277 lines
31 KiB
XML
277 lines
31 KiB
XML
<document id="1DDFBC36C2AE9798178A5B136BEA225B" ID-CLB-Dataset="49983" ID-DOI="10.5281/zenodo.180883" ID-GBIF-Dataset="1ee405b6-83e1-4586-8ad0-5d36a840e026" ID-ISSN="1175-5326" ID-Zenodo-Dep="180883" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459818752024" checkinUser="plazi" docAuthor="Arnold, Nicholas & Bour, Roger" docDate="2008" docId="03926D377958FFEF5580257EFCF7995E" docLanguage="en" docName="zt01705p050.pdf" docOrigin="Zootaxa 1705" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Nactus soniae Arnold & Bour, 2008, n. sp." docType="treatment" docVersion="10" lastPageNumber="43" masterDocId="FFAB154F7959FFEC55172161FFBF9B1C" masterDocTitle="A new Nactus gecko (Gekkonidae) and a new Leiolopisma skink (Scincidae) from La Réunion, Indian Ocean, based on recent fossil remains and ancient DNA sequence" masterLastPageNumber="50" masterPageNumber="40" pageNumber="41" updateTime="1732567773391" updateUser="ExternalLinkService">
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<mods:title id="E5422A11A95C181D942D6972383AB6A8">A new Nactus gecko (Gekkonidae) and a new Leiolopisma skink (Scincidae) from La Réunion, Indian Ocean, based on recent fossil remains and ancient DNA sequence</mods:title>
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<mods:name id="2C41215D03C616683F1A219932033FBC" type="personal">
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<mods:namePart id="AADB94532BDD528A83AC892FF2C660EF">Arnold, Nicholas</mods:namePart>
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<mods:name id="67297791BE1ABF25385BB7EDC105BDD3" type="personal">
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<mods:roleTerm id="0A5123FF25E93C8CCC79469D3E9B889C">Author</mods:roleTerm>
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<mods:namePart id="1F99C7FDA017031CAD43398F389FBBB3">Bour, Roger</mods:namePart>
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<mods:typeOfResource id="C96DAF89A2B8F025AB712C9E8F217820">text</mods:typeOfResource>
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<mods:title id="7AC37B6F19E1B6D6336905A224CD6382">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8E5890ABF70E31C94B1BA18923EBB3A2">
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<mods:date id="D86C26881C0020FAA4797AE376904DCA">2008</mods:date>
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<mods:detail id="4A25D1ED02D82A01BA58BE0125CB77EC" type="volume">
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<mods:number id="040524D2D0DCFDB0347AB75F903D2A8A">1705</mods:number>
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<mods:start id="68F39F6EFF31331E074A59B33598C54A">40</mods:start>
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<mods:classification id="0EE075A2B6307A2FCB48C1C6019D47D7">journal article</mods:classification>
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<mods:identifier id="78D10D01657A06BF003BA372B6832391" type="CLB-Dataset">49983</mods:identifier>
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<mods:identifier id="1C60F66784A732BC13026F302A5FFBC7" type="DOI">10.5281/zenodo.180883</mods:identifier>
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<mods:identifier id="972063371353C600C0AF08143646AB88" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C88CED66F85418A6B3B05160518D5DBB" type="Zenodo-Dep">180883</mods:identifier>
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<treatment id="03926D377958FFEF5580257EFCF7995E" ID-DOI="http://doi.org/10.5281/zenodo.6228591" ID-GBIF-Taxon="119355283" ID-Zenodo-Dep="6228591" LSID="urn:lsid:plazi:treatment:03926D377958FFEF5580257EFCF7995E" httpUri="http://treatment.plazi.org/id/03926D377958FFEF5580257EFCF7995E" lastPageId="3" lastPageNumber="43" pageId="1" pageNumber="41" scope_family="Gekkonidae">
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<subSubSection id="C3218FAA7958FFED5580257EFE349F25" box="[151,395,1055,1081]" pageId="1" pageNumber="41" type="nomenclature">
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<paragraph id="8B84DC217958FFED5580257EFE349F25" blockId="1.[151,395,1055,1081]" box="[151,395,1055,1081]" pageId="1" pageNumber="41">
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<heading id="D0CC6B4D7958FFED5580257EFE349F25" bold="true" box="[151,395,1055,1081]" fontSize="11" level="1" pageId="1" pageNumber="41" reason="1">
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<emphasis id="B94F00337958FFED5580257EFE349F25" bold="true" box="[151,395,1055,1081]" pageId="1" pageNumber="41">
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<taxonomicName id="4C3BA7A27958FFED5580257EFEFE9F25" ID-CoL="45JTQ" box="[151,321,1055,1081]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="species" species="soniae" status="sp. nov.">
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<emphasis id="B94F00337958FFED5580257EFEFE9F25" bold="true" box="[151,321,1055,1081]" italics="true" pageId="1" pageNumber="41">Nactus soniae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27CBD487958FFED545F257EFE349F25" box="[328,395,1055,1081]" pageId="1" pageNumber="41" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3218FAA7958FFED5580250DFBC79F9F" box="[151,1144,1132,1155]" pageId="1" pageNumber="41" type="reference_group">
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<paragraph id="8B84DC217958FFED5580250DFBC79F9F" blockId="1.[151,1144,1132,1155]" box="[151,1144,1132,1155]" pageId="1" pageNumber="41">
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<treatmentCitationGroup id="AB2BFB0F7958FFED5580250DFBC79F9F" box="[151,1144,1132,1155]" pageId="1" pageNumber="41">
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<taxonomicName id="4C3BA7A27958FFED5580250DFEE09F9F" box="[151,351,1132,1155]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="species" species="borbonicus">
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<emphasis id="B94F00337958FFED5580250DFEE09F9F" box="[151,351,1132,1155]" italics="true" pageId="1" pageNumber="41">Nactus borbonicus</emphasis>
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</taxonomicName>
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(nomen nudum) in
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<treatmentCitation id="0A9AFA307958FFED5721250DFCBF9F9F" author="Probst" box="[566,768,1132,1155]" page="144" pageId="1" pageNumber="41" year="1997">
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<bibRefCitation id="EFAAA1D07958FFED5721250DFCBF9F9F" author="Probst" box="[566,768,1132,1155]" pageId="1" pageNumber="50" refString="Probst, J. M. (1997) Animaux de La Reunion. Azalees Editions, Sainte-Marie, La Reunion, pp. 1 - 167." type="book chapter" year="1997">Probst (1997: 144)</bibRefCitation>
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</treatmentCitation>
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, and in
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<bibRefCitation id="EFAAA1D07958FFED5643250DFBCB9F9F" author="Probst" box="[852,1140,1132,1155]" pageId="1" pageNumber="50" refString="Probst, J. M. & Brial, P. (2002) Recits anciens de naturalistes a l'ile Bourbon. Association Nature et Patrimoine, Le Port, La Reunion, pp. 1 - 112." type="book chapter" year="2002">Probst and Brial (2002:24)</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3218FAA7958FFED558025D3FB1B9E00" pageId="1" pageNumber="41" type="etymology">
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<paragraph id="8B84DC217958FFED558025D3FB1B9E00" blockId="1.[151,1436,1202,2028]" pageId="1" pageNumber="41">
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<emphasis id="B94F00337958FFED558025D3FE9C9FD0" bold="true" box="[151,291,1202,1228]" pageId="1" pageNumber="41">Etymology.</emphasis>
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Named after Sonia Ribes-Beaudemoulin, present curator of the Muséum d’Histoire Naturelle, Saint-Denis,
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<collectingCountry id="F32C9CB17958FFED542625BBFE7E9FE8" box="[305,449,1242,1268]" name="Réunion" pageId="1" pageNumber="41">La Réunion</collectingCountry>
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. She organised a dozen expeditions to collect important subfossil remains of vertebrates, especially tortoises, birds and bats from the marshes of l’Ermitage near St-Gilles.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3218FAA7958FFED55D1244BFBBE9E58" box="[198,1025,1322,1348]" pageId="1" pageNumber="41" type="distribution">
