics="true" pageId="34" pageNumber="485">N</emphasis>
= 15).
</paragraph>
</subSubSection>
<subSubSection id="C321652A3108FFB8FF0DB571D195FB43" pageId="34" pageNumber="485" type="distribution">
<paragraph id="8B8436A13108FFB8FF0DB571D195FB43" blockId="34.[151,1437,182,1260]" pageId="34" pageNumber="485">
<emphasis id="B94FEAB33108FFB8FF0DB571D779FBF3" bold="true" box="[199,591,1091,1116]" pageId="34" pageNumber="485">Distribution and natural history.</emphasis>
<taxonomicName id="4C3B4D223108FFB8FD9CB576D605FBF3" authority="Garg &amp; Biju, 2017" authorityName="Garg &amp; Biju" authorityYear="2017" box="[598,819,1091,1116]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="34" pageNumber="485" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
<emphasis id="B94FEAB33108FFB8FD9CB576D605FBF3" box="[598,819,1091,1116]" italics="true" pageId="34" pageNumber="485">Fejervarya neilcoxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57C83108FFB8FCF1B571D6A3FBF3" box="[827,917,1091,1116]" pageId="34" pageNumber="485" rank="species">
<emphasis id="B94FEAB33108FFB8FCF1B571D6A3FBF3" bold="true" box="[827,917,1091,1116]" pageId="34" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
is currently known only from its 
<typeStatus id="548088033108FFB8FADAB576D076FBF3" box="[1296,1344,1092,1116]" pageId="34" pageNumber="485">type</typeStatus>
locality Parambikulam, south of Palghat gap in the 
<collectingRegion id="49FFF8433108FFB8FD42B55AD7D5FBD0" box="[648,739,1128,1151]" country="Sri Lanka" name="Western" pageId="34" pageNumber="485">Western</collectingRegion>
Ghats state of 
<collectingRegion id="49FFF8433108FFB8FC59B554D6E8FBD0" box="[915,990,1126,1151]" country="India" name="Kerala" pageId="34" pageNumber="485">Kerala</collectingRegion>
(
<figureCitation id="13002A243108FFB8FC25B554D103FBD0" box="[1007,1077,1126,1151]" captionStart="FIGURE 3" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[151,1436,197,1739]" captionTargetId="figure@6.[151,1436,193,1744]" captionTargetPageId="6" captionText="FIGURE 3. Geographical distribution of five species in Fejervarya rufescens group of the Western Ghats, Peninsular India. The Western Ghats biodiversity hotspot region is shown in orange colour. Locality details are referenced in Table 3." httpUri="https://zenodo.org/record/829839/files/figure.png" pageId="34" pageNumber="485">Fig. 3</figureCitation>
). The 
<typeStatus id="548088033108FFB8FB4BB55AD187FB2F" box="[1153,1201,1128,1152]" pageId="34" pageNumber="485">type</typeStatus>
series was observed near roadside vegetation with temporary pools of water. The specific site was close to a large moss-covered rock cutting surrounded with primary forest. Calling males were found close to water puddles and they stopped calling with any slight movement around them. Collections were made between 20:00–21:00 hours.
</paragraph>
</subSubSection>
</treatment>
</document>