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<document id="07051B2C015481266410044476E98B29" ID-DOI="10.11646/zootaxa.5297.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8009123" ID-ZooBank="6E7C1770-D107-4FF7-A2DC-F2A693E581F1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686032543022" checkinUser="plazi" docAuthor="Marin, Ivan, Palatov, Dmitry &amp; Copilaș-Ciocianu, Denis" docDate="2023" docId="03927E02FFC8A40FFF55FD668CCBFA28" docLanguage="en" docName="zootaxa.5297.4.2.pdf" docOrigin="Zootaxa 5297 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Litorogammarus dursi Marin &amp; Palatov &amp; Copilaș-Ciocianu 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="498" masterDocId="FFAB067AFFC0A400FFC2FF9D895FFFE9" masterDocTitle="The remarkable Ponto-Caspian amphipod diversity of the lower Durso River (SW Caucasus) with the description of Litorogammarus dursi gen. et sp. nov." masterLastPageNumber="517" masterPageNumber="483" pageNumber="491" updateTime="1686033058541" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FAF654944E6F721D280C69C4D61717C8">The remarkable Ponto-Caspian amphipod diversity of the lower Durso River (SW Caucasus) with the description of Litorogammarus dursi gen. et sp. nov.</mods:title>
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<mods:namePart id="549F0893CCCAD362930ED1FDA2957AED">Marin, Ivan</mods:namePart>
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<mods:affiliation id="DF8EA4CE65510F53105CEF746B863633">A. N. Severtsov Institute of Ecology and Evolution of RAS, 119071, Moscow, Russia &amp; coralliodecapoda @ mail. ru; vanomarin @ yahoo. com; https: // orcid. org / 0000 - 0003 - 0552 - 8456</mods:affiliation>
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<mods:namePart id="DD47B296DFEF76A2B73CAFDA55010CD9">Palatov, Dmitry</mods:namePart>
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<mods:affiliation id="FF6FD9EA58DB088F8C9610BACB90FDE1">A. N. Severtsov Institute of Ecology and Evolution of RAS, 119071, Moscow, Russia &amp; triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
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<mods:namePart id="2D295E8035EE761657964E649BDCBE53">Copilaș-Ciocianu, Denis</mods:namePart>
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<mods:affiliation id="0C0DDB854917921FC99816B35B2818BA">Laboratory of Evolutionary Ecology of Hydrobionts, Nature Research Centre, Vilnius, Lithuania &amp; denis. copilas-ciocianu @ gamtc. lt; https: // orcid. org / 0000 - 0002 - 6374 - 2365</mods:affiliation>
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<taxonomicName id="4C3BB497FFC8A408FF55FD6688F7FCFC" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[151,424,762,789]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="491" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFC8A408FF55FD6688F7FCFC" bold="true" box="[151,424,762,789]" italics="true" pageId="8" pageNumber="491">Litorogammarus dursi</emphasis>
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<paragraph id="8B84CF14FFC8A408FF55FC8289A7FCD0" blockId="8.[151,525,762,825]" box="[151,248,799,825]" pageId="8" pageNumber="491">
<figureCitation id="1300D391FFC8A408FF55FC8289A7FCD0" box="[151,248,799,825]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="9.[151,250,1943,1967]" captionStartId-1="10.[151,250,1943,1967]" captionStartId-2="11.[151,250,1943,1967]" captionStartId-3="12.[151,250,1943,1967]" captionStartId-4="13.[151,250,1938,1962]" captionTargetBox-0="[184,1402,186,1916]" captionTargetBox-1="[185,1407,185,1910]" captionTargetBox-2="[186,1398,184,1916]" captionTargetBox-3="[183,1405,184,1917]" captionTargetBox-4="[151,1435,181,1914]" captionTargetId-0="figure-17@9.[181,1405,181,1919]" captionTargetId-1="figure-35@10.[181,1407,181,1919]" captionTargetId-2="figure-17@11.[181,1407,181,1919]" captionTargetId-3="figure-35@12.[179,1408,181,1920]" captionTargetId-4="figure-17@13.[151,1435,181,1914]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." captionText-1="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." captionText-2="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." captionText-3="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." captionText-4="FIGURE 8. Litorogammarus dursi sp. nov., Ô (LEMMI): a—head; b—urosomal segments, lateral view; c—urosomal segment and telson, dorsal view; d—urosomal, lateral view; e—retinacula of pleopod I." figureDoi-0="http://doi.org/10.5281/zenodo.8009137" figureDoi-1="http://doi.org/10.5281/zenodo.8009139" figureDoi-2="http://doi.org/10.5281/zenodo.8009143" figureDoi-3="http://doi.org/10.5281/zenodo.8009147" figureDoi-4="http://doi.org/10.5281/zenodo.8009154" httpUri-0="https://zenodo.org/record/8009137/files/figure.png" httpUri-1="https://zenodo.org/record/8009139/files/figure.png" httpUri-2="https://zenodo.org/record/8009143/files/figure.png" httpUri-3="https://zenodo.org/record/8009147/files/figure.png" httpUri-4="https://zenodo.org/record/8009154/files/figure.png" pageId="8" pageNumber="491">Figs 4–8</figureCitation>
</paragraph>
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<emphasis id="B94F1306FFC8A408FF55FCFD88DDFC93" bold="true" box="[151,386,864,890]" pageId="8" pageNumber="491">Material examined.</emphasis>
<materialsCitation id="3B53C549FFC8A408FE49FCFD8B08FC0F" collectingDate="2021-07-20" collectionCode="ZMMU" collectorName="Palatov D. M. &amp; Marin I. N." country="Russia" county="southwestern Caucasus" latitude="44.67889" location="Durso river" longLatPrecision="1" longitude="37.561554" municipality="Novorossiysk area" pageId="8" pageNumber="491" specimenCount="1" stateProvince="Krasnodar" typeStatus="holotype">
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: Ô (bl. 
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), 
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Mb-1243— 
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, 
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, 
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, 
<collectingMunicipality id="6BE0556EFFC8A408FE97FC198B77FC76" box="[341,552,900,927]" pageId="8" pageNumber="491">Novorossiysk area</collectingMunicipality>
, lower part of 
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, about 30–40 meters from its mouth, 
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<geoCoordinate id="EE0FA9D3FFC8A408FF55FC35886AFC2B" box="[151,309,936,962]" degrees="37" direction="east" minutes="33" orientation="longitude" pageId="8" pageNumber="491" precision="1" seconds="41.6" value="37.561554">37°33’41.6”E</geoCoordinate>
, a small expansion of the river with a weak current, among aquatic vegetation, 
<collectingMethod id="527AB703FFC8A408FB03FC358C00FC2A" box="[1217,1375,936,963]" pageId="8" pageNumber="491">with hand net</collectingMethod>
, coll. 
<collectorName id="26CEAAC2FFC8A408FF55FC518872FC0E" box="[151,301,972,999]" pageId="8" pageNumber="491">Palatov D.M.</collectorName>
&amp; 
<collectorName id="26CEAAC2FFC8A408FE93FC518896FC0E" box="[337,457,972,999]" pageId="8" pageNumber="491">Marin I.N.</collectorName>
, 
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<collectingDate id="EFC1103CFFC8A408FE14FC518B08FC0F" box="[470,599,972,998]" pageId="8" pageNumber="491" value="2021-07-20">20.07.2021</collectingDate>
</date>
</materialsCitation>
. 
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<typeStatus id="548071B6FFC8A408FDA0FC508B8CFC0E" box="[610,723,973,999]" pageId="8" pageNumber="491">Paratypes</typeStatus>
: Ô (bl. 
<quantity id="4CC362F1FFC8A408FCE4FC518ADFFC0F" box="[806,896,972,999]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="491" unit="mm" value="8.5">8.5 mm</quantity>
), 
<specimenCount id="9D3D049DFFC8A408FC51FC508AF7FC0E" box="[915,936,973,999]" pageId="8" pageNumber="491" type="female">♀</specimenCount>
(bl. 
<quantity id="4CC362F1FFC8A408FC1EFC508D7EFC0F" box="[988,1057,972,999]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="491" unit="mm" value="7.0">7 mm</quantity>
), 
<collectionCode id="ED2A57D1FFC8A408FBF1FC508DD5FC0E" box="[1075,1162,973,999]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="491" type="Museum">ZMMU</collectionCode>
Mb-1244, same locality
</materialsCitation>
<materialsCitation id="3B53C549FFC8A408FF55FC6D88F7FBE3" box="[151,424,1008,1035]" collectingDate="2021-07-20" collectionCode="ZMMU" collectorName="Palatov D. M. &amp; Marin I. N." country="Russia" county="southwestern Caucasus" latitude="44.67889" location="Durso river" longLatPrecision="1" longitude="37.561554" municipality="Novorossiysk area" pageId="8" pageNumber="491" specimenCount="1" stateProvince="Krasnodar" typeStatus="paratype">and data as for holotype</materialsCitation>
. 
<materialsCitation id="3B53C549FFC8A408FE73FC6D8D73FBE3" box="[433,1068,1008,1036]" collectingDate="2021-07-20" collectionCode="OO, LEMMI" collectorName="Palatov D. M. &amp; Marin I. N." country="Russia" county="southwestern Caucasus" latitude="44.67889" location="Durso river" longLatPrecision="1" longitude="37.561554" municipality="Novorossiysk area" pageId="8" pageNumber="491" specimenCount="3" specimenCount-female="3" stateProvince="Krasnodar">
<typeStatus id="548071B6FFC8A408FE73FC6D8BD0FBE3" box="[433,655,1008,1034]" isOtherTypeStatus="true" pageId="8" pageNumber="491">Additional material</typeStatus>
: 
<specimenCount id="9D3D049DFFC8A408FD58FC6D8B8CFBE3" box="[666,723,1008,1034]" pageId="8" pageNumber="491" type="female">3♀♀</specimenCount>
, 3 
<collectionCode id="ED2A57D1FFC8A408FD2FFC6C8A44FBE5" box="[749,795,1009,1036]" pageId="8" pageNumber="491">ÔÔ</collectionCode>
, 
<collectionCode id="ED2A57D1FFC8A408FCE4FC6C8AD9FBE2" box="[806,902,1009,1035]" pageId="8" pageNumber="491">LEMMI</collectionCode>
, same location
</materialsCitation>
<materialsCitation id="3B53C549FFC8A408FBF1FC6D8C43FBE3" box="[1075,1308,1008,1035]" collectingDate="2021-07-20" collectionCode="ZMMU" collectorName="Palatov D. M. &amp; Marin I. N." country="Russia" county="southwestern Caucasus" latitude="44.67889" location="Durso river" longLatPrecision="1" longitude="37.561554" municipality="Novorossiysk area" pageId="8" pageNumber="491" specimenCount="1" stateProvince="Krasnodar">and data as holotype</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3219C9FFFC8A408FF05FB898847FBBB" pageId="8" pageNumber="491" type="etymology">
<paragraph id="8B84CF14FFC8A408FF05FB898847FBBB" blockId="8.[151,1437,864,2043]" pageId="8" pageNumber="491">
<emphasis id="B94F1306FFC8A408FF05FB898811FBC7" bold="true" box="[199,334,1044,1070]" pageId="8" pageNumber="491">Etymology.</emphasis>
The species is named after the Durso River, in the lower course of which the new species was discovered.
</paragraph>
</subSubSection>
<subSubSection id="C3219C9FFFC8A408FF05FBC188A2FAA7" pageId="8" pageNumber="491" type="diagnosis">
<paragraph id="8B84CF14FFC8A408FF05FBC188A2FAA7" blockId="8.[151,1437,864,2043]" pageId="8" pageNumber="491">
<emphasis id="B94F1306FFC8A408FF05FBC1881DFB9F" bold="true" box="[199,322,1116,1142]" pageId="8" pageNumber="491">Diagnosis.</emphasis>
<emphasis id="B94F1306FFC8A408FE8BFBC188D9FB9F" box="[329,390,1116,1142]" italics="true" pageId="8" pageNumber="491">Head</emphasis>
with deeply concave anteroventral lobe. 
<emphasis id="B94F1306FFC8A408FC98FBC18AC4FB9F" box="[858,923,1116,1142]" italics="true" pageId="8" pageNumber="491">Pleon</emphasis>
with urosomite I with 2 robust setae on each margin and 2 median spine-like setae posterodorsally; urosomite II with 2 robust setae on each margin and 2 submedian spine-like setae (lateral spine stronger than median) dorsally; urosomite III with a row of 3–4 robust setae and 2 simple median setae. 
<emphasis id="B94F1306FFC8A408FDDCFB558BD0FB0B" box="[542,655,1224,1250]" italics="true" pageId="8" pageNumber="491">Antenna I</emphasis>
with 6-segmented accessory flagellum. 
<emphasis id="B94F1306FFC8A408FBA3FB558D83FB0B" box="[1121,1244,1224,1250]" italics="true" pageId="8" pageNumber="491">Antenna II</emphasis>
without calceoli and aesthetascs. 
<emphasis id="B94F1306FFC8A408FE93FB7088E1FAEF" box="[337,446,1260,1286]" italics="true" pageId="8" pageNumber="491">Upper lip</emphasis>
with concave distal margin. 
<emphasis id="B94F1306FFC8A408FCC2FB718A29FAEF" box="[768,886,1260,1286]" italics="true" pageId="8" pageNumber="491">Mandibles</emphasis>
with large and flattened incisor teeth. Outer lobe of 
<emphasis id="B94F1306FFC8A408FF7BFA8D887FFAC3" box="[185,288,1296,1322]" italics="true" pageId="8" pageNumber="491">maxilla I</emphasis>
very wide. 
<emphasis id="B94F1306FFC8A408FE6FFA8D8B79FAC3" box="[429,550,1296,1322]" italics="true" pageId="8" pageNumber="491">Maxilliped</emphasis>
with bluntly rounded outer and inner lobes. 
<emphasis id="B94F1306FFC8A408FBF0FA8D8DB8FAC3" box="[1074,1255,1296,1322]" italics="true" pageId="8" pageNumber="491">Epimeral plates</emphasis>
with sharp and produced posteroventral angles.
</paragraph>
</subSubSection>
<subSubSection id="C3219C9FFFC8A40FFF05FAC58CCBFA28" lastPageId="15" lastPageNumber="498" pageId="8" pageNumber="491" type="description">
<paragraph id="8B84CF14FFC8A408FF05FAC588EAF9CF" blockId="8.[151,1437,864,2043]" pageId="8" pageNumber="491">
<emphasis id="B94F1306FFC8A408FF05FAC58806FA9B" bold="true" box="[199,345,1368,1394]" pageId="8" pageNumber="491">Description.</emphasis>
<emphasis id="B94F1306FFC8A408FEA6FAC588C1FA9B" box="[356,414,1368,1394]" italics="true" pageId="8" pageNumber="491">Body</emphasis>
length 
<quantity id="4CC362F1FFC8A408FE3BFAC58B35FA9A" box="[505,618,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="8" pageNumber="491" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
. 
<emphasis id="B94F1306FFC8A408FDBAFAC58BEAFA9B" box="[632,693,1368,1394]" italics="true" pageId="8" pageNumber="491">Head</emphasis>
with deeply concave anteroventral lobe (
<figureCitation id="1300D391FFC8A408FB59FAC58DAFFA9B" box="[1179,1264,1368,1394]" captionStart="FIGURE 8" captionStartId="13.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-17@13.[151,1435,181,1914]" captionTargetPageId="13" captionText="FIGURE 8. Litorogammarus dursi sp. nov., Ô (LEMMI): a—head; b—urosomal segments, lateral view; c—urosomal segment and telson, dorsal view; d—urosomal, lateral view; e—retinacula of pleopod I." figureDoi="http://doi.org/10.5281/zenodo.8009154" httpUri="https://zenodo.org/record/8009154/files/figure.png" pageId="8" pageNumber="491">
Fig. 8 
<emphasis id="B94F1306FFC8A408FB20FAC48DAFFA9B" box="[1250,1264,1369,1394]" italics="true" pageId="8" pageNumber="491">a</emphasis>
</figureCitation>
); eye elongate reniform, well pigmented. 
<emphasis id="B94F1306FFC8A408FE16FAE08B16FA7F" box="[468,585,1405,1430]" italics="true" pageId="8" pageNumber="491">Metasoma</emphasis>
unarmed, with some short setules near dorsal articulation of segments. 
<emphasis id="B94F1306FFC8A408FF55FA3C880EFA53" box="[151,337,1441,1466]" italics="true" pageId="8" pageNumber="491">Urosomites I–III</emphasis>
(
<figureCitation id="1300D391FFC8A408FEA2FA3D88EFFA53" box="[352,432,1440,1466]" captionStart="FIGURE 8" captionStartId="13.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-17@13.[151,1435,181,1914]" captionTargetPageId="13" captionText="FIGURE 8. Litorogammarus dursi sp. nov., Ô (LEMMI): a—head; b—urosomal segments, lateral view; c—urosomal segment and telson, dorsal view; d—urosomal, lateral view; e—retinacula of pleopod I." figureDoi="http://doi.org/10.5281/zenodo.8009154" httpUri="https://zenodo.org/record/8009154/files/figure.png" pageId="8" pageNumber="491">
Fig. 8 
<emphasis id="B94F1306FFC8A408FE60FA3D88EFFA53" box="[418,432,1440,1466]" italics="true" pageId="8" pageNumber="491">b</emphasis>
</figureCitation>
) smooth, free, without any elevations; 
<emphasis id="B94F1306FFC8A408FCA6FA3C8ABCFA53" box="[868,995,1441,1466]" italics="true" pageId="8" pageNumber="491">urosomite I</emphasis>
with 2 robust setae on each margin and 2 median spine-like setae posterodorsally; 
<emphasis id="B94F1306FFC8A408FDB4FA588BA0FA37" box="[630,767,1477,1502]" italics="true" pageId="8" pageNumber="491">urosomite II</emphasis>
with 2 robust setae on each margin and 2 submedian spine-like setae (lateral spine stronger than median) dorsally; 
<emphasis id="B94F1306FFC8A408FCCCFA748AFEF9EB" box="[782,929,1513,1538]" italics="true" pageId="8" pageNumber="491">urosomite III</emphasis>
with a row of 3–4 robust setae and 2 simple median setae (
<figureCitation id="1300D391FFC8A408FEFAF99188FBF9CF" box="[312,420,1548,1575]" captionStart="FIGURE 8" captionStartId="13.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-17@13.[151,1435,181,1914]" captionTargetPageId="13" captionText="FIGURE 8. Litorogammarus dursi sp. nov., Ô (LEMMI): a—head; b—urosomal segments, lateral view; c—urosomal segment and telson, dorsal view; d—urosomal, lateral view; e—retinacula of pleopod I." figureDoi="http://doi.org/10.5281/zenodo.8009154" httpUri="https://zenodo.org/record/8009154/files/figure.png" pageId="8" pageNumber="491">
Fig. 8 
<emphasis id="B94F1306FFC8A408FEB8F99188FBF9CF" box="[378,420,1548,1575]" italics="true" pageId="8" pageNumber="491">b–d</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8B84CF14FFC8A408FF05F9AD8B5AF917" blockId="8.[151,1437,864,2043]" pageId="8" pageNumber="491">
<emphasis id="B94F1306FFC8A408FF05F9AD8869F9A3" box="[199,310,1584,1610]" italics="true" pageId="8" pageNumber="491">Antenna I</emphasis>
(
<figureCitation id="1300D391FFC8A408FE85F9AD88C6F9A3" box="[327,409,1584,1610]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="8" pageNumber="491">
Fig. 4 
<emphasis id="B94F1306FFC8A408FE49F9AC88C6F9A3" box="[395,409,1585,1610]" italics="true" pageId="8" pageNumber="491">a</emphasis>
</figureCitation>
) longer and slender than 
<emphasis id="B94F1306FFC8A408FD7FF9AC8A6CF9A3" box="[701,819,1585,1610]" italics="true" pageId="8" pageNumber="491">antenna II</emphasis>
, especially in males, close to 1/3 of body length, with robust peduncle segments, articles ratio is 1.0:0.87:0.56; primary flagellum long with 26–30 segments, with aesthetascs, with 6-segmented accessory flagellum (
<figureCitation id="1300D391FFC8A408FD25F9E58A66F97B" box="[743,825,1656,1682]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="8" pageNumber="491">
Fig. 4 
<emphasis id="B94F1306FFC8A408FCE9F9E58A66F97B" box="[811,825,1656,1682]" italics="true" pageId="8" pageNumber="491">b</emphasis>
</figureCitation>
); both primary and accessory flagellum covered with short distal setae. 
<emphasis id="B94F1306FFC8A408FE9CF901888AF95F" box="[350,469,1692,1718]" italics="true" pageId="8" pageNumber="491">Antenna II</emphasis>
(
<figureCitation id="1300D391FFC8A408FE21F9018B12F95F" box="[483,589,1692,1718]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="8" pageNumber="491">
Fig. 4 
<emphasis id="B94F1306FFC8A408FDE7F9008B12F95F" box="[549,589,1692,1718]" italics="true" pageId="8" pageNumber="491">c, d</emphasis>
</figureCitation>
) sexually dimorphic: strong and densely setose with distally curved setae in males, with strong, forward pointing gland cone; peduncle articles II–V relatively stout, with ratio about 1.0:2.2:2.8, without calceoli and aesthetascs.
</paragraph>
<paragraph id="8B84CF14FFC8A40EFF05F8948D5AFE44" blockId="8.[151,1437,864,2043]" lastBlockId="14.[151,1437,150,2013]" lastPageId="14" lastPageNumber="497" pageId="8" pageNumber="491">
<emphasis id="B94F1306FFC8A408FF05F894886AF8CB" box="[199,309,1800,1826]" italics="true" pageId="8" pageNumber="491">Upper lip</emphasis>
(
<emphasis id="B94F1306FFC8A408FE87F89588C9F8CB" box="[325,406,1800,1826]" italics="true" pageId="8" pageNumber="491">labrum</emphasis>
) (
<figureCitation id="1300D391FFC8A408FE72F8958B5EF8CB" box="[432,513,1800,1827]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FE31F8948B5EF8CB" box="[499,513,1801,1826]" italics="true" pageId="8" pageNumber="491">a</emphasis>
</figureCitation>
) with concave distal margin, covered with minute setae. 
<emphasis id="B94F1306FFC8A408FB4BF8948DA8F8CB" box="[1161,1271,1800,1826]" italics="true" pageId="8" pageNumber="491">Lower lip</emphasis>
(
<emphasis id="B94F1306FFC8A408FAC5F8958C0AF8CB" box="[1287,1365,1800,1826]" italics="true" pageId="8" pageNumber="491">labium</emphasis>
) (
<figureCitation id="1300D391FFC8A408FAADF89589ECF8AF" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FF67F8B189ECF8AF" box="[165,179,1836,1862]" italics="true" pageId="8" pageNumber="491">b</emphasis>
</figureCitation>
) outer lobes covered with thin setae, bearing feebly developed inner lobes. 
<emphasis id="B94F1306FFC8A408FC3EF8B18D2DF8AF" box="[1020,1138,1836,1862]" italics="true" pageId="8" pageNumber="491">Mandibles</emphasis>
slightly asymmetrical; 
<emphasis id="B94F1306FFC8A408FABAF8B189A0F883" italics="true" pageId="8" pageNumber="491">left mandible</emphasis>
(
<figureCitation id="1300D391FFC8A408FECDF8CD8802F883" box="[271,349,1872,1899]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FE93F8CC8802F883" box="[337,349,1873,1898]" italics="true" pageId="8" pageNumber="491">c</emphasis>
</figureCitation>
): incisor with 9–10 small horizontally flattened teeth (
<figureCitation id="1300D391FFC8A408FC03F8CD8D4DF883" box="[961,1042,1872,1899]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FBC6F8CD8D4DF883" box="[1028,1042,1872,1898]" italics="true" pageId="8" pageNumber="491">d</emphasis>
</figureCitation>
), 
<emphasis id="B94F1306FFC8A408FBEBF8CD8D8FF883" box="[1065,1232,1872,1898]" italics="true" pageId="8" pageNumber="491">lacinia mobilis</emphasis>
with 4 teeth; with a row of 10 serrated setae between lacinia and molar, few spatulate setae and a long seta at base of molar (
<figureCitation id="1300D391FFC8A408FAFEF8E98CD3F867" box="[1340,1420,1908,1935]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FABCF8E98CD3F867" box="[1406,1420,1908,1934]" italics="true" pageId="8" pageNumber="491">d</emphasis>
</figureCitation>
), proximal article of palp without setae; article II with a tuft of long submarginal simple setae, distal article with 10 sets of 4–10 A-setae, 15 C-setae, 9–10 D-setae, almost equal in length, and 6 E-setae (
<figureCitation id="1300D391FFC8A408FB91F8218DFCF83F" box="[1107,1187,1980,2007]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FB57F8218DFCF83F" box="[1173,1187,1980,2006]" italics="true" pageId="8" pageNumber="491">d</emphasis>
</figureCitation>
); 
<emphasis id="B94F1306FFC8A408FB79F8218C3FF83F" box="[1211,1376,1980,2006]" italics="true" pageId="8" pageNumber="491">right mandible</emphasis>
(
<figureCitation id="1300D391FFC8A408FAADF82189EEF813" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="8" pageNumber="491">
Fig. 5 
<emphasis id="B94F1306FFC8A408FF67F87C89EEF813" box="[165,177,2017,2042]" italics="true" pageId="8" pageNumber="491">e</emphasis>
</figureCitation>
): 
<emphasis id="B94F1306FFC8A408FF08F87C882CF813" box="[202,371,2017,2042]" italics="true" pageId="8" pageNumber="491">incisor process</emphasis>
with 9 teeth, 
<emphasis id="B94F1306FFC8A408FDCEF87D8BEBF813" box="[524,692,2016,2042]" italics="true" pageId="8" pageNumber="491">lacinia mobilis</emphasis>
bifurcate, toothed, with a row of 6 serrated setae between lacinia and molar (
<figureCitation id="1300D391FFCEA40EFED5FF0A883DFF58" box="[279,354,150,177]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="14" pageNumber="497">
Fig. 5 
<emphasis id="B94F1306FFCEA40EFE98FF0A883DFF58" box="[346,354,151,177]" italics="true" pageId="14" pageNumber="497">f</emphasis>
</figureCitation>
). 
<emphasis id="B94F1306FFCEA40EFEBBFF0A88BFFF59" box="[377,480,151,177]" italics="true" pageId="14" pageNumber="497">Maxilla I</emphasis>
(
<figureCitation id="1300D391FFCEA40EFE32FF0A8B1FFF59" box="[496,576,150,177]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="14" pageNumber="497">
Fig. 
<quantity id="4CC362F1FFCEA40EFDE6FF0B8B1FFF59" box="[548,576,150,177]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="14" pageNumber="497" unit="g" value="5.0">
5 
<emphasis id="B94F1306FFCEA40EFDF0FF0A8B1FFF59" box="[562,576,151,176]" italics="true" pageId="14" pageNumber="497">g</emphasis>
</quantity>
</figureCitation>
) inner lobe broadening distally, distal margin straight, armed with numerous simple setae; outer lobe bluntly rounded distally, with about 20 hairbrush-like apical setae; palp asymmetrical, with a convex outer, and slightly concave inner margins, distal article about 3 times longer than basal right palp wide, with simple distal spines and lateral setae. 
<emphasis id="B94F1306FFCEA40EFD4BFE9E8BA2FEF5" box="[649,765,259,285]" italics="true" pageId="14" pageNumber="497">Maxilla II</emphasis>
(
<figureCitation id="1300D391FFCEA40EFCCDFE9E8A3DFEF4" box="[783,866,258,285]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="14" pageNumber="497">
Fig. 5 
<emphasis id="B94F1306FFCEA40EFC96FE9E8A3DFEF4" box="[852,866,259,285]" italics="true" pageId="14" pageNumber="497">h</emphasis>
</figureCitation>
) inner lobe broad, distally rounded, with slightly produced blunt ventral margin, armed with numerous setae medially and apically; outer lobe wider than inner, expanding distally, distal margin blunt, rounded with apical setae. 
<emphasis id="B94F1306FFCEA40EFCB0FED68AB4FE8C" box="[882,1003,331,357]" italics="true" pageId="14" pageNumber="497">Maxilliped</emphasis>
(
<figureCitation id="1300D391FFCEA40EFC38FED68D1CFE8D" box="[1018,1091,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[185,1407,185,1910]" captionTargetId="figure-35@10.[181,1407,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—upper lip; b—lower lip; c—left mandible; d—same, incisor process and pars incisiva; e—right mandible; f—same, incisor process and pars incisiva; g—left maxilla II; h—maxilla I; i—maxilliped." figureDoi="http://doi.org/10.5281/zenodo.8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="14" pageNumber="497">
Fig. 5 
<emphasis id="B94F1306FFCEA40EFBF9FED68D1CFE8D" box="[1083,1091,331,356]" italics="true" pageId="14" pageNumber="497">i</emphasis>
</figureCitation>
): inner and outer plates bluntly rounded and slightly broadening distally, densely setose, inner lobe with 2–3 short and stout apical spines; palp 4- articulated, covered with numerous long simple setae, terminal article hooked.
</paragraph>
<caption id="DF449F9CFFC9A409FF55F80A8A56F811" ID-DOI="http://doi.org/10.5281/zenodo.8009137" ID-Zenodo-Dep="8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="9" pageNumber="492" startId="9.[151,250,1943,1967]" targetBox="[184,1402,186,1916]" targetPageId="9" targetType="figure">
<paragraph id="8B84CF14FFC9A409FF55F80A8A56F811" blockId="9.[151,1436,1943,2040]" pageId="9" pageNumber="492">
<emphasis id="B94F1306FFC9A409FF55F80A884CF846" bold="true" box="[151,275,1943,1967]" pageId="9" pageNumber="492">FIGURE 4.</emphasis>
<taxonomicName id="4C3BB497FFC9A409FED8F80588A2F846" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[282,509,1943,1967]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="492" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFC9A409FED8F80588A2F846" box="[282,509,1943,1967]" italics="true" pageId="9" pageNumber="492">Litorogammarus dursi</emphasis>
</taxonomicName>
<emphasis id="B94F1306FFC9A409FDC6F8058B04F859" bold="true" box="[516,603,1944,1968]" pageId="9" pageNumber="492">
<taxonomicNameLabel id="A27CAE7DFFC9A409FDC6F8058B08F859" box="[516,599,1944,1968]" pageId="9" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype Ô, ZMMU Mb-1243 (
<emphasis id="B94F1306FFC9A409FC61F8058D44F846" box="[931,1051,1944,1968]" italics="true" pageId="9" pageNumber="492">a–c, e, f, i, j</emphasis>
), paratype ♀, ZMMU Mb-1244 (
<emphasis id="B94F1306FFC9A409FAB2F80A8991F83A" italics="true" pageId="9" pageNumber="492">d, g, h, k, l</emphasis>
): 
<emphasis id="B94F1306FFC9A409FF21F82189AFF83A" box="[227,240,1980,2003]" italics="true" pageId="9" pageNumber="492">a</emphasis>
—antenna I; 
<emphasis id="B94F1306FFC9A409FEB1F82688DFF83A" box="[371,384,1979,2003]" italics="true" pageId="9" pageNumber="492">b</emphasis>
—accessory flagellum of antenna I; 
<emphasis id="B94F1306FFC9A409FD2FF8218A4EF83A" box="[749,785,1979,2003]" italics="true" pageId="9" pageNumber="492">c, d</emphasis>
—antenna II; 
<emphasis id="B94F1306FFC9A409FC5EF8218A9FF83A" box="[924,960,1980,2003]" italics="true" pageId="9" pageNumber="492">e, g</emphasis>
—gnathopod I; 
<emphasis id="B94F1306FFC9A409FB9DF8218D39F83D" box="[1119,1126,1980,2004]" italics="true" pageId="9" pageNumber="492">f</emphasis>
, 
<emphasis id="B94F1306FFC9A409FBB0F8268D20F83A" box="[1138,1151,1979,2003]" italics="true" pageId="9" pageNumber="492">h</emphasis>
—palmar margin of chela of GnI; 
<emphasis id="B94F1306FFC9A409FF0EF87D885EF81E" box="[204,257,2015,2039]" italics="true" pageId="9" pageNumber="492">i, k—</emphasis>
gnathopod II; 
<emphasis id="B94F1306FFC9A409FE53F87D88C7F81E" box="[401,408,2016,2039]" italics="true" pageId="9" pageNumber="492">j</emphasis>
, 
<emphasis id="B94F1306FFC9A409FE67F84288F3F81E" box="[421,428,2015,2039]" italics="true" pageId="9" pageNumber="492">l</emphasis>
—palmar margin of chela of GnII.
</paragraph>
</caption>
<caption id="DF449F9CFFCAA40AFF55F80A88EBF81E" ID-DOI="http://doi.org/10.5281/zenodo.8009139" ID-Zenodo-Dep="8009139" httpUri="https://zenodo.org/record/8009139/files/figure.png" pageId="10" pageNumber="493" startId="10.[151,250,1943,1967]" targetBox="[185,1407,185,1910]" targetPageId="10" targetType="figure">
<paragraph id="8B84CF14FFCAA40AFF55F80A88EBF81E" blockId="10.[151,1437,1943,2039]" pageId="10" pageNumber="493">
<emphasis id="B94F1306FFCAA40AFF55F80A8849F859" bold="true" box="[151,278,1943,1968]" pageId="10" pageNumber="493">FIGURE 5.</emphasis>
<taxonomicName id="4C3BB497FFCAA40AFEDDF8058B59F846" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[287,518,1943,1967]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="493" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFCAA40AFEDDF8058B59F846" box="[287,518,1943,1967]" italics="true" pageId="10" pageNumber="493">Litorogammarus dursi</emphasis>
</taxonomicName>
<emphasis id="B94F1306FFCAA40AFDD2F8058B35F859" bold="true" box="[528,618,1944,1968]" pageId="10" pageNumber="493">
<taxonomicNameLabel id="A27CAE7DFFCAA40AFDD2F8058B3AF859" box="[528,613,1944,1968]" pageId="10" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype Ô, ZMMU Mb-1243: 
<emphasis id="B94F1306FFCAA40AFC7DF8058A93F846" box="[959,972,1944,1967]" italics="true" pageId="10" pageNumber="493">a</emphasis>
—upper lip; 
<emphasis id="B94F1306FFCAA40AFB91F80A8D3FF846" box="[1107,1120,1943,1967]" italics="true" pageId="10" pageNumber="493">b</emphasis>
—lower lip; 
<emphasis id="B94F1306FFCAA40AFB25F8058DADF846" box="[1255,1266,1944,1967]" italics="true" pageId="10" pageNumber="493">c</emphasis>
—left mandible; 
<emphasis id="B94F1306FFCAA40AFF55F82689E5F83A" box="[151,186,1979,2003]" italics="true" pageId="10" pageNumber="493">d—</emphasis>
same, incisor process and pars incisiva; 
<emphasis id="B94F1306FFCAA40AFD98F8218B3AF83A" box="[602,613,1980,2003]" italics="true" pageId="10" pageNumber="493">e</emphasis>
—right mandible; 
<emphasis id="B94F1306FFCAA40AFCE6F8218A1EF83D" box="[804,833,1980,2004]" italics="true" pageId="10" pageNumber="493">f—</emphasis>
same, incisor process and pars incisiva; 
<emphasis id="B94F1306FFCAA40AFB23F8218DB1F83A" box="[1249,1262,1980,2003]" italics="true" pageId="10" pageNumber="493">g</emphasis>
—left maxilla II; 
<emphasis id="B94F1306FFCAA40AFF55F84289E5F81E" box="[151,186,2015,2039]" italics="true" pageId="10" pageNumber="493">h—</emphasis>
maxilla I; 
<emphasis id="B94F1306FFCAA40AFEE0F87D881FF81E" box="[290,320,2016,2039]" italics="true" pageId="10" pageNumber="493">i—</emphasis>
maxilliped.
</paragraph>
</caption>
<caption id="DF449F9CFFCBA40BFF55F80A89B4F81E" ID-DOI="http://doi.org/10.5281/zenodo.8009143" ID-Zenodo-Dep="8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="11" pageNumber="494" startId="11.[151,250,1943,1967]" targetBox="[186,1398,184,1916]" targetPageId="11" targetType="figure">
<paragraph id="8B84CF14FFCBA40BFF55F80A89B4F81E" blockId="11.[151,1436,1943,2039]" pageId="11" pageNumber="494">
<emphasis id="B94F1306FFCBA40BFF55F80A884DF846" bold="true" box="[151,274,1943,1967]" pageId="11" pageNumber="494">FIGURE 6.</emphasis>
<taxonomicName id="4C3BB497FFCBA40BFEDAF80588A4F846" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[280,507,1943,1967]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="494" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFCBA40BFEDAF80588A4F846" box="[280,507,1943,1967]" italics="true" pageId="11" pageNumber="494">Litorogammarus dursi</emphasis>
</taxonomicName>
<emphasis id="B94F1306FFCBA40BFDC3F8058B0EF859" bold="true" box="[513,593,1944,1968]" pageId="11" pageNumber="494">
<taxonomicNameLabel id="A27CAE7DFFCBA40BFDC3F8058B0EF859" box="[513,593,1944,1968]" pageId="11" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype Ô, ZMMU Mb-1243: 
<emphasis id="B94F1306FFCBA40BFC58F8058AF8F846" box="[922,935,1944,1967]" italics="true" pageId="11" pageNumber="494">a</emphasis>
—pereopod III; 
<emphasis id="B94F1306FFCBA40BFB88F80A8D08F846" box="[1098,1111,1943,1967]" italics="true" pageId="11" pageNumber="494">b</emphasis>
—dactylus of PIII; 
<emphasis id="B94F1306FFCBA40BFADBF8058C7BF846" box="[1305,1316,1944,1967]" italics="true" pageId="11" pageNumber="494">c</emphasis>
—pereopod IV; 
<emphasis id="B94F1306FFCBA40BFF7CF8268994F83A" box="[190,203,1979,2003]" italics="true" pageId="11" pageNumber="494">d</emphasis>
—dactylus of PIV; 
<emphasis id="B94F1306FFCBA40BFE4DF82188C5F83A" box="[399,410,1980,2003]" italics="true" pageId="11" pageNumber="494">e</emphasis>
—pereopod V; 
<emphasis id="B94F1306FFCBA40BFDF4F8268B0BF83A" box="[566,596,1979,2003]" italics="true" pageId="11" pageNumber="494">f—</emphasis>
dactylus of PV; 
<emphasis id="B94F1306FFCBA40BFD35F8218A45F83A" box="[759,794,1980,2003]" italics="true" pageId="11" pageNumber="494">g—</emphasis>
pereopod VI; 
<emphasis id="B94F1306FFCBA40BFC65F8268A95F83A" box="[935,970,1979,2003]" italics="true" pageId="11" pageNumber="494">h—</emphasis>
dactylus of PVI; 
<emphasis id="B94F1306FFCBA40BFBB5F8218DCAF83A" box="[1143,1173,1980,2003]" italics="true" pageId="11" pageNumber="494">i—</emphasis>
pereopod VII; 
<emphasis id="B94F1306FFCBA40BFAE8F8218C17F83A" box="[1322,1352,1980,2003]" italics="true" pageId="11" pageNumber="494">j—</emphasis>
dactylus of PVII.
</paragraph>
</caption>
<caption id="DF449F9CFFCCA40CFF55F80A89E7F81E" ID-DOI="http://doi.org/10.5281/zenodo.8009147" ID-Zenodo-Dep="8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="12" pageNumber="495" startId="12.[151,250,1943,1967]" targetBox="[183,1405,184,1917]" targetPageId="12" targetType="figure">
<paragraph id="8B84CF14FFCCA40CFF55F80A89E7F81E" blockId="12.[151,1437,1943,2039]" pageId="12" pageNumber="495">
<emphasis id="B94F1306FFCCA40CFF55F80A884CF859" bold="true" box="[151,275,1943,1968]" pageId="12" pageNumber="495">FIGURE 7.</emphasis>
<taxonomicName id="4C3BB497FFCCA40CFEDBF80588A2F846" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[281,509,1943,1967]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="495" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFCCA40CFEDBF80588A2F846" box="[281,509,1943,1967]" italics="true" pageId="12" pageNumber="495">Litorogammarus dursi</emphasis>
</taxonomicName>
<emphasis id="B94F1306FFCCA40CFDC1F8058B0CF859" bold="true" box="[515,595,1944,1968]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="A27CAE7DFFCCA40CFDC1F8058B0CF859" box="[515,595,1944,1968]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype Ô, ZMMU Mb-1243 (
<emphasis id="B94F1306FFCCA40CFC62F8058D5AF846" box="[928,1029,1943,1968]" italics="true" pageId="12" pageNumber="495">a–f, h, j, k</emphasis>
), paratype ♀, ZMMU Mb-1244 (
<emphasis id="B94F1306FFCCA40CFA9BF8058CD2F846" box="[1369,1421,1943,1967]" italics="true" pageId="12" pageNumber="495">g, i, l</emphasis>
): 
<emphasis id="B94F1306FFCCA40CFF55F82189E3F83A" box="[151,188,1980,2003]" italics="true" pageId="12" pageNumber="495">a–c</emphasis>
—epimeral plates I–III; 
<emphasis id="B94F1306FFCCA40CFE76F826889EF83A" box="[436,449,1979,2003]" italics="true" pageId="12" pageNumber="495">d</emphasis>
—telson; 
<emphasis id="B94F1306FFCCA40CFDE7F8218B18F83A" box="[549,583,1980,2003]" italics="true" pageId="12" pageNumber="495">e—</emphasis>
pleopod I; 
<emphasis id="B94F1306FFCCA40CFD74F8218B8BF83D" box="[694,724,1980,2004]" italics="true" pageId="12" pageNumber="495">f—</emphasis>
retinacula of pleopod I; 
<emphasis id="B94F1306FFCCA40CFC09F8268AADF83A" box="[971,1010,1979,2003]" italics="true" pageId="12" pageNumber="495">h, g</emphasis>
—uropod I; 
<emphasis id="B94F1306FFCCA40CFBB3F8218DD2F83A" box="[1137,1165,1980,2003]" italics="true" pageId="12" pageNumber="495">i, j</emphasis>
—pleopod II; 
<emphasis id="B94F1306FFCCA40CFADCF8268C0AF83A" box="[1310,1365,1979,2003]" italics="true" pageId="12" pageNumber="495">k, l—</emphasis>
uropod III.
</paragraph>
</caption>
<caption id="DF449F9CFFCDA40DFF55F80F8AEBF824" ID-DOI="http://doi.org/10.5281/zenodo.8009154" ID-Zenodo-Dep="8009154" httpUri="https://zenodo.org/record/8009154/files/figure.png" pageId="13" pageNumber="496" startId="13.[151,250,1938,1962]" targetBox="[151,1435,181,1914]" targetPageId="13" targetType="figure">
<paragraph id="8B84CF14FFCDA40DFF55F80F8AEBF824" blockId="13.[151,1436,1938,1998]" pageId="13" pageNumber="496">
<emphasis id="B94F1306FFCDA40DFF55F80F884DF843" bold="true" box="[151,274,1938,1962]" pageId="13" pageNumber="496">FIGURE 8.</emphasis>
<taxonomicName id="4C3BB497FFCDA40DFEDAF80E88A4F843" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[280,507,1938,1962]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="496" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFCDA40DFEDAF80E88A4F843" box="[280,507,1938,1962]" italics="true" pageId="13" pageNumber="496">Litorogammarus dursi</emphasis>
</taxonomicName>
<emphasis id="B94F1306FFCDA40DFDC3F80F8B07F843" bold="true" box="[513,600,1938,1962]" pageId="13" pageNumber="496">
<taxonomicNameLabel id="A27CAE7DFFCDA40DFDC3F80F8B0CF843" box="[513,595,1938,1962]" pageId="13" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
Ô (LEMMI): 
<emphasis id="B94F1306FFCDA40DFD2AF80E8BAAF843" box="[744,757,1939,1962]" italics="true" pageId="13" pageNumber="496">a</emphasis>
—head; 
<emphasis id="B94F1306FFCDA40DFC89F80F8A07F843" box="[843,856,1938,1962]" italics="true" pageId="13" pageNumber="496">b</emphasis>
—urosomal segments, lateral view; 
<emphasis id="B94F1306FFCDA40DFB00F80E8DBBF843" box="[1218,1252,1939,1962]" italics="true" pageId="13" pageNumber="496">c—</emphasis>
urosomal segment and telson, dorsal view; 
<emphasis id="B94F1306FFCDA40DFE4CF82B88C4F827" box="[398,411,1974,1998]" italics="true" pageId="13" pageNumber="496">d</emphasis>
—urosomal, lateral view; 
<emphasis id="B94F1306FFCDA40DFD61F82A8BF1F827" box="[675,686,1975,1998]" italics="true" pageId="13" pageNumber="496">e</emphasis>
—retinacula of pleopod I.
</paragraph>
</caption>
<paragraph id="8B84CF14FFCEA40EFF05FE2A8CCEFD24" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="497">
<emphasis id="B94F1306FFCEA40EFF05FE2A880BFE39" box="[199,340,439,465]" italics="true" pageId="14" pageNumber="497">Gnathopod I</emphasis>
(
<figureCitation id="1300D391FFCEA40EFEA1FE2A88A3FE38" box="[355,508,439,465]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFE67FE2A88A3FE38" box="[421,508,439,465]" italics="true" pageId="14" pageNumber="497">e, f, g, h</emphasis>
</figureCitation>
) with coxal plate distally slightly widened, concave, with 1 distal and 1–2 subdistal setae on anterior margin; basis sub-linear, with numerous long anterior and posterior setae; ischium about as long as wide, equal to merus; merus about 1.5 times longer than wide, with medium setae on posterior margin; carpus short, almost triangular, with a cluster of long setae on posterior margin; propodus (palm) pyriform in males (
<figureCitation id="1300D391FFCEA40EFB3EFDBE8C17FDD5" box="[1276,1352,547,573]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFAFEFDBE8C17FDD5" box="[1340,1352,547,572]" italics="true" pageId="14" pageNumber="497">e</emphasis>
</figureCitation>
), about 2–2.5 times as long as wide, with oblique distoventral part of palmar margin in males and straight in females; palmar margin crenellated in its proximal (angular) part, with 5–7 palmar marginal spines and 1 mid-palmar spine in male (
<figureCitation id="1300D391FFCEA40EFF5DFD1289B5FD40" box="[159,234,655,681]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFF20FD1289B5FD40" box="[226,234,655,681]" italics="true" pageId="14" pageNumber="497">f</emphasis>
</figureCitation>
), and with 1 palmar angle spine and 1 short mid-palmar spine in females (
<figureCitation id="1300D391FFCEA40EFBE9FD128D24FD40" box="[1067,1147,655,681]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFBAFFD128D24FD40" box="[1133,1147,655,681]" italics="true" pageId="14" pageNumber="497">h</emphasis>
</figureCitation>
); dactylus reaching about 40% length of propodus in males and less than 30% in females, posterior margin convex, with 2 long medial setae.
</paragraph>
<paragraph id="8B84CF14FFCEA40EFF05FD4A8A9CFBDC" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="497">
<emphasis id="B94F1306FFCEA40EFF05FD4A883FFD19" box="[199,352,727,753]" italics="true" pageId="14" pageNumber="497">Gnathopod II</emphasis>
(
<figureCitation id="1300D391FFCEA40EFEB2FD4A8B5BFD18" box="[368,516,727,753]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFE71FD4A8B5BFD18" box="[435,516,727,753]" italics="true" pageId="14" pageNumber="497">i, j, k, l</emphasis>
</figureCitation>
) significantly larger in males; coxal plate subrectangular, with convex dorsal and ventral margins, with 1–2 setae on anterior margin subdorsally, without setae on lower margin and facial setae; basis sub-linear, with several long anterior setae and posterior setae; ischium about as long as wide, equal to merus; merus about as long as wide, with several long setae on posterior margin; carpus triangular, with slightly marked ventral lobe, armed with a cluster of long setae on posterior margin; propodus (palm) deeply trapezoidal, about 1.7 times as long as wide, with oblique distoventral part of palmar margin in males (
<figureCitation id="1300D391FFCEA40EFC14FC168D7FFC4D" box="[982,1056,907,933]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFBDAFC168D7FFC4D" box="[1048,1056,907,932]" italics="true" pageId="14" pageNumber="497">i</emphasis>
</figureCitation>
) and rectangular elongated, about 2.5 times as long a wide, with straight palmar margin in females (
<figureCitation id="1300D391FFCEA40EFCA8FC328AE5FC21" box="[874,954,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 
<quantity id="4CC362F1FFCEA40EFC5CFC328AE5FC21" box="[926,954,943,969]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="14" pageNumber="497" unit="g" value="4.0">
4 
<emphasis id="B94F1306FFCEA40EFC6EFC328AE5FC21" box="[940,954,943,968]" italics="true" pageId="14" pageNumber="497">g</emphasis>
</quantity>
</figureCitation>
); palmar margin crenellated in its proximal (angular) part, with row of 9–10 thin marginal spines and 4 mid-palmar spines in males (
<figureCitation id="1300D391FFCEA40EFBB1FC4E8DE2FC05" box="[1139,1213,979,1005]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFB77FC4E8DE2FC05" box="[1205,1213,979,1004]" italics="true" pageId="14" pageNumber="497">j</emphasis>
</figureCitation>
) and with 3 angular and 1 mid-palmar spines in females (
<figureCitation id="1300D391FFCEA40EFDFCFC6A8BCFFBF8" box="[574,656,1015,1041]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="14" pageNumber="497">
Fig. 4 
<emphasis id="B94F1306FFCEA40EFD40FC6A8BCFFBF8" box="[642,656,1015,1041]" italics="true" pageId="14" pageNumber="497">h</emphasis>
</figureCitation>
); dactylus reaching about 40% length of propodus in males and less than 30% in females, posterior margin convex, with 2 long medial setae.
</paragraph>
<paragraph id="8B84CF14FFCEA40EFF05FBA28AC4F87C" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="497">
<emphasis id="B94F1306FFCEA40EFF05FBA28802FBB1" box="[199,349,1087,1113]" italics="true" pageId="14" pageNumber="497">Pereiopod III</emphasis>
(
<figureCitation id="1300D391FFCEA40EFEA9FBA288E5FBB1" box="[363,442,1087,1113]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFE6EFBA288E5FBB1" box="[428,442,1087,1112]" italics="true" pageId="14" pageNumber="497">a</emphasis>
</figureCitation>
) with segments covered with numerous anterior and posterior stout and small setae; coxal plate III almost rectangular, concave distally, with 1 seta on anteroventral margin; basis sub-linear, about 3.5 times as long as wide; ischium as long as wide; merus about 3 times as long as wide; carpus about 4 times as long as wide, with anterior margin smooth with distodorsal setae only; propodus about 5 times as long as wide, with smooth anterior margin; dactylus (
<figureCitation id="1300D391FFCEA40EFE06FB528B4AFB00" box="[452,533,1231,1257]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFDC5FB528B4AFB00" box="[519,533,1231,1257]" italics="true" pageId="14" pageNumber="497">d</emphasis>
</figureCitation>
) relatively stout, curved, with 1 seta at hinge of unguis. 
<emphasis id="B94F1306FFCEA40EFB5EFB528C6DFB01" box="[1180,1330,1231,1257]" italics="true" pageId="14" pageNumber="497">Pereiopod IV</emphasis>
(
<figureCitation id="1300D391FFCEA40EFA81FB528CCDFB01" box="[1347,1426,1231,1257]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFA44FB528CCDFB01" box="[1414,1426,1231,1256]" italics="true" pageId="14" pageNumber="497">c</emphasis>
</figureCitation>
) with segments covered with numerous anterior and posterior stout and small simple setae; coxal plate IV almost quadrate, e distal margin, concave, with 4–5 small setae along lower anterior margin; basis sub-linear, about 3.5 times as long as wide; ischium as long as wide; merus about 3 times as long as wide; carpus about 4 times as long as wide, with smooth anterior margin and distodorsal spines only; propodus about 4 times as long as wide, with smooth anterior margin; dactylus (
<figureCitation id="1300D391FFCEA40EFE79FA1E8B55FA74" box="[443,522,1411,1437]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFE3EFA1E8B55FA74" box="[508,522,1411,1437]" italics="true" pageId="14" pageNumber="497">d</emphasis>
</figureCitation>
) stout, curved, with 1 simple seta at hinge of unguis. 
<emphasis id="B94F1306FFCEA40EFB96FA1E8D81FA75" box="[1108,1246,1411,1437]" italics="true" pageId="14" pageNumber="497">Pereiopod V</emphasis>
(
<figureCitation id="1300D391FFCEA40EFB2EFA1E8C66FA75" box="[1260,1337,1411,1437]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFAEFFA1E8C66FA75" box="[1325,1337,1411,1436]" italics="true" pageId="14" pageNumber="497">e</emphasis>
</figureCitation>
) slightly shorter than PpVI–VII; coxal plates V bilobate, with 1 small seta on anterior lobe and 1 long spine on posterior lobe; basis relatively wide, about 1.5 times as long as wide, with dorsal margin slightly convex, armed 5–6 small stout spines, with well-developed ventral lobe armed with short spines, posteroventral angle bluntly rounded; ischium subquadrate, about as long as wide; merus about 2–2.5 times as long as wide; carpus about 3 times as long as wide; carpal and meral segments with strong spines and spiniform setae along anterior and posterior margins; propodus about 7–8 times as long as wide, with clusters of small spines along anterior margin and long simple spines along posterior margin; dactylus (
<figureCitation id="1300D391FFCEA40EFE16F9E28B7EF970" box="[468,545,1663,1689]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFDDBF9E28B7EF970" box="[537,545,1663,1689]" italics="true" pageId="14" pageNumber="497">f</emphasis>
</figureCitation>
) stout, curved, with 1 seta at hinge of unguis. 
<emphasis id="B94F1306FFCEA40EFB83F9E28C4AF971" box="[1089,1301,1663,1689]" italics="true" pageId="14" pageNumber="497">Pereiopods VI–VII</emphasis>
(
<figureCitation id="1300D391FFCEA40EFAEAF9E28CCCF971" box="[1320,1427,1663,1689]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFAAFF9E28CCCF971" box="[1389,1427,1663,1689]" italics="true" pageId="14" pageNumber="497">f, g</emphasis>
</figureCitation>
) almost similar in shape, PVII slightly stronger than PVI; coxal plates VI–VII bilobate, with anterior lobe bearing 1–2 spines or unarmed, with 1–2 spines on posterior lobe; basis wider in proximal part and narrowing distally, with relatively short spines along anterior margin and short spiniform setae along posterior margin; ischium subquadrate, about as long as wide; merus about 2.5 times as long as wide; carpus about 3.5–4 times as long as wide; propodus about 6 and 5 times as long as wide in PVI and PVII, respectively; carpal, meral and propodal segments with strong spines and stout setae along anterior and posterior margins; dactylus (
<figureCitation id="1300D391FFCEA40EFC6AF8CA8D4FF899" box="[936,1040,1879,1905]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[186,1398,184,1916]" captionTargetId="figure-17@11.[181,1407,181,1919]" captionTargetPageId="11" captionText="FIGURE 6. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243: a—pereopod III; b—dactylus of PIII; c—pereopod IV; d—dactylus of PIV; e—pereopod V; f—dactylus of PV; g—pereopod VI; h—dactylus of PVI; i—pereopod VII; j—dactylus of PVII." figureDoi="http://doi.org/10.5281/zenodo.8009143" httpUri="https://zenodo.org/record/8009143/files/figure.png" pageId="14" pageNumber="497">
Fig. 6 
<emphasis id="B94F1306FFCEA40EFC29F8CA8D4FF899" box="[1003,1040,1879,1905]" italics="true" pageId="14" pageNumber="497">h, j</emphasis>
</figureCitation>
) stout, curved, with 1 seta at hinge of unguis. 
<emphasis id="B94F1306FFCEA40EFECCF8E688D7F87C" box="[270,392,1915,1941]" italics="true" pageId="14" pageNumber="497">Coxal gills</emphasis>
balloon-shaped, present on GnII and PpIII–VI.
</paragraph>
<paragraph id="8B84CF14FFCEA40FFF05F80288D3FF10" blockId="14.[151,1437,150,2013]" lastBlockId="15.[151,1437,151,1473]" lastPageId="15" lastPageNumber="498" pageId="14" pageNumber="497">
<emphasis id="B94F1306FFCEA40EFF05F8028822F851" box="[199,381,1951,1977]" italics="true" pageId="14" pageNumber="497">Epimeral plate I</emphasis>
(
<figureCitation id="1300D391FFCEA40EFE4EF8028883F851" box="[396,476,1951,1977]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="14" pageNumber="497">
Fig. 7 
<emphasis id="B94F1306FFCEA40EFE0CF8028883F851" box="[462,476,1951,1976]" italics="true" pageId="14" pageNumber="497">a</emphasis>
</figureCitation>
) with triangular posteroventral corner, posterior margin with 1 spine and 1 stout setae, ventral margin slightly convex. 
<emphasis id="B94F1306FFCEA40EFE34F85E8BEAF835" box="[502,693,1987,2013]" italics="true" pageId="14" pageNumber="497">Epimeral plate II</emphasis>
(
<figureCitation id="1300D391FFCEA40EFD00F85E8A4EF834" box="[706,785,1987,2013]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="14" pageNumber="497">
Fig. 7 
<emphasis id="B94F1306FFCEA40EFCC1F85E8A4EF834" box="[771,785,1987,2013]" italics="true" pageId="14" pageNumber="497">b</emphasis>
</figureCitation>
) ventrally convex, anterior part with 4 submarginal spines, posterior corner triangularly produced, bearing several small setae on posterior margin. 
<emphasis id="B94F1306FFCFA40FFB4FFF0A8C02FF59" box="[1165,1373,151,177]" italics="true" pageId="15" pageNumber="498">Epimeral plate III</emphasis>
(
<figureCitation id="1300D391FFCFA40FFAADFF0A89EEFF3D" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 7 
<emphasis id="B94F1306FFCFA40FFF67FF2689EEFF3D" box="[165,177,187,212]" italics="true" pageId="15" pageNumber="498">c</emphasis>
</figureCitation>
) almost straight ventrally, with 4 submarginal spine-like setae in its anterior part, with produced bluntly pointed posteroventral corner.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FE9E8AF9FEA8" blockId="15.[151,1437,151,1473]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FE9E8836FEF5" box="[199,361,259,285]" italics="true" pageId="15" pageNumber="498">Pleopods I–III</emphasis>
(
<figureCitation id="1300D391FFCFA40FFEBBFE9E8898FEF5" box="[377,455,259,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 7 
<emphasis id="B94F1306FFCFA40FFE79FE9E8898FEF5" box="[443,455,259,284]" italics="true" pageId="15" pageNumber="498">e</emphasis>
</figureCitation>
) similar in shape; both rami almost subequal, with plumose setae, basal segment with 2 coupling slender hooks and 1–2 long setae in retinacules (
<figureCitation id="1300D391FFCFA40FFCDDFEBA8A31FEA8" box="[799,878,295,321]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Figs 7 
<emphasis id="B94F1306FFCFA40FFCA4FEBA8A31FEA8" box="[870,878,295,321]" italics="true" pageId="15" pageNumber="498">f</emphasis>
</figureCitation>
, 
<figureCitation id="1300D391FFCFA40FFCB9FEBA8ACAFEA9" box="[891,917,295,321]" captionStart="FIGURE 8" captionStartId="13.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-17@13.[151,1435,181,1914]" captionTargetPageId="13" captionText="FIGURE 8. Litorogammarus dursi sp. nov., Ô (LEMMI): a—head; b—urosomal segments, lateral view; c—urosomal segment and telson, dorsal view; d—urosomal, lateral view; e—retinacula of pleopod I." figureDoi="http://doi.org/10.5281/zenodo.8009154" httpUri="https://zenodo.org/record/8009154/files/figure.png" pageId="15" pageNumber="498">
8 
<emphasis id="B94F1306FFCFA40FFC4BFEBA8ACAFEA9" box="[905,917,295,320]" italics="true" pageId="15" pageNumber="498">e</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FED68AC0FD18" blockId="15.[151,1437,151,1473]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FED6886EFE8D" box="[199,305,331,357]" italics="true" pageId="15" pageNumber="498">Uropod I</emphasis>
(
<figureCitation id="1300D391FFCFA40FFE86FED688E8FE8C" box="[324,439,331,357]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 
<quantity id="4CC362F1FFCFA40FFEBEFED688C5FE8D" box="[380,410,331,357]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="15" pageNumber="498" unit="g" value="7.0">
7 
<emphasis id="B94F1306FFCFA40FFE48FED688C5FE8D" box="[394,410,331,356]" italics="true" pageId="15" pageNumber="498">g</emphasis>
</quantity>
, h
</figureCitation>
) with peduncle armed with 1 large proximoventral spine, with 3–4 inner and 3 outer basiofacial spines, with 2 subdistal spines; inner ramus equal to outer ramus; outer ramus (exopodite) with 2 dorsal and 4 subdistal dorsal spines; rami equal in size, inner ramus (endopodite) with 3 dorsal and 3 apical spines, outer ramus (exopodite) with 3 dorsal and 3 apical spines. 
<emphasis id="B94F1306FFCFA40FFD32FE2A8A3EFE39" box="[752,865,439,465]" italics="true" pageId="15" pageNumber="498">Uropod II</emphasis>
(
<figureCitation id="1300D391FFCFA40FFCB3FE2A8A8CFE39" box="[881,979,439,465]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 7 
<emphasis id="B94F1306FFCFA40FFC76FE2A8A8CFE39" box="[948,979,439,464]" italics="true" pageId="15" pageNumber="498">i, j</emphasis>
</figureCitation>
) smaller than uropod I; peduncle with 3 inner and 2 outer basiofacial spines, with 2 subdistal dorsal spines; outer ramus (exopodite) equal to inner ramus (endopodite); inner ramus with 3 dorsal, 1 mesial and 5 apical spines; outer ramus with 2 dorsal, 1 mesial and 4 apical spines. 
<emphasis id="B94F1306FFCFA40FFF2EFDBE8838FDD5" box="[236,359,547,573]" italics="true" pageId="15" pageNumber="498">Uropod III</emphasis>
(
<figureCitation id="1300D391FFCFA40FFEB5FDBE8880FDD4" box="[375,479,547,573]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 7 
<emphasis id="B94F1306FFCFA40FFE79FDBE8880FDD4" box="[443,479,547,573]" italics="true" pageId="15" pageNumber="498">k, l</emphasis>
</figureCitation>
) of parviramous 
<typeStatus id="548071B6FFCFA40FFD66FDBE8B87FDD4" box="[676,728,547,573]" pageId="15" pageNumber="498">type</typeStatus>
; peduncle about 1.2 times longer than wide, with 2 clusters of subdistal spines; inner ramus (endopodite) reduced, scale-like, about 6 times shorter than outer ramus (endopodite) in males, with a row of 5–7 subdistal spines and several small setae; outer ramus (endopodite) 2-segmented, with proximal segment armed with 7 groups of spines and/or short setae along outer margin, with 4 groups of short spines and plumose setae along inner margin; distal segment short, but distinct, about 6–7 times shorter than proximal segment, about 2 times longer than subdistal spines accompanying it.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FD668BF3FCB4" blockId="15.[151,1437,151,1473]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FD668850FCFC" box="[199,271,763,789]" italics="true" pageId="15" pageNumber="498">Telson</emphasis>
(
<figureCitation id="1300D391FFCFA40FFEDDFD668830FCFC" box="[287,367,763,789]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 7 
<emphasis id="B94F1306FFCFA40FFEA3FD668830FCFC" box="[353,367,763,789]" italics="true" pageId="15" pageNumber="498">d</emphasis>
</figureCitation>
) entirely cleft; lobes elongated, suboval, about 1.8 times as long as wide, each lobe armed with a group of 3–4 strong apical spines and 2 small slender setae, and with 1, rarely 2 groups of 2–3 dorsal submedian strong spines, accompanying with simple setae.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FCFB8AF9FC04" blockId="15.[151,1437,151,1473]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FCFB88E6FC69" bold="true" box="[199,441,870,896]" pageId="15" pageNumber="498">Sexual dimorphism.</emphasis>
The stronger and densely setose antenna II and larger propodus of gnathopod II easily distinguish males from females (
<figureCitation id="1300D391FFCFA40FFDC7FC168B30FC4C" box="[517,623,907,933]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="15" pageNumber="498">
Fig. 4 
<emphasis id="B94F1306FFCFA40FFD85FC168B30FC4C" box="[583,623,907,933]" italics="true" pageId="15" pageNumber="498">c, d</emphasis>
</figureCitation>
); propodus (palm) of gnathopod I is subrectangular in females (
<figureCitation id="1300D391FFCFA40FFAFEFC168CD3FC4D" box="[1340,1420,907,933]" captionStart="FIGURE 4" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[184,1402,186,1916]" captionTargetId="figure-17@9.[181,1405,181,1919]" captionTargetPageId="9" captionText="FIGURE 4. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–c, e, f, i, j), paratype ♀, ZMMU Mb-1244 (d, g, h, k, l): a—antenna I; b—accessory flagellum of antenna I; c, d—antenna II; e, g—gnathopod I; f, h—palmar margin of chela of GnI; i, k—gnathopod II; j, l—palmar margin of chela of GnII." figureDoi="http://doi.org/10.5281/zenodo.8009137" httpUri="https://zenodo.org/record/8009137/files/figure.png" pageId="15" pageNumber="498">
Fig. 
<quantity id="4CC362F1FFCFA40FFAB2FC168CD3FC4D" box="[1392,1420,907,933]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="15" pageNumber="498" unit="g" value="4.0">
4 
<emphasis id="B94F1306FFCFA40FFABCFC168CD3FC4D" box="[1406,1420,907,932]" italics="true" pageId="15" pageNumber="498">g</emphasis>
</quantity>
</figureCitation>
). Coxal gills are quite linear in females. Uropod III is also larger in males than in females (
<figureCitation id="1300D391FFCFA40FFB54FC328DA2FC20" box="[1174,1277,943,969]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[183,1405,184,1917]" captionTargetId="figure-35@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 7. Litorogammarus dursi sp. nov., holotype Ô, ZMMU Mb-1243 (a–f, h, j, k), paratype ♀, ZMMU Mb-1244 (g, i, l): a–c—epimeral plates I–III; d—telson; e—pleopod I; f—retinacula of pleopod I; h, g—uropod I; i, j—pleopod II; k, l—uropod III." figureDoi="http://doi.org/10.5281/zenodo.8009147" httpUri="https://zenodo.org/record/8009147/files/figure.png" pageId="15" pageNumber="498">
Fig. 7 
<emphasis id="B94F1306FFCFA40FFB1BFC328DA2FC20" box="[1241,1277,943,969]" italics="true" pageId="15" pageNumber="498">k, l</emphasis>
</figureCitation>
); inner ramus (endopodite) is about 4–5 times in females vs. about 6 times in males.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FC6B8D41FBF8" blockId="15.[151,1437,151,1473]" box="[199,1054,1014,1041]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FC6B8810FBF9" bold="true" box="[199,335,1014,1040]" pageId="15" pageNumber="498">Coloration.</emphasis>
Body and appendages translucent. Eyes cornea silver or brown.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FB868D8DFBDC" blockId="15.[151,1437,151,1473]" box="[199,1234,1050,1079]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FB868860FBDD" bold="true" box="[199,319,1050,1077]" pageId="15" pageNumber="498">Body size.</emphasis>
The largest collected 
<specimenCount id="9D3D049DFFCFA40FFDF4FB868B14FBDC" box="[566,587,1051,1077]" pageId="15" pageNumber="498" type="female">♀</specimenCount>
has tbl. 9.0 mm; the largest collected Ô has tbl. 11.0 mm.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FBA38AEBFBB0" blockId="15.[151,1437,151,1473]" box="[199,948,1086,1113]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FBA38B46FBB0" bold="true" box="[199,537,1086,1113]" pageId="15" pageNumber="498">GenBank accession number.</emphasis>
OQ407693–OQ407694 (this study).
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FBFF8DF2FAE4" blockId="15.[151,1437,151,1473]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FBFF88A9FB95" bold="true" box="[199,502,1122,1149]" pageId="15" pageNumber="498">Habitat and distribution.</emphasis>
The species is presently known only from its 
<typeStatus id="548071B6FFCFA40FFBCBFBFE8D66FB94" box="[1033,1081,1123,1149]" pageId="15" pageNumber="498">type</typeStatus>
locality—the lower part of the small coastal Durso River flowing into the Black Sea, southwestern Caucasus, 
<collectingCountry id="F32C8F84FFCFA40FFBCDFB1A8D02FB48" box="[1039,1117,1159,1185]" name="Russia" pageId="15" pageNumber="498">Russia</collectingCountry>
. All studied specimens were collected in the area of the river expansion with a relatively weak water current and numerous coastal aquatic vegetation, where this species is probably living. No specimens of the new species were collected in the river part with stronger currents or estuarine part with brackish water, under pebble, stones or boulders.
</paragraph>
<paragraph id="8B84CF14FFCFA40FFF05FA8B8CCBFA28" blockId="15.[151,1437,151,1473]" pageId="15" pageNumber="498">
<emphasis id="B94F1306FFCFA40FFF05FA8B88EFFAD8" bold="true" box="[199,432,1302,1329]" pageId="15" pageNumber="498">Taxonomic remark.</emphasis>
<taxonomicName id="4C3BB497FFCFA40FFE78FA8A8BECFAD8" authority="Marin &amp; Palatov &amp; Copilaș-Ciocianu, 2023" authorityName="Marin &amp; Palatov &amp; Copilaș-Ciocianu" authorityYear="2023" box="[442,691,1303,1329]" class="Malacostraca" family="Talitridae" genus="Litorogammarus" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="498" phylum="Arthropoda" rank="species" species="dursi" status="sp. nov.">
<emphasis id="B94F1306FFCFA40FFE78FA8A8BECFAD8" box="[442,691,1303,1329]" italics="true" pageId="15" pageNumber="498">Litorogammarus dursi</emphasis>
</taxonomicName>
<emphasis id="B94F1306FFCFA40FFD7EFA8A8A48FAD8" bold="true" box="[700,791,1303,1329]" pageId="15" pageNumber="498">
<taxonomicNameLabel id="A27CAE7DFFCFA40FFD7EFA8A8A48FAD8" box="[700,791,1303,1329]" pageId="15" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possess very unusual mouthparts, such as concave distal margin of the upper lip, large and flattened incisor teeth of mandible, very wide outer lobe of maxilla I and bluntly rounded outer and inner lobe of maxilliped, as well as combination of other features, such as armature of urosome and form of EpI–III easily separate this species from the congeners and all other 
<taxonomicName id="4C3BB497FFCFA40FFBDDFA1E8DBEFA74" authorityName="Stebbing" authorityYear="1899" box="[1055,1249,1411,1437]" class="Malacostraca" family="Gammaridae" genus="Echinogammarus" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="B94F1306FFCFA40FFBDDFA1E8DBEFA74" box="[1055,1249,1411,1437]" italics="true" pageId="15" pageNumber="498">Echinogammarus</emphasis>
</taxonomicName>
-like gammarids. Distally produced anterolateral lobe of the head also allow to distinguish the species among the congeners (
<tableCitation id="C6B9FAAFFFCFA40FFAF3FA3A8CDAFA28" box="[1329,1413,1447,1473]" captionStart="TABLE 3" captionStartId="16.[152,244,151,175]" captionText="TABLE 3. Morphological comparison of the species belonging the genus Litorogammarus gen. nov. Asterisk denotes sequences obtained in this study." pageId="15" pageNumber="498">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>