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<paragraph id="8B84DC217958FFED55D1244BFBBE9E58" blockId="1.[151,1436,1202,2028]" box="[198,1025,1322,1348]" pageId="1" pageNumber="41">
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<emphasis id="B94F00337958FFED55D1244BFED89E58" bold="true" box="[198,359,1322,1348]" pageId="1" pageNumber="41">Distribution.</emphasis>
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Known only from
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<collectingCountry id="F32C9CB17958FFED5750244BFD6C9E58" box="[583,723,1322,1348]" name="Réunion" pageId="1" pageNumber="41">La Réunion</collectingCountry>
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, southwest Indian Ocean.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3218FAA7958FFED55D12433FD089D10" pageId="1" pageNumber="41" type="materials_examined">
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<paragraph id="8B84DC217958FFED55D12433FAD09E88" blockId="1.[151,1436,1202,2028]" pageId="1" pageNumber="41">
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<emphasis id="B94F00337958FFED55D12433FE8B9E70" bold="true" box="[198,308,1362,1388]" pageId="1" pageNumber="41">Material</emphasis>
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(
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<figureCitation id="1300C0A47958FFED54512433FE0B9E70" box="[326,436,1362,1388]" captionStart-0="FIGURE 2 a, 2 b" captionStart-1="FIGURE 3" captionStartId-0="4.[151,255,1867,1891]" captionStartId-1="6.[151,255,1392,1416]" captionTargetBox-0="[264,1322,194,1842]" captionTargetBox-1="[301,1293,196,1371]" captionTargetId-0="figure@4.[264,1323,983,1844]" captionTargetId-1="figure@6.[301,1294,196,1372]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 2 a, 2 b. Nactus soniae n. sp., material from Grotte au Sable, St-Gilles, La Réunion. a. Holotype. Frontal bone (dorsal and ventral views). Scale in mm. b. Paratypes. Left maxilla and right dentary (lateral views). Scales in mm." captionText-1="FIGURE 3. Nactus soniae n. sp. Scanning electron microscope photograph of holotype frontal bone (dorsal view)." httpUri-0="https://zenodo.org/record/180885/files/figure.png" httpUri-1="https://zenodo.org/record/180887/files/figure.png" pageId="1" pageNumber="41">Figs 2–3</figureCitation>
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).
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<typeStatus id="548062837958FFED54DB2433FD819E70" box="[460,574,1362,1388]" pageId="1" pageNumber="41" type="holotype">Holotype</typeStatus>
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: Grotte au Sable, St-Gilles,
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<collectingCountry id="F32C9CB17958FFED56852433FB9C9E70" box="[914,1059,1362,1388]" name="Réunion" pageId="1" pageNumber="41">La Réunion</collectingCountry>
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; frontal bone;
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<collectionCode id="ED2A44E47958FFED51C22433FA929E70" box="[1237,1325,1362,1388]" pageId="1" pageNumber="41">BMNH</collectionCode>
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R16534.
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<typeStatus id="548062837958FFED5580241BFEB19E88" box="[151,270,1402,1428]" pageId="1" pageNumber="41" type="paratype">Paratypes</typeStatus>
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: Grotte au Sable, St-Gilles,
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<collectingCountry id="F32C9CB17958FFED5744241BFD5F9E88" box="[595,736,1402,1428]" name="Réunion" pageId="1" pageNumber="41">La Réunion</collectingCountry>
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; 1 frontal, 1 maxilla, 1 dentary;
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<collectionCode id="ED2A44E47958FFED514E241BFB0E9E88" box="[1113,1201,1402,1428]" pageId="1" pageNumber="41">BMNH</collectionCode>
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R16535-16537.
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</paragraph>
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<paragraph id="8B84DC217958FFED55D124C3FD089D10" blockId="1.[151,1436,1202,2028]" pageId="1" pageNumber="41">
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<collectionCode id="ED2A44E47958FFED55D124C3FEB99EA0" LSID="urn:lsid:biocol.org:col:15536" box="[198,262,1442,1468]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="1" pageNumber="41">DNA</collectionCode>
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sequence. Grotte au Sable, St-Gilles,
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<collectingCountry id="F32C9CB17958FFED57F324C3FCC89EA0" box="[740,887,1442,1468]" name="Réunion" pageId="1" pageNumber="41">La Réunion</collectingCountry>
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. 681 base pairs (bp) of mitochondrial
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<collectionCode id="ED2A44E47958FFED504B24C3FA239EA0" LSID="urn:lsid:biocol.org:col:15536" box="[1372,1436,1442,1468]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="1" pageNumber="41">DNA</collectionCode>
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sequence extracted from a frontal bone, comprising 305 bp of cytochrome and 374 bp of 12S rRNA genes (J. J. Austin and E. N. Arnold, unpublished data).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3218FAA7958FFED55D1277BFBE19CF0" pageId="1" pageNumber="41" type="diagnosis">
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<paragraph id="8B84DC217958FFED55D1277BFCD39DC8" blockId="1.[151,1436,1202,2028]" pageId="1" pageNumber="41">
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<emphasis id="B94F00337958FFED55D1277BFEF99D28" bold="true" box="[198,326,1562,1588]" pageId="1" pageNumber="41">Diagnosis.</emphasis>
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A small
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<taxonomicName id="4C3BA7A27958FFED54A6277AFDBD9D28" box="[433,514,1563,1588]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B94F00337958FFED54A6277AFDBD9D28" box="[433,514,1563,1588]" italics="true" pageId="1" pageNumber="41">Nactus</emphasis>
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</taxonomicName>
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, estimated from available bones to be about
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<quantity id="4CC371C47958FFED5119277BFBE59D28" box="[1038,1114,1562,1588]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="41" unit="mm" value="45.0">45mm</quantity>
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from snout to vent. Frontal not elongate, with only weakly developed anterior lateral submaxillary flanges. Maxilla with dorsal lamina inset medially, so outer surface of tooth-bearing body of bone is rounded; its posterior section comparatively short. Tooth counts low with 29 teeth in maxilla and
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<quantity id="4CC371C47958FFED561C27F3FCF79DB0" box="[779,840,1682,1708]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="1" pageNumber="41" unit="in" value="24.0">24 in</quantity>
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dentary of available material. Also has distinctive mitochondrial DNA sequence (12S rRNA and cytochrome b).
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</paragraph>
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<paragraph id="8B84DC217958FFED55D12783FBE19CF0" blockId="1.[151,1436,1202,2028]" pageId="1" pageNumber="41">
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Differs from other
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<taxonomicName id="4C3BA7A27958FFED54B12782FE479DE0" box="[422,504,1763,1788]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B94F00337958FFED54B12782FE479DE0" box="[422,504,1763,1788]" italics="true" pageId="1" pageNumber="41">Nactus</emphasis>
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</taxonomicName>
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in conformation of maxilla and from other Mascarene species in the following additional features:
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<taxonomicName id="4C3BA7A27958FFED5492266AFDF49C38" box="[389,587,1803,1828]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="species" species="serpensinsula">
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<emphasis id="B94F00337958FFED5492266AFDF49C38" box="[389,587,1803,1828]" italics="true" pageId="1" pageNumber="41">N. serpensinsula</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3BA7A27958FFED579D266AFCB79C38" box="[650,776,1803,1828]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="species" species="durrelli">
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<emphasis id="B94F00337958FFED579D266AFCB79C38" box="[650,776,1803,1828]" italics="true" pageId="1" pageNumber="41">N. durrelli</emphasis>
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</taxonomicName>
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of Mauritius—smaller size and lower tooth counts;
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<taxonomicName id="4C3BA7A27958FFED5096266AFEF39C50" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="species" species="coindemerensis">
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<emphasis id="B94F00337958FFED5096266AFEF39C50" italics="true" pageId="1" pageNumber="41">N. coindemerensis</emphasis>
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</taxonomicName>
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of Mauritius—larger size; two undescribed subfossil species from Rodrigues (E. N. Arnold, J. J. Austin and C. G. Jones, unpublished observations)—lower tooth counts, only weakly developed anterior lateral submaxillary flanges on frontal bone, also lacking the elongation of the frontal found in one of the Rodrigues species. Distinguished from sympatric
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<taxonomicName id="4C3BA7A27958FFED57B926CAFC9E9CD8" box="[686,801,1963,1988]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="41" phylum="Chordata" rank="genus">
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<emphasis id="B94F00337958FFED57B926CAFC9E9CD8" box="[686,801,1963,1988]" italics="true" pageId="1" pageNumber="41">Phelsuma</emphasis>
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</taxonomicName>
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geckos (
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<figureCitation id="1300C0A47958FFED569D26CBFC719CD8" box="[906,974,1962,1988]" captionStart="FIGURE 1" captionStartId="2.[151,255,1026,1050]" captionTargetBox="[264,1323,194,1002]" captionTargetId="figure@2.[264,1323,194,1002]" captionTargetPageId="2" captionText="FIGURE 1. Phelsuma sp., material from Grotte au Sable, St-Gilles, La Réunion. Above: frontals (dorsal and ventral views). Below: left maxilla (medial view). Scale in mm." httpUri="https://zenodo.org/record/180884/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
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) by much narrower frontal bone with a longer tubular section, a higher dorsal lamina on the maxilla and coarser dentition.
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</paragraph>
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</subSubSection>
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<caption id="DF448CA9795BFFEE55802563FD509F20" httpUri="https://zenodo.org/record/180884/files/figure.png" pageId="2" pageNumber="42" targetBox="[264,1323,194,1002]" targetPageId="2">
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<paragraph id="8B84DC21795BFFEE55802563FD509F20" blockId="2.[151,1436,1026,1084]" pageId="2" pageNumber="42">
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<emphasis id="B94F0033795BFFEE55802563FEAA9F06" bold="true" box="[151,277,1026,1050]" pageId="2" pageNumber="42">FIGURE 1</emphasis>
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.
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<taxonomicName id="4C3BA7A2795BFFEE54322562FE319F06" box="[293,398,1027,1050]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="genus">
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<emphasis id="B94F0033795BFFEE54322562FE319F06" box="[293,398,1027,1050]" italics="true" pageId="2" pageNumber="42">Phelsuma</emphasis>
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</taxonomicName>
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sp., material from Grotte au Sable, St-Gilles, La Réunion. Above: frontals (dorsal and ventral views). Below: left maxilla (medial view). Scale in mm.
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</paragraph>
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</caption>
|
||
<subSubSection id="C3218FAA795BFFEF55D12511FCF7995E" lastPageId="3" lastPageNumber="43" pageId="2" pageNumber="42" type="description">
|
||
<paragraph id="8B84DC21795BFFEE55D12511FDE49D5E" blockId="2.[151,1436,1136,2002]" pageId="2" pageNumber="42">
|
||
<emphasis id="B94F0033795BFFEE55D12511FEE19F96" bold="true" box="[198,350,1136,1162]" pageId="2" pageNumber="42">Description.</emphasis>
|
||
Available material is very fragmentary with only three skull elements represented. Frontal somewhat expanded anteriorly, and strongly expanded posteriorly, with an intervening narrow waist between the orbits. The tubular section of the frontal is relatively long and the sides of the expanded posterior section of the bone are thickened below. Dorsal surface slightly but distinctly concave, with a depressed anterior facet or facets that would lie under the prefrontals in a complete skull. The shape of the anterior border of the subdermal surface of the frontal, bordering this facet posteriorly is convex anteriorly with a pointed projection on each side. Small lateral flanges are visible on the frontal anteriorly which would lie under the posterior dorsal projection of the lamina of each maxilla. The three
|
||
<taxonomicName id="4C3BA7A2795BFFEE57E624E9FCE09EBD" box="[753,863,1416,1441]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="species" species="soniae">
|
||
<emphasis id="B94F0033795BFFEE57E624E9FCE09EBD" box="[753,863,1416,1441]" italics="true" pageId="2" pageNumber="42">N. soniae</emphasis>
|
||
</taxonomicName>
|
||
frontals examined (including one sacrificed during DNA extraction) show considerable variation in form, including how broad they are, the shape of the anterior border of the dermal surface which is convex and pointed in the
|
||
<typeStatus id="54806283795BFFEE56D624B9FB989EEE" box="[961,1063,1496,1522]" pageId="2" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
but more gently rounded in the others, and the precise size of the small anterior lateral flanges. The lateral borders of the frontal anterior to its narrow waist may also be rather wavy.
|
||
</paragraph>
|
||
<paragraph id="8B84DC21795BFFEE55D12731FB709C9E" blockId="2.[151,1436,1136,2002]" pageId="2" pageNumber="42">
|
||
Maxilla generally similar to other
|
||
<taxonomicName id="4C3BA7A2795BFFEE57412731FD189D75" box="[598,679,1616,1641]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94F0033795BFFEE57412731FD189D75" box="[598,679,1616,1641]" italics="true" pageId="2" pageNumber="42">Nactus</emphasis>
|
||
</taxonomicName>
|
||
, especially
|
||
<emphasis id="B94F0033795BFFEE56252731FC439D75" box="[818,1020,1616,1641]" italics="true" pageId="2" pageNumber="42">
|
||
<taxonomicName id="4C3BA7A2795BFFEE56252731FC479D75" box="[818,1016,1616,1641]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="species" species="serpensinsula">N. serpensinsula</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with a high dorsal lamina that rises to an acute point posteriorly, where it contacts the frontal, overlapping its anterior lateral flange. A row of four foramina in a horizontal line on the lateral surface of the tooth-bearing body of the bone and two more where the body and the dorsal lamina join. Differs from
|
||
<taxonomicName id="4C3BA7A2795BFFEE57CF27A9FC249DFD" box="[728,923,1736,1761]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="species" species="serpensinsula">
|
||
<emphasis id="B94F0033795BFFEE57CF27A9FC249DFD" box="[728,923,1736,1761]" italics="true" pageId="2" pageNumber="42">N. serpensinsula</emphasis>
|
||
</taxonomicName>
|
||
and other
|
||
<taxonomicName id="4C3BA7A2795BFFEE510227A9FBD89DFD" box="[1045,1127,1736,1761]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94F0033795BFFEE510227A9FBD89DFD" box="[1045,1127,1736,1761]" italics="true" pageId="2" pageNumber="42">Nactus</emphasis>
|
||
</taxonomicName>
|
||
in the dorsal lamina being inset medially on the tooth-bearing body of the maxilla so, in transverse section, the outer face of this is rounded before the lamina rises. The body of the maxilla posterior to the lamina is relatively short. There are 24 teeth, a similar count to that found in Mauritian
|
||
<taxonomicName id="4C3BA7A2795BFFEE57E92621FC689C45" box="[766,983,1856,1881]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="species" species="coindemerensis">
|
||
<emphasis id="B94F0033795BFFEE57E92621FC689C45" box="[766,983,1856,1881]" italics="true" pageId="2" pageNumber="42">N. coindemerensis</emphasis>
|
||
</taxonomicName>
|
||
which has about 26, but considerably lower than those in
|
||
<taxonomicName id="4C3BA7A2795BFFEE54692609FDFE9C9D" box="[382,577,1896,1921]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="species" species="serpensinsula">
|
||
<emphasis id="B94F0033795BFFEE54692609FDFE9C9D" box="[382,577,1896,1921]" italics="true" pageId="2" pageNumber="42">N. serpensinsula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3BA7A2795BFFEE576D2609FD4A9C9D" box="[634,757,1896,1921]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="species" species="durrelli">
|
||
<emphasis id="B94F0033795BFFEE576D2609FD4A9C9D" box="[634,757,1896,1921]" italics="true" pageId="2" pageNumber="42">N. durrelli</emphasis>
|
||
</taxonomicName>
|
||
which have about 36–38 teeth in adults.
|
||
</paragraph>
|
||
<paragraph id="8B84DC21795BFFEF55D126F1FCA29A36" blockId="2.[151,1436,1136,2002]" lastBlockId="3.[151,1436,152,578]" lastPageId="3" lastPageNumber="43" pageId="2" pageNumber="42">
|
||
Dentary is similar to that of most other
|
||
<taxonomicName id="4C3BA7A2795BFFEE578426F1FD5B9CB5" box="[659,740,1936,1961]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94F0033795BFFEE578426F1FD5B9CB5" box="[659,740,1936,1961]" italics="true" pageId="2" pageNumber="42">Nactus</emphasis>
|
||
</taxonomicName>
|
||
species, being rather slender, with a short ventral groove at the anterior extremity; six foramina in a roughly horizontal line on lateral surface which has a tridentate posterior border, medial surface with a depressed splenial scar and a V-shaped posterior border in which the upper edge has a pronounced step. The bone, which is from a relatively small individual, has 29 teeth, a count again similar to that found in Mauritian
|
||
<emphasis id="B94F0033795AFFEF57262189FCAE9A1D" box="[561,785,232,257]" italics="true" pageId="3" pageNumber="43">
|
||
<taxonomicName id="4C3BA7A2795AFFEF57262189FCB29A1D" box="[561,781,232,257]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="coindemerensis">N. coindemerensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but considerably lower than those in
|
||
<taxonomicName id="4C3BA7A2795AFFEF51C02189FA239A1D" box="[1239,1436,232,257]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="serpensinsula">
|
||
<emphasis id="B94F0033795AFFEF51C02189FA239A1D" box="[1239,1436,232,257]" italics="true" pageId="3" pageNumber="43">N. serpensinsula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3BA7A2795AFFEF55DF2071FEFC9A35" box="[200,323,272,297]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="durrelli">
|
||
<emphasis id="B94F0033795AFFEF55DF2071FEFC9A35" box="[200,323,272,297]" italics="true" pageId="3" pageNumber="43">N. durrelli</emphasis>
|
||
</taxonomicName>
|
||
which have about 39–40 teeth in adults.
|
||
</paragraph>
|
||
<paragraph id="8B84DC21795AFFEF55D12059FACA9AD6" blockId="3.[151,1436,152,578]" pageId="3" pageNumber="43">
|
||
The size of the frontals, the elements apparently from the largest individuals represented in the available sample, indicate that
|
||
<taxonomicName id="4C3BA7A2795AFFEF54842001FDBC9A65" box="[403,515,352,377]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="soniae">
|
||
<emphasis id="B94F0033795AFFEF54842001FDBC9A65" box="[403,515,352,377]" italics="true" pageId="3" pageNumber="43">N. soniae</emphasis>
|
||
</taxonomicName>
|
||
was around
|
||
<quantity id="4CC371C4795AFFEF578B2001FD589A66" box="[668,743,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="43" unit="mm" value="45.0">45mm</quantity>
|
||
from snout to vent. It is consequently intermediate in size between
|
||
<taxonomicName id="4C3BA7A2795AFFEF541620E9FE669ABD" box="[257,473,392,417]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="coindemerensis">
|
||
<emphasis id="B94F0033795AFFEF541620E9FE669ABD" box="[257,473,392,417]" italics="true" pageId="3" pageNumber="43">N. coindemerensis</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<collectingCountry id="F32C9CB1795AFFEF571420E9FDCA9ABE" box="[515,629,392,418]" name="Mauritius" pageId="3" pageNumber="43">Mauritius</collectingCountry>
|
||
, which grows to about
|
||
<quantity id="4CC371C4795AFFEF569A20E9FC679ABE" box="[909,984,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="43" unit="mm" value="36.0">36mm</quantity>
|
||
from snout to vent, and
|
||
<taxonomicName id="4C3BA7A2795AFFEF51E920E9FF779AD5" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="serpensinsula">
|
||
<emphasis id="B94F0033795AFFEF51E920E9FF779AD5" italics="true" pageId="3" pageNumber="43">N. serpensinsula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3BA7A2795AFFEF541620D1FEC49AD5" box="[257,379,432,457]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="durrelli">
|
||
<emphasis id="B94F0033795AFFEF541620D1FEC49AD5" box="[257,379,432,457]" italics="true" pageId="3" pageNumber="43">N. durrelli</emphasis>
|
||
</taxonomicName>
|
||
from the same island which reach at least
|
||
<quantity id="4CC371C4795AFFEF567920D1FC069AD6" box="[878,953,432,458]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="43" unit="mm" value="67.0">67mm</quantity>
|
||
and
|
||
<quantity id="4CC371C4795AFFEF56E520D1FB839AD6" box="[1010,1084,432,458]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="43" unit="mm" value="55.0">55mm</quantity>
|
||
(
|
||
<bibRefCitation id="EFAAA1D0795AFFEF515B20D1FAD79AD6" author="Arnold" box="[1100,1384,432,458]" pageId="3" pageNumber="50" refString="Arnold, E. N. & Jones, C. J. (1994) The night geckos of the genus Nactus in the Mascarene islands with a description of the distinctive population on Round island. Dodo, 30, 119 - 131." type="journal article" year="1994">Arnold and Jones, 1994</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84DC21795AFFEF55D120B6FCF7995E" blockId="3.[151,1436,152,578]" pageId="3" pageNumber="43">
|
||
<emphasis id="B94F0033795AFFEF55D120B6FEC89AED" bold="true" box="[198,375,471,497]" pageId="3" pageNumber="43">Relationships.</emphasis>
|
||
DNA confirms that
|
||
<taxonomicName id="4C3BA7A2795AFFEF576420B9FD5A9AED" box="[627,741,472,497]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="soniae">
|
||
<emphasis id="B94F0033795AFFEF576420B9FD5A9AED" box="[627,741,472,497]" italics="true" pageId="3" pageNumber="43">N. soniae</emphasis>
|
||
</taxonomicName>
|
||
is a
|
||
<taxonomicName id="4C3BA7A2795AFFEF563420B9FCCB9AED" box="[803,884,472,497]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94F0033795AFFEF563420B9FCCB9AED" box="[803,884,472,497]" italics="true" pageId="3" pageNumber="43">Nactus</emphasis>
|
||
</taxonomicName>
|
||
and indicates it is most closely related to the diminutive
|
||
<taxonomicName id="4C3BA7A2795AFFEF54092361FE499905" box="[286,502,512,537]" class="Reptilia" family="Gekkonidae" genus="Nactus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="43" phylum="Chordata" rank="species" species="coindemerensis">
|
||
<emphasis id="B94F0033795AFFEF54092361FE499905" box="[286,502,512,537]" italics="true" pageId="3" pageNumber="43">N. coindemerensis</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<collectingCountry id="F32C9CB1795AFFEF57082361FD2E9906" box="[543,657,512,538]" name="Mauritius" pageId="3" pageNumber="43">Mauritius</collectingCountry>
|
||
(J. J. Austin and E. N. Arnold, unpublished data), something also supported by its relatively small size and low tooth counts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |