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<mods:title id="E7620EBA5450E99B82B841CCD0C1330E">A new species of the genus Lucasioides Kwon, 1993 (Isopoda, Oniscidea, Agnaridae) from Siberia, Russia</mods:title>
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<mods:namePart id="B6D3A772613BA180D30AF2032B213B51">Nefediev, Pavel S.</mods:namePart>
<mods:affiliation id="6F3EC6DB47D9256A4EBA50ADEB657B58">Altai State University, Prospekt Lenina 61, 656049 Barnaul, Russia. &amp; Biological Institute, Tomsk State University, Prospekt Lenina, 36, 634050 Tomsk, Russia &amp; Tigirek State Nature Reserve, Nikitina Street 111, 656049 Barnaul, Russia</mods:affiliation>
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<mods:affiliation id="9B46B86A5FB2D9E583CB3981928CDFBD">I. D. Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, Borok, 109, 152742 Yaroslavl Region, Russia. turba 13 @ mail. ru; http: // orcid. org / 0000 - 0001 - 9441 - 2791 &amp; Cherepovets State University, Prospekt Lunacharskogo 5, 162600 Cherepovets, Vologda Region, Russia</mods:affiliation>
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<emphasis id="B94FEAA33852FFC7FF4FFC4EFE4B8C98" bold="true" box="[151,397,973,1000]" italics="true" pageId="2" pageNumber="142">Lucasioides altaicus</emphasis>
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<figureCitation id="13002A343852FFC7FF4FFC71FF3F8B7C" box="[151,249,1010,1036]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1297,1321]" captionStartId-1="4.[151,250,1260,1284]" captionStartId-2="5.[151,250,1545,1569]" captionStartId-3="6.[151,250,1020,1044]" captionStartId-4="7.[151,250,1208,1232]" captionStartId-5="7.[151,250,1970,1994]" captionStartId-6="8.[151,250,1204,1228]" captionTargetBox-0="[170,1416,184,1266]" captionTargetBox-1="[160,1423,212,1225]" captionTargetBox-2="[157,1430,243,1513]" captionTargetBox-3="[157,1432,185,991]" captionTargetBox-4="[298,1288,185,1177]" captionTargetBox-5="[288,1300,1331,1942]" captionTargetBox-6="[297,1289,182,1176]" captionTargetId-0="figure-26@3.[164,1423,180,1272]" captionTargetId-1="figure-30@4.[151,1436,200,1236]" captionTargetId-2="figure-66@5.[151,1436,236,1521]" captionTargetId-3="figure-17@6.[151,1437,180,995]" captionTargetId-4="figure-26@7.[294,1293,180,1183]" captionTargetId-5="figure-49@7.[283,1304,1328,1946]" captionTargetId-6="figure-17@8.[293,1294,180,1179]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." captionText-1="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." captionText-2="FIGURE 3. Lucasioides altaicus sp. nov., female paratype: A—pereionites 6, 7, pleon and telson; B—nodulis lateralis on pereonite 7; C—pereionites 1–3; D—nodulis lateralis on pereonite 2." captionText-3="FIGURE 4. Lucasioides altaicus sp. nov., female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped." captionText-4="FIGURE 5. Lucasioides altaicus sp. nov., male paratype: A, B—pereopod 6; C, D—pereopod 7." captionText-5="FIGURE 6. Lucasioides altaicus sp. nov., male paratype: A—exopod of pleopod 1; B—endopod of pleopod 2; C—exopod of pleopod 2; D—endopod of pleopod 2; E—exopod of pleopod 3; F—exopod of pleopod 4; G—exopod of pleopod 5." captionText-6="FIGURE 7. Lucasioides altaicus sp. nov., male paratype: A—exopod and endopod of pleopod 2; B—exopod of pleopod 1; C, D—endopod of pleopod 2." figureDoi-0="http://doi.org/10.5281/zenodo.4422840" figureDoi-1="http://doi.org/10.5281/zenodo.4422842" figureDoi-2="http://doi.org/10.5281/zenodo.4422844" figureDoi-3="http://doi.org/10.5281/zenodo.4422846" figureDoi-4="http://doi.org/10.5281/zenodo.4422848" figureDoi-5="http://doi.org/10.5281/zenodo.4422850" figureDoi-6="http://doi.org/10.5281/zenodo.4422852" httpUri-0="https://zenodo.org/record/4422840/files/figure.png" httpUri-1="https://zenodo.org/record/4422842/files/figure.png" httpUri-2="https://zenodo.org/record/4422844/files/figure.png" httpUri-3="https://zenodo.org/record/4422846/files/figure.png" httpUri-4="https://zenodo.org/record/4422848/files/figure.png" httpUri-5="https://zenodo.org/record/4422850/files/figure.png" httpUri-6="https://zenodo.org/record/4422852/files/figure.png" pageId="2" pageNumber="142">Figs 1–7</figureCitation>
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<emphasis id="B94FEAA33852FFC7FF4FFBB5FEF48B3E" box="[151,306,1078,1102]" italics="true" pageId="2" pageNumber="142">Mongoloniscus</emphasis>
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sp. (pro parte)— 
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<bibRefCitation id="EFAA4B403852FFC7FE39FBB5FD2C8B3E" author="Gongalsky, K. B. &amp; Nefediev, P. S. &amp; Turbanov, I. S. &amp; Slatina, V. Yu. &amp; Lebedev, Yu. M." box="[481,746,1078,1102]" pageId="2" pageNumber="142" pagination="26 - 27" refId="ref4633" refString="Gongalsky, K. B., Nefediev, P. S., Turbanov, I. S., Slatina, V. Yu. &amp; Lebedev, Yu. M. (2017) Terrestrial isopods (Isopoda, Oniscidea, Agnaridae) from the northernmost limit of isopod distribution in southern Siberia, Russia. In: Abstract Book of 10 th International Symposium on Terrestrial Isopod Biology. Hungarian Biological Society, Budapest, 26 - 27." type="book chapter" year="2017">
Gongalsky 
<emphasis id="B94FEAA33852FFC7FD8DFBB4FD478B3E" box="[597,641,1078,1102]" italics="true" pageId="2" pageNumber="142">et al</emphasis>
. 2017: 26
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.
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, ³ (
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, Mc-1420), 
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, Charyshskoye Dis-trict, near Komendantka, 
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University (ASU) Field Station “Goluboi Utios”, 
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, 
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, 
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a.s.l., N slope, plot 2, 
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, 
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<emphasis id="B94FEAA33852FFC7FD40FB47FC978BAE" box="[664,849,1220,1246]" italics="true" pageId="2" pageNumber="142">Populus tremula</emphasis>
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, 
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<emphasis id="B94FEAA33852FFC7FCB8FB46FBC98BAE" box="[864,1039,1220,1246]" italics="true" pageId="2" pageNumber="142">Pinus sylvestris</emphasis>
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, 
<taxonomicName id="4C3B4D323852FFC7FBC6FB46FAEE8BAE" box="[1054,1320,1220,1247]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="chamaedrifolia">
<emphasis id="B94FEAA33852FFC7FBC6FB46FAEE8BAE" box="[1054,1320,1220,1247]" italics="true" pageId="2" pageNumber="142">Spiraea chamaedrifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FAEFFB46FECB8A72" class="Magnoliopsida" family="Saxifragaceae" genus="Bergenia" kingdom="Animalia" order="Saxifragales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="crassifolia">
<emphasis id="B94FEAA33852FFC7FAEFFB46FECB8A72" italics="true" pageId="2" pageNumber="142">Bergenia crassifolia</emphasis>
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, hand sampling, 
<date id="FF8510713852FFC7FE16FB6AFDA78A72" box="[462,609,1256,1283]" pageId="2" pageNumber="142" value="2016-07-13">13 July 2016</date>
, leg. 
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.Yu. Slatina, Kh.Kh. Nedoev, S. 
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. Niyazov.
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<typeStatus id="548088133852FFC7FF1FFA8EFEFE8A57" box="[199,312,1293,1319]" pageId="2" pageNumber="142" type="paratype">Paratypes</typeStatus>
: 1 ³ (
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, Mc-1421); 1 ³, 
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(KG), 
<collectingCountry id="F32C76213852FFC7FCFEFA8FFCB58A56" box="[806,883,1292,1318]" name="Russia" pageId="2" pageNumber="142">Russia</collectingCountry>
, southwestern Siberia, 
<collectingRegion id="49FFF8533852FFC7FBA3FA8FFB378A56" box="[1147,1265,1292,1318]" country="Russia" name="Altay" pageId="2" pageNumber="142">Altai Krai</collectingRegion>
, Charyshskoye District, near Komendantka, ASU Field Station “Goluboi Utios”, 
<geoCoordinate id="EE0F50763852FFC7FCB4FAB1FBC18A3D" box="[876,1031,1330,1357]" degrees="51" direction="north" minutes="21" orientation="latitude" pageId="2" pageNumber="142" precision="1" seconds="44.3" value="51.362305">51°21ʹ44.3ʺN</geoCoordinate>
, 
<geoCoordinate id="EE0F50763852FFC7FBCAFAB1FB6C8A3D" box="[1042,1194,1330,1357]" degrees="83" direction="east" minutes="37" orientation="longitude" pageId="2" pageNumber="142" precision="1" seconds="42.6" value="83.6285">83°37ʹ42.6ʺE</geoCoordinate>
, 
<quantity id="4CC39B543852FFC7FB6CFAB3FB3C8A3B" box="[1204,1274,1328,1355]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="142" unit="m" value="620.0">620 m</quantity>
a.s.l., N slope, plot 1, 
<taxonomicName id="4C3B4D323852FFC7FF3DFAD6FE988A1E" authorityName="A.W.Roth" authorityYear="1788" box="[229,350,1364,1390]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="B94FEAA33852FFC7FF3DFAD6FE988A1E" box="[229,350,1364,1390]" italics="true" pageId="2" pageNumber="142">B. pendula</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FEB3FAD6FE1A8A1E" box="[363,476,1364,1390]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="tremula">
<emphasis id="B94FEAA33852FFC7FEB3FAD6FE1A8A1E" box="[363,476,1364,1390]" italics="true" pageId="2" pageNumber="142">P. tremula</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FE32FAD6FDAB8A1E" box="[490,621,1364,1390]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="B94FEAA33852FFC7FE32FAD6FDAB8A1E" box="[490,621,1364,1390]" italics="true" pageId="2" pageNumber="142">P. sylvestris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FDA3FAD6FC878A1E" box="[635,833,1364,1391]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="chamaedrifolia">
<emphasis id="B94FEAA33852FFC7FDA3FAD6FC878A1E" box="[635,833,1364,1391]" italics="true" pageId="2" pageNumber="142">S. chamaedrifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FC97FAD6FC228A1E" box="[847,996,1364,1390]" class="Magnoliopsida" family="Saxifragaceae" genus="Bergenia" kingdom="Animalia" order="Saxifragales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="crassifolia">
<emphasis id="B94FEAA33852FFC7FC97FAD6FC228A1E" box="[847,996,1364,1390]" italics="true" pageId="2" pageNumber="142">B. crassifolia</emphasis>
</taxonomicName>
, soil sampling, sample 4 (
<quantity id="4CC39B543852FFC7FADFFAD6FAAB8A1F" box="[1287,1389,1365,1391]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="142" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
), 
<date id="FF8510713852FFC7FA58FAD6FEC18AE3" pageId="2" pageNumber="142" value="2018-07-12">12 July 2018</date>
, leg. 
<collectionCode id="ED2AAE743852FFC7FEE7FAFAFE958AE3" box="[319,339,1401,1427]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="142" type="Museum">V</collectionCode>
.Yu. Slatina, Kh.Kh. Nedoev; 3 ³³, 
<specimenCount id="9D3DFD383852FFC7FD3CFAFBFCE48AE3" box="[740,802,1400,1427]" pageId="2" pageNumber="142" type="female">5 ♀♀</specimenCount>
(
<collectionCode id="ED2AAE743852FFC7FCE8FAFAFC4A8AE3" box="[816,908,1401,1427]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="142" type="Museum">ZMMU</collectionCode>
, Mc-1422); 
<specimenCount id="9D3DFD383852FFC7FBCBFAFAFBFA8AE3" box="[1043,1084,1401,1427]" pageId="2" pageNumber="142" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED2AAE743852FFC7FB91FAFAFB638AE3" box="[1097,1189,1401,1427]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="142" type="Museum">ZMMU</collectionCode>
, Mc-1424), 1 ³ (KG), 4 ³³ (ASU), location, date, and collectors same as 
<typeStatus id="548088133852FFC7FD3CFA1FFC8C8AC6" box="[740,842,1436,1462]" pageId="2" pageNumber="142" type="holotype">holotype</typeStatus>
; 1 ³, 
<specimenCount id="9D3DFD383852FFC7FC57FA1EFC7F8AC7" box="[911,953,1437,1463]" pageId="2" pageNumber="142" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED2AAE743852FFC7FC11FA1EFBE38AC7" box="[969,1061,1437,1463]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="142" type="Museum">ZMMU</collectionCode>
, Mc-1423), 
<collectingCountry id="F32C76213852FFC7FB77FA1FFB3A8AC6" box="[1199,1276,1436,1462]" name="Russia" pageId="2" pageNumber="142">Russia</collectingCountry>
, southwestern Siberia, 
<collectingRegion id="49FFF8533852FFC7FF2BFA43FEAE8AAA" box="[243,360,1472,1498]" country="Russia" name="Altay" pageId="2" pageNumber="142">Altai Krai</collectingRegion>
, Charyshskoye District, near Komendantka, ASU Field Station “Goluboi Utios”, 
<geoCoordinate id="EE0F50763852FFC7FB24FA41FA5E8AAD" box="[1276,1432,1474,1501]" degrees="51" direction="north" minutes="21" orientation="latitude" pageId="2" pageNumber="142" precision="1" seconds="20.3" value="51.355637">51°21ʹ20.3ʺN</geoCoordinate>
, 
<geoCoordinate id="EE0F50763852FFC7FF4FFA65FEE98971" box="[151,303,1510,1537]" degrees="83" direction="east" minutes="37" orientation="longitude" pageId="2" pageNumber="142" precision="1" seconds="37.0" value="83.626945">83°37ʹ37.0ʺE</geoCoordinate>
, 
<taxonomicName id="4C3B4D323852FFC7FEE6FA66FDCE8A8E" box="[318,520,1508,1535]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="chamaedrifolia">
<emphasis id="B94FEAA33852FFC7FEE6FA66FDCE8A8E" box="[318,520,1508,1535]" italics="true" pageId="2" pageNumber="142">S. chamaedrifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FDC1FA66FCDA8A8E" authorityName="Lamarck" authorityYear="1785" box="[537,796,1508,1535]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B94FEAA33852FFC7FDC1FA66FCDA8A8E" box="[537,796,1508,1535]" italics="true" pageId="2" pageNumber="142">Caragana arborescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323852FFC7FCF5FA66FC338A8E" authorityName="Linnaeus" authorityYear="1753" box="[813,1013,1509,1534]" class="Magnoliopsida" family="Caprifoliaceae" genus="Lonicera" kingdom="Plantae" order="Dipsacales" pageId="2" pageNumber="142" phylum="Tracheophyta" rank="species" species="tatarica">
<emphasis id="B94FEAA33852FFC7FCF5FA66FC338A8E" box="[813,1013,1509,1534]" italics="true" pageId="2" pageNumber="142">Lonicera tatarica</emphasis>
</taxonomicName>
on southern slope, 
<quantity id="4CC39B543852FFC7FB03FA66FAE38A8F" box="[1243,1317,1509,1535]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="142" unit="m" value="480.0">480 m</quantity>
a.s.l., soil sampling, sample 1 (
<quantity id="4CC39B543852FFC7FEA7F98AFE208953" box="[383,486,1545,1571]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="142" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
), 
<date id="FF8510713852FFC7FE21F98BFD0E8953" box="[505,712,1544,1571]" pageId="2" pageNumber="142" value="2018-09-06">6 September 2018</date>
, leg. Kh.Kh. Nedoev.
</paragraph>
</subSubSection>
<subSubSection id="C321653A3852FFC7FF1FF9AFFE36891A" pageId="2" pageNumber="142" type="diagnosis">
<paragraph id="8B8436B13852FFC7FF1FF9AFFE36891A" blockId="2.[151,1437,1148,2039]" pageId="2" pageNumber="142">
<emphasis id="B94FEAA33852FFC7FF1FF9AFFE848936" bold="true" box="[199,322,1580,1606]" pageId="2" pageNumber="142">Diagnosis.</emphasis>
Male exopod of pleopod 1 with concave tip and lower lobe is longer than the upper one; telson is elongated and shows sharp tip.
</paragraph>
</subSubSection>
<subSubSection id="C321653A3852FFC6FF1FF9F7FDED8889" lastPageId="3" lastPageNumber="143" pageId="2" pageNumber="142" type="description">
<paragraph id="8B8436B13852FFC7FF1FF9F7FA858817" blockId="2.[151,1437,1148,2039]" pageId="2" pageNumber="142">
<emphasis id="B94FEAA33852FFC7FF1FF9F7FE9F89FE" bold="true" box="[199,345,1652,1678]" pageId="2" pageNumber="142">Description.</emphasis>
Maximum body length: male 8.0 mm (
<typeStatus id="548088133852FFC7FCD6F9F7FCB489FE" box="[782,882,1652,1678]" pageId="2" pageNumber="142" type="holotype">holotype</typeStatus>
<quantity id="4CC39B543852FFC7FCA0F9F6FC1489FF" box="[888,978,1653,1679]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="142" unit="mm" value="7.2">7.2 mm</quantity>
), female 
<quantity id="4CC39B543852FFC7FBEFF9F7FB5689FF" box="[1079,1168,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="142" unit="mm" value="6.7">6.7 mm</quantity>
. 
<emphasis id="B94FEAA33852FFC7FB42F9F7FB1289FE" box="[1178,1236,1652,1678]" italics="true" pageId="2" pageNumber="142">Body</emphasis>
dark grey-brown, with two rows of light grey-brown spots in medial part of coxal plates of pereonal segments 2–7 (
<figureCitation id="13002A343852FFC7FB37F91BFA8C89C3" box="[1263,1354,1688,1715]" captionStart="FIGURE 1" captionStartId="3.[151,250,1297,1321]" captionTargetBox="[170,1416,184,1266]" captionTargetId="figure-26@3.[164,1423,180,1272]" captionTargetPageId="3" captionText="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." figureDoi="http://doi.org/10.5281/zenodo.4422840" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="2" pageNumber="142">Fig. 1A</figureCitation>
). Body relatively elongated, pleon forming no continuous margin with pereon (
<figureCitation id="13002A343852FFC7FC6EF93FFBC889A7" box="[950,1038,1724,1751]" captionStart="FIGURE 1" captionStartId="3.[151,250,1297,1321]" captionTargetBox="[170,1416,184,1266]" captionTargetId="figure-26@3.[164,1423,180,1272]" captionTargetPageId="3" captionText="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." figureDoi="http://doi.org/10.5281/zenodo.4422840" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="2" pageNumber="142">Fig. 1A</figureCitation>
). Dorsal surface of tergites smooth. Posterior edges of coxal plates of pereonites 1–3 straight, those of pereonites 4–7 curved (
<figureCitation id="13002A343852FFC7FB48F963FB2D898B" box="[1168,1259,1760,1787]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="2" pageNumber="142">Fig. 2A</figureCitation>
). 
<emphasis id="B94FEAA33852FFC7FB27F963FAFA898A" box="[1279,1340,1760,1786]" italics="true" pageId="2" pageNumber="142">Head</emphasis>
covered with scattered, sharp, triangular, dorsal setae (
<figureCitation id="13002A343852FFC7FD4DF887FD2A886F" box="[661,748,1796,1823]" captionStart="FIGURE 1" captionStartId="3.[151,250,1297,1321]" captionTargetBox="[170,1416,184,1266]" captionTargetId="figure-26@3.[164,1423,180,1272]" captionTargetPageId="3" captionText="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." figureDoi="http://doi.org/10.5281/zenodo.4422840" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="2" pageNumber="142">Fig. 1B</figureCitation>
). 
<emphasis id="B94FEAA33852FFC7FCD9F887FC73886E" box="[769,949,1796,1822]" italics="true" pageId="2" pageNumber="142">Noduli laterales</emphasis>
on pereonites 1–4 located at approximately same distance to edge of coxal plates (
<figureCitation id="13002A343852FFC7FD89F8ABFD698833" box="[593,687,1832,1859]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="2" pageNumber="142">Figs 2A</figureCitation>
, 
<figureCitation id="13002A343852FFC7FD65F8AAFD0B8833" box="[701,717,1833,1859]" captionStart="FIGURE 3" captionStartId="5.[151,250,1545,1569]" captionTargetBox="[157,1430,243,1513]" captionTargetId="figure-66@5.[151,1436,236,1521]" captionTargetPageId="5" captionText="FIGURE 3. Lucasioides altaicus sp. nov., female paratype: A—pereionites 6, 7, pleon and telson; B—nodulis lateralis on pereonite 7; C—pereionites 1–3; D—nodulis lateralis on pereonite 2." figureDoi="http://doi.org/10.5281/zenodo.4422844" httpUri="https://zenodo.org/record/4422844/files/figure.png" pageId="2" pageNumber="142">3</figureCitation>
). Cephalic lobes poorly developed. Distal edge of median lobe rounded (
<figureCitation id="13002A343852FFC7FEDAF8CFFE9B8817" box="[258,349,1868,1895]" captionStart="FIGURE 1" captionStartId="3.[151,250,1297,1321]" captionTargetBox="[170,1416,184,1266]" captionTargetId="figure-26@3.[164,1423,180,1272]" captionTargetPageId="3" captionText="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." figureDoi="http://doi.org/10.5281/zenodo.4422840" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="2" pageNumber="142">Figs 1B</figureCitation>
, 
<figureCitation id="13002A343852FFC7FEB1F8CEFE4B8817" box="[361,397,1869,1895]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="2" pageNumber="142">2B</figureCitation>
). Telson with an elongated distal part, distal corner forming a triangle (
<figureCitation id="13002A343852FFC7FB70F8CFFAC28817" box="[1192,1284,1868,1895]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="2" pageNumber="142">Figs 2C</figureCitation>
, 
<figureCitation id="13002A343852FFC7FAC8F8CEFAF38817" box="[1296,1333,1869,1895]" captionStart="FIGURE 3" captionStartId="5.[151,250,1545,1569]" captionTargetBox="[157,1430,243,1513]" captionTargetId="figure-66@5.[151,1436,236,1521]" captionTargetPageId="5" captionText="FIGURE 3. Lucasioides altaicus sp. nov., female paratype: A—pereionites 6, 7, pleon and telson; B—nodulis lateralis on pereonite 7; C—pereionites 1–3; D—nodulis lateralis on pereonite 2." figureDoi="http://doi.org/10.5281/zenodo.4422844" httpUri="https://zenodo.org/record/4422844/files/figure.png" pageId="2" pageNumber="142">3A</figureCitation>
).
</paragraph>
<paragraph id="8B8436B13852FFC7FF1FF8F3FD848887" blockId="2.[151,1437,1148,2039]" pageId="2" pageNumber="142">
<emphasis id="B94FEAA33852FFC7FF1FF8F3FEFC88FA" box="[199,314,1904,1930]" italics="true" pageId="2" pageNumber="142">Antennula</emphasis>
with three articles (
<figureCitation id="13002A343852FFC7FDCEF8F3FDB688FB" box="[534,624,1904,1931]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="2" pageNumber="142">Fig. 2D</figureCitation>
); first article wide and long; second article 0.5 times as long as first; third article narrow as almost as long as first article, bearing tuft of setae at apex. 
<emphasis id="B94FEAA33852FFC7FC26F816FB9D88DF" box="[1022,1115,1941,1967]" italics="true" pageId="2" pageNumber="142">Antenna</emphasis>
of medium length, reaching pereonite 3; flagellum with two articles, proximal article slightly shorter than distal one (
<figureCitation id="13002A343852FFC7FBA3F83BFB1088A3" box="[1147,1238,1976,2003]" captionStart="FIGURE 1" captionStartId="3.[151,250,1297,1321]" captionTargetBox="[170,1416,184,1266]" captionTargetId="figure-26@3.[164,1423,180,1272]" captionTargetPageId="3" captionText="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." figureDoi="http://doi.org/10.5281/zenodo.4422840" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="2" pageNumber="142">Figs 1C</figureCitation>
, 
<figureCitation id="13002A343852FFC7FB3BF83AFAC088A3" box="[1251,1286,1977,2003]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="2" pageNumber="142">2E</figureCitation>
). Aesthetascs absent from apical articles of antenna.
</paragraph>
<caption id="DF4466393853FFC6FF4FFA92FAEB8A59" ID-DOI="http://doi.org/10.5281/zenodo.4422840" ID-Zenodo-Dep="4422840" box="[151,1325,1297,1321]" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="3" pageNumber="143" startId="3.[151,250,1297,1321]" targetBox="[170,1416,184,1266]" targetPageId="3">
<paragraph id="8B8436B13853FFC6FF4FFA92FAEB8A59" blockId="3.[151,1325,1297,1321]" box="[151,1325,1297,1321]" pageId="3" pageNumber="143">
<emphasis id="B94FEAA33853FFC6FF4FFA92FED58A59" bold="true" box="[151,275,1297,1321]" pageId="3" pageNumber="143">FIGURE 1.</emphasis>
<taxonomicName id="4C3B4D323853FFC6FEC1FA92FE2F8A59" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[281,489,1297,1321]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33853FFC6FEC1FA92FE2F8A59" box="[281,489,1297,1321]" italics="true" pageId="3" pageNumber="143">Lucasioides altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33853FFC6FE37FA92FD868A59" bold="true" box="[495,576,1297,1321]" pageId="3" pageNumber="143">
<taxonomicNameLabel id="A27C57D83853FFC6FE37FA92FD868A59" box="[495,576,1297,1321]" pageId="3" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A—male paratype, dorsal view; female paratype: B—head; C—antenna.
</paragraph>
</caption>
<paragraph id="8B8436B13853FFC6FF1FFAF8FCAD89C5" blockId="3.[151,1437,1403,2042]" pageId="3" pageNumber="143">
<emphasis id="B94FEAA33853FFC6FF1FFAF8FEA28AE5" box="[199,356,1403,1429]" italics="true" pageId="3" pageNumber="143">Left mandible</emphasis>
(
<figureCitation id="13002A343853FFC6FEAEFAF8FE168AE5" box="[374,464,1403,1429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1020,1044]" captionTargetBox="[157,1432,185,991]" captionTargetId="figure-17@6.[151,1437,180,995]" captionTargetPageId="6" captionText="FIGURE 4. Lucasioides altaicus sp. nov., female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4422846" httpUri="https://zenodo.org/record/4422846/files/figure.png" pageId="3" pageNumber="143">Fig. 4A</figureCitation>
) with molar process carrying two teeth and 
<emphasis id="B94FEAA33853FFC6FC0AFAF8FBBD8AE5" box="[978,1147,1403,1429]" italics="true" pageId="3" pageNumber="143">lacinia mobilis</emphasis>
with straight edge. Lobe covered by setae with three plumose penicils followed by row of 18–20 penicils located basal to 
<emphasis id="B94FEAA33853FFC6FB27FA1CFA8B8AC9" box="[1279,1357,1439,1465]" italics="true" pageId="3" pageNumber="143">lacinia</emphasis>
. Right mandible smaller than left mandible, molar process with thee teeth and 
<emphasis id="B94FEAA33853FFC6FC60FA40FB998AAD" box="[952,1119,1475,1501]" italics="true" pageId="3" pageNumber="143">lacinia mobilis</emphasis>
with two teeth and hairy bilobate lobe with row of 18–20 penicils (
<figureCitation id="13002A343853FFC6FD8DFA64FD6B8971" box="[597,685,1511,1537]" captionStart="FIGURE 4" captionStartId="6.[151,250,1020,1044]" captionTargetBox="[157,1432,185,991]" captionTargetId="figure-17@6.[151,1437,180,995]" captionTargetPageId="6" captionText="FIGURE 4. Lucasioides altaicus sp. nov., female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4422846" httpUri="https://zenodo.org/record/4422846/files/figure.png" pageId="3" pageNumber="143">Fig. 4B</figureCitation>
). 
<emphasis id="B94FEAA33853FFC6FD19FA64FCE88971" box="[705,814,1511,1537]" italics="true" pageId="3" pageNumber="143">Maxillula</emphasis>
(
<figureCitation id="13002A343853FFC6FCE5FA64FC558971" box="[829,915,1511,1537]" captionStart="FIGURE 4" captionStartId="6.[151,250,1020,1044]" captionTargetBox="[157,1432,185,991]" captionTargetId="figure-17@6.[151,1437,180,995]" captionTargetPageId="6" captionText="FIGURE 4. Lucasioides altaicus sp. nov., female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4422846" httpUri="https://zenodo.org/record/4422846/files/figure.png" pageId="3" pageNumber="143">Fig. 4C</figureCitation>
) medial corner of inner endite with two strong hirsute setae.Apical edge of outer endite bearing nine spines divided into two groups; five medial spines slender and with split tips, and four lateral spines stouter and with simple tips. Tip of endite covered with brush of dense setae. 
<emphasis id="B94FEAA33853FFC6FF4FF9D0FF28891D" box="[151,238,1619,1645]" italics="true" pageId="3" pageNumber="143">Maxilla</emphasis>
with bilobate edge, medial half of apical edge of inner lobe with brush of short dense setae (
<figureCitation id="13002A343853FFC6FB2CF9D0FA89891D" box="[1268,1359,1619,1645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1020,1044]" captionTargetBox="[157,1432,185,991]" captionTargetId="figure-17@6.[151,1437,180,995]" captionTargetPageId="6" captionText="FIGURE 4. Lucasioides altaicus sp. nov., female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4422846" httpUri="https://zenodo.org/record/4422846/files/figure.png" pageId="3" pageNumber="143">Fig. 4D</figureCitation>
). Inner margin with a subapical tubercle. Maxilliped with outer corner of endite with two acute tips and large spine near inner corner (
<figureCitation id="13002A343853FFC6FEF7F918FE4089C5" box="[303,390,1691,1717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1020,1044]" captionTargetBox="[157,1432,185,991]" captionTargetId="figure-17@6.[151,1437,180,995]" captionTargetPageId="6" captionText="FIGURE 4. Lucasioides altaicus sp. nov., female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4422846" httpUri="https://zenodo.org/record/4422846/files/figure.png" pageId="3" pageNumber="143">Fig. 4E</figureCitation>
). Basal article of endopod with large spine.
</paragraph>
<paragraph id="8B8436B13853FFC6FF1FF93CFBAD89A9" blockId="3.[151,1437,1403,2042]" box="[199,1131,1727,1754]" pageId="3" pageNumber="143">
<emphasis id="B94FEAA33853FFC6FF1FF93CFEF889A9" box="[199,318,1727,1753]" italics="true" pageId="3" pageNumber="143">Pereopods</emphasis>
(
<figureCitation id="13002A343853FFC6FE95F93CFE5589A9" box="[333,403,1727,1753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="3" pageNumber="143">Figs 2</figureCitation>
F–H, 5). Ventral edge of male pereopod 7 carpus slightly curved.
</paragraph>
<paragraph id="8B8436B13853FFC6FF1FF960FE5A8851" blockId="3.[151,1437,1403,2042]" pageId="3" pageNumber="143">
<emphasis id="B94FEAA33853FFC6FF1FF960FEEE898D" box="[199,296,1763,1789]" italics="true" pageId="3" pageNumber="143">Uropods</emphasis>
(
<figureCitation id="13002A343853FFC6FEE0F960FE57898D" box="[312,401,1763,1789]" captionStart="FIGURE 1" captionStartId="3.[151,250,1297,1321]" captionTargetBox="[170,1416,184,1266]" captionTargetId="figure-26@3.[164,1423,180,1272]" captionTargetPageId="3" captionText="FIGURE 1. Lucasioides altaicus sp. nov., A—male paratype, dorsal view; female paratype: B—head; C—antenna." figureDoi="http://doi.org/10.5281/zenodo.4422840" httpUri="https://zenodo.org/record/4422840/files/figure.png" pageId="3" pageNumber="143">Fig. 1A</figureCitation>
, 
<figureCitation id="13002A343853FFC6FE45F960FE78898D" box="[413,446,1763,1789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="3" pageNumber="143">2C</figureCitation>
) colored as dorsal surface of body. Exopods elongated, with widest part at 1/3 proximal distance (
<figureCitation id="13002A343853FFC6FEDAF884FE9B8851" box="[258,349,1799,1825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1260,1284]" captionTargetBox="[160,1423,212,1225]" captionTargetId="figure-30@4.[151,1436,200,1236]" captionTargetPageId="4" captionText="FIGURE 2. Lucasioides altaicus sp. nov., female paratype:A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="3" pageNumber="143">Figs 2C</figureCitation>
, 
<figureCitation id="13002A343853FFC6FEB1F884FE498851" box="[361,399,1799,1825]" captionStart="FIGURE 3" captionStartId="5.[151,250,1545,1569]" captionTargetBox="[157,1430,243,1513]" captionTargetId="figure-66@5.[151,1436,236,1521]" captionTargetPageId="5" captionText="FIGURE 3. Lucasioides altaicus sp. nov., female paratype: A—pereionites 6, 7, pleon and telson; B—nodulis lateralis on pereonite 7; C—pereionites 1–3; D—nodulis lateralis on pereonite 2." figureDoi="http://doi.org/10.5281/zenodo.4422844" httpUri="https://zenodo.org/record/4422844/files/figure.png" pageId="3" pageNumber="143">3A</figureCitation>
).
</paragraph>
<paragraph id="8B8436B13853FFC6FF1FF8A9FDED8889" blockId="3.[151,1437,1403,2042]" pageId="3" pageNumber="143">
<emphasis id="B94FEAA33853FFC6FF1FF8A9FEC48835" bold="true" box="[199,258,1834,1861]" italics="true" pageId="3" pageNumber="143">Male</emphasis>
. Shape of genital papilla typical of the genus. All exopods at lateral margin with lungs. Exopod of 
<emphasis id="B94FEAA33853FFC6FA9AF8A8FF638819" italics="true" pageId="3" pageNumber="143">pleopod 1</emphasis>
(
<figureCitation id="13002A343853FFC6FF6CF8CCFED78819" box="[180,273,1871,1897]" captionStart="FIGURE 6" captionStartId="7.[151,250,1970,1994]" captionTargetBox="[288,1300,1331,1942]" captionTargetId="figure-49@7.[283,1304,1328,1946]" captionTargetPageId="7" captionText="FIGURE 6. Lucasioides altaicus sp. nov., male paratype: A—exopod of pleopod 1; B—endopod of pleopod 2; C—exopod of pleopod 2; D—endopod of pleopod 2; E—exopod of pleopod 3; F—exopod of pleopod 4; G—exopod of pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.4422850" httpUri="https://zenodo.org/record/4422850/files/figure.png" pageId="3" pageNumber="143">Figs 6A</figureCitation>
, 
<figureCitation id="13002A343853FFC6FEC4F8D3FEFB881A" box="[284,317,1872,1898]" captionStart="FIGURE 7" captionStartId="8.[151,250,1204,1228]" captionTargetBox="[297,1289,182,1176]" captionTargetId="figure-17@8.[293,1294,180,1179]" captionTargetPageId="8" captionText="FIGURE 7. Lucasioides altaicus sp. nov., male paratype: A—exopod and endopod of pleopod 2; B—exopod of pleopod 1; C, D—endopod of pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.4422852" httpUri="https://zenodo.org/record/4422852/files/figure.png" pageId="3" pageNumber="143">7B</figureCitation>
) with long hind lobe bearing deep hollow and 9–10 setae at apex; inner margin slightly concave and devoid of setae. Endopod of pleopod 1 with a dorsal furrow and row of spines (
<figureCitation id="13002A343853FFC6FBD7F8F0FBAD88FD" box="[1039,1131,1907,1933]" captionStart="FIGURE 6" captionStartId="7.[151,250,1970,1994]" captionTargetBox="[288,1300,1331,1942]" captionTargetId="figure-49@7.[283,1304,1328,1946]" captionTargetPageId="7" captionText="FIGURE 6. Lucasioides altaicus sp. nov., male paratype: A—exopod of pleopod 1; B—endopod of pleopod 2; C—exopod of pleopod 2; D—endopod of pleopod 2; E—exopod of pleopod 3; F—exopod of pleopod 4; G—exopod of pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.4422850" httpUri="https://zenodo.org/record/4422850/files/figure.png" pageId="3" pageNumber="143">Figs 6B</figureCitation>
, 
<figureCitation id="13002A343853FFC6FBAEF8F0FB7B88FE" box="[1142,1213,1907,1934]" captionStart="FIGURE 7" captionStartId="8.[151,250,1204,1228]" captionTargetBox="[297,1289,182,1176]" captionTargetId="figure-17@8.[293,1294,180,1179]" captionTargetPageId="8" captionText="FIGURE 7. Lucasioides altaicus sp. nov., male paratype: A—exopod and endopod of pleopod 2; B—exopod of pleopod 1; C, D—endopod of pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.4422852" httpUri="https://zenodo.org/record/4422852/files/figure.png" pageId="3" pageNumber="143">7C, D</figureCitation>
). 
<emphasis id="B94FEAA33853FFC6FB09F8F0FA8588FD" box="[1233,1347,1907,1933]" italics="true" pageId="3" pageNumber="143">Pleopod 2</emphasis>
exopod triangular, with concave outer margin bearing 12 setae (
<figureCitation id="13002A343853FFC6FCC8F814FCAA88C1" box="[784,876,1943,1969]" captionStart="FIGURE 6" captionStartId="7.[151,250,1970,1994]" captionTargetBox="[288,1300,1331,1942]" captionTargetId="figure-49@7.[283,1304,1328,1946]" captionTargetPageId="7" captionText="FIGURE 6. Lucasioides altaicus sp. nov., male paratype: A—exopod of pleopod 1; B—endopod of pleopod 2; C—exopod of pleopod 2; D—endopod of pleopod 2; E—exopod of pleopod 3; F—exopod of pleopod 4; G—exopod of pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.4422850" httpUri="https://zenodo.org/record/4422850/files/figure.png" pageId="3" pageNumber="143">Figs 6C</figureCitation>
, 
<figureCitation id="13002A343853FFC6FCA1F814FC5988C1" box="[889,927,1943,1969]" captionStart="FIGURE 7" captionStartId="8.[151,250,1204,1228]" captionTargetBox="[297,1289,182,1176]" captionTargetId="figure-17@8.[293,1294,180,1179]" captionTargetPageId="8" captionText="FIGURE 7. Lucasioides altaicus sp. nov., male paratype: A—exopod and endopod of pleopod 2; B—exopod of pleopod 1; C, D—endopod of pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.4422852" httpUri="https://zenodo.org/record/4422852/files/figure.png" pageId="3" pageNumber="143">7A</figureCitation>
); endopod much longer than exopod, narrow and parallel-sided (
<figureCitation id="13002A343853FFC6FEA8F838FE0888A5" box="[368,462,1979,2005]" captionStart="FIGURE 6" captionStartId="7.[151,250,1970,1994]" captionTargetBox="[288,1300,1331,1942]" captionTargetId="figure-49@7.[283,1304,1328,1946]" captionTargetPageId="7" captionText="FIGURE 6. Lucasioides altaicus sp. nov., male paratype: A—exopod of pleopod 1; B—endopod of pleopod 2; C—exopod of pleopod 2; D—endopod of pleopod 2; E—exopod of pleopod 3; F—exopod of pleopod 4; G—exopod of pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.4422850" httpUri="https://zenodo.org/record/4422850/files/figure.png" pageId="3" pageNumber="143">Figs 6D</figureCitation>
, 
<figureCitation id="13002A343853FFC6FE03F838FDC688A5" box="[475,512,1979,2005]" captionStart="FIGURE 7" captionStartId="8.[151,250,1204,1228]" captionTargetBox="[297,1289,182,1176]" captionTargetId="figure-17@8.[293,1294,180,1179]" captionTargetPageId="8" captionText="FIGURE 7. Lucasioides altaicus sp. nov., male paratype: A—exopod and endopod of pleopod 2; B—exopod of pleopod 1; C, D—endopod of pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.4422852" httpUri="https://zenodo.org/record/4422852/files/figure.png" pageId="3" pageNumber="143">7A</figureCitation>
). 
<emphasis id="B94FEAA33853FFC6FDCDF838FD7688A5" box="[533,688,1979,2005]" italics="true" pageId="3" pageNumber="143">Pleopods 3–4</emphasis>
exopods (
<figureCitation id="13002A343853FFC6FCFDF838FCAE88A6" box="[805,872,1979,2006]" captionStart="FIGURE 6" captionStartId="7.[151,250,1970,1994]" captionTargetBox="[288,1300,1331,1942]" captionTargetId="figure-49@7.[283,1304,1328,1946]" captionTargetPageId="7" captionText="FIGURE 6. Lucasioides altaicus sp. nov., male paratype: A—exopod of pleopod 1; B—endopod of pleopod 2; C—exopod of pleopod 2; D—endopod of pleopod 2; E—exopod of pleopod 3; F—exopod of pleopod 4; G—exopod of pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.4422850" httpUri="https://zenodo.org/record/4422850/files/figure.png" pageId="3" pageNumber="143">Fig. 6</figureCitation>
E–G) trapezoidal, slightly decreasing in size from 3 to 4. 
<emphasis id="B94FEAA33853FFC6FF3DF85CFE918889" box="[229,343,2015,2041]" italics="true" pageId="3" pageNumber="143">Pleopod 5</emphasis>
exopod triangular.
</paragraph>
</subSubSection>
<subSubSection id="C321653A3854FFC1FF1FFF14FB908FC2" box="[199,1110,151,178]" pageId="4" pageNumber="144" type="etymology">
<paragraph id="8B8436B13854FFC1FF1FFF14FB908FC2" blockId="4.[199,1110,151,178]" box="[199,1110,151,178]" pageId="4" pageNumber="144">
<emphasis id="B94FEAA33854FFC1FF1FFF14FE888FC1" bold="true" box="[199,334,151,177]" pageId="4" pageNumber="144">Etymology.</emphasis>
The specific epithet refers to the 
<emphasis id="B94FEAA33854FFC1FD1DFF14FC908FC0" box="[709,854,151,177]" italics="true" pageId="4" pageNumber="144">locus typicus</emphasis>
(the 
<collectingRegion id="49FFF8533854FFC1FC57FF14FC018FC1" box="[911,967,151,177]" country="Russia" name="Altay" pageId="4" pageNumber="144">Altai</collectingRegion>
Mountains).
