162 lines
16 KiB
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162 lines
16 KiB
XML
<document id="3738DC6F41FE236B85075976316D7A3B" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude & Tabuce, Rodolphe" docDate="2021" docId="03923C45FF87FF8E33F1FDA9FB5D1343" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Otodus (Carcharocles)" docType="treatment" docVersion="6" lastPageNumber="125" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="125" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="0A1CB661C11E312FCD88702646083DDA">
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<mods:title id="6D360CC8675C5A06F571EA5F4AB68EE3">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="0392E446F4F0C2B5EB6EB443BB1394CD">Zouhri, Samir</mods:namePart>
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</mods:name>
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<mods:roleTerm id="2FD3D2975A7CDBD66CD907A8B90B9BCB">Author</mods:roleTerm>
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<mods:namePart id="10AE4855CCEFDA545F50D8F83FE023C9">Gingerich, Philip D.</mods:namePart>
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</mods:name>
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<mods:name id="BE2572216E837D7F610D7FE45843ACEA" type="personal">
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<mods:roleTerm id="CD8AC189F0F404666230E6D21D01F8BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="370702E160888C442707496FD3E08708">Khalloufi, Bouziane</mods:namePart>
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</mods:name>
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<mods:name id="BF47145BEE4C137490438958A34C7732" type="personal">
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<mods:roleTerm id="DC7ACB92E3664B4634F9ECA7BC65C675">Author</mods:roleTerm>
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<mods:namePart id="78A7C66D43AA9E4C20D325D16D273332">Bourdon, Estelle</mods:namePart>
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<mods:roleTerm id="95853DDB4B857C38F33596D45E011FD1">Author</mods:roleTerm>
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<mods:namePart id="45EDF61521D3CD7124AEC3A525949E52">Adnet, Sylvain</mods:namePart>
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<mods:namePart id="B958969327F5BBC4F2B26CEF2846F8D1">Jouve, Stéphane</mods:namePart>
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</mods:role>
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<mods:namePart id="804EE83F63472F329E875854E41D475F">Elboudali, Najia</mods:namePart>
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</mods:name>
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<mods:name id="E4B0F0597FCCE258A12FBEE1895F037A" type="personal">
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<mods:namePart id="F0EC2A129524215EF7CC5A3E03315DED">Amane, Ayoub</mods:namePart>
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</mods:name>
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<mods:roleTerm id="D60EF699C23A060B9CC42000EA13B199">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CCCB05A5FBE2206B6F81CEF9A74B3B98">Rage, Jean-Claude</mods:namePart>
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</mods:name>
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<mods:name id="FD815ADF035DF21921C7EE18A09EC9F4" type="personal">
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<mods:roleTerm id="4820400D282B89D4A49D80DFF37634B1">Author</mods:roleTerm>
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<mods:namePart id="30CA243890B180D2FA6F3DD86B5E47A2">Tabuce, Rodolphe</mods:namePart>
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<mods:typeOfResource id="14596463DC66DB7CDD40D65A878D83E2">text</mods:typeOfResource>
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<mods:titleInfo id="C985440C46008A44F6CAACC9CEB22F2E">
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<mods:title id="31DB6CB03A6DE03270953558AA05EF5C">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="CD47A7DF818DB21616B28710A669A724">
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<mods:date id="A597115798E18D4A75FBEEA87601EE3F">2021</mods:date>
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<mods:detail id="D6249B41610C811A6072E6898A4D36A3" type="pubDate">
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<mods:number id="A6CB6A9B75817925E991E03F88FC251F">2021-03-11</mods:number>
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<mods:detail id="504DDA22A9FF80037AAADB3D67063B36" type="volume">
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<mods:number id="743965E83013AB476007AD5373BE526E">43</mods:number>
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</mods:detail>
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<mods:detail id="C99D082D4FDBC218DAB14B775AD67D84" type="issue">
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<mods:number id="9F0D5E32F0B9C80D9609D3C14996403B">5</mods:number>
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</mods:detail>
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<mods:start id="891164B62339533F02E4C89C11A5117D">121</mods:start>
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<mods:end id="3BD4D9E2D2C5908CE5C26057A3146FDC">150</mods:end>
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</mods:extent>
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<mods:classification id="068475599F2200EC961696A05E3ABDE8">journal article</mods:classification>
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<mods:identifier id="0D8E4FD5DAFA89AA2B9AE0C7B30B1880" type="CLB-Dataset">7746</mods:identifier>
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<mods:identifier id="78B5C7AE59BF2D8144DD3F89B7476543" type="DOI">10.5252/geodiversitas2021v43a5</mods:identifier>
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<mods:identifier id="6A7ED7CC418F65E9C12EB73FA14B90FC" type="GBIF-Dataset">e42658a2-0e15-4e90-9bc5-0d00cbcbd781</mods:identifier>
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<mods:identifier id="F649ADA3BEE0D6314CAE614EC4E0633C" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="56D5FD4879B2842C12EA4406D583BE06" type="Zenodo-Dep">4605963</mods:identifier>
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<mods:identifier id="5639B0CB0D78AA924FADE04266DE4CCC" type="ZooBank">urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C</mods:identifier>
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</mods:mods>
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<treatment id="03923C45FF87FF8E33F1FDA9FB5D1343" ID-DOI="http://doi.org/10.5281/zenodo.4606611" ID-GBIF-Taxon="180569479" ID-Zenodo-Dep="4606611" LSID="urn:lsid:plazi:treatment:03923C45FF87FF8E33F1FDA9FB5D1343" httpUri="http://treatment.plazi.org/id/03923C45FF87FF8E33F1FDA9FB5D1343" lastPageNumber="125" pageId="6" pageNumber="125">
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<subSubSection id="C321DED8FF87FF8E33F1FDA9FB6714FE" box="[1001,1266,630,656]" pageId="6" pageNumber="125" type="nomenclature">
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<paragraph id="8B848D53FF87FF8E33F1FDA9FB6714FE" blockId="6.[1001,1266,630,656]" box="[1001,1266,630,656]" pageId="6" pageNumber="125">
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<heading id="D0CC3A3FFF87FF8E33F1FDA9FB6714FE" box="[1001,1266,630,656]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="125" reason="2">
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<taxonomicName id="4C3BF6D0FF87FF8E33F1FDA9FB5814FE" authority="" box="[1001,1229,630,656]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="subGenus" subGenus="Carcharocles">
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<emphasis id="B94F5141FF87FF8E33F1FDA9FBA614FE" box="[1001,1075,630,656]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
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(
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<emphasis id="B94F5141FF87FF8E345BFDA9FB5114FE" box="[1091,1220,630,656]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
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)
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</taxonomicName>
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sp.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF87FF8E3335FD70FBBE148F" pageId="6" pageNumber="125" type="materials_examined">
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<paragraph id="8B848D53FF87FF8E3335FD70FBBE148F" blockId="6.[813,1455,687,737]" pageId="6" pageNumber="125">
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EXAMINED MATERIAL. —
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<materialsCitation id="3B53870EFF87FF8E3431FD70FBBE148F" ID-GBIF-Occurrence="3053068319" pageId="6" pageNumber="125">A dozen isolated teeth, figured material includes FSAC Bouj-323.</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF87FF8E3335FCC9FAD01580" pageId="6" pageNumber="125" type="description">
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<paragraph id="8B848D53FF87FF8E3335FCC9FC281540" blockId="6.[811,1456,790,1007]" box="[813,957,790,816]" pageId="6" pageNumber="125">DESCRIPTION</paragraph>
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<paragraph id="8B848D53FF87FF8E3335FCEAFAD01580" blockId="6.[811,1456,790,1007]" pageId="6" pageNumber="125">
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Teeth are some smaller than those of previous species and up to
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<quantity id="4CC320B6FF87FF8E3351FC8AFC15151E" box="[841,896,853,880]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="125" unit="cm" value="5.0">5 cm</quantity>
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in height (
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<figureCitation id="130091D6FF87FF8E33E2FC8AFBE4151E" box="[1018,1137,853,880]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2D, E</figureCitation>
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). Compared to
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<taxonomicName id="4C3BF6D0FF87FF8E353CFC8AFB8515E1" baseAuthorityName="Jaekel" baseAuthorityYear="1895" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="sokolowi" subGenus="Carcharocles">
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<emphasis id="B94F5141FF87FF8E353CFC8AFAFB1501" box="[1316,1390,853,879]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
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(
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<emphasis id="B94F5141FF87FF8E3566FC8AFC1315E1" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
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) cf.
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<emphasis id="B94F5141FF87FF8E33A3FCAAFB8515E1" box="[955,1040,885,911]" italics="true" pageId="6" pageNumber="125">sokolowi</emphasis>
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</taxonomicName>
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, teeth of this second species display a flatter root with subrectangular basal extremities of lobes, a more slender cusp with finer serrations and two cusplets which are smaller and lower than in other coeval species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF87FF8E3335FBCAFB5D1343" pageId="6" pageNumber="125" type="discussion">
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<paragraph id="8B848D53FF87FF8E3335FBCAFC041243" blockId="6.[813,1457,1045,1325]" box="[813,913,1045,1071]" pageId="6" pageNumber="125">REMARKS</paragraph>
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<paragraph id="8B848D53FF87FF8E3335FBEBFB5D1343" blockId="6.[813,1457,1045,1325]" pageId="6" pageNumber="125">
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These characters are different from the coeval small megatoothed shark species
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<taxonomicName id="4C3BF6D0FF87FF8E3408FB8BFAFB1200" baseAuthorityName="Blainville" baseAuthorityYear="1818" box="[1040,1390,1108,1135]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="auriculatus" subGenus="Carcharocles">
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<emphasis id="B94F5141FF87FF8E3408FB8BFBCF1200" box="[1040,1114,1108,1134]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
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(
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<emphasis id="B94F5141FF87FF8E3472FB8BFB7F1200" box="[1130,1258,1108,1134]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
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)
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<emphasis id="B94F5141FF87FF8E34E0FB8BFAFB1200" box="[1272,1390,1108,1134]" italics="true" pageId="6" pageNumber="125">auriculatus</emphasis>
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</taxonomicName>
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<emphasis id="B94F5141FF87FF8E3576FB8BFAE71200" box="[1390,1394,1108,1134]" italics="true" pageId="6" pageNumber="125">,</emphasis>
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commonly reported in worldwide middle Eocene deposits (e.g.,
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<bibRefCitation id="EFAAF0A2FF87FF8E3335FB4BFB9012C0" author="WARD J. W. & WIEST R. L." box="[813,1029,1172,1198]" pageId="6" pageNumber="125" pagination="81 - 88" refId="ref26141" refString="WARD J. W. & WIEST R. L. 1990. - A checklist of Palaeocene and Eocene sharks and rays (Chondrichthyes) from the Pamunkey Group, Maryland and Virginia, USA. Tertiary Research 12: 81 - 88." type="journal article" year="1990">Ward & Wiest 1990</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF87FF8E3409FB4BFB3712C0" author="DUTHEIL D. B." box="[1041,1186,1172,1198]" pageId="6" pageNumber="125" pagination="27 - 36" refId="ref20730" refString="DUTHEIL D. B. 1991. - A checklist of Neoselachii (Pisces, Chondrichtyes) from Paleogene of Paris Basin, France. Tertiary Research 13: 27 - 36." type="journal article" year="1991">Dutheil 1991</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF87FF8E34B6FB4BFAB112C0" author="LONG D. J." box="[1198,1316,1172,1198]" pageId="6" pageNumber="125" pagination="11 - 32" refId="ref22908" refString="LONG D. J. 1992. - Sharks from the La Meseta Formation (Eocene), Seymour Island, Antarctic Peninsula. Journal of Vertebrate Paleontology 12 (1): 11 - 32. https: // doi. org / 10.1080 / 02 724634.1992. 10011428" type="journal article" year="1992">Long 1992</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF87FF8E3528FB4BFC5712A0" author="CAPPETTA H. & STRINGER G." pageId="6" pageNumber="125" pagination="51 - 56" refId="ref20300" refString="CAPPETTA H. & STRINGER G. 2002. - A new batoid genus (Neoselachii: Myliobatiformes) from the Yazoo (late Eocene) of Louisiana, USA. Tertiary Research 21: 51 - 56." type="journal article" year="2002">Cappetta & Stringer 2002</bibRefCitation>
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,
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<bibRefCitation id="EFAAF0A2FF87FF8E33C9FB6BFB5B12A0" author="CAPPETTA H. & CASE G. R." box="[977,1230,1204,1230]" pageId="6" pageNumber="125" pagination="43 - 103" refId="ref20339" refString="CAPPETTA H. & CASE G. R. 2016. - A Selachian Fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA. Palaeontographica Abteilung A 307 (1 - 6): 43 - 103." type="journal article" year="2016">Cappetta & Case 2016</bibRefCitation>
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). Similar otodontid morphology is observed in the Lutetian material of
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<collectingCountry id="F32CCDC3FF87FF8E3560FB0BFA251280" box="[1400,1456,1236,1262]" name="Togo" pageId="6" pageNumber="125">Togo</collectingCountry>
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(Pers. Observ.) where co-occurrence of two representatives of megatoothed sharks is also noticed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |