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<document id="19E7DD96FAAB1717A094AC790B63E811" ID-DOI="10.11646/zootaxa.3860.3.1" ID-GBIF-Dataset="073bdbbd-acbe-49af-8bd6-8a92a7ca97c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="228955" ID-ZooBank="46E66BEA-2679-42AF-B5EC-322D773B52F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460904162693" checkinUser="plazi" docAuthor="Poore, Gary C. B. &amp; Andreakis, Nikos" docDate="2014" docId="039194508B78FF941F890671FDF9FE02" docLanguage="en" docName="zt03860p225.pdf" docOrigin="Zootaxa 3860 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agononida auscerta Poore &amp; Andreakis 2012" docType="treatment" docVersion="17" lastPageNumber="209" masterDocId="FFA8EC288B7EFF9C1F1E0065FFC5FFF7" masterDocTitle="More species of the Agononida incerta complex revealed by molecules and morphology (Crustacea: Decapoda: Anomura: Munididae)" masterLastPageNumber="225" masterPageNumber="201" pageNumber="207" updateTime="1698360624884" updateUser="plazi">
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<mods:title id="5C1F231C6EA83411EC0BF1C72121FFCE">More species of the Agononida incerta complex revealed by molecules and morphology (Crustacea: Decapoda: Anomura: Munididae)</mods:title>
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<mods:namePart id="D530118CC8E5CA6BFA2A3577F93E96DF">Poore, Gary C. B.</mods:namePart>
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<mods:namePart id="8AE9BA8C7A979A58352532033FFA2DB6">Andreakis, Nikos</mods:namePart>
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<mods:title id="1263C85C5D35AF331D7FD68004BC0A4C">Zootaxa</mods:title>
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<mods:date id="FA22C8CE2849F36FD410C9E686949CEA">2014</mods:date>
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<treatment id="039194508B78FF941F890671FDF9FE02" ID-DOI="http://doi.org/10.5281/zenodo.3511591" ID-GBIF-Taxon="119587812" ID-Zenodo-Dep="3511591" LSID="urn:lsid:plazi:treatment:039194508B78FF941F890671FDF9FE02" httpUri="http://treatment.plazi.org/id/039194508B78FF941F890671FDF9FE02" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
<subSubSection id="C32276CD8B78FF9A1F890671FD1DF963" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="8B8725468B78FF9A1F890671FD09F9D9" blockId="6.[151,716,1556,1615]" box="[151,716,1556,1582]" pageId="6" pageNumber="207">
<heading id="D0CF922A8B78FF9A1F890671FD09F9D9" bold="true" box="[151,716,1556,1582]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<taxonomicName id="4C385EC58B78FF9A1F890671FD09F9D9" authority="Poore &amp; Andreakis, 2012" authorityName="Poore &amp; Andreakis" authorityYear="2012" box="[151,716,1556,1582]" class="Malacostraca" family="Galatheidae" genus="Agononida" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="auscerta">
<emphasis id="B94CF9548B78FF9A1F890671FD09F9D9" bold="true" box="[151,716,1556,1582]" pageId="6" pageNumber="207">
<emphasis id="B94CF9548B78FF9A1F890671FE4FF9D9" bold="true" box="[151,394,1556,1582]" italics="true" pageId="6" pageNumber="207">Agononida auscerta</emphasis>
<bibRefCitation id="EFA958B78B78FF9A1E8F0671FD09F9D9" author="Poore" box="[401,716,1556,1582]" pageId="6" pageNumber="207" refString="Poore, G. C. B. &amp; Andreakis, N. (2012) The Agononida incerta species complex unravelled (Crustacea: Decapoda: Anomura: Munididae). Zootaxa, 3429, 1 - 29." type="journal article" year="2012">Poore &amp; Andreakis, 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8725468B78FF9A1F890653FEE0F9B8" blockId="6.[151,716,1556,1615]" box="[151,293,1590,1615]" pageId="6" pageNumber="207">
(
<figureCitation id="130339C38B78FF9A1F810653FF23F9B8" box="[159,230,1590,1615]" captionStart="FIGURE 2" captionStartId="5.[151,250,1821,1843]" captionTargetBox="[156,1420,1268,1800]" captionTargetId="figure@5.[438,867,1351,1801]" captionTargetPageId="5" captionText="FIGURE 2. Colour photographs of living animals by T. - Y. Chan unless otherwise noted. a, Agononida africerta, paratype, Mozambique, MNHN IU- 2008 - 10190; b, A. auscerta, PNG, MNHN IU- 2011 - 2426; c, A. incerta, Taiwan (from Baba et al. 2009 and Poore &amp; Andreakis 2012); d, A. indocerta, Western Australia, D. Evans (from Jones &amp; Morgan 2002 and Poore &amp; Andreakis 2012); e, A. madagascerta n. sp., paratype, Madagascar, MNHN IU- 2009 - 1891; f, A. norfocerta, PNG, MNHN IU- 2011 - 891; g, A. polycerta n. sp., French Polynesia, TARASOC stn DW 3377 (specimen not located), L. Corbari; h, A. rubrizonata, paratype, Taiwan (from Poore &amp; Andreakis 2012); i, A. tasmancerta, New South Wales (specimen not located), Image # 741 reproduced from 35 mm slide courtesy of The Neville Coleman Collection." httpUri="https://zenodo.org/record/228957/files/figure.png" pageId="6" pageNumber="207">Figs 2</figureCitation>
b, 4a)
</paragraph>
<paragraph id="8B8725468B78FF9A1F89061BFD1DF963" blockId="6.[151,851,1662,1715]" box="[151,728,1662,1685]" pageId="6" pageNumber="207">
<treatmentCitationGroup id="AB2802688B78FF9A1F89061BFD1DF963" box="[151,728,1662,1685]" pageId="6" pageNumber="207">
<taxonomicName id="4C385EC58B78FF9A1F89061BFEC3F963" box="[151,262,1662,1684]" class="Malacostraca" family="Galatheidae" genus="Agononida" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="B94CF9548B78FF9A1F89061BFEC3F963" box="[151,262,1662,1684]" italics="true" pageId="6" pageNumber="207">Agononida</emphasis>
</taxonomicName>
sp.
<taxonomicName id="4C385EC58B78FF9A1E31061BFE65F962" box="[303,416,1662,1685]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="incerta">
aff.
<emphasis id="B94CF9548B78FF9A1E4606E5FE65F962" box="[344,416,1664,1685]" italics="true" pageId="6" pageNumber="207">incerta</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A9903578B78FF9A1EA3061AFD57F963" author="Poore" box="[445,658,1662,1685]" page="18" pageId="6" pageNumber="207" year="2008">
<bibRefCitation id="EFA958B78B78FF9A1EA3061AFDAAF963" author="Poore" box="[445,623,1662,1685]" pageId="6" pageNumber="207" refString="Poore, G. C. B., McCallum, A. W. &amp; Taylor, J. (2008) Decapod Crustacea of the continental margin of southwestern and central Western Australia: preliminary identