treatments-xml/data/03/92/81/0392814CE909FFC3FF4EF88DFD117CF3.xml

<document id="FBDCAB45B66BBB0D4AA12632460C6D16" ID-CLB-Dataset="24730" ID-DOI="10.11646/zootaxa.4702.1.6" ID-GBIF-Dataset="45baa47f-f6a0-4a0d-afc3-476b4489b5b5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3562827" ID-ZooBank="833FCD58-12A5-4D9A-A91C-895EA854CD98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575479089760" checkinUser="plazi" docAuthor="Hata, Harutaka &amp; Motomura, Hiroyuki" docDate="2019" docId="0392814CE909FFC3FF4EF88DFD117CF3" docLanguage="en" docName="zootaxa.4702.1.6.pdf" docOrigin="Zootaxa 4702 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sardinella alcyone Hata &amp; Motomura 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FFABF934E909FFC6FFD9FFF6FFC17C0F" masterDocTitle="Sardinella alcyone n. sp., a new sardine (Teleostei: Clupeiformes: Clupeidae) from the northwestern Pacific Ocean" masterLastPageNumber="25" masterPageNumber="19" pageNumber="19" updateTime="1698771943697" updateUser="ExternalLinkService">
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<mods:title id="FD101B916EA9F7EEA604E3F73BADE3C1">Sardinella alcyone n. sp., a new sardine (Teleostei: Clupeiformes: Clupeidae) from the northwestern Pacific Ocean</mods:title>
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<mods:namePart id="4E43CFA39DFF1ACCC4731FD54343BFD3">Hata, Harutaka</mods:namePart>
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<treatment id="0392814CE909FFC3FF4EF88DFD117CF3" ID-GBIF-Taxon="160618261" LSID="urn:lsid:plazi:treatment:0392814CE909FFC3FF4EF88DFD117CF3" httpUri="http://treatment.plazi.org/id/0392814CE909FFC3FF4EF88DFD117CF3" lastPageId="5" lastPageNumber="23" pageId="0" pageNumber="19">
<subSubSection id="C32163D1E909FFC6FF4EF88DFBB47BB6" pageId="0" pageNumber="19" type="nomenclature">
<paragraph id="8B84305AE909FFC6FF4EF88DFE027B9A" blockId="0.[151,1141,1915,2013]" box="[151,451,1915,1942]" pageId="0" pageNumber="19">
<heading id="D0CC8736E909FFC6FF4EF88DFE027B9A" bold="true" box="[151,451,1915,1942]" fontSize="11" level="1" pageId="0" pageNumber="19" reason="1">
<emphasis id="B94FEC48E909FFC6FF4EF88DFE027B9A" bold="true" box="[151,451,1915,1942]" pageId="0" pageNumber="19">
<taxonomicName id="4C3B4BD9E909FFC6FF4EF88DFEB87B99" authority="Hata &amp; Motomura, 2019" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[151,377,1915,1942]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="0" pageNumber="19" phylum="Chordata" rank="species" species="alcyone" status="sp. nov.">
<emphasis id="B94FEC48E909FFC6FF4EF88DFEB87B99" bold="true" box="[151,377,1915,1942]" italics="true" pageId="0" pageNumber="19">Sardinella alcyone</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C5133E909FFC6FE59F88DFE027B9A" box="[384,451,1915,1941]" pageId="0" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B84305AE909FFC6FF4EF869FBB47BB6" blockId="0.[151,1141,1915,2013]" box="[151,1141,1950,1977]" pageId="0" pageNumber="19">
[New English name: Blueback 
<taxonomicName id="4C3B4BD9E909FFC6FE2CF869FDAB7BB6" authorityName="Valenciennes" authorityYear="1847" box="[501,618,1951,1977]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="0" pageNumber="19" phylum="Chordata" rank="genus">Sardinella</taxonomicName>
; new standard Japanese name: Shiosai-iwashi]
</paragraph>
</subSubSection>
<subSubSection id="C32163D1E909FFC6FF4EF835FEBC7BD2" box="[151,381,1987,2013]" pageId="0" pageNumber="19" type="description">
<paragraph id="8B84305AE909FFC6FF4EF835FEBC7BD2" blockId="0.[151,1141,1915,2013]" box="[151,381,1987,2013]" pageId="0" pageNumber="19">
<figureCitation id="13002CDFE909FFC6FF4EF835FEDE7BD2" box="[151,287,1987,2013]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1119,1144]" captionStartId-1="2.[151,250,1401,1426]" captionStartId-2="3.[151,250,924,949]" captionStartId-3="5.[151,250,1322,1347]" captionTargetBox-0="[151,1436,525,1094]" captionTargetBox-1="[447,1159,579,1376]" captionTargetBox-2="[159,1429,189,892]" captionTargetBox-3="[161,1422,281,1289]" captionTargetId-0="figure@1.[151,1436,525,1094]" captionTargetId-1="figure@2.[408,1179,525,1376]" captionTargetId-2="figure@3.[151,1436,181,899]" captionTargetId-3="figure@5.[151,1436,273,1297]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionText-0="FIGURE 1. Holotype of Sardinella alcyone n. sp. KAUM–I. 108333, 96.1 mm SL, Okinawa Island, Ryukyu Islands, Japan." captionText-1="FIGURE 2. Stained lateral body scale removed from Sardinella alcyone n. sp. (KAUM–I. 108335, paratype, 72.2 mm SL, Okinawa Island, Ryukyu Islands, Japan)." captionText-2="FIGURE 3. Relationships of total gill-raker numbers (TGR) on (A) first gill arch (1GA), (B) second gill arch (2GA), (C) third gill arch (3GA), and (D) fourth gill arch (4GA) relative to standard length in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." captionText-3="FIGURE 4. Relationships of (A) pectoral-fin length (as % of standard length; SL) to SL, (B) pelvic-fin length (as % of SL) to SL, (C) upper-jaw length (as % of SL) to SL, (D) lower-jaw length (as % of SL) to SL, and (E) pre-anal-fin length (as % of SL) to SL in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi-0="http://doi.org/10.5281/zenodo.3562829" figureDoi-1="http://doi.org/10.5281/zenodo.3562831" figureDoi-2="http://doi.org/10.5281/zenodo.3562833" figureDoi-3="http://doi.org/10.5281/zenodo.3562835" httpUri-0="https://zenodo.org/record/3562829/files/figure.png" httpUri-1="https://zenodo.org/record/3562831/files/figure.png" httpUri-2="https://zenodo.org/record/3562833/files/figure.png" httpUri-3="https://zenodo.org/record/3562835/files/figure.png" pageId="0" pageNumber="19">Figures 1–4</figureCitation>
; 
<tableCitation id="C6B905E1E909FFC6FEF3F835FEBC7BD2" box="[298,381,1987,2013]" captionStart="TABLE 1" captionStartId="3.[151,244,1102,1129]" captionTargetBox="[163,1384,1144,1974]" captionText="TABLE 1. Counts and measurements of specimens of Sardinella alcyone n. sp." httpUri="http://table.plazi.org/id/DF4460D2E90AFFC5FF4EFBB8FBD57866" pageId="0" pageNumber="19" tableUuid="DF4460D2E90AFFC5FF4EFBB8FBD57866">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32163D1E908FFC7FF4EFF6FFCE77DF8" pageId="1" pageNumber="20" type="materials_examined">
<paragraph id="8B84305AE908FFC7FF4EFF6FFB6B7CD8" blockId="1.[151,1437,153,503]" pageId="1" pageNumber="20">
<materialsCitation id="3B533A07E908FFC7FF4EFF6FFB6B7CD8" ID-GBIF-Occurrence="2462935307" collectingDate="2017-03-12" collectionCode="KAUM-I" collectorName="K. Shibukawa &amp; M. Nakae." country="Japan" county="Ryukyu Islands" elevation="1" latitude="26.634722" location="Nasada River" longLatPrecision="20" longitude="128.00194" municipality="Okinawa Island" pageId="1" pageNumber="20" specimenCount="1" typeStatus="holotype">
<emphasis id="B94FEC48E908FFC7FF4EFF6FFECA7CBC" bold="true" box="[151,267,153,179]" pageId="1" pageNumber="20">
<typeStatus id="54808EF8E908FFC7FF4EFF6FFEC77CBC" box="[151,262,153,179]" pageId="1" pageNumber="20" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED2AA89FE908FFC7FEC8FF6FFE437CBC" box="[273,386,153,179]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 108333, 
<quantity id="4CC39DBFE908FFC7FE34FF6FFD917CBC" box="[493,592,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="9.61" pageId="1" pageNumber="20" unit="mm" value="96.1">96.1 mm</quantity>
SL, mouth of 
<location id="8EE46681E908FFC7FD29FF6FFC487CBC" LSID="urn:lsid:plazi:treatment:0392814CE909FFC3FF4EF88DFD117CF3:8EE46681E908FFC7FD29FF6FFC487CBC" box="[752,905,153,179]" country="Japan" county="Ryukyu Islands" latitude="26.634722" longLatPrecision="20" longitude="128.00194" municipality="Okinawa Island" name="Nasada River" pageId="1" pageNumber="20">Nasada River</location>
, 
<location id="8EE46681E908FFC7FC4AFF6FFC137CBC" LSID="urn:lsid:plazi:treatment:0392814CE909FFC3FF4EF88DFD117CF3:8EE46681E908FFC7FC4AFF6FFC137CBC" box="[915,978,153,179]" country="Japan" county="Ryukyu Islands" latitude="26.634722" longLatPrecision="20" longitude="128.00194" municipality="Okinawa Island" name="Nago" pageId="1" pageNumber="20">Nago</location>
, 
<collectingMunicipality id="6BE0AA20E908FFC7FC04FF6FFB517CBC" box="[989,1168,153,179]" pageId="1" pageNumber="20">Okinawa Island</collectingMunicipality>
, 
<collectingCounty id="62E548D6E908FFC7FB42FF6FFA8A7CBC" box="[1179,1355,153,179]" pageId="1" pageNumber="20">Ryukyu Islands</collectingCounty>
, 
<collectingCountry id="F32C70CAE908FFC7FA8CFF6FFA597CBC" box="[1365,1432,153,179]" name="Japan" pageId="1" pageNumber="20">Japan</collectingCountry>
, 
<geoCoordinate id="EE0F569DE908FFC7FF4EFF4AFEDE7CD8" box="[151,287,188,215]" degrees="26" direction="north" minutes="38" orientation="latitude" pageId="1" pageNumber="20" precision="15" seconds="05" value="26.634722">26°38′05″N</geoCoordinate>
, 
<geoCoordinate id="EE0F569DE908FFC7FEF3FF4AFE7C7CD8" box="[298,445,188,215]" degrees="128" direction="east" minutes="00" orientation="longitude" pageId="1" pageNumber="20" precision="15" seconds="07" value="128.00194">128°00′07″E</geoCoordinate>
, 
<elevation id="0016D769E908FFC7FE10FF4BFDFA7CD8" box="[457,571,189,215]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="20" unit="m" value="1.0">
<quantity id="4CC39DBFE908FFC7FE10FF4BFE357CD8" box="[457,500,189,215]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="20" unit="m" value="1.0">1 m</quantity>
depth
</elevation>
, 
<date id="FF85169AE908FFC7FD9EFF4BFD217CD8" box="[583,736,189,215]" pageId="1" pageNumber="20" value="2017-03-12">
<collectingDate id="EFC1EF72E908FFC7FD9EFF4BFD217CD8" box="[583,736,189,215]" pageId="1" pageNumber="20" value="2017-03-12">12 Mar. 2017</collectingDate>
</date>
, 
<collectingMethod id="527A484DE908FFC7FD32FF4BFCA77CD8" box="[747,870,189,215]" pageId="1" pageNumber="20">casting net</collectingMethod>
, 
<collectorName id="26CE558CE908FFC7FCA8FF4BFBCE7CD8" box="[881,1039,189,215]" pageId="1" pageNumber="20">K. Shibukawa</collectorName>
&amp; 
<collectorName id="26CE558CE908FFC7FBEDFF4BFB6B7CD8" box="[1076,1194,189,215]" pageId="1" pageNumber="20">M. Nakae.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B84305AE908FFC7FF1EFF17FCE77DF8" blockId="1.[151,1437,153,503]" pageId="1" pageNumber="20">
<materialsCitation id="3B533A07E908FFC7FF1EFF17FCE37DF8" ID-GBIF-Occurrence="2462935308" collectingDate="2000-11-09" collectingDateMax="2017-03-11" collectingDateMin="2000-11-09" collectionCode="KAUM-I, URM-P" collectorName="K. Shibukawa &amp; M. Nakae." country="Japan" county="Nakaoji" elevation="30" latitude="26.634722" location="Island" longLatPrecision="20" longitude="128.00194" municipality="Haneji" pageId="1" pageNumber="20" specimenCode="SPMN-NI 44303, SPMN-PI 44304, SPMN-PI 44305, NI 44306, URM-P 41020, URM-P 41021, URM-P 41022, URM-P 41023, BMNH 1979.3" specimenCount="18" stateProvince="Tainan" typeStatus="paratype">
<emphasis id="B94FEC48E908FFC7FF1EFF17FE877CF4" bold="true" box="[199,326,225,251]" pageId="1" pageNumber="20">
<typeStatus id="54808EF8E908FFC7FF1EFF17FE807CF4" box="[199,321,225,251]" pageId="1" pageNumber="20" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D3DFBD3E908FFC7FE97FF17FE2B7CF4" box="[334,490,225,251]" pageId="1" pageNumber="20" type="generic">18 specimens</specimenCount>
, 
<quantity id="4CC39DBFE908FFC7FE2FFF17FD697CF4" box="[502,680,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="8.819999999999999" metricValueMax="10.98" metricValueMin="6.659999999999999" pageId="1" pageNumber="20" unit="mm" value="88.19999999999999" valueMax="109.8" valueMin="66.6">66.6–109.8 mm</quantity>
SL. 
<emphasis id="B94FEC48E908FFC7FD06FF16FC837CF4" bold="true" box="[735,834,224,251]" pageId="1" pageNumber="20">
<collectingCountry id="F32C70CAE908FFC7FD06FF16FCFC7CF4" box="[735,829,224,251]" name="Japan" pageId="1" pageNumber="20">JAPAN</collectingCountry>
:
</emphasis>
<collectionCode id="ED2AA89FE908FFC7FC93FF17FC7A7CF4" box="[842,955,225,251]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 108334, 
<quantity id="4CC39DBFE908FFC7FBF0FF17FB4F7CF4" box="[1065,1166,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="7.84" pageId="1" pageNumber="20" unit="mm" value="78.4">78.4 mm</quantity>
SL, 
<collectionCode id="ED2AA89FE908FFC7FB1CFF17FAF77CF4" box="[1221,1334,225,251]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 108336, 
<quantity id="4CC39DBFE908FFC7FF4EFEF3FF3D7D10" box="[151,252,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="7.93" pageId="1" pageNumber="20" unit="mm" value="79.3">79.3 mm</quantity>
SL, 
<specimenCode id="DB9D9821E908FFC7FEEAFEF3FE397D10" box="[307,504,261,287]" collectionCode="SPMN-NI" pageId="1" pageNumber="20">SPMN-NI 44303</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FDDDFEF3FDAA7D10" box="[516,619,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="7.74" pageId="1" pageNumber="20" unit="mm" value="77.4">77.4 mm</quantity>
, 
<specimenCode id="DB9D9821E908FFC7FDAEFEF3FCF67D10" box="[631,823,261,287]" collectionCode="SPMN-PI" pageId="1" pageNumber="20">SPMN-PI 44304</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FC9AFEF3FC697D10" box="[835,936,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="7.39" pageId="1" pageNumber="20" unit="mm" value="73.9">73.9 mm</quantity>
SL, 
<specimenCode id="DB9D9821E908FFC7FC06FEF3FB5F7D10" box="[991,1182,260,287]" collectionCode="SPMN-PI" pageId="1" pageNumber="20">SPMN-PI 44305</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FB73FEF3FACE7D10" box="[1194,1295,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="7.99" pageId="1" pageNumber="20" unit="mm" value="79.9">79.9 mm</quantity>
SL, SPMN- 
<specimenCode id="DB9D9821E908FFC7FF4EFEDFFEC77D4C" box="[151,262,297,323]" collectionCode="NI" country="Japan" name="Nagao Institute" pageId="1" pageNumber="20">NI 44306</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FECAFEDFFEB87D4C" box="[275,377,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="6.659999999999999" pageId="1" pageNumber="20" unit="mm" value="66.6">66.6 mm</quantity>
SL, collected with the 
<typeStatus id="54808EF8E908FFC7FD5DFEDFFD2B7D4C" box="[644,746,297,323]" pageId="1" pageNumber="20" type="holotype">holotype</typeStatus>
; 
<collectionCode id="ED2AA89FE908FFC7FD2FFEDFFCA67D4C" box="[758,871,297,323]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 108335, 
<quantity id="4CC39DBFE908FFC7FC0FFEDFFBFD7D4C" box="[982,1084,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="7.22" pageId="1" pageNumber="20" unit="mm" value="72.2">72.2 mm</quantity>
SL, same locality as holotype, collected in 
<date id="FF85169AE908FFC7FEFCFEBBFE007D68" box="[293,449,333,359]" pageId="1" pageNumber="20" value="2017-03-11">
<collectingDate id="EFC1EF72E908FFC7FEFCFEBBFE007D68" box="[293,449,333,359]" pageId="1" pageNumber="20" value="2017-03-11">11 Mar. 2017</collectingDate>
</date>
; 
<specimenCode id="DB9D9821E908FFC7FE09FEBBFDBB7D68" box="[464,634,333,359]" collectionCode="URM-P" pageId="1" pageNumber="20">URM-P 41020</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FD5EFEBAFD2C7D68" box="[647,749,332,359]" metricMagnitude="-2" metricUnit="m" metricValue="8.15" pageId="1" pageNumber="20" unit="mm" value="81.5">81.5 mm</quantity>
SL, 
<specimenCode id="DB9D9821E908FFC7FCFEFEBBFC107D68" box="[807,977,333,359]" collectionCode="URM-P" pageId="1" pageNumber="20">URM-P 41021</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FC06FEBBFB847D68" box="[991,1093,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="9.08" pageId="1" pageNumber="20" unit="mm" value="90.8">90.8 mm</quantity>
SL, 
<specimenCode id="DB9D9821E908FFC7FBA6FEBBFAE87D68" box="[1151,1321,333,359]" collectionCode="URM-P" pageId="1" pageNumber="20">URM-P 41022</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FAEFFEBBFA5C7D68" box="[1334,1437,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="8.94" pageId="1" pageNumber="20" unit="mm" value="89.4">89.4 mm</quantity>
SL, 
<specimenCode id="DB9D9821E908FFC7FF11FE87FEB07D84" box="[200,369,369,395]" collectionCode="URM-P" pageId="1" pageNumber="20">URM-P 41023</specimenCode>
, 
<quantity id="4CC39DBFE908FFC7FEA7FE87FE257D84" box="[382,484,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="9.82" pageId="1" pageNumber="20" unit="mm" value="98.2">98.2 mm</quantity>
SL, 
<collectionCode id="ED2AA89FE908FFC7FDC5FE87FDB57D84" box="[540,628,369,395]" pageId="1" pageNumber="20">URM-P</collectionCode>
<quantity id="4CC39DBFE908FFC7FDA4FE87FC867D84" box="[637,839,368,395]" metricMagnitude="4" metricUnit="m" metricValue="4.10241065" pageId="1" pageNumber="20" unit="mm" value="4.10241065E7">41024, 106.5 mm</quantity>
SL, 
<collectionCode id="ED2AA89FE908FFC7FCA6FE87FC167D84" box="[895,983,369,395]" pageId="1" pageNumber="20">URM-P</collectionCode>
<quantity id="4CC39DBFE908FFC7FC39FE86FB687D84" box="[992,1193,368,395]" metricMagnitude="4" metricUnit="m" metricValue="4.10251035" pageId="1" pageNumber="20" unit="mm" value="4.10251035E7">41025, 103.5 mm</quantity>
SL, 
<collectingCounty id="62E548D6E908FFC7FB3BFE87FAFF7D84" box="[1250,1342,369,395]" pageId="1" pageNumber="20">Nakaoji</collectingCounty>
, 
<collectingMunicipality id="6BE0AA20E908FFC7FA92FE87FA587D84" box="[1355,1433,369,395]" pageId="1" pageNumber="20">Haneji</collectingMunicipality>
, Okinawa 
<location id="8EE46681E908FFC7FEDBFE63FE887DA0" LSID="urn:lsid:plazi:treatment:0392814CE909FFC3FF4EF88DFD117CF3:8EE46681E908FFC7FEDBFE63FE887DA0" box="[258,329,405,431]" country="Japan" county="Nakaoji" municipality="Haneji" name="Island" pageId="1" pageNumber="20" stateProvince="Tainan">Island</location>
, 
<location id="8EE46681E908FFC7FE8DFE63FDC27DA0" LSID="urn:lsid:plazi:treatment:0392814CE909FFC3FF4EF88DFD117CF3:8EE46681E908FFC7FE8DFE63FDC27DA0" box="[340,515,405,431]" country="Japan" county="Nakaoji" municipality="Haneji" name="Ryukyu Islands" pageId="1" pageNumber="20" stateProvince="Tainan">Ryukyu Islands</location>
, 
<date id="FF85169AE908FFC7FDD7FE63FD587DA0" box="[526,665,405,431]" pageId="1" pageNumber="20" value="2000-11-09">
<collectingDate id="EFC1EF72E908FFC7FDD7FE63FD587DA0" box="[526,665,405,431]" pageId="1" pageNumber="20" value="2000-11-09">9 Nov. 2000</collectingDate>
</date>
. 
<emphasis id="B94FEC48E908FFC7FD7BFE62FCDC7DA0" bold="true" box="[674,797,404,431]" pageId="1" pageNumber="20">
<collectingCountry id="F32C70CAE908FFC7FD7BFE62FCD97DA0" box="[674,792,404,431]" name="Taiwan" pageId="1" pageNumber="20">TAIWAN</collectingCountry>
:
</emphasis>
<specimenCode id="DB9D9821E908FFC7FCFAFE63FC0D7DA0" box="[803,972,405,431]" collectionCode="BMNH" pageId="1" pageNumber="20">BMNH 1979.3</specimenCode>
.21.333, 
<quantity id="4CC39DBFE908FFC7FBF2FE63FB5D7DA0" box="[1067,1180,405,431]" metricMagnitude="-1" metricUnit="m" metricValue="1.098" pageId="1" pageNumber="20" unit="mm" value="109.8">109.8 mm</quantity>
SL, 
<collectingRegion id="49FFFEB8E908FFC7FB16FE63FADF7DA0" box="[1231,1310,405,431]" country="Taiwan" name="Tainan" pageId="1" pageNumber="20">Tainan</collectingRegion>
; 
<collectionCode id="ED2AA89FE908FFC7FAF1FE63FA587DA0" box="[1320,1433,405,431]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 113218, 
<quantity id="4CC39DBFE908FFC7FF20FE4FFE9E7DDC" box="[249,351,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="7.77" pageId="1" pageNumber="20" unit="mm" value="77.7">77.7 mm</quantity>
SL, 
<collectionCode id="ED2AA89FE908FFC7FE4FFE4FFDC67DDC" box="[406,519,441,467]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 113219, 
<quantity id="4CC39DBFE908FFC7FDACFE4FFD1B7DDC" box="[629,730,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="8.36" pageId="1" pageNumber="20" unit="mm" value="83.6">83.6 mm</quantity>
SL, 
<collectionCode id="ED2AA89FE908FFC7FCCBFE4FFC427DDC" box="[786,899,441,467]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 113782, 
<quantity id="4CC39DBFE908FFC7FC29FE4EFB947DDC" box="[1008,1109,440,467]" metricMagnitude="-2" metricUnit="m" metricValue="9.05" pageId="1" pageNumber="20" unit="mm" value="90.5">90.5 mm</quantity>
SL, 
<collectionCode id="ED2AA89FE908FFC7FB54FE4FFB3F7DDC" box="[1165,1278,441,467]" pageId="1" pageNumber="20">KAUM–I</collectionCode>
. 113784, 
<quantity id="4CC39DBFE908FFC7FAB2FE4FFF027DF8" metricMagnitude="-2" metricUnit="m" metricValue="8.03" pageId="1" pageNumber="20" unit="mm" value="80.3">80.3 mm</quantity>
SL, off 
<location id="8EE46681E908FFC7FEF9FE2BFE607DF8" LSID="urn:lsid:plazi:treatment:0392814CE909FFC3FF4EF88DFD117CF3:8EE46681E908FFC7FEF9FE2BFE607DF8" box="[288,417,477,503]" country="Japan" county="Nakaoji" municipality="Haneji" name="Dong-gang" pageId="1" pageNumber="20" stateProvince="Tainan">Dong-gang</location>
, 
<collectingRegion id="49FFFEB8E908FFC7FE75FE2BFDAD7DF8" box="[428,620,477,503]" country="Taiwan" name="Pingtung" pageId="1" pageNumber="20">Pingtung County</collectingRegion>
, 
<elevation id="0016D769E908FFC7FDAFFE2AFCE37DF8" box="[630,802,476,503]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="1" pageNumber="20" unit="m" value="30.0" valueMax="50.0" valueMin="10.0">
<quantity id="4CC39DBFE908FFC7FDAFFE2AFD187DF8" box="[630,729,476,503]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="1" pageNumber="20" unit="m" value="30.0" valueMax="50.0" valueMin="10.0">10–50 m</quantity>
depth
</elevation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF4460D2E908FFC7FF4EFBA9FA467878" ID-DOI="http://doi.org/10.5281/zenodo.3562829" ID-Zenodo-Dep="3562829" box="[151,1415,1119,1144]" httpUri="https://zenodo.org/record/3562829/files/figure.png" pageId="1" pageNumber="20" startId="1.[151,250,1119,1144]" targetBox="[151,1436,525,1094]" targetPageId="1">
<paragraph id="8B84305AE908FFC7FF4EFBA9FA467878" blockId="1.[151,1415,1119,1144]" box="[151,1415,1119,1144]" pageId="1" pageNumber="20">
<emphasis id="B94FEC48E908FFC7FF4EFBA9FED27877" bold="true" box="[151,275,1119,1144]" pageId="1" pageNumber="20">FIGURE 1.</emphasis>
Holotype of 
<taxonomicName id="4C3B4BD9E908FFC7FE40FB96FD967877" authority="Hata &amp; Motomura, 2019" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[409,599,1120,1144]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="alcyone" status="sp. nov.">
<emphasis id="B94FEC48E908FFC7FE40FB96FD967877" box="[409,599,1120,1144]" italics="true" pageId="1" pageNumber="20">Sardinella alcyone</emphasis>
</taxonomicName>
<emphasis id="B94FEC48E908FFC7FD84FB96FD567877" bold="true" box="[605,663,1120,1144]" pageId="1" pageNumber="20">
<taxonomicNameLabel id="A27C5133E908FFC7FD84FB96FD567877" box="[605,663,1120,1144]" pageId="1" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
KAUM–I. 108333, 96.1 mm SL, Okinawa Island, Ryukyu Islands, Japan.
</paragraph>
</caption>
<subSubSection id="C32163D1E908FFC7FF1EFB3FFB8C7A27" pageId="1" pageNumber="20" type="diagnosis">
<paragraph id="8B84305AE908FFC7FF1EFB3FFB8C7A27" blockId="1.[151,1437,1225,2045]" pageId="1" pageNumber="20">
<emphasis id="B94FEC48E908FFC7FF1EFB3FFE8378EB" bold="true" box="[199,322,1225,1252]" pageId="1" pageNumber="20">Diagnosis.</emphasis>
A species of 
<taxonomicName id="4C3B4BD9E908FFC7FE0BFB3CFD8778EB" authorityName="Valenciennes" authorityYear="1847" box="[466,582,1226,1252]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="1" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B94FEC48E908FFC7FE0BFB3CFD8778EB" box="[466,582,1226,1252]" italics="true" pageId="1" pageNumber="20">Sardinella</emphasis>
</taxonomicName>
with the following combination of characters: caudal fin with black posterior margin; lateral body scales with centrally discontinuous vertical striae, and few perforations and pores posteriorly; 38–41 (modally 39) scale rows in longitudinal series; body scales deciduous; black spot on dorsal-fin origin; pelvic fin with one unbranched and seven branched rays; gill rakers 34–42 (39) in upper series on 1st gill arch, 64–72 (67) in lower series, 99–112 (103) in total; gill rakers 33–40 (35) in upper series on 2nd gill arch, 64–76 (74) in lower series, 97–115 (101) in total; gill rakers 31–38 (33) in upper series on 3rd gill arch, 48–60 (57) in lower series, 79–98 (84) in total; gill rakers 24–31 (27) in upper series on 4th gill arch, 37–47 (39) in lower, 62–77 (68) in total; gill rakers 25–31 (27) on hind face of third gill arch; 17 or 18 (18) + 12–14 (13) = 30–32 (31) scutes on ventral of body; pectoral fin long, 20.7–23.4% SL; pelvic fin long, 11.9–13.1% SL; upper jaw moderately long, 10.8–12.4% SL; lower jaw moderately long, 11.8–13.4% SL; pre-anal-fin length 77.4–82.3% SL.
</paragraph>
</subSubSection>
<subSubSection id="C32163D1E908FFC4FF1EF9C7FB617AF3" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="description">
<paragraph id="8B84305AE908FFC4FF1EF9C7FD7F7DF8" blockId="1.[151,1437,1225,2045]" lastBlockId="2.[151,1437,153,503]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<emphasis id="B94FEC48E908FFC7FF1EF9C7FE987A43" bold="true" box="[199,345,1585,1612]" pageId="1" pageNumber="20">Description.</emphasis>
Counts and measurements, expressed as percentages of SL, are given in 
<tableCitation id="C6B905E1E908FFC7FB53F9C4FB1C7A43" box="[1162,1245,1586,1612]" captionStart="TABLE 1" captionStartId="3.[151,244,1102,1129]" captionTargetBox="[163,1384,1144,1974]" captionText="TABLE 1. Counts and measurements of specimens of Sardinella alcyone n. sp." httpUri="http://table.plazi.org/id/DF4460D2E90AFFC5FF4EFBB8FBD57866" pageId="1" pageNumber="20" tableUuid="DF4460D2E90AFFC5FF4EFBB8FBD57866">Table 1</tableCitation>
. Data for the 
<typeStatus id="54808EF8E908FFC7FAAEF9C4FF1C7A7F" pageId="1" pageNumber="20" type="holotype">holotype</typeStatus>
are presented first, followed by 
<typeStatus id="54808EF8E908FFC7FD9DF9A1FD657A7E" box="[580,676,1623,1649]" pageId="1" pageNumber="20" type="paratype">paratype</typeStatus>
data in parentheses. Body oblong, compressed, deepest at dorsal-fin origin. Dorsal profile of body elevated from snout tip to dorsal-fin origin, thereafter decreasing to uppermost point of caudal-fin base. Ventral profile of body curved downward from lower-jaw tip to pelvic-fin insertion, thereafter rounded to ventralmost point of caudal-fin base. Anus on ventral midline, slightly anterior to anal-fin origin, posterior to midpoint of body. Mouth terminal, small, posterior tip of maxilla not reaching vertical through anterior margin of iris. Premaxilla and hypomaxilla without teeth. Ventral margin of maxilla toothed. Lower jaw with several conical teeth anteriorly. Posterior ramus of lower jaw elevated. Second supramaxilla symmetrical. Nostrils close to each other, anterior to orbit. Orbit elliptical, eye and iris round. Eyes covered with adipose eyelid posteriorly. Interorbital space flat. ten (seven to 11) bony striae on top of head. Gill rakers long, slender, with small asperities on anterior surface. Pseudobranchial filaments present. Gill opening with two fleshy outgrowths on posterior margin, a large papilla on ventral margin. Posterior margins of preopercle and opercle smooth. Scales cycloid, thin, deciduous, except for robust ventral scutes. Scales on lateral body surface with several centrally discontinuous vertical striae, few perforations and pores posteriorly (
<figureCitation id="13002CDFE90BFFC4FD95FF6FFD517CBC" box="[588,656,153,179]" captionStart="FIGURE 2" captionStartId="2.[151,250,1401,1426]" captionTargetBox="[447,1159,579,1376]" captionTargetId="figure@2.[408,1179,525,1376]" captionTargetPageId="2" captionText="FIGURE 2. Stained lateral body scale removed from Sardinella alcyone n. sp. (KAUM–I. 108335, paratype, 72.2 mm SL, Okinawa Island, Ryukyu Islands, Japan)." figureDoi="http://doi.org/10.5281/zenodo.3562831" httpUri="https://zenodo.org/record/3562831/files/figure.png" pageId="2" pageNumber="21">Fig. 2</figureCitation>
). Bases of dorsal and anal fins with low scaly sheaths. Predorsal scales paired. No elongate, wing-like scales present beneath normal paired scales. No scales on head and fins, except for a broad triangular sheath of scales on caudal fin. Abdomen from isthmus to anus with 31 (30 to 32) scutes. Predorsal scutes absent. No lateral line. Anteriormost point of pectoral-fin insertion anterior to posteriormost point of opercle. Upper, posterior and ventral margins of pectoral fin nearly straight. Posterior tip of pectoral fin pointed. Pectoralfin axillary scale present. Posteriormost dorsal-fin ray not filamentous. Anteriormost point of pelvic-fin insertion located directly below base of 7th (7th–10th) dorsal-fin ray. Posterior tip of depressed pelvic fin reaching between vertical through posterior end of dorsal-fin base and anus. Pelvic-fin axillary scale present. Anal-fin origin posterior to vertical through posteriormost point of dorsal-fin base. Two posteriormost anal-fin rays enlarged. Caudal fin forked. Posterior tips of caudal-fin lobes pointed.
</paragraph>
<caption id="DF4460D2E90BFFC4FF4EFA8FFDF679BA" ID-DOI="http://doi.org/10.5281/zenodo.3562831" ID-Zenodo-Dep="3562831" httpUri="https://zenodo.org/record/3562831/files/figure.png" pageId="2" pageNumber="21" startId="2.[151,250,1401,1426]" targetBox="[447,1159,579,1376]" targetPageId="2">
<paragraph id="8B84305AE90BFFC4FF4EFA8FFDF679BA" blockId="2.[151,1436,1401,1462]" pageId="2" pageNumber="21">
<emphasis id="B94FEC48E90BFFC4FF4EFA8FFED4799D" bold="true" box="[151,277,1401,1426]" pageId="2" pageNumber="21">FIGURE 2.</emphasis>
Stained lateral body scale removed from 
<taxonomicName id="4C3B4BD9E90BFFC4FD15FA8FFC4D799E" authority="Hata &amp; Motomura, 2019" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[716,908,1401,1425]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="alcyone" status="sp. nov.">
<emphasis id="B94FEC48E90BFFC4FD15FA8FFC4D799E" box="[716,908,1401,1425]" italics="true" pageId="2" pageNumber="21">Sardinella alcyone</emphasis>
</taxonomicName>
<emphasis id="B94FEC48E90BFFC4FC4CFA8FFC11799E" bold="true" box="[917,976,1401,1425]" pageId="2" pageNumber="21">
<taxonomicNameLabel id="A27C5133E90BFFC4FC4CFA8FFC11799E" box="[917,976,1401,1425]" pageId="2" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(KAUM–I. 108335, paratype, 72.2 mm SL, Okinawa Island, Ryukyu Islands, Japan).
</paragraph>
</caption>
<paragraph id="8B84305AE90BFFC4FF1EF9FEFCA17ABD" blockId="2.[151,1437,1544,1932]" pageId="2" pageNumber="21">
<emphasis id="B94FEC48E90BFFC4FF1EF9FEFE037A2D" box="[199,450,1544,1570]" italics="true" pageId="2" pageNumber="21">Coloration when fresh</emphasis>
. [Based on color photographs of seven specimens (KAUM–I. 113219, 108333, 108334, 108335, 108336, and SPMN-PI 44303, 44304)]. Dorsum bluish-green (or bluish-black). Lateral surface of body yellowish-silver, upper part metallic blueish. Black spot on dorsal-fin origin. Dorsal fin yellow, melanophores scattered on upper part of fin. Pectoral, pelvic, and anal fins yellowish (or translucent whitish). Caudal fin yellowish, posterior margin black. Black melanophores scattered on caudal-fin rays.
</paragraph>
<paragraph id="8B84305AE90BFFC4FF1EF94AFB617AF3" blockId="2.[151,1437,1544,1932]" pageId="2" pageNumber="21">
<emphasis id="B94FEC48E90BFFC4FF1EF94AFDD57AD9" box="[199,532,1724,1750]" italics="true" pageId="2" pageNumber="21">Color of preserved specimens</emphasis>
. Body dark brown dorsally, elsewhere yellowish-silver. Black spot on dorsal-fin origin and distal tips of caudal fin lobes. Melanophores scattered on upper part of dorsal fin.
</paragraph>
</subSubSection>
<subSubSection id="C32163D1E90BFFC4FF1EF8F3FCDC7B4B" pageId="2" pageNumber="21" type="distribution">
<paragraph id="8B84305AE90BFFC4FF1EF8F3FCDC7B4B" blockId="2.[151,1437,1544,1932]" pageId="2" pageNumber="21">
<emphasis id="B94FEC48E90BFFC4FF1EF8F3FEA07B2F" bold="true" box="[199,353,1797,1824]" pageId="2" pageNumber="21">Distribution.</emphasis>
Currently known only from Okinawa Island, Ryukyu Islands, 
<collectingCountry id="F32C70CAE90BFFC4FBF4F8F1FBAC7B2E" box="[1069,1133,1799,1825]" name="Japan" pageId="2" pageNumber="21">Japan</collectingCountry>
and southwestern 
<collectingCountry id="F32C70CAE90BFFC4FA9AF8F0FA587B2F" box="[1347,1433,1798,1824]" name="Taiwan" pageId="2" pageNumber="21">Taiwan</collectingCountry>
. The species inhabits coastal shallow waters and estuaries.
</paragraph>
</subSubSection>
<subSubSection id="C32163D1E90BFFC4FF1EF8B8FE2B7B83" pageId="2" pageNumber="21" type="etymology">
<paragraph id="8B84305AE90BFFC4FF1EF8B8FE2B7B83" blockId="2.[151,1437,1544,1932]" pageId="2" pageNumber="21">
<emphasis id="B94FEC48E90BFFC4FF1EF8B8FE8F7B67" bold="true" box="[199,334,1870,1896]" pageId="2" pageNumber="21">Etymology.</emphasis>
The specific name 
<taxonomicName id="4C3B4BD9E90BFFC4FDFDF8B8FDBA7B67" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[548,635,1870,1896]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="alcyone">
<emphasis id="B94FEC48E90BFFC4FDFDF8B8FDBA7B67" box="[548,635,1870,1896]" italics="true" pageId="2" pageNumber="21">alcyone</emphasis>
</taxonomicName>
is derived from Greek meaning “kingfisher”, in reference to the brilliant blueish dorsum of the species.
</paragraph>
</subSubSection>
<caption id="DF4460D2E90AFFC5FF4EFC6AFE577FF2" ID-DOI="http://doi.org/10.5281/zenodo.3562833" ID-Zenodo-Dep="3562833" httpUri="https://zenodo.org/record/3562833/files/figure.png" pageId="3" pageNumber="22" startId="3.[151,250,924,949]" targetBox="[159,1429,189,892]" targetPageId="3">
<paragraph id="8B84305AE90AFFC5FF4EFC6AFE577FF2" blockId="3.[151,1437,924,1021]" pageId="3" pageNumber="22">
<emphasis id="B94FEC48E90AFFC5FF4EFC6AFED27FBA" bold="true" box="[151,275,924,949]" pageId="3" pageNumber="22">FIGURE 3.</emphasis>
Relationships of total gill-raker numbers (TGR) on (A) first gill arch (1GA), (B) second gill arch (2GA), (C) third gill arch (3GA), and (D) fourth gill arch (4GA) relative to standard length in 
<taxonomicName id="4C3B4BD9E90AFFC5FC6AFC37FBB37FD6" authority="Hata &amp; Motomura, 2019" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[947,1138,961,985]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="alcyone" status="sp. nov.">
<emphasis id="B94FEC48E90AFFC5FC6AFC37FBB37FD6" box="[947,1138,961,985]" italics="true" pageId="3" pageNumber="22">Sardinella alcyone</emphasis>
</taxonomicName>
<emphasis id="B94FEC48E90AFFC5FBA0FC37FB727FD6" bold="true" box="[1145,1203,961,985]" pageId="3" pageNumber="22">
<taxonomicNameLabel id="A27C5133E90AFFC5FBA0FC37FB727FD6" box="[1145,1203,961,985]" pageId="3" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(closed circles) and 
<taxonomicName id="4C3B4BD9E90AFFC5FA50FC37FF287FF2" authorityName="Hata &amp; Motomura" authorityYear="2019" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="pacifica">
<emphasis id="B94FEC48E90AFFC5FA50FC37FF287FF2" italics="true" pageId="3" pageNumber="22">S. pacifica</emphasis>
</taxonomicName>
(open triangles).
</paragraph>
</caption>
<caption id="DF4460D2E90AFFC5FF4EFBB8FBD57866" ID-Table-UUID="DF4460D2E90AFFC5FF4EFBB8FBD57866" box="[151,1044,1102,1130]" httpUri="http://table.plazi.org/id/DF4460D2E90AFFC5FF4EFBB8FBD57866" pageId="3" pageNumber="22" startId="3.[151,244,1102,1129]" targetBox="[163,1384,1144,1974]" targetIsTable="true" targetPageId="3">
<paragraph id="8B84305AE90AFFC5FF4EFBB8FBD57866" blockId="3.[151,1044,1102,1130]" box="[151,1044,1102,1130]" pageId="3" pageNumber="22">
<emphasis id="B94FEC48E90AFFC5FF4EFBB8FED07866" bold="true" box="[151,273,1102,1129]" pageId="3" pageNumber="22">
<tableCitation id="C6B905E1E90AFFC5FF4EFBB8FEC87866" box="[151,265,1102,1129]" captionStart="TABLE 1" captionStartId="4.[163,247,163,188]" captionTargetBox="[163,1384,204,1597]" captionText="TABLE 1. (Continued)" pageId="3" pageNumber="22">TABLE 1</tableCitation>
.
</emphasis>
Counts and measurements of specimens of 
<taxonomicName id="4C3B4BD9E90AFFC5FD25FBB9FC0F7866" authority="Hata &amp; Motomura, 2019" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[764,974,1103,1129]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="alcyone" status="sp. nov.">
<emphasis id="B94FEC48E90AFFC5FD25FBB9FC0F7866" box="[764,974,1103,1129]" italics="true" pageId="3" pageNumber="22">Sardinella alcyone</emphasis>
</taxonomicName>
<emphasis id="B94FEC48E90AFFC5FC0CFBB9FBD57866" bold="true" box="[981,1044,1103,1129]" pageId="3" pageNumber="22">
<taxonomicNameLabel id="A27C5133E90AFFC5FC0CFBB9FBD57866" box="[981,1044,1103,1129]" pageId="3" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</caption>
<paragraph id="8B84305AE90AFFC5FCCDFB8EFA937BB9" pageId="3" pageNumber="22">
<table id="F93BC2FAE90A0039FF7AFB8EFAA97BB9" box="[163,1384,1144,1974]" colsContinueIn="4.[163,1384,204,1597]" gridcols="4" gridrows="21" pageId="3" pageNumber="22">
<tr id="350B3218E90A0039FF7AFB8EFAA9789D" box="[163,1384,1144,1170]" gridrow="0" pageId="3" pageNumber="22" rowspan-0="1" rowspan-3="1">
<th id="76DA5B64E90A0039FD3CFB8EFC61789D" box="[741,928,1144,1170]" gridcol="1" gridrow="0" pageId="3" pageNumber="22">Holotype</th>
<th id="76DA5B64E90A0039FBDBFB8EFB76789D" box="[1026,1207,1144,1170]" gridcol="2" gridrow="0" pageId="3" pageNumber="22">Paratypes</th>
</tr>
<tr id="350B3218E90A0039FF7AFB57FAA978B5" box="[163,1384,1185,1210]" gridrow="1" pageId="3" pageNumber="22" rowspan-0="1" rowspan-3="1">
<td id="76DA5B64E90A0039FD3CFB57FC6178B5" box="[741,928,1185,1210]" gridcol="1" gridrow="1" pageId="3" pageNumber="22">Japan</td>
<td id="76DA5B64E90A0039FBDBFB57FB7678B5" box="[1026,1207,1185,1210]" gridcol="2" gridrow="1" pageId="3" pageNumber="22">Japan and Taiwan</td>
</tr>
<tr id="350B3218E90A0039FF7AFB3CFAA978ED" box="[163,1384,1226,1250]" gridrow="2" pageId="3" pageNumber="22" rowspan-0="1" rowspan-3="1">
<td id="76DA5B64E90A0039FD3CFB3CFC6178ED" box="[741,928,1226,1250]" gridcol="1" gridrow="2" pageId="3" pageNumber="22">KAUM–I. 108333</td>
<td id="76DA5B64E90A0039FBDBFB3CFB7678ED" box="[1026,1207,1226,1250]" gridcol="2" gridrow="2" pageId="3" pageNumber="22">
<emphasis id="B94FEC48E90AFFC5FBE5FB3CFB8878EE" box="[1084,1097,1226,1249]" italics="true" pageId="3" pageNumber="22">n</emphasis>
= 18
</td>
</tr>
<tr id="350B3218E90A0039FF7AFB07FAA97905" box="[163,1384,1265,1290]" gridrow="3" pageId="3" pageNumber="22" rowspan-3="1">
<th id="76DA5B64E90A0039FF7AFB07FDD37905" box="[163,530,1265,1290]" gridcol="0" gridrow="3" pageId="3" pageNumber="22">Standard length (SL; mm)</th>
<td id="76DA5B64E90A0039FD3CFB07FC617905" box="[741,928,1265,1290]" gridcol="1" gridrow="3" pageId="3" pageNumber="22">96.1</td>
<td id="76DA5B64E90A0039FBDBFB07FB767905" box="[1026,1207,1265,1290]" gridcol="2" gridrow="3" pageId="3" pageNumber="22">66.6–109.8</td>
</tr>
<tr id="350B3218E90A0039FF7AFAEFFAA9793D" box="[163,1384,1305,1330]" gridrow="4" pageId="3" pageNumber="22" rowspan-1="1" rowspan-2="1">
<th id="76DA5B64E90A0039FF7AFAEFFDD3793D" box="[163,530,1305,1330]" gridcol="0" gridrow="4" pageId="3" pageNumber="22">Counts</th>
<td id="76DA5B64E90A0039FAFAFAEFFAA9793D" box="[1315,1384,1305,1330]" gridcol="3" gridrow="4" pageId="3" pageNumber="22">Modes</td>
</tr>
<tr id="350B3218E90A0039FF7AFAB4FAA97954" box="[163,1384,1346,1371]" gridrow="5" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AFAB4FDD37954" box="[163,530,1346,1371]" gridcol="0" gridrow="5" pageId="3" pageNumber="22">Dorsal-fin rays (unbranched)</th>
<td id="76DA5B64E90A0039FD3CFAB4FC617954" box="[741,928,1346,1371]" gridcol="1" gridrow="5" pageId="3" pageNumber="22">4</td>
<td id="76DA5B64E90A0039FBDBFAB4FB767954" box="[1026,1207,1346,1371]" gridcol="2" gridrow="5" pageId="3" pageNumber="22">4–5</td>
<td id="76DA5B64E90A0039FAFAFAB4FAA97954" box="[1315,1384,1346,1371]" gridcol="3" gridrow="5" pageId="3" pageNumber="22">4</td>
</tr>
<tr id="350B3218E90A0039FF7AFA9CFAA9798C" box="[163,1384,1386,1411]" gridrow="6" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AFA9CFDD3798C" box="[163,530,1386,1411]" gridcol="0" gridrow="6" pageId="3" pageNumber="22">Dorsal-fin rays (branched)</th>
<td id="76DA5B64E90A0039FD3CFA9CFC61798C" box="[741,928,1386,1411]" gridcol="1" gridrow="6" pageId="3" pageNumber="22">15</td>
<td id="76DA5B64E90A0039FBDBFA9CFB76798C" box="[1026,1207,1386,1411]" gridcol="2" gridrow="6" pageId="3" pageNumber="22">14–16</td>
<td id="76DA5B64E90A0039FAFAFA9CFAA9798C" box="[1315,1384,1386,1411]" gridcol="3" gridrow="6" pageId="3" pageNumber="22">15</td>
</tr>
<tr id="350B3218E90A0039FF7AFA64FAA979A4" box="[163,1384,1426,1451]" gridrow="7" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AFA64FDD379A4" box="[163,530,1426,1451]" gridcol="0" gridrow="7" pageId="3" pageNumber="22">Anal-fin rays (unbranched)</th>
<td id="76DA5B64E90A0039FD3CFA64FC6179A4" box="[741,928,1426,1451]" gridcol="1" gridrow="7" pageId="3" pageNumber="22">3</td>
<td id="76DA5B64E90A0039FBDBFA64FB7679A4" box="[1026,1207,1426,1451]" gridcol="2" gridrow="7" pageId="3" pageNumber="22">3–4</td>
<td id="76DA5B64E90A0039FAFAFA64FAA979A4" box="[1315,1384,1426,1451]" gridcol="3" gridrow="7" pageId="3" pageNumber="22">3</td>
</tr>
<tr id="350B3218E90A0039FF7AFA4CFAA979DB" box="[163,1384,1466,1492]" gridrow="8" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AFA4CFDD379DB" box="[163,530,1466,1492]" gridcol="0" gridrow="8" pageId="3" pageNumber="22">Anal-fin rays (branched)</th>
<td id="76DA5B64E90A0039FD3CFA4CFC6179DB" box="[741,928,1466,1492]" gridcol="1" gridrow="8" pageId="3" pageNumber="22">18</td>
<td id="76DA5B64E90A0039FBDBFA4CFB7679DB" box="[1026,1207,1466,1492]" gridcol="2" gridrow="8" pageId="3" pageNumber="22">16–19</td>
<td id="76DA5B64E90A0039FAFAFA4CFAA979DB" box="[1315,1384,1466,1492]" gridcol="3" gridrow="8" pageId="3" pageNumber="22">18</td>
</tr>
<tr id="350B3218E90A0039FF7AFA15FAA979F3" box="[163,1384,1507,1532]" gridrow="9" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AFA15FDD379F3" box="[163,530,1507,1532]" gridcol="0" gridrow="9" pageId="3" pageNumber="22">Pectoral-fin rays (unbranched)</th>
<td id="76DA5B64E90A0039FD3CFA15FC6179F3" box="[741,928,1507,1532]" gridcol="1" gridrow="9" pageId="3" pageNumber="22">1</td>
<td id="76DA5B64E90A0039FBDBFA15FB7679F3" box="[1026,1207,1507,1532]" gridcol="2" gridrow="9" pageId="3" pageNumber="22">1</td>
<td id="76DA5B64E90A0039FAFAFA15FAA979F3" box="[1315,1384,1507,1532]" gridcol="3" gridrow="9" pageId="3" pageNumber="22">1</td>
</tr>
<tr id="350B3218E90A0039FF7AF9FDFAA97A2B" box="[163,1384,1547,1572]" gridrow="10" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF9FDFDD37A2B" box="[163,530,1547,1572]" gridcol="0" gridrow="10" pageId="3" pageNumber="22">Pectoral-fin rays (branched)</th>
<td id="76DA5B64E90A0039FD3CF9FDFC617A2B" box="[741,928,1547,1572]" gridcol="1" gridrow="10" pageId="3" pageNumber="22">14</td>
<td id="76DA5B64E90A0039FBDBF9FDFB767A2B" box="[1026,1207,1547,1572]" gridcol="2" gridrow="10" pageId="3" pageNumber="22">13–16</td>
<td id="76DA5B64E90A0039FAFAF9FDFAA97A2B" box="[1315,1384,1547,1572]" gridcol="3" gridrow="10" pageId="3" pageNumber="22">14</td>
</tr>
<tr id="350B3218E90A0039FF7AF9C5FAA97A43" box="[163,1384,1587,1612]" gridrow="11" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF9C5FDD37A43" box="[163,530,1587,1612]" gridcol="0" gridrow="11" pageId="3" pageNumber="22">Pelvic-fin rays (unbranched)</th>
<td id="76DA5B64E90A0039FD3CF9C5FC617A43" box="[741,928,1587,1612]" gridcol="1" gridrow="11" pageId="3" pageNumber="22">1</td>
<td id="76DA5B64E90A0039FBDBF9C5FB767A43" box="[1026,1207,1587,1612]" gridcol="2" gridrow="11" pageId="3" pageNumber="22">1</td>
<td id="76DA5B64E90A0039FAFAF9C5FAA97A43" box="[1315,1384,1587,1612]" gridcol="3" gridrow="11" pageId="3" pageNumber="22">1</td>
</tr>
<tr id="350B3218E90A0039FF7AF9ADFAA97A7B" box="[163,1384,1627,1652]" gridrow="12" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF9ADFDD37A7B" box="[163,530,1627,1652]" gridcol="0" gridrow="12" pageId="3" pageNumber="22">Pelvic-fin rays (branched)</th>
<td id="76DA5B64E90A0039FD3CF9ADFC617A7B" box="[741,928,1627,1652]" gridcol="1" gridrow="12" pageId="3" pageNumber="22">7</td>
<td id="76DA5B64E90A0039FBDBF9ADFB767A7B" box="[1026,1207,1627,1652]" gridcol="2" gridrow="12" pageId="3" pageNumber="22">7</td>
<td id="76DA5B64E90A0039FAFAF9ADFAA97A7B" box="[1315,1384,1627,1652]" gridcol="3" gridrow="12" pageId="3" pageNumber="22">7</td>
</tr>
<tr id="350B3218E90A0039FF7AF972FAA97A92" box="[163,1384,1668,1693]" gridrow="13" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF972FDD37A92" box="[163,530,1668,1693]" gridcol="0" gridrow="13" pageId="3" pageNumber="22">Gill rakers on 1st gill arch (upper)</th>
<td id="76DA5B64E90A0039FD3CF972FC617A92" box="[741,928,1668,1693]" gridcol="1" gridrow="13" pageId="3" pageNumber="22">39</td>
<td id="76DA5B64E90A0039FBDBF972FB767A92" box="[1026,1207,1668,1693]" gridcol="2" gridrow="13" pageId="3" pageNumber="22">34–42</td>
<td id="76DA5B64E90A0039FAFAF972FAA97A92" box="[1315,1384,1668,1693]" gridcol="3" gridrow="13" pageId="3" pageNumber="22">39</td>
</tr>
<tr id="350B3218E90A0039FF7AF95AFAA97ACA" box="[163,1384,1708,1733]" gridrow="14" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF95AFDD37ACA" box="[163,530,1708,1733]" gridcol="0" gridrow="14" pageId="3" pageNumber="22">Gill rakers on 1st gill arch (lower)</th>
<td id="76DA5B64E90A0039FD3CF95AFC617ACA" box="[741,928,1708,1733]" gridcol="1" gridrow="14" pageId="3" pageNumber="22">68</td>
<td id="76DA5B64E90A0039FBDBF95AFB767ACA" box="[1026,1207,1708,1733]" gridcol="2" gridrow="14" pageId="3" pageNumber="22">64–72</td>
<td id="76DA5B64E90A0039FAFAF95AFAA97ACA" box="[1315,1384,1708,1733]" gridcol="3" gridrow="14" pageId="3" pageNumber="22">67</td>
</tr>
<tr id="350B3218E90A0039FF7AF922FAA97AE2" box="[163,1384,1748,1773]" gridrow="15" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF922FDD37AE2" box="[163,530,1748,1773]" gridcol="0" gridrow="15" pageId="3" pageNumber="22">Gill rakers on 1st gill arch (total)</th>
<td id="76DA5B64E90A0039FD3CF922FC617AE2" box="[741,928,1748,1773]" gridcol="1" gridrow="15" pageId="3" pageNumber="22">107</td>
<td id="76DA5B64E90A0039FBDBF922FB767AE2" box="[1026,1207,1748,1773]" gridcol="2" gridrow="15" pageId="3" pageNumber="22">99–112</td>
<td id="76DA5B64E90A0039FAFAF922FAA97AE2" box="[1315,1384,1748,1773]" gridcol="3" gridrow="15" pageId="3" pageNumber="22">103</td>
</tr>
<tr id="350B3218E90A0039FF7AF90AFAA97B1A" box="[163,1384,1788,1813]" gridrow="16" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF90AFDD37B1A" box="[163,530,1788,1813]" gridcol="0" gridrow="16" pageId="3" pageNumber="22">Gill rakers on 2nd gill arch (upper)</th>
<td id="76DA5B64E90A0039FD3CF90AFC617B1A" box="[741,928,1788,1813]" gridcol="1" gridrow="16" pageId="3" pageNumber="22">37</td>
<td id="76DA5B64E90A0039FBDBF90AFB767B1A" box="[1026,1207,1788,1813]" gridcol="2" gridrow="16" pageId="3" pageNumber="22">33–40</td>
<td id="76DA5B64E90A0039FAFAF90AFAA97B1A" box="[1315,1384,1788,1813]" gridcol="3" gridrow="16" pageId="3" pageNumber="22">35</td>
</tr>
<tr id="350B3218E90A0039FF7AF8D3FAA97B31" box="[163,1384,1829,1854]" gridrow="17" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF8D3FDD37B31" box="[163,530,1829,1854]" gridcol="0" gridrow="17" pageId="3" pageNumber="22">Gill rakers on 2nd gill arch (lower)</th>
<td id="76DA5B64E90A0039FD3CF8D3FC617B31" box="[741,928,1829,1854]" gridcol="1" gridrow="17" pageId="3" pageNumber="22">74</td>
<td id="76DA5B64E90A0039FBDBF8D3FB767B31" box="[1026,1207,1829,1854]" gridcol="2" gridrow="17" pageId="3" pageNumber="22">64–76</td>
<td id="76DA5B64E90A0039FAFAF8D3FAA97B31" box="[1315,1384,1829,1854]" gridcol="3" gridrow="17" pageId="3" pageNumber="22">74</td>
</tr>
<tr id="350B3218E90A0039FF7AF8BBFAA97B69" box="[163,1384,1869,1894]" gridrow="18" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF8BBFDD37B69" box="[163,530,1869,1894]" gridcol="0" gridrow="18" pageId="3" pageNumber="22">Gill rakers on 2nd gill arch (total)</th>
<td id="76DA5B64E90A0039FD3CF8BBFC617B69" box="[741,928,1869,1894]" gridcol="1" gridrow="18" pageId="3" pageNumber="22">111</td>
<td id="76DA5B64E90A0039FBDBF8BBFB767B69" box="[1026,1207,1869,1894]" gridcol="2" gridrow="18" pageId="3" pageNumber="22">97–115</td>
<td id="76DA5B64E90A0039FAFAF8BBFAA97B69" box="[1315,1384,1869,1894]" gridcol="3" gridrow="18" pageId="3" pageNumber="22">101</td>
</tr>
<tr id="350B3218E90A0039FF7AF883FAA97B81" box="[163,1384,1909,1934]" gridrow="19" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF883FDD37B81" box="[163,530,1909,1934]" gridcol="0" gridrow="19" pageId="3" pageNumber="22">Gill rakers on 3rd gill arch (upper)</th>
<td id="76DA5B64E90A0039FD3CF883FC617B81" box="[741,928,1909,1934]" gridcol="1" gridrow="19" pageId="3" pageNumber="22">35</td>
<td id="76DA5B64E90A0039FBDBF883FB767B81" box="[1026,1207,1909,1934]" gridcol="2" gridrow="19" pageId="3" pageNumber="22">31–38</td>
<td id="76DA5B64E90A0039FAFAF883FAA97B81" box="[1315,1384,1909,1934]" gridcol="3" gridrow="19" pageId="3" pageNumber="22">33</td>
</tr>
<tr id="350B3218E90A0039FF7AF86BFAA97BB9" box="[163,1384,1949,1974]" gridrow="20" pageId="3" pageNumber="22">
<th id="76DA5B64E90A0039FF7AF86BFDD37BB9" box="[163,530,1949,1974]" gridcol="0" gridrow="20" pageId="3" pageNumber="22">Gill rakers on 3rd gill arch (lower)</th>
<td id="76DA5B64E90A0039FD3CF86BFC617BB9" box="[741,928,1949,1974]" gridcol="1" gridrow="20" pageId="3" pageNumber="22">57</td>
<td id="76DA5B64E90A0039FBDBF86BFB767BB9" box="[1026,1207,1949,1974]" gridcol="2" gridrow="20" pageId="3" pageNumber="22">48–60</td>
<td id="76DA5B64E90A0039FAFAF86BFAA97BB9" box="[1315,1384,1949,1974]" gridcol="3" gridrow="20" pageId="3" pageNumber="22">57</td>
</tr>
</table>
</paragraph>
<paragraph id="8B84305AE90AFFC5FB54F830FA507BD1" blockId="3.[1165,1425,1990,2014]" box="[1165,1425,1990,2014]" pageId="3" pageNumber="22">
<tableNote id="76DD31D4E90AFFC5FB54F830FA507BD1" box="[1165,1425,1990,2014]" pageId="3" pageNumber="22" targetBox="[163,1384,1144,1974]" targetPageId="3">...Continued on next page</tableNote>
</paragraph>
<caption id="DF4460D2E90DFFC2FF7AFF55FE507CB3" box="[163,401,163,188]" pageId="4" pageNumber="23" startId="4.[163,247,163,188]" targetBox="[163,1384,204,1597]" targetIsTable="true" targetPageId="4">
<paragraph id="8B84305AE90DFFC2FF7AFF55FE507CB3" blockId="4.[163,401,163,188]" box="[163,401,163,188]" pageId="4" pageNumber="23">
<emphasis id="B94FEC48E90DFFC2FF7AFF55FED17CB3" bold="true" box="[163,272,163,188]" pageId="4" pageNumber="23">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B84305AE90DFFC2FCCDFF3AFA9D7A32" pageId="4" pageNumber="23">
<table id="F93BC2FAE90D0039FF7AFF3AFAA97A32" box="[163,1384,204,1597]" colsContinueFrom="3.[163,1384,1144,1974]" gridcols="4" gridrows="35" pageId="4" pageNumber="23">
<tr id="350B3218E90D0039FF7AFF3AFAA97CEA" box="[163,1384,204,229]" gridrow="0" pageId="4" pageNumber="23" rowspan-0="1" rowspan-3="1">
<th id="76DA5B64E90D0039FD3CFF3AFC617CEA" box="[741,928,204,229]" gridcol="1" gridrow="0" pageId="4" pageNumber="23">Holotype</th>
<th id="76DA5B64E90D0039FBDBFF3AFB767CEA" box="[1026,1207,204,229]" gridcol="2" gridrow="0" pageId="4" pageNumber="23">Paratypes</th>
</tr>
<tr id="350B3218E90D0039FF7AFF02FAA97D02" box="[163,1384,244,269]" gridrow="1" pageId="4" pageNumber="23" rowspan-0="1" rowspan-3="1">
<td id="76DA5B64E90D0039FD3CFF02FC617D02" box="[741,928,244,269]" gridcol="1" gridrow="1" pageId="4" pageNumber="23">Japan</td>
<td id="76DA5B64E90D0039FBDBFF02FB767D02" box="[1026,1207,244,269]" gridcol="2" gridrow="1" pageId="4" pageNumber="23">Japan and Taiwan</td>
</tr>
<tr id="350B3218E90D0039FF7AFEEBFAA97D3A" box="[163,1384,285,309]" gridrow="2" pageId="4" pageNumber="23" rowspan-0="1" rowspan-3="1">
<td id="76DA5B64E90D0039FD3CFEEBFC617D3A" box="[741,928,285,309]" gridcol="1" gridrow="2" pageId="4" pageNumber="23">KAUM–I. 108333</td>
<td id="76DA5B64E90D0039FBDBFEEBFB767D3A" box="[1026,1207,285,309]" gridcol="2" gridrow="2" pageId="4" pageNumber="23">
<emphasis id="B94FEC48E90DFFC2FBE5FEE8FB887D3A" box="[1084,1097,286,309]" italics="true" pageId="4" pageNumber="23">n</emphasis>
= 18
</td>
</tr>
<tr id="350B3218E90D0039FF7AFEB2FAA97D52" box="[163,1384,324,349]" gridrow="3" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFEB2FDA87D52" box="[163,617,324,349]" gridcol="0" gridrow="3" pageId="4" pageNumber="23">Gill rakers on 3rd gill arch (total)</th>
<td id="76DA5B64E90D0039FD3CFEB2FC617D52" box="[741,928,324,349]" gridcol="1" gridrow="3" pageId="4" pageNumber="23">92</td>
<td id="76DA5B64E90D0039FBDBFEB2FB767D52" box="[1026,1207,324,349]" gridcol="2" gridrow="3" pageId="4" pageNumber="23">79–98</td>
<td id="76DA5B64E90D0039FAFDFEB2FAA97D52" box="[1316,1384,324,349]" gridcol="3" gridrow="3" pageId="4" pageNumber="23">84</td>
</tr>
<tr id="350B3218E90D0039FF7AFE9BFAA97D89" box="[163,1384,365,390]" gridrow="4" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFE9BFDA87D89" box="[163,617,365,390]" gridcol="0" gridrow="4" pageId="4" pageNumber="23">Gill rakers on 4th gill arch (upper)</th>
<td id="76DA5B64E90D0039FD3CFE9BFC617D89" box="[741,928,365,390]" gridcol="1" gridrow="4" pageId="4" pageNumber="23">27</td>
<td id="76DA5B64E90D0039FBDBFE9BFB767D89" box="[1026,1207,365,390]" gridcol="2" gridrow="4" pageId="4" pageNumber="23">24–31</td>
<td id="76DA5B64E90D0039FAFDFE9BFAA97D89" box="[1316,1384,365,390]" gridcol="3" gridrow="4" pageId="4" pageNumber="23">27</td>
</tr>
<tr id="350B3218E90D0039FF7AFE63FAA97DA1" box="[163,1384,405,430]" gridrow="5" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFE63FDA87DA1" box="[163,617,405,430]" gridcol="0" gridrow="5" pageId="4" pageNumber="23">Gill rakers on 4th gill arch (lower)</th>
<td id="76DA5B64E90D0039FD3CFE63FC617DA1" box="[741,928,405,430]" gridcol="1" gridrow="5" pageId="4" pageNumber="23">40</td>
<td id="76DA5B64E90D0039FBDBFE63FB767DA1" box="[1026,1207,405,430]" gridcol="2" gridrow="5" pageId="4" pageNumber="23">37–47</td>
<td id="76DA5B64E90D0039FAFDFE63FAA97DA1" box="[1316,1384,405,430]" gridcol="3" gridrow="5" pageId="4" pageNumber="23">39</td>
</tr>
<tr id="350B3218E90D0039FF7AFE4BFAA97DD9" box="[163,1384,445,470]" gridrow="6" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFE4BFDA87DD9" box="[163,617,445,470]" gridcol="0" gridrow="6" pageId="4" pageNumber="23">Gill rakers on 4th gill arch (total)</th>
<td id="76DA5B64E90D0039FD3CFE4BFC617DD9" box="[741,928,445,470]" gridcol="1" gridrow="6" pageId="4" pageNumber="23">67</td>
<td id="76DA5B64E90D0039FBDBFE4BFB767DD9" box="[1026,1207,445,470]" gridcol="2" gridrow="6" pageId="4" pageNumber="23">62–77</td>
<td id="76DA5B64E90D0039FAFDFE4BFAA97DD9" box="[1316,1384,445,470]" gridcol="3" gridrow="6" pageId="4" pageNumber="23">68</td>
</tr>
<tr id="350B3218E90D0039FF7AFE13FAA97DF1" box="[163,1384,485,510]" gridrow="7" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFE13FDA87DF1" box="[163,617,485,510]" gridcol="0" gridrow="7" pageId="4" pageNumber="23">Gill rakers on posterior face of 3rd gill arch</th>
<td id="76DA5B64E90D0039FD3CFE13FC617DF1" box="[741,928,485,510]" gridcol="1" gridrow="7" pageId="4" pageNumber="23">26</td>
<td id="76DA5B64E90D0039FBDBFE13FB767DF1" box="[1026,1207,485,510]" gridcol="2" gridrow="7" pageId="4" pageNumber="23">25–31</td>
<td id="76DA5B64E90D0039FAFDFE13FAA97DF1" box="[1316,1384,485,510]" gridcol="3" gridrow="7" pageId="4" pageNumber="23">27</td>
</tr>
<tr id="350B3218E90D0039FF7AFDF8FAA97E28" box="[163,1384,526,551]" gridrow="8" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFDF8FDA87E28" box="[163,617,526,551]" gridcol="0" gridrow="8" pageId="4" pageNumber="23">Prepelvic scutes</th>
<td id="76DA5B64E90D0039FD3CFDF8FC617E28" box="[741,928,526,551]" gridcol="1" gridrow="8" pageId="4" pageNumber="23">18</td>
<td id="76DA5B64E90D0039FBDBFDF8FB767E28" box="[1026,1207,526,551]" gridcol="2" gridrow="8" pageId="4" pageNumber="23">17–18</td>
<td id="76DA5B64E90D0039FAFDFDF8FAA97E28" box="[1316,1384,526,551]" gridcol="3" gridrow="8" pageId="4" pageNumber="23">18</td>
</tr>
<tr id="350B3218E90D0039FF7AFDC0FAA97E40" box="[163,1384,566,591]" gridrow="9" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFDC0FDA87E40" box="[163,617,566,591]" gridcol="0" gridrow="9" pageId="4" pageNumber="23">Postpelvic scutes</th>
<td id="76DA5B64E90D0039FD3CFDC0FC617E40" box="[741,928,566,591]" gridcol="1" gridrow="9" pageId="4" pageNumber="23">13</td>
<td id="76DA5B64E90D0039FBDBFDC0FB767E40" box="[1026,1207,566,591]" gridcol="2" gridrow="9" pageId="4" pageNumber="23">12–14</td>
<td id="76DA5B64E90D0039FAFDFDC0FAA97E40" box="[1316,1384,566,591]" gridcol="3" gridrow="9" pageId="4" pageNumber="23">13</td>
</tr>
<tr id="350B3218E90D0039FF7AFDA8FAA97E78" box="[163,1384,606,631]" gridrow="10" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFDA8FDA87E78" box="[163,617,606,631]" gridcol="0" gridrow="10" pageId="4" pageNumber="23">Total scutes</th>
<td id="76DA5B64E90D0039FD3CFDA8FC617E78" box="[741,928,606,631]" gridcol="1" gridrow="10" pageId="4" pageNumber="23">31</td>
<td id="76DA5B64E90D0039FBDBFDA8FB767E78" box="[1026,1207,606,631]" gridcol="2" gridrow="10" pageId="4" pageNumber="23">30–32</td>
<td id="76DA5B64E90D0039FAFDFDA8FAA97E78" box="[1316,1384,606,631]" gridcol="3" gridrow="10" pageId="4" pageNumber="23">31</td>
</tr>
<tr id="350B3218E90D0039FF7AFD70FAA97E90" box="[163,1384,646,671]" gridrow="11" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFD70FDA87E90" box="[163,617,646,671]" gridcol="0" gridrow="11" pageId="4" pageNumber="23">Scale rows in longitudinal series</th>
<td id="76DA5B64E90D0039FD3CFD70FC617E90" box="[741,928,646,671]" gridcol="1" gridrow="11" pageId="4" pageNumber="23">38</td>
<td id="76DA5B64E90D0039FBDBFD70FB767E90" box="[1026,1207,646,671]" gridcol="2" gridrow="11" pageId="4" pageNumber="23">38–41</td>
<td id="76DA5B64E90D0039FAFDFD70FAA97E90" box="[1316,1384,646,671]" gridcol="3" gridrow="11" pageId="4" pageNumber="23">39</td>
</tr>
<tr id="350B3218E90D0039FF7AFD58FAA97EC8" box="[163,1384,686,711]" gridrow="12" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFD58FDA87EC8" box="[163,617,686,711]" gridcol="0" gridrow="12" pageId="4" pageNumber="23">Pseudobranchial filaments</th>
<td id="76DA5B64E90D0039FD3CFD58FC617EC8" box="[741,928,686,711]" gridcol="1" gridrow="12" pageId="4" pageNumber="23">18</td>
<td id="76DA5B64E90D0039FBDBFD58FB767EC8" box="[1026,1207,686,711]" gridcol="2" gridrow="12" pageId="4" pageNumber="23">16–20</td>
<td id="76DA5B64E90D0039FAFDFD58FAA97EC8" box="[1316,1384,686,711]" gridcol="3" gridrow="12" pageId="4" pageNumber="23">17</td>
</tr>
<tr id="350B3218E90D0039FF7AFD2EFAA97EFF" box="[163,1384,728,752]" gridrow="13" pageId="4" pageNumber="23" rowspan-1="1" rowspan-2="1">
<th id="76DA5B64E90D0039FF7AFD2EFDA87EFF" box="[163,617,728,752]" gridcol="0" gridrow="13" pageId="4" pageNumber="23">Measurements (%SL)</th>
<td id="76DA5B64E90D0039FAFDFD2EFAA97EFF" box="[1316,1384,728,752]" gridcol="3" gridrow="13" pageId="4" pageNumber="23">Means</td>
</tr>
<tr id="350B3218E90D0039FF7AFD09FAA97F17" box="[163,1384,767,792]" gridrow="14" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFD09FDA87F17" box="[163,617,767,792]" gridcol="0" gridrow="14" pageId="4" pageNumber="23">Head length</th>
<td id="76DA5B64E90D0039FD3CFD09FC617F17" box="[741,928,767,792]" gridcol="1" gridrow="14" pageId="4" pageNumber="23">26.6</td>
<td id="76DA5B64E90D0039FBDBFD09FB767F17" box="[1026,1207,767,792]" gridcol="2" gridrow="14" pageId="4" pageNumber="23">24.7–28.6</td>
<td id="76DA5B64E90D0039FAFDFD09FAA97F17" box="[1316,1384,767,792]" gridcol="3" gridrow="14" pageId="4" pageNumber="23">27.1</td>
</tr>
<tr id="350B3218E90D0039FF7AFCD1FAA97F4F" box="[163,1384,807,832]" gridrow="15" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFCD1FDA87F4F" box="[163,617,807,832]" gridcol="0" gridrow="15" pageId="4" pageNumber="23">Body depth</th>
<td id="76DA5B64E90D0039FD3CFCD1FC617F4F" box="[741,928,807,832]" gridcol="1" gridrow="15" pageId="4" pageNumber="23">34.3</td>
<td id="76DA5B64E90D0039FBDBFCD1FB767F4F" box="[1026,1207,807,832]" gridcol="2" gridrow="15" pageId="4" pageNumber="23">26.4–36.8</td>
<td id="76DA5B64E90D0039FAFDFCD1FAA97F4F" box="[1316,1384,807,832]" gridcol="3" gridrow="15" pageId="4" pageNumber="23">31.1</td>
</tr>
<tr id="350B3218E90D0039FF7AFCB9FAA97F67" box="[163,1384,847,872]" gridrow="16" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFCB9FDA87F67" box="[163,617,847,872]" gridcol="0" gridrow="16" pageId="4" pageNumber="23">Pre-dorsal-fin length</th>
<td id="76DA5B64E90D0039FD3CFCB9FC617F67" box="[741,928,847,872]" gridcol="1" gridrow="16" pageId="4" pageNumber="23">46.0</td>
<td id="76DA5B64E90D0039FBDBFCB9FB767F67" box="[1026,1207,847,872]" gridcol="2" gridrow="16" pageId="4" pageNumber="23">43.4–46.8</td>
<td id="76DA5B64E90D0039FAFDFCB9FAA97F67" box="[1316,1384,847,872]" gridcol="3" gridrow="16" pageId="4" pageNumber="23">45.4</td>
</tr>
<tr id="350B3218E90D0039FF7AFC8EFAA97F9E" box="[163,1384,888,913]" gridrow="17" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFC8EFDA87F9E" box="[163,617,888,913]" gridcol="0" gridrow="17" pageId="4" pageNumber="23">Snout tip to pectoral-fin insertion</th>
<td id="76DA5B64E90D0039FD3CFC8EFC617F9E" box="[741,928,888,913]" gridcol="1" gridrow="17" pageId="4" pageNumber="23">27.1</td>
<td id="76DA5B64E90D0039FBDBFC8EFB767F9E" box="[1026,1207,888,913]" gridcol="2" gridrow="17" pageId="4" pageNumber="23">26.5–29.1</td>
<td id="76DA5B64E90D0039FAFDFC8EFAA97F9E" box="[1316,1384,888,913]" gridcol="3" gridrow="17" pageId="4" pageNumber="23">27.7</td>
</tr>
<tr id="350B3218E90D0039FF7AFC56FAA97FB6" box="[163,1384,928,953]" gridrow="18" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFC56FDA87FB6" box="[163,617,928,953]" gridcol="0" gridrow="18" pageId="4" pageNumber="23">Snout tip to pelvic-fin insertion</th>
<td id="76DA5B64E90D0039FD3CFC56FC617FB6" box="[741,928,928,953]" gridcol="1" gridrow="18" pageId="4" pageNumber="23">55.2</td>
<td id="76DA5B64E90D0039FBDBFC56FB767FB6" box="[1026,1207,928,953]" gridcol="2" gridrow="18" pageId="4" pageNumber="23">51.9–56.4</td>
<td id="76DA5B64E90D0039FAFDFC56FAA97FB6" box="[1316,1384,928,953]" gridcol="3" gridrow="18" pageId="4" pageNumber="23">54.4</td>
</tr>
<tr id="350B3218E90D0039FF7AFC3EFAA97FEE" box="[163,1384,968,993]" gridrow="19" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFC3EFDA87FEE" box="[163,617,968,993]" gridcol="0" gridrow="19" pageId="4" pageNumber="23">Pre-anal-fin length</th>
<td id="76DA5B64E90D0039FD3CFC3EFC617FEE" box="[741,928,968,993]" gridcol="1" gridrow="19" pageId="4" pageNumber="23">81.6</td>
<td id="76DA5B64E90D0039FBDBFC3EFB767FEE" box="[1026,1207,968,993]" gridcol="2" gridrow="19" pageId="4" pageNumber="23">77.4–82.3</td>
<td id="76DA5B64E90D0039FAFDFC3EFAA97FEE" box="[1316,1384,968,993]" gridcol="3" gridrow="19" pageId="4" pageNumber="23">80.3</td>
</tr>
<tr id="350B3218E90D0039FF7AFC06FAA97806" box="[163,1384,1008,1033]" gridrow="20" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFC06FDA87806" box="[163,617,1008,1033]" gridcol="0" gridrow="20" pageId="4" pageNumber="23">Dorsal-fin base length</th>
<td id="76DA5B64E90D0039FD3CFC06FC617806" box="[741,928,1008,1033]" gridcol="1" gridrow="20" pageId="4" pageNumber="23">15.8</td>
<td id="76DA5B64E90D0039FBDBFC06FB767806" box="[1026,1207,1008,1033]" gridcol="2" gridrow="20" pageId="4" pageNumber="23">14.3–17.1</td>
<td id="76DA5B64E90D0039FAFDFC06FAA97806" box="[1316,1384,1008,1033]" gridcol="3" gridrow="20" pageId="4" pageNumber="23">15.8</td>
</tr>
<tr id="350B3218E90D0039FF7AFBEFFAA9783D" box="[163,1384,1049,1074]" gridrow="21" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFBEFFDA8783D" box="[163,617,1049,1074]" gridcol="0" gridrow="21" pageId="4" pageNumber="23">Anal-fin base length</th>
<td id="76DA5B64E90D0039FD3CFBEFFC61783D" box="[741,928,1049,1074]" gridcol="1" gridrow="21" pageId="4" pageNumber="23">15.0</td>
<td id="76DA5B64E90D0039FBDBFBEFFB76783D" box="[1026,1207,1049,1074]" gridcol="2" gridrow="21" pageId="4" pageNumber="23">14.9–18.4</td>
<td id="76DA5B64E90D0039FAFDFBEFFAA9783D" box="[1316,1384,1049,1074]" gridcol="3" gridrow="21" pageId="4" pageNumber="23">16.4</td>
</tr>
<tr id="350B3218E90D0039FF7AFBB7FAA97855" box="[163,1384,1089,1114]" gridrow="22" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFBB7FDA87855" box="[163,617,1089,1114]" gridcol="0" gridrow="22" pageId="4" pageNumber="23">Caudal-peduncle length</th>
<td id="76DA5B64E90D0039FD3CFBB7FC617855" box="[741,928,1089,1114]" gridcol="1" gridrow="22" pageId="4" pageNumber="23">6.7</td>
<td id="76DA5B64E90D0039FBDBFBB7FB767855" box="[1026,1207,1089,1114]" gridcol="2" gridrow="22" pageId="4" pageNumber="23">6.1–9.6</td>
<td id="76DA5B64E90D0039FAFDFBB7FAA97855" box="[1316,1384,1089,1114]" gridcol="3" gridrow="22" pageId="4" pageNumber="23">7.4</td>
</tr>
<tr id="350B3218E90D0039FF7AFB9FFAA9788D" box="[163,1384,1129,1154]" gridrow="23" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFB9FFDA8788D" box="[163,617,1129,1154]" gridcol="0" gridrow="23" pageId="4" pageNumber="23">Caudal-peduncle depth</th>
<td id="76DA5B64E90D0039FD3CFB9FFC61788D" box="[741,928,1129,1154]" gridcol="1" gridrow="23" pageId="4" pageNumber="23">9.6</td>
<td id="76DA5B64E90D0039FBDBFB9FFB76788D" box="[1026,1207,1129,1154]" gridcol="2" gridrow="23" pageId="4" pageNumber="23">8.2–11.8</td>
<td id="76DA5B64E90D0039FAFDFB9FFAA9788D" box="[1316,1384,1129,1154]" gridcol="3" gridrow="23" pageId="4" pageNumber="23">9.9</td>
</tr>
<tr id="350B3218E90D0039FF7AFB67FAA978A0" box="[163,1384,1169,1199]" gridrow="24" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFB67FDA878A0" box="[163,617,1169,1199]" gridcol="0" gridrow="24" pageId="4" pageNumber="23">D–P1</th>
<td id="76DA5B64E90D0039FD3CFB67FC6178A0" box="[741,928,1169,1199]" gridcol="1" gridrow="24" pageId="4" pageNumber="23">34.1</td>
<td id="76DA5B64E90D0039FBDBFB67FB7678A0" box="[1026,1207,1169,1199]" gridcol="2" gridrow="24" pageId="4" pageNumber="23">32.6–34.9</td>
<td id="76DA5B64E90D0039FAFDFB67FAA978A0" box="[1316,1384,1169,1199]" gridcol="3" gridrow="24" pageId="4" pageNumber="23">32.7</td>
</tr>
<tr id="350B3218E90D0039FF7AFB4CFAA978D8" box="[163,1384,1210,1239]" gridrow="25" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFB4CFDA878D8" box="[163,617,1210,1239]" gridcol="0" gridrow="25" pageId="4" pageNumber="23">D–P2</th>
<td id="76DA5B64E90D0039FD3CFB4CFC6178D8" box="[741,928,1210,1239]" gridcol="1" gridrow="25" pageId="4" pageNumber="23">33.1</td>
<td id="76DA5B64E90D0039FBDBFB4CFB7678D8" box="[1026,1207,1210,1239]" gridcol="2" gridrow="25" pageId="4" pageNumber="23">25.6–36.0</td>
<td id="76DA5B64E90D0039FAFDFB4CFAA978D8" box="[1316,1384,1210,1239]" gridcol="3" gridrow="25" pageId="4" pageNumber="23">30.2</td>
</tr>
<tr id="350B3218E90D0039FF7AFB14FAA978F4" box="[163,1384,1250,1275]" gridrow="26" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFB14FDA878F4" box="[163,617,1250,1275]" gridcol="0" gridrow="26" pageId="4" pageNumber="23">D–A</th>
<td id="76DA5B64E90D0039FD3CFB14FC6178F4" box="[741,928,1250,1275]" gridcol="1" gridrow="26" pageId="4" pageNumber="23">44.0</td>
<td id="76DA5B64E90D0039FBDBFB14FB7678F4" box="[1026,1207,1250,1275]" gridcol="2" gridrow="26" pageId="4" pageNumber="23">39.6–47.0</td>
<td id="76DA5B64E90D0039FAFDFB14FAA978F4" box="[1316,1384,1250,1275]" gridcol="3" gridrow="26" pageId="4" pageNumber="23">42.5</td>
</tr>
<tr id="350B3218E90D0039FF7AFAFCFAA97927" box="[163,1384,1290,1320]" gridrow="27" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFAFCFDA87927" box="[163,617,1290,1320]" gridcol="0" gridrow="27" pageId="4" pageNumber="23">P1–P2</th>
<td id="76DA5B64E90D0039FD3CFAFCFC617927" box="[741,928,1290,1320]" gridcol="1" gridrow="27" pageId="4" pageNumber="23">29.4</td>
<td id="76DA5B64E90D0039FBDBFAFCFB767927" box="[1026,1207,1290,1320]" gridcol="2" gridrow="27" pageId="4" pageNumber="23">25.6–29.3</td>
<td id="76DA5B64E90D0039FAFDFAFCFAA97927" box="[1316,1384,1290,1320]" gridcol="3" gridrow="27" pageId="4" pageNumber="23">28.2</td>
</tr>
<tr id="350B3218E90D0039FF7AFAC4FAA9795F" box="[163,1384,1330,1360]" gridrow="28" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFAC4FDA8795F" box="[163,617,1330,1360]" gridcol="0" gridrow="28" pageId="4" pageNumber="23">P2–A</th>
<td id="76DA5B64E90D0039FD3CFAC4FC61795F" box="[741,928,1330,1360]" gridcol="1" gridrow="28" pageId="4" pageNumber="23">30.4</td>
<td id="76DA5B64E90D0039FBDBFAC4FB76795F" box="[1026,1207,1330,1360]" gridcol="2" gridrow="28" pageId="4" pageNumber="23">27.5–30.9</td>
<td id="76DA5B64E90D0039FAFDFAC4FAA9795F" box="[1316,1384,1330,1360]" gridcol="3" gridrow="28" pageId="4" pageNumber="23">28.7</td>
</tr>
<tr id="350B3218E90D0039FF7AFAADFAA9797B" box="[163,1384,1371,1396]" gridrow="29" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFAADFDA8797B" box="[163,617,1371,1396]" gridcol="0" gridrow="29" pageId="4" pageNumber="23">Pectoral-fin length</th>
<td id="76DA5B64E90D0039FD3CFAADFC61797B" box="[741,928,1371,1396]" gridcol="1" gridrow="29" pageId="4" pageNumber="23">21.9</td>
<td id="76DA5B64E90D0039FBDBFAADFB76797B" box="[1026,1207,1371,1396]" gridcol="2" gridrow="29" pageId="4" pageNumber="23">20.7–23.4</td>
<td id="76DA5B64E90D0039FAFDFAADFAA9797B" box="[1316,1384,1371,1396]" gridcol="3" gridrow="29" pageId="4" pageNumber="23">21.8</td>
</tr>
<tr id="350B3218E90D0039FF7AFA75FAA97993" box="[163,1384,1411,1436]" gridrow="30" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFA75FDA87993" box="[163,617,1411,1436]" gridcol="0" gridrow="30" pageId="4" pageNumber="23">Pelvic-fin length</th>
<td id="76DA5B64E90D0039FD3CFA75FC617993" box="[741,928,1411,1436]" gridcol="1" gridrow="30" pageId="4" pageNumber="23">12.1</td>
<td id="76DA5B64E90D0039FBDBFA75FB767993" box="[1026,1207,1411,1436]" gridcol="2" gridrow="30" pageId="4" pageNumber="23">11.9–13.1</td>
<td id="76DA5B64E90D0039FAFDFA75FAA97993" box="[1316,1384,1411,1436]" gridcol="3" gridrow="30" pageId="4" pageNumber="23">12.4</td>
</tr>
<tr id="350B3218E90D0039FF7AFA5DFAA979CB" box="[163,1384,1451,1476]" gridrow="31" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFA5DFDA879CB" box="[163,617,1451,1476]" gridcol="0" gridrow="31" pageId="4" pageNumber="23">Interorbital width</th>
<td id="76DA5B64E90D0039FD3CFA5DFC6179CB" box="[741,928,1451,1476]" gridcol="1" gridrow="31" pageId="4" pageNumber="23">4.5</td>
<td id="76DA5B64E90D0039FBDBFA5DFB7679CB" box="[1026,1207,1451,1476]" gridcol="2" gridrow="31" pageId="4" pageNumber="23">4.1–4.9</td>
<td id="76DA5B64E90D0039FAFDFA5DFAA979CB" box="[1316,1384,1451,1476]" gridcol="3" gridrow="31" pageId="4" pageNumber="23">4.6</td>
</tr>
<tr id="350B3218E90D0039FF7AFA25FAA979E3" box="[163,1384,1491,1516]" gridrow="32" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFA25FDA879E3" box="[163,617,1491,1516]" gridcol="0" gridrow="32" pageId="4" pageNumber="23">Postorbital length</th>
<td id="76DA5B64E90D0039FD3CFA25FC6179E3" box="[741,928,1491,1516]" gridcol="1" gridrow="32" pageId="4" pageNumber="23">11.8</td>
<td id="76DA5B64E90D0039FBDBFA25FB7679E3" box="[1026,1207,1491,1516]" gridcol="2" gridrow="32" pageId="4" pageNumber="23">11.2–12.4</td>
<td id="76DA5B64E90D0039FAFDFA25FAA979E3" box="[1316,1384,1491,1516]" gridcol="3" gridrow="32" pageId="4" pageNumber="23">11.8</td>
</tr>
<tr id="350B3218E90D0039FF7AFA0AFAA97A1A" box="[163,1384,1532,1557]" gridrow="33" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AFA0AFDA87A1A" box="[163,617,1532,1557]" gridcol="0" gridrow="33" pageId="4" pageNumber="23">Upper-jaw length</th>
<td id="76DA5B64E90D0039FD3CFA0AFC617A1A" box="[741,928,1532,1557]" gridcol="1" gridrow="33" pageId="4" pageNumber="23">11.9</td>
<td id="76DA5B64E90D0039FBDBFA0AFB767A1A" box="[1026,1207,1532,1557]" gridcol="2" gridrow="33" pageId="4" pageNumber="23">10.8–12.4</td>
<td id="76DA5B64E90D0039FAFDFA0AFAA97A1A" box="[1316,1384,1532,1557]" gridcol="3" gridrow="33" pageId="4" pageNumber="23">11.6</td>
</tr>
<tr id="350B3218E90D0039FF7AF9D2FAA97A32" box="[163,1384,1572,1597]" gridrow="34" pageId="4" pageNumber="23">
<th id="76DA5B64E90D0039FF7AF9D2FDA87A32" box="[163,617,1572,1597]" gridcol="0" gridrow="34" pageId="4" pageNumber="23">Mandible length</th>
<td id="76DA5B64E90D0039FD3CF9D2FC617A32" box="[741,928,1572,1597]" gridcol="1" gridrow="34" pageId="4" pageNumber="23">12.2</td>
<td id="76DA5B64E90D0039FBDBF9D2FB767A32" box="[1026,1207,1572,1597]" gridcol="2" gridrow="34" pageId="4" pageNumber="23">11.8–13.4</td>
<td id="76DA5B64E90D0039FAFDF9D2FAA97A32" box="[1316,1384,1572,1597]" gridcol="3" gridrow="34" pageId="4" pageNumber="23">12.5</td>
</tr>
</table>
</paragraph>
<subSubSection id="C32163D1E90DFFC3FF1EF942FD117CF3" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23" type="discussion">
<paragraph id="8B84305AE90DFFC3FF1EF942FD117CF3" blockId="4.[151,1437,1715,2031]" lastBlockId="5.[151,1437,153,252]" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
<emphasis id="B94FEC48E90DFFC2FF1EF942FEFA7AC1" bold="true" box="[199,315,1716,1742]" pageId="4" pageNumber="23">Remarks.</emphasis>
<taxonomicName id="4C3B4BD9E90DFFC2FE9BF942FDD57AC1" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[322,532,1716,1742]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="alcyone">
<emphasis id="B94FEC48E90DFFC2FE9BF942FDD57AC1" box="[322,532,1716,1742]" italics="true" pageId="4" pageNumber="23">Sardinella alcyone</emphasis>
</taxonomicName>
is assignable to the genus 
<taxonomicName id="4C3B4BD9E90DFFC2FC98F942FC747AC1" authorityName="Valenciennes" authorityYear="1847" box="[833,949,1716,1742]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B94FEC48E90DFFC2FC98F942FC747AC1" box="[833,949,1716,1742]" italics="true" pageId="4" pageNumber="23">Sardinella</emphasis>
</taxonomicName>
, defined by 
<bibRefCitation id="EFAA4DABE90DFFC2FB98F942FACD7AC1" author="Whitehead, P. J. P." box="[1089,1292,1715,1742]" pageId="4" pageNumber="23" pagination="1 - 303" refId="ref3757" refString="Whitehead, P. J. P. (1985) FAO species catalogue vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. Part 1 - Chirocentridae, Clupeidae and Pristigasteridae. FAO Fisheries Synopsis, No. 125, 7 (1), 1 - 303." type="journal article" year="1985">Whitehead (1985)</bibRefCitation>
and 
<bibRefCitation id="EFAA4DABE90DFFC2FA9DF943FEE07AFD" author="Munroe, T. A. &amp; Wongratana, T. &amp; Nizinski, M. S." pageId="4" pageNumber="23" pagination="1775 - 1821" refId="ref3521" refString="Munroe, T. A., Wongratana, T. &amp; Nizinski, M. S. (1999) Order Clupeiformes. Clupeidae. Herrings (also, sardines, sprat, pilchard, and menhades). In: Carpenter, K. E. &amp; Niem, V. H. (Eds.), FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 3. Batoid Fishes, Chimaeras and Bony Fishes Part 1 (Elopidae to Linophrynidae). FAO, Rome, pp. 1775 - 1821." type="book chapter" year="1999">