</paragraph>
</subSubSection>
<caption id="DF4466393854FFC1FF4FFB6FFD4D8A57" ID-DOI="http://doi.org/10.5281/zenodo.4422842" ID-Zenodo-Dep="4422842" httpUri="https://zenodo.org/record/4422842/files/figure.png" pageId="4" pageNumber="144" startId="4.[151,250,1260,1284]" targetBox="[160,1423,212,1225]" targetPageId="4">
<paragraph id="8B8436B13854FFC1FF4FFB6FFD4D8A57" blockId="4.[151,1437,1260,1320]" pageId="4" pageNumber="144">
<emphasis id="B94FEAA33854FFC1FF4FFB6FFED48A74" bold="true" box="[151,274,1260,1284]" pageId="4" pageNumber="144">FIGURE 2.</emphasis>
<taxonomicName id="4C3B4D323854FFC1FECFFB6FFE238A74" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[279,485,1260,1284]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33854FFC1FECFFB6FFE238A74" box="[279,485,1260,1284]" italics="true" pageId="4" pageNumber="144">Lucasioides altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33854FFC1FE32FB6FFDFC8A74" bold="true" box="[490,570,1260,1284]" pageId="4" pageNumber="144">
<taxonomicNameLabel id="A27C57D83854FFC1FE32FB6FFDFC8A74" box="[490,570,1260,1284]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype: A—pereion edge; B—head; C—telson; D—antennula; E—antenna; F—pereopod 1; G—pereopod 6; H—pereopod 7.
</paragraph>
</caption>
<subSubSection id="C321653A3854FFC1FF1FFAF9FB9D8954" pageId="4" pageNumber="144" type="distribution">
<paragraph id="8B8436B13854FFC1FF1FFAF9FB9D8954" blockId="4.[151,1437,1402,2041]" pageId="4" pageNumber="144">
<emphasis id="B94FEAA33854FFC1FF1FFAF9FE328AE4" bold="true" box="[199,500,1402,1428]" pageId="4" pageNumber="144">Distribution and habitat.</emphasis>
The species has been collected in the lowlands of 
<collectingRegion id="49FFF8533854FFC1FBEBFAF9FBAD8AE4" box="[1075,1131,1402,1428]" country="Russia" name="Altay" pageId="4" pageNumber="144">Altai</collectingRegion>
Mountains in the environs of Charyshskoye Village. The area supports a steppe habitat with 
<taxonomicName id="4C3B4D323854FFC1FCABFA1CFBFC8AC8" box="[883,1082,1438,1465]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="chamaedrifolia">
<emphasis id="B94FEAA33854FFC1FCABFA1CFBFC8AC8" box="[883,1082,1438,1465]" italics="true" pageId="4" pageNumber="144">S. chamaedrifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323854FFC1FB90FA1CFB368AC8" authorityName="Lamarck" authorityYear="1785" box="[1096,1264,1438,1465]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="B94FEAA33854FFC1FB90FA1CFB368AC8" box="[1096,1264,1438,1465]" italics="true" pageId="4" pageNumber="144">C. arborescens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323854FFC1FAFEFA1CFA5B8AC8" authorityName="Linnaeus" authorityYear="1753" box="[1318,1437,1439,1464]" class="Magnoliopsida" family="Caprifoliaceae" genus="Lonicera" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="tatarica">
<emphasis id="B94FEAA33854FFC1FAFEFA1CFA5B8AC8" box="[1318,1437,1439,1464]" italics="true" pageId="4" pageNumber="144">L. tatarica</emphasis>
</taxonomicName>
bushes and scattered 
<taxonomicName id="4C3B4D323854FFC1FE5AFA40FDC38AAC" box="[386,517,1474,1500]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="B94FEAA33854FFC1FE5AFA40FDC38AAC" box="[386,517,1474,1500]" italics="true" pageId="4" pageNumber="144">P. sylvestris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323854FFC1FDCBFA40FD4D8AAC" authorityName="A.W.Roth" authorityYear="1788" box="[531,651,1474,1500]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="B94FEAA33854FFC1FDCBFA40FD4D8AAC" box="[531,651,1474,1500]" italics="true" pageId="4" pageNumber="144">B. pendula</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323854FFC1FD18FA40FCF78AAC" box="[704,817,1474,1500]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="tremula">
<emphasis id="B94FEAA33854FFC1FD18FA40FCF78AAC" box="[704,817,1474,1500]" italics="true" pageId="4" pageNumber="144">P. tremula</emphasis>
</taxonomicName>
stands with 
<taxonomicName id="4C3B4D323854FFC1FC64FA40FB448AAC" box="[956,1154,1474,1501]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="chamaedrifolia">
<emphasis id="B94FEAA33854FFC1FC64FA40FB448AAC" box="[956,1154,1474,1501]" italics="true" pageId="4" pageNumber="144">S. chamaedrifolia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323854FFC1FB6FFA40FA8A8AAC" box="[1207,1356,1474,1500]" class="Magnoliopsida" family="Saxifragaceae" genus="Bergenia" kingdom="Animalia" order="Saxifragales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="crassifolia">
<emphasis id="B94FEAA33854FFC1FB6FFA40FA8A8AAC" box="[1207,1356,1474,1500]" italics="true" pageId="4" pageNumber="144">B. crassifolia</emphasis>
</taxonomicName>
, growing on the southern and northern rocky slopes of a mountain, respectively. The above is the first record of the genus 
<taxonomicName id="4C3B4D323854FFC1FF4FF989FE688954" authority="Kwon, 1993" authorityName="Kwon" authorityYear="1993" box="[151,430,1546,1573]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FF4FF989FEDB8954" box="[151,285,1546,1572]" italics="true" pageId="4" pageNumber="144">Lucasioides</emphasis>
<bibRefCitation id="EFAA4B403854FFC1FEFCF988FE688954" author="Kwon, D." box="[292,430,1546,1573]" pageId="4" pageNumber="144" pagination="133 - 158" refId="ref4911" refString="Kwon, D. (1993) Terrestrial Isopoda (Crustacea) from Korea. Korean Journal of Zoology, 36, 133 - 158." type="journal article" year="1993">Kwon, 1993</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F32C76213854FFC1FE2BF989FD868954" box="[499,576,1546,1572]" name="Russia" pageId="4" pageNumber="144">Russia</collectingCountry>
, showing its presently northwestern range limit.
</paragraph>
</subSubSection>
<subSubSection id="C321653A3854FFC3FF1FF9ADFD668A98" lastPageId="6" lastPageNumber="146" pageId="4" pageNumber="144" type="discussion">
<paragraph id="8B8436B13854FFC0FF1FF9ADFA4A8FA5" blockId="4.[151,1437,1402,2041]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="145" pageId="4" pageNumber="144">
<emphasis id="B94FEAA33854FFC1FF1FF9ADFEFD8938" bold="true" box="[199,315,1582,1608]" pageId="4" pageNumber="144">Remarks.</emphasis>
The new species is assigned to 
<taxonomicName id="4C3B4D323854FFC1FD7BF9ADFCEF8938" authorityName="Kwon" authorityYear="1993" box="[675,809,1582,1608]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FD7BF9ADFCEF8938" box="[675,809,1582,1608]" italics="true" pageId="4" pageNumber="144">Lucasioides</emphasis>
</taxonomicName>
since it has five pairs of pleopodal lungs; the flagellum of antenna consisting of two articles, and the position of the 
<emphasis id="B94FEAA33854FFC1FCE6F9D1FC29891C" box="[830,1007,1618,1644]" italics="true" pageId="4" pageNumber="144">noduli laterales</emphasis>
distinctive for the genus. This species seems to be particularly close to 
<taxonomicName id="4C3B4D323854FFC1FDCCF9F4FCFB89E0" authorityName="Borutzky" authorityYear="1978" box="[532,829,1654,1680]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="B94FEAA33854FFC1FDCCF9F4FCFB89E0" box="[532,829,1654,1680]" italics="true" pageId="4" pageNumber="144">Desertoniscus mongolicus</emphasis>
</taxonomicName>
due to the similar shape of most body parts. Yet the taxonomic position of 
<taxonomicName id="4C3B4D323854FFC1FE48F918FDF489C4" authorityName="Borutzky" authorityYear="1978" box="[400,562,1690,1716]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="B94FEAA33854FFC1FE48F918FDF489C4" box="[400,562,1690,1716]" italics="true" pageId="4" pageNumber="144">D. mongolicus</emphasis>
</taxonomicName>
within 
<taxonomicName id="4C3B4D323854FFC1FD5DF918FCE289C4" authorityName="Verhoeff" authorityYear="1930" box="[645,804,1691,1716]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FD5DF918FCE289C4" box="[645,804,1691,1716]" italics="true" pageId="4" pageNumber="144">Desertoniscus</emphasis>
</taxonomicName>
needs to be revised. In the original description, 
<bibRefCitation id="EFAA4B403854FFC1FAECF919FF2489A8" author="Borutzky, E. V." pageId="4" pageNumber="144" pagination="3 - 21" refId="ref4293" refString="Borutzky, E. V. (1978) On the genus Desertoniscus Verhoeff, 1930. Sbornik Trudov Zoologicheskogo Muzeya MGU, 16, 3 - 21. [in Russian]" type="journal article" year="1978">Borutzky (1978)</bibRefCitation>
noted that 
<taxonomicName id="4C3B4D323854FFC1FE87F93CFDC489A8" authorityName="Borutzky" authorityYear="1978" box="[351,514,1726,1752]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="B94FEAA33854FFC1FE87F93CFDC489A8" box="[351,514,1726,1752]" italics="true" pageId="4" pageNumber="144">D. mongolicus</emphasis>
</taxonomicName>
occupies a position intermediate between the genera 
<taxonomicName id="4C3B4D323854FFC1FB81F93CFB3E89A8" authorityName="Verhoeff" authorityYear="1930" box="[1113,1272,1727,1752]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FB81F93CFB3E89A8" box="[1113,1272,1727,1752]" italics="true" pageId="4" pageNumber="144">Desertoniscus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323854FFC1FAF6F93DFF1B898C" authorityName="Verhoeff" authorityYear="1930" class="Malacostraca" family="Trachelipodidae" genus="Mongoloniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FAF6F93DFF1B898C" italics="true" pageId="4" pageNumber="144">Mongoloniscus</emphasis>
</taxonomicName>
, but in fact both 
<taxonomicName id="4C3B4D323854FFC1FE4FF960FDFF898C" authorityName="Borutzky" authorityYear="1978" box="[407,569,1762,1788]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="B94FEAA33854FFC1FE4FF960FDFF898C" box="[407,569,1762,1788]" italics="true" pageId="4" pageNumber="144">D. mongolicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323854FFC1FDB5F960FD24898C" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[621,738,1762,1788]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33854FFC1FDB5F960FD24898C" box="[621,738,1762,1788]" italics="true" pageId="4" pageNumber="144">L. altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33854FFC1FD30F961FC86898C" bold="true" box="[744,832,1762,1788]" pageId="4" pageNumber="144">
<taxonomicNameLabel id="A27C57D83854FFC1FD30F961FC86898C" box="[744,832,1762,1788]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also bear characteristics of both 
<taxonomicName id="4C3B4D323854FFC1FB71F961FAA8898C" authorityName="Verhoeff" authorityYear="1917" box="[1193,1390,1762,1788]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FB71F961FAA8898C" box="[1193,1390,1762,1788]" italics="true" pageId="4" pageNumber="144">Protracheoniscus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323854FFC1FF4FF885FEDB8850" authorityName="Kwon" authorityYear="1993" box="[151,285,1798,1824]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FF4FF885FEDB8850" box="[151,285,1798,1824]" italics="true" pageId="4" pageNumber="144">Lucasioides</emphasis>
</taxonomicName>
(e.g., the position of the 
<emphasis id="B94FEAA33854FFC1FDE4F885FD2B8850" box="[572,749,1798,1824]" italics="true" pageId="4" pageNumber="144">noduli laterales</emphasis>
). However, these two species differ in the shape of the male exopod of pleopod 1: 
<taxonomicName id="4C3B4D323854FFC1FE52F8A8FDEA8834" authorityName="Borutzky" authorityYear="1978" box="[394,556,1834,1860]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="B94FEAA33854FFC1FE52F8A8FDEA8834" box="[394,556,1834,1860]" italics="true" pageId="4" pageNumber="144">D. mongolicus</emphasis>
</taxonomicName>
has a longer upper lobe at the tip, while in 
<taxonomicName id="4C3B4D323854FFC1FBD2F8A8FB468834" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[1034,1152,1834,1860]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33854FFC1FBD2F8A8FB468834" box="[1034,1152,1834,1860]" italics="true" pageId="4" pageNumber="144">L. altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33854FFC1FB5EF8A9FB188834" bold="true" box="[1158,1246,1834,1860]" pageId="4" pageNumber="144">
<taxonomicNameLabel id="A27C57D83854FFC1FB5EF8A9FB188834" box="[1158,1246,1834,1860]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the lower lobe is longer; 
<taxonomicName id="4C3B4D323854FFC1FF35F8CCFE578818" authorityName="Borutzky" authorityYear="1978" box="[237,401,1870,1896]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="B94FEAA33854FFC1FF35F8CCFE578818" box="[237,401,1870,1896]" italics="true" pageId="4" pageNumber="144">D. mongolicus</emphasis>
</taxonomicName>
has antenna reaching only the posterior edge of tergite I, while they are longer in 
<taxonomicName id="4C3B4D323854FFC1FAFEF8CCFA5B8818" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[1318,1437,1870,1896]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33854FFC1FAFEF8CCFA5B8818" box="[1318,1437,1870,1896]" italics="true" pageId="4" pageNumber="144">L. altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33854FFC1FF4FF8F1FF3688FC" bold="true" box="[151,240,1906,1932]" pageId="4" pageNumber="144">
<taxonomicNameLabel id="A27C57D83854FFC1FF4FF8F1FF3688FC" box="[151,240,1906,1932]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and reaching the posterior edge of tergite II. Within the genus 
<taxonomicName id="4C3B4D323854FFC1FC7AF8F1FBEE88FC" authorityName="Kwon" authorityYear="1993" box="[930,1064,1906,1932]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33854FFC1FC7AF8F1FBEE88FC" box="[930,1064,1906,1932]" italics="true" pageId="4" pageNumber="144">Lucasioides</emphasis>
</taxonomicName>
, there are no species which could be mixed up with the new species. The morphologically closest species are 
<taxonomicName id="4C3B4D323854FFC1FC30F814FAC888C0" authority="(Arcangeli, 1927)" baseAuthorityName="Arcangeli" baseAuthorityYear="1927" box="[1000,1294,1942,1968]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="isseli">
<emphasis id="B94FEAA33854FFC1FC30F814FB8688C0" box="[1000,1088,1942,1968]" italics="true" pageId="4" pageNumber="144">L. isseli</emphasis>
(
<bibRefCitation id="EFAA4B403854FFC1FB96F815FAC388C0" author="Arcangeli, A." box="[1102,1285,1942,1968]" pageId="4" pageNumber="144" pagination="211 - 269" refId="ref4177" refString="Arcangeli, A. (1927) Isopodi terrestri raccolti nell'Estreme Oriente dal Prof. Filippo Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria del R. Istituto Superiore Agrario in Portici, 20, 211 - 269." type="journal article" year="1927">Arcangeli, 1927</bibRefCitation>
)
</taxonomicName>
from 
<collectingCountry id="F32C76213854FFC1FA8AF815FA5E88C0" box="[1362,1432,1942,1968]" name="China" pageId="4" pageNumber="144">China</collectingCountry>
, 
<taxonomicName id="4C3B4D323854FFC1FF4FF838FE2D88A4" authority="(Nunomura, 1987)" baseAuthorityName="Nunomura" baseAuthorityYear="1987" box="[151,491,1978,2004]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="sinuosus">
<emphasis id="B94FEAA33854FFC1FF4FF838FED288A4" box="[151,276,1979,2004]" italics="true" pageId="4" pageNumber="144">L. sinuosus</emphasis>
(Nunomura, 1987)
</taxonomicName>
from 
<collectingCountry id="F32C76213854FFC1FDF6F838FDB488A5" box="[558,626,1979,2005]" name="South Korea" pageId="4" pageNumber="144">Korea</collectingCountry>
and 
<collectingCountry id="F32C76213854FFC1FD70F838FD2D88A5" box="[680,747,1979,2005]" name="Japan" pageId="4" pageNumber="144">Japan</collectingCountry>
, 
<taxonomicName id="4C3B4D323854FFC1FD2DF838FB9F88A4" authority="Nunomura 2008" authorityName="Nunomura" authorityYear="2008" box="[757,1113,1978,2005]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="toyamaensis">
<emphasis id="B94FEAA33854FFC1FD2DF838FC5A88A4" box="[757,924,1979,2004]" italics="true" pageId="4" pageNumber="144">L. toyamaensis</emphasis>
<bibRefCitation id="EFAA4B403854FFC1FC7BF838FB9F88A4" author="Nunomura, N." box="[931,1113,1978,2005]" pageId="4" pageNumber="144" pagination="3 - 11" refId="ref5058" refString="Nunomura, N. (2008) Two new species of the genus Lucasioides (Crustacea, Isopod) from mole's tunnels of Nanto-shi, Toyama, middle Japan. Bulletin of the Toyama Science Museum, 31, 3 - 11." type="journal article" year="2008">Nunomura 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323854FFC1FBBEF838FA5E88A4" authority="Nunomura 2008" authorityName="Nunomura" authorityYear="2008" box="[1126,1432,1978,2005]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="sagarai">
<emphasis id="B94FEAA33854FFC1FBBEF838FB1F88A4" box="[1126,1241,1979,2004]" italics="true" pageId="4" pageNumber="144">L. sagarai</emphasis>
<bibRefCitation id="EFAA4B403854FFC1FB07F838FA5E88A4" author="Nunomura, N." box="[1247,1432,1978,2005]" pageId="4" pageNumber="144" pagination="3 - 11" refId="ref5058" refString="Nunomura, N. (2008) Two new species of the genus Lucasioides (Crustacea, Isopod) from mole's tunnels of Nanto-shi, Toyama, middle Japan. Bulletin of the Toyama Science Museum, 31, 3 - 11." type="journal article" year="2008">Nunomura 2008</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C3B4D323854FFC1FF1FF85CFDD78888" authority="Nunomura 2010" authorityName="Nunomura" authorityYear="2010" box="[199,529,2014,2041]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="144" phylum="Arthropoda" rank="species" species="yokahatai">
<emphasis id="B94FEAA33854FFC1FF1FF85CFE958888" box="[199,339,2014,2040]" italics="true" pageId="4" pageNumber="144">L. yokahatai</emphasis>
<bibRefCitation id="EFAA4B403854FFC1FE82F85CFDD78888" author="Nunomura, N." box="[346,529,2014,2041]" pageId="4" pageNumber="144" pagination="33 - 37" refId="ref5101" refString="Nunomura, N. (2010) A new species of the genus Lucasioides (Crustacea: Isopoda) from nest material of Mogera imaizumii, Toyama-ken, central Japan. Bulletin of the Toyama Science Museum, 33, 33 - 37." type="journal article" year="2010">Nunomura 2010</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F32C76213854FFC1FD8EF85CFD508889" box="[598,662,2015,2041]" name="Japan" pageId="4" pageNumber="144">Japan</collectingCountry>
(
<bibRefCitation id="EFAA4B403854FFC1FD7CF85DFCA68888" author="Nunomura, N." box="[676,864,2014,2040]" pageId="4" pageNumber="144" pagination="3 - 11" refId="ref5058" refString="Nunomura, N. (2008) Two new species of the genus Lucasioides (Crustacea, Isopod) from mole's tunnels of Nanto-shi, Toyama, middle Japan. Bulletin of the Toyama Science Museum, 31, 3 - 11." type="journal article" year="2008">Nunomura 2008</bibRefCitation>
, 
<bibRefCitation id="EFAA4B403854FFC1FCB3F85DFC6E8888" author="Nunomura, N." box="[875,936,2014,2040]" pageId="4" pageNumber="144" pagination="33 - 37" refId="ref5101" refString="Nunomura, N. (2010) A new species of the genus Lucasioides (Crustacea: Isopoda) from nest material of Mogera imaizumii, Toyama-ken, central Japan. Bulletin of the Toyama Science Museum, 33, 33 - 37." type="journal article" year="2010">2010</bibRefCitation>
). The new species readily differs from all by the sharp and elongated telson, and in the shape of the exopod of pleopod 1. A molecular analysis is needed for the complex of those small species of 
<taxonomicName id="4C3B4D323855FFC0FDCFFF38FD498FA5" authorityName="Schmidt" authorityYear="2003" box="[535,655,187,213]" class="Malacostraca" family="Agnaridae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="family">Agnaridae</taxonomicName>
, and a complete taxonomic revision of these sister genera is needed.
</paragraph>
<caption id="DF4466393855FFC0FF4FF98AFCFA8936" ID-DOI="http://doi.org/10.5281/zenodo.4422844" ID-Zenodo-Dep="4422844" httpUri="https://zenodo.org/record/4422844/files/figure.png" pageId="5" pageNumber="145" startId="5.[151,250,1545,1569]" targetBox="[157,1430,243,1513]" targetPageId="5">
<paragraph id="8B8436B13855FFC0FF4FF98AFCFA8936" blockId="5.[151,1436,1545,1606]" pageId="5" pageNumber="145">
<emphasis id="B94FEAA33855FFC0FF4FF98AFED58951" bold="true" box="[151,275,1545,1569]" pageId="5" pageNumber="145">FIGURE 3.</emphasis>
<taxonomicName id="4C3B4D323855FFC0FEC2F98AFE2C8951" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[282,490,1545,1569]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33855FFC0FEC2F98AFE2C8951" box="[282,490,1545,1569]" italics="true" pageId="5" pageNumber="145">Lucasioides altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33855FFC0FE29F989FD848952" bold="true" box="[497,578,1546,1570]" pageId="5" pageNumber="145">
<taxonomicNameLabel id="A27C57D83855FFC0FE29F989FD848952" box="[497,578,1546,1570]" pageId="5" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype: A—pereionites 6, 7, pleon and telson; B—nodulis lateralis on pereonite 7; C—pereionites 1–3; D—nodulis lateralis on pereonite 2.
</paragraph>
</caption>
<paragraph id="8B8436B13855FFC0FF1FF914FEFF88FA" blockId="5.[151,1437,1687,1930]" pageId="5" pageNumber="145">
<emphasis id="B94FEAA33855FFC0FF1FF914FE0889C1" bold="true" box="[199,462,1687,1713]" pageId="5" pageNumber="145">Phylogenetic analysis.</emphasis>
A total of 11 nucleotide sequences were used to assess both species and intraspecific variation (
<figureCitation id="13002A343855FFC0FED2F93FFE8989A6" box="[266,335,1724,1750]" captionStart="FIGURE 8" captionStartId="8.[151,250,1942,1966]" captionTargetBox="[158,1428,1366,1911]" captionTargetId="figure-51@8.[151,1436,1359,1917]" captionTargetPageId="8" captionText="FIGURE 8. Phylogenetic reconstruction of selected species of family Agnaridae, based on the analysis of COI mtDNA gene sequences using the Bayesian analysis (BA) with the GTR + G + I evolutionary model for Maximum-Likelihood (ML), using Kimura2-parameter (K2P) model; bootstrap support for branch nodes shown for 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.4422854" httpUri="https://zenodo.org/record/4422854/files/figure.png" pageId="5" pageNumber="145">Fig. 8</figureCitation>
): two of them were obtained from 
<taxonomicName id="4C3B4D323855FFC0FD0FF93FFC8889A6" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[727,846,1724,1750]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33855FFC0FD0FF93FFC8889A6" box="[727,846,1724,1750]" italics="true" pageId="5" pageNumber="145">L. altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33855FFC0FC8DF93FFC6989A6" bold="true" box="[853,943,1724,1750]" pageId="5" pageNumber="145">
<taxonomicNameLabel id="A27C57D83855FFC0FC8DF93FFC6989A6" box="[853,943,1724,1750]" pageId="5" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and further nine revealed earlier from 
<taxonomicName id="4C3B4D323855FFC0FAB2F93FFD83898A" authority="(C. Koch, 1841)" baseAuthorityName="C. Koch" baseAuthorityYear="1841" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="B94FEAA33855FFC0FAB2F93FFE40898A" italics="true" pageId="5" pageNumber="145">Protracheoniscus politus</emphasis>
(C. Koch, 1841)
</taxonomicName>
from 
<collectingRegion id="49FFF8533855FFC0FD52F963FD33898A" box="[650,757,1760,1786]" country="Hungary" name="Budapest" pageId="5" pageNumber="145">Budapest</collectingRegion>
, 
<collectingCountry id="F32C76213855FFC0FCD9F963FCA2898A" box="[769,868,1760,1786]" name="Hungary" pageId="5" pageNumber="145">Hungary</collectingCountry>
, 
<taxonomicName id="4C3B4D323855FFC0FCB7F963FABA898A" authority="Gongalsky et Turbanov, 2018" authorityName="Gongalsky et Turbanov" authorityYear="2018" box="[879,1404,1760,1786]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="pokarzhevskii">
<emphasis id="B94FEAA33855FFC0FCB7F963FBE3898A" box="[879,1061,1760,1786]" italics="true" pageId="5" pageNumber="145">P. pokarzhevskii</emphasis>
<bibRefCitation id="EFAA4B403855FFC0FBF5F963FABA898A" author="Gongalsky, K. B. &amp; Turbanov, I. S. &amp; Medvedev, D. A. &amp; Volkova, J. S." box="[1069,1404,1760,1786]" pageId="5" pageNumber="145" pagination="189 - 205" refId="ref4716" refString="Gongalsky, K. B., Turbanov, I. S., Medvedev, D. A. &amp; Volkova, J. S. (2018) Description of a new species of the genus Protracheoniscus Verhoeff, 1917 and redescription of Protracheoniscus kryszanovskii Borutzky, 1957 from the southeast of European Russia (Isopoda, Oniscidea, Agnaridae). ZooKeys, 801, 189 - 205. https: // doi. org / 10.3897 / zookeys. 801.23167" type="journal article" year="2018">Gongalsky et Turbanov, 2018</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3B4D323855FFC0FA50F963FE25886E" authority="Borutzky, 1957" authorityName="Borutzky" authorityYear="1957" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="kryszanovskii">
<emphasis id="B94FEAA33855FFC0FA50F963FEE8886E" italics="true" pageId="5" pageNumber="145">P. kryszanovskii</emphasis>
Borutzky, 1957
</taxonomicName>
, 
<taxonomicName id="4C3B4D323855FFC0FE35F887FC90886E" authority="Demianowicz, 1932" authorityName="Demianowicz" authorityYear="1932" box="[493,854,1796,1822]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="nogaicus">
<emphasis id="B94FEAA33855FFC0FE35F887FDAA886D" box="[493,620,1796,1821]" italics="true" pageId="5" pageNumber="145">P. nogaicus</emphasis>
Demianowicz, 1932
</taxonomicName>
, 
<taxonomicName id="4C3B4D323855FFC0FCB8F887FBA9886E" authority="(Dollfus, 1903)" baseAuthorityName="Dollfus" baseAuthorityYear="1903" box="[864,1135,1796,1822]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B94FEAA33855FFC0FCB8F887FC7B886D" box="[864,957,1796,1821]" italics="true" pageId="5" pageNumber="145">P. major</emphasis>
(Dollfus, 1903)
</taxonomicName>
and 
<taxonomicName id="4C3B4D323855FFC0FB7CF887FE908832" authority="Gongalsky, 2017" authorityName="Gongalsky" authorityYear="2017" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="zaitsevi">
<emphasis id="B94FEAA33855FFC0FB7CF887FA5A886D" box="[1188,1436,1796,1821]" italics="true" pageId="5" pageNumber="145">Desertoniscus zaitsevi</emphasis>
Gongalsky, 2017
</taxonomicName>
from the Republic of 
<collectingRegion id="49FFF8533855FFC0FD8AF8ABFD058832" box="[594,707,1832,1858]" country="Russia" name="Kalmykiya" pageId="5" pageNumber="145">Kalmykia</collectingRegion>
, 
<collectingCountry id="F32C76213855FFC0FD08F8ABFCDD8832" box="[720,795,1832,1858]" name="Russia" pageId="5" pageNumber="145">Russia</collectingCountry>
(
<bibRefCitation id="EFAA4B403855FFC0FCF2F8ABFBED8832" author="Gongalsky, K. B. &amp; Turbanov, I. S. &amp; Medvedev, D. A. &amp; Volkova, J. S." box="[810,1067,1832,1858]" pageId="5" pageNumber="145" pagination="189 - 205" refId="ref4716" refString="Gongalsky, K. B., Turbanov, I. S., Medvedev, D. A. &amp; Volkova, J. S. (2018) Description of a new species of the genus Protracheoniscus Verhoeff, 1917 and redescription of Protracheoniscus kryszanovskii Borutzky, 1957 from the southeast of European Russia (Isopoda, Oniscidea, Agnaridae). ZooKeys, 801, 189 - 205. https: // doi. org / 10.3897 / zookeys. 801.23167" type="journal article" year="2018">
Gongalsky 
<emphasis id="B94FEAA33855FFC0FC75F8ABFC198832" box="[941,991,1832,1858]" italics="true" pageId="5" pageNumber="145">et al</emphasis>
. 2018
</bibRefCitation>
). The loci of the 
<emphasis id="B94FEAA33855FFC0FB34F8ABFADA8832" box="[1260,1308,1832,1858]" italics="true" pageId="5" pageNumber="145">COI</emphasis>
mtDNA of 
<taxonomicName id="4C3B4D323855FFC0FF4FF8CFFDE78816" authority="(Brandt, 1833)" baseAuthorityName="Brandt" baseAuthorityYear="1833" box="[151,545,1868,1894]" class="Malacostraca" family="Trachelipodidae" genus="Hemilepistus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="klugii">
<emphasis id="B94FEAA33855FFC0FF4FF8CFFEB48816" box="[151,370,1868,1894]" italics="true" pageId="5" pageNumber="145">Hemilepistus klugii</emphasis>
(Brandt, 1833)
</taxonomicName>
and 
<taxonomicName id="4C3B4D323855FFC0FD83F8CFFCB98816" authority="Lincoln, 1970" authorityName="Lincoln" authorityYear="1970" box="[603,895,1868,1894]" class="Malacostraca" family="Trachelipodidae" genus="Hemilepistus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="145" phylum="Arthropoda" rank="species" species="schirasi">
<emphasis id="B94FEAA33855FFC0FD83F8CFFD1E8816" box="[603,728,1868,1894]" italics="true" pageId="5" pageNumber="145">H. schirasi</emphasis>
Lincoln, 1970
</taxonomicName>
from 
<collectingCountry id="F32C76213855FFC0FC1FF8CFFC328816" box="[967,1012,1868,1894]" name="Iran" pageId="5" pageNumber="145">Iran</collectingCountry>
(
<bibRefCitation id="EFAA4B403855FFC0FBDCF8CFFB3C8816" author="Dimitriou, A. C. &amp; Taiti, S. &amp; Schmalfuss, H. &amp; Sfenthourakis, S." box="[1028,1274,1868,1894]" pageId="5" pageNumber="145" pagination="163 - 176" refId="ref4455" refString="Dimitriou, A. C., Taiti, S., Schmalfuss, H. &amp; Sfenthourakis, S. (2018) A molecular phylogeny of Porcellionidae (Isopoda, Oniscidea) reveals inconsistencies with present taxonomy. ZooKeys, 801, 163 - 176. https: // doi. org / 10.3897 / zookeys. 801.23566" type="journal article" year="2018">
Dimitriou 
<emphasis id="B94FEAA33855FFC0FBA5F8CFFB768816" box="[1149,1200,1868,1894]" italics="true" pageId="5" pageNumber="145">et al</emphasis>
. 2018
</bibRefCitation>
) were used as outgroup taxa.
</paragraph>
<caption id="DF4466393856FFC3FF4FFC7FFEF28B48" ID-DOI="http://doi.org/10.5281/zenodo.4422846" ID-Zenodo-Dep="4422846" httpUri="https://zenodo.org/record/4422846/files/figure.png" pageId="6" pageNumber="146" startId="6.[151,250,1020,1044]" targetBox="[157,1432,185,991]" targetPageId="6">
<paragraph id="8B8436B13856FFC3FF4FFC7FFEF28B48" blockId="6.[151,1437,1020,1080]" pageId="6" pageNumber="146">
<emphasis id="B94FEAA33856FFC3FF4FFC7FFED58B64" bold="true" box="[151,275,1020,1044]" pageId="6" pageNumber="146">FIGURE 4.</emphasis>
<taxonomicName id="4C3B4D323856FFC3FEC1FC7FFE2F8B64" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[281,489,1020,1044]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33856FFC3FEC1FC7FFE2F8B64" box="[281,489,1020,1044]" italics="true" pageId="6" pageNumber="146">Lucasioides altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33856FFC3FE37FC7FFD868B64" bold="true" box="[495,576,1020,1044]" pageId="6" pageNumber="146">
<taxonomicNameLabel id="A27C57D83856FFC3FE37FC7FFD868B64" box="[495,576,1020,1044]" pageId="6" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype: A—left mandible; B—right mandible; C—maxillula; D—maxilla; E—maxilliped.
</paragraph>
</caption>
<paragraph id="8B8436B13856FFC3FF1FFB09FCB88A44" blockId="6.[151,1437,1162,1513]" pageId="6" pageNumber="146">
An analysis of the average values of pairwise sequence divergences (
<emphasis id="B94FEAA33856FFC3FC1EFB08FC128BD4" box="[966,980,1163,1188]" italics="true" pageId="6" pageNumber="146">p</emphasis>
-distances) of partial 
<emphasis id="B94FEAA33856FFC3FB18FB08FB368BD5" box="[1216,1264,1163,1189]" italics="true" pageId="6" pageNumber="146">COI</emphasis>
mtDNA (
<tableCitation id="C6B9030A3856FFC3FA84FB09FF608BB8" captionStart="TABLE 2" captionStartId="6.[151,239,1560,1584]" captionTargetPageId="6" captionText="TABLE 2. Estimates of pairwise sequence divergence (uncorrected p-distances) of the gene COI mtDNA haplotypes between Lucasioides altaicus sp. nov. and eight species of Agnaridae." httpUri="http://table.plazi.org/id/DF4466393856FFC3FF4FF99BFC458924" pageId="6" pageNumber="146" tableUuid="DF4466393856FFC3FF4FF99BFC458924">Table 2</tableCitation>
) shows that the intraspecific difference of two 
<emphasis id="B94FEAA33856FFC3FD67FB2DFD2F8BB8" box="[703,745,1198,1224]" italics="true" pageId="6" pageNumber="146">loci</emphasis>
of 
<taxonomicName id="4C3B4D323856FFC3FCC8FB2CFC4E8BB8" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[784,904,1198,1224]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33856FFC3FCC8FB2CFC4E8BB8" box="[784,904,1198,1224]" italics="true" pageId="6" pageNumber="146">L. altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33856FFC3FC48FB2DFC2C8BB8" bold="true" box="[912,1002,1198,1224]" pageId="6" pageNumber="146">
<taxonomicNameLabel id="A27C57D83856FFC3FC48FB2DFC2C8BB8" box="[912,1002,1198,1224]" pageId="6" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is 0%, which indicates their complete identity. Within the considered species, 
<emphasis id="B94FEAA33856FFC3FD82FB50FDAE8B9C" box="[602,616,1235,1260]" italics="true" pageId="6" pageNumber="146">p</emphasis>
-distances range from 15.7% between 
<taxonomicName id="4C3B4D323856FFC3FBC4FB50FB558B9C" authority="Gongalsky &amp; Nefediev &amp; Turbanov, 2021" authorityName="Gongalsky &amp; Nefediev &amp; Turbanov" authorityYear="2021" box="[1052,1171,1234,1260]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="altaicus" status="sp. nov.">
<emphasis id="B94FEAA33856FFC3FBC4FB50FB558B9C" box="[1052,1171,1234,1260]" italics="true" pageId="6" pageNumber="146">L. altaicus</emphasis>
</taxonomicName>
<emphasis id="B94FEAA33856FFC3FB43FB51FB308B9C" bold="true" box="[1179,1270,1234,1260]" pageId="6" pageNumber="146">
<taxonomicNameLabel id="A27C57D83856FFC3FB43FB51FB308B9C" box="[1179,1270,1234,1260]" pageId="6" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C3B4D323856FFC3FAF7FB50FF208A60" authorityName="Borutzky" authorityYear="1957" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="kryszanovskii">
<emphasis id="B94FEAA33856FFC3FAF7FB50FF208A60" italics="true" pageId="6" pageNumber="146">P. kryszanovskii</emphasis>
</taxonomicName>
up to 24.9% between 
<taxonomicName id="4C3B4D323856FFC3FE33FB74FD938A60" baseAuthorityName="C. Koch" baseAuthorityYear="1841" box="[491,597,1270,1296]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="B94FEAA33856FFC3FE33FB74FD938A60" box="[491,597,1270,1296]" italics="true" pageId="6" pageNumber="146">P. politus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B4D323856FFC3FD57FB74FD368A60" baseAuthorityName="Dollfus" baseAuthorityYear="1903" box="[655,752,1271,1296]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B94FEAA33856FFC3FD57FB74FD368A60" box="[655,752,1271,1296]" italics="true" pageId="6" pageNumber="146">P. major</emphasis>
</taxonomicName>
. Similar genetic distances correspond to possible variations within the 
<emphasis id="B94FEAA33856FFC3FED7FA98FEF98A45" box="[271,319,1307,1333]" italics="true" pageId="6" pageNumber="146">COI</emphasis>
mtDNA gene for woodlice (
<bibRefCitation id="EFAA4B403856FFC3FDA6FA99FCB48A44" author="Dimitriou, A. C. &amp; Taiti, S. &amp; Schmalfuss, H. &amp; Sfenthourakis, S." box="[638,882,1306,1333]" pageId="6" pageNumber="146" pagination="163 - 176" refId="ref4455" refString="Dimitriou, A. C., Taiti, S., Schmalfuss, H. &amp; Sfenthourakis, S. (2018) A molecular phylogeny of Porcellionidae (Isopoda, Oniscidea) reveals inconsistencies with present taxonomy. ZooKeys, 801, 163 - 176. https: // doi. org / 10.3897 / zookeys. 801.23566" type="journal article" year="2018">
Dimitriou 
<emphasis id="B94FEAA33856FFC3FD2EFA98FCE18A44" box="[758,807,1306,1332]" italics="true" pageId="6" pageNumber="146">et al</emphasis>
. 2018
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8436B13856FFC3FF1FFABCFD668A98" blockId="6.[151,1437,1162,1513]" pageId="6" pageNumber="146">
It seems noteworthy that 
<taxonomicName id="4C3B4D323856FFC3FE3FFABCFD988A28" authorityName="Gongalsky" authorityYear="2017" box="[487,606,1343,1368]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="zaitsevi">
<emphasis id="B94FEAA33856FFC3FE3FFABCFD988A28" box="[487,606,1343,1368]" italics="true" pageId="6" pageNumber="146">D. zaitsevi</emphasis>
</taxonomicName>
falls within the 
<emphasis id="B94FEAA33856FFC3FCC0FABCFCE08A28" box="[792,806,1343,1368]" italics="true" pageId="6" pageNumber="146">p</emphasis>
-distances range for the genus 
<taxonomicName id="4C3B4D323856FFC3FB59FABDFA838A28" authorityName="Verhoeff" authorityYear="1917" box="[1153,1349,1342,1368]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33856FFC3FB59FABDFA838A28" box="[1153,1349,1342,1368]" italics="true" pageId="6" pageNumber="146">Protracheoniscus</emphasis>
</taxonomicName>
, whose average values of pairwise sequence divergences reach 18.0% with 
<taxonomicName id="4C3B4D323856FFC3FC4FFAE0FC308A0C" baseAuthorityName="Dollfus" baseAuthorityYear="1903" box="[919,1014,1379,1404]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B94FEAA33856FFC3FC4FFAE0FC308A0C" box="[919,1014,1379,1404]" italics="true" pageId="6" pageNumber="146">P. major</emphasis>
</taxonomicName>
and 23.4% with 
<taxonomicName id="4C3B4D323856FFC3FB61FAE0FAFC8A0C" authorityName="Demianowicz" authorityYear="1932" box="[1209,1338,1379,1404]" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="species" species="nogaicus">
<emphasis id="B94FEAA33856FFC3FB61FAE0FAFC8A0C" box="[1209,1338,1379,1404]" italics="true" pageId="6" pageNumber="146">P. nogaicus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C3B4D323856FFC3FA91FAE0FED48AD0" authorityName="Verhoeff" authorityYear="1917" class="Malacostraca" family="Trachelipodidae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33856FFC3FA91FAE0FED48AD0" italics="true" pageId="6" pageNumber="146">Protracheoniscus</emphasis>
</taxonomicName>
appears to be paraphyletic (
<figureCitation id="13002A343856FFC3FD8AFA05FD518AD0" box="[594,663,1414,1440]" captionStart="FIGURE 8" captionStartId="8.[151,250,1942,1966]" captionTargetBox="[158,1428,1366,1911]" captionTargetId="figure-51@8.[151,1436,1359,1917]" captionTargetPageId="8" captionText="FIGURE 8. Phylogenetic reconstruction of selected species of family Agnaridae, based on the analysis of COI mtDNA gene sequences using the Bayesian analysis (BA) with the GTR + G + I evolutionary model for Maximum-Likelihood (ML), using Kimura2-parameter (K2P) model; bootstrap support for branch nodes shown for 1,000 replications." figureDoi="http://doi.org/10.5281/zenodo.4422854" httpUri="https://zenodo.org/record/4422854/files/figure.png" pageId="6" pageNumber="146">Fig. 8</figureCitation>
). However, this aspect and the question of the validity of the genera 
<taxonomicName id="4C3B4D323856FFC3FF4FFA28FE218AB4" authority="Verhoeff, 1930" authorityName="Verhoeff" authorityYear="1930" box="[151,487,1450,1476]" class="Malacostraca" family="Trachelipodidae" genus="Desertoniscus" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33856FFC3FF4FFA28FEF08AB4" box="[151,310,1451,1476]" italics="true" pageId="6" pageNumber="146">Desertoniscus</emphasis>
Verhoeff, 1930
</taxonomicName>
and 
<taxonomicName id="4C3B4D323856FFC3FDF8FA29FCFC8AB4" authority="Kwon, 1993" authorityName="Kwon" authorityYear="1993" box="[544,826,1450,1477]" class="Malacostraca" family="Trachelipodidae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEAA33856FFC3FDF8FA29FD608AB4" box="[544,678,1450,1476]" italics="true" pageId="6" pageNumber="146">Lucasioides</emphasis>
<bibRefCitation id="EFAA4B403856FFC3FD76FA28FCFC8AB4" author="Kwon, D." box="[686,826,1450,1477]" pageId="6" pageNumber="146" pagination="133 - 158" refId="ref4911" refString="Kwon, D. (1993) Terrestrial Isopoda (Crustacea) from Korea. Korean Journal of Zoology, 36, 133 - 158." type="journal article" year="1993">Kwon, 1993</bibRefCitation>
</taxonomicName>
is to be left outside present paper, requiring a special study based on a more representative material.
</paragraph>
</subSubSection>
</treatment>
</document>