Munroe 
<emphasis id="B94FEC48E90DFFC2FF4EF92FFF0E7AFD" box="[151,207,1752,1778]" italics="true" pageId="4" pageNumber="23">et al.</emphasis>
(1999)
</bibRefCitation>
, due to its compressed body, abdomen covered with prominently keeled scutes, paired predorsal scales, a symmetrical second supramaxilla, toothless hypomaxilla, two enlarged posteriormost anal-fin rays, the dorsal fin without filamentous rays, and two fleshy outgrowths on the hind margin of the gill opening. The new species most closely resembles 
<taxonomicName id="4C3B4BD9E90DFFC2FEBFF8B2FDFA7B51" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[358,571,1860,1886]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="pacifica">
<emphasis id="B94FEC48E90DFFC2FEBFF8B2FDFA7B51" box="[358,571,1860,1886]" italics="true" pageId="4" pageNumber="23">Sardinella pacifica</emphasis>
</taxonomicName>
, sharing centrally discontinuous striae on the lateral body scales, a dark spot on the dorsal-fin origin, more than 63 lower gill rakers on the first gill arch, 8 pelvic-fin rays, 17 or 18 prepelvic scutes, and more than 13 pseudobranchial filaments (
<bibRefCitation id="EFAA4DABE90DFFC2FD35F87AFC6B7BA9" author="Whitehead, P. J. P." box="[748,938,1931,1958]" pageId="4" pageNumber="23" pagination="1 - 303" refId="ref3757" refString="Whitehead, P. J. P. (1985) FAO species catalogue vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. Part 1 - Chirocentridae, Clupeidae and Pristigasteridae. FAO Fisheries Synopsis, No. 125, 7 (1), 1 - 303." type="journal article" year="1985">Whitehead 1985</bibRefCitation>
; 
<bibRefCitation id="EFAA4DABE90DFFC2FC6FF87BFB507BA9" author="Munroe, T. A. &amp; Wongratana, T. &amp; Nizinski, M. S." box="[950,1169,1932,1959]" pageId="4" pageNumber="23" pagination="1775 - 1821" refId="ref3521" refString="Munroe, T. A., Wongratana, T. &amp; Nizinski, M. S. (1999) Order Clupeiformes. Clupeidae. Herrings (also, sardines, sprat, pilchard, and menhades). In: Carpenter, K. E. &amp; Niem, V. H. (Eds.), FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 3. Batoid Fishes, Chimaeras and Bony Fishes Part 1 (Elopidae to Linophrynidae). FAO, Rome, pp. 1775 - 1821." type="book chapter" year="1999">
Munroe 
<emphasis id="B94FEC48E90DFFC2FBCFF87BFB8E7BA9" box="[1046,1103,1932,1958]" italics="true" pageId="4" pageNumber="23">et al.</emphasis>
1999
</bibRefCitation>
; 
<bibRefCitation id="EFAA4DABE90DFFC2FB47F87AFA9D7BA9" author="Stern, N. &amp; Rinkevich, B. &amp; Goren, M." box="[1182,1372,1932,1958]" pageId="4" pageNumber="23" pagination="2282 - 2305" refId="ref3699" refString="Stern, N., Rinkevich, B. &amp; Goren, M. (2016) Integrative approach revises the frequently misidentified species of Sardinella (Clupeidae) of the Indo-West Pacific Ocean. Journal of Fish Biology, 89, 2282 - 2305. https: // doi. org / 10.1111 / jfb. 13114" type="journal article" year="2016">
Stern 
<emphasis id="B94FEC48E90DFFC2FB39F87BFAD87BA9" box="[1248,1305,1932,1958]" italics="true" pageId="4" pageNumber="23">et al.</emphasis>
2016
</bibRefCitation>
; 
<bibRefCitation id="EFAA4DABE90DFFC2FABEF87BFEB77BC5" author="Hata, H. &amp; Motomura, H." pageId="4" pageNumber="23" pagination="274 - 280" refId="ref3404" refString="Hata, H. &amp; Motomura, H. (2019 a) A new species of sardine, Sardinella electra (Talaostei: Clupeiformes: Clupeidae), from the Ryukyu Islands, Japan. Zootaxa, 4565 (2), 274 - 280. http: // dx. doi. org / 10.11646 / zootaxa. 4565.2.11" type="journal article" year="2019">Hata &amp; Motomura 2019a</bibRefCitation>
, b). However, 
<taxonomicName id="4C3B4BD9E90DFFC2FDC1F847FD4B7BC5" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[536,650,1968,1995]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="alcyone">
<emphasis id="B94FEC48E90DFFC2FDC1F847FD4B7BC5" box="[536,650,1968,1995]" italics="true" pageId="4" pageNumber="23">S. alcyone</emphasis>
</taxonomicName>
is distinguished from the latter by the number of gill-rakers on the first to fourth gill arches (
<tableCitation id="C6B905E1E90DFFC2FE5CF822FE1D7BE1" box="[389,476,2004,2030]" captionStart="TABLE 1" captionStartId="3.[151,244,1102,1129]" captionTargetBox="[163,1384,1144,1974]" captionText="TABLE 1. Counts and measurements of specimens of Sardinella alcyone n. sp." httpUri="http://table.plazi.org/id/DF4460D2E90AFFC5FF4EFBB8FBD57866" pageId="4" pageNumber="23" tableUuid="DF4460D2E90AFFC5FF4EFBB8FBD57866">Table 1</tableCitation>
; 
<figureCitation id="13002CDFE90DFFC2FE32F822FDEF7BE1" box="[491,558,2004,2030]" captionStart="FIGURE 3" captionStartId="3.[151,250,924,949]" captionTargetBox="[159,1429,189,892]" captionTargetId="figure@3.[151,1436,181,899]" captionTargetPageId="3" captionText="FIGURE 3. Relationships of total gill-raker numbers (TGR) on (A) first gill arch (1GA), (B) second gill arch (2GA), (C) third gill arch (3GA), and (D) fourth gill arch (4GA) relative to standard length in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi="http://doi.org/10.5281/zenodo.3562833" httpUri="https://zenodo.org/record/3562833/files/figure.png" pageId="4" pageNumber="23">Fig. 3</figureCitation>
), and greater pectoral fin (20.7–23.4% SL vs. 18.2–20.8% in 
<taxonomicName id="4C3B4BD9E90DFFC2FB30F823FAA17BE1" authorityName="Hata &amp; Motomura" authorityYear="2019" box="[1257,1376,2004,2031]" class="Actinopterygii" family="Clupeidae" genus="Sardinella" kingdom="Animalia" order="Clupeiformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="pacifica">
<emphasis id="B94FEC48E90DFFC2FB30F823FAA17BE1" box="[1257,1376,2004,2031]" italics="true" pageId="4" pageNumber="23">S. pacifica</emphasis>
</taxonomicName>
; 
<figureCitation id="13002CDFE90DFFC3FAA9F822FF7C7CBC" captionStart="FIGURE 4" captionStartId="5.[151,250,1322,1347]" captionTargetBox="[161,1422,281,1289]" captionTargetId="figure@5.[151,1436,273,1297]" captionTargetPageId="5" captionText="FIGURE 4. Relationships of (A) pectoral-fin length (as % of standard length; SL) to SL, (B) pelvic-fin length (as % of SL) to SL, (C) upper-jaw length (as % of SL) to SL, (D) lower-jaw length (as % of SL) to SL, and (E) pre-anal-fin length (as % of SL) to SL in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi="http://doi.org/10.5281/zenodo.3562835" httpUri="https://zenodo.org/record/3562835/files/figure.png" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">Fig. 4A</figureCitation>
), pelvic fin (11.9–13.1% SL vs. 10.3–11.9%; 
<figureCitation id="13002CDFE90CFFC3FD65FF6FFCD57CBC" box="[700,788,153,179]" captionStart="FIGURE 4" captionStartId="5.[151,250,1322,1347]" captionTargetBox="[161,1422,281,1289]" captionTargetId="figure@5.[151,1436,273,1297]" captionTargetPageId="5" captionText="FIGURE 4. Relationships of (A) pectoral-fin length (as % of standard length; SL) to SL, (B) pelvic-fin length (as % of SL) to SL, (C) upper-jaw length (as % of SL) to SL, (D) lower-jaw length (as % of SL) to SL, and (E) pre-anal-fin length (as % of SL) to SL in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi="http://doi.org/10.5281/zenodo.3562835" httpUri="https://zenodo.org/record/3562835/files/figure.png" pageId="5" pageNumber="24">Fig. 4B</figureCitation>
), maxilla (10.8–12.4% SL vs. 9.3–10.9%; 
<figureCitation id="13002CDFE90CFFC3FB28FF6FFA897CBC" box="[1265,1352,153,179]" captionStart="FIGURE 4" captionStartId="5.[151,250,1322,1347]" captionTargetBox="[161,1422,281,1289]" captionTargetId="figure@5.[151,1436,273,1297]" captionTargetPageId="5" captionText="FIGURE 4. Relationships of (A) pectoral-fin length (as % of standard length; SL) to SL, (B) pelvic-fin length (as % of SL) to SL, (C) upper-jaw length (as % of SL) to SL, (D) lower-jaw length (as % of SL) to SL, and (E) pre-anal-fin length (as % of SL) to SL in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi="http://doi.org/10.5281/zenodo.3562835" httpUri="https://zenodo.org/record/3562835/files/figure.png" pageId="5" pageNumber="24">Fig. 4C</figureCitation>
), lower jaw (11.8–13.4% SL vs. 10.4–11.6%; 
<figureCitation id="13002CDFE90CFFC3FD91FF4BFD617CD8" box="[584,672,189,215]" captionStart="FIGURE 4" captionStartId="5.[151,250,1322,1347]" captionTargetBox="[161,1422,281,1289]" captionTargetId="figure@5.[151,1436,273,1297]" captionTargetPageId="5" captionText="FIGURE 4. Relationships of (A) pectoral-fin length (as % of standard length; SL) to SL, (B) pelvic-fin length (as % of SL) to SL, (C) upper-jaw length (as % of SL) to SL, (D) lower-jaw length (as % of SL) to SL, and (E) pre-anal-fin length (as % of SL) to SL in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi="http://doi.org/10.5281/zenodo.3562835" httpUri="https://zenodo.org/record/3562835/files/figure.png" pageId="5" pageNumber="24">Fig. 4D</figureCitation>
) and pre-anal-fin length (77.4–82.3% SL vs. 72.9–79.3%; 
<figureCitation id="13002CDFE90CFFC3FAE6FF4BFA557CD8" box="[1343,1428,189,215]" captionStart="FIGURE 4" captionStartId="5.[151,250,1322,1347]" captionTargetBox="[161,1422,281,1289]" captionTargetId="figure@5.[151,1436,273,1297]" captionTargetPageId="5" captionText="FIGURE 4. Relationships of (A) pectoral-fin length (as % of standard length; SL) to SL, (B) pelvic-fin length (as % of SL) to SL, (C) upper-jaw length (as % of SL) to SL, (D) lower-jaw length (as % of SL) to SL, and (E) pre-anal-fin length (as % of SL) to SL in Sardinella alcyone n. sp. (closed circles) and S. pacifica (open triangles)." figureDoi="http://doi.org/10.5281/zenodo.3562835" httpUri="https://zenodo.org/record/3562835/files/figure.png" pageId="5" pageNumber="24">Fig. 4E</figureCitation>
) proportions (
<bibRefCitation id="EFAA4DABE90CFFC3FEFEFF17FD897CF4" author="Hata, H. &amp; Motomura, H." box="[295,584,225,251]" pageId="5" pageNumber="24" pagination="75 - 83" refId="ref3465" refString="Hata, H. &amp; Motomura, H. (2019 b) A new species of sardine, Sardinella pacifica (Teleostei, Clupeiformes, Clupeidae), from the Philippines. ZooKeys, 829 (3), 75 - 83. https: // doi. org / 10.3897 / zookeys. 829.30688" type="journal article" year="2019">Hata &amp; Motomura 2019b</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
</treatment>
</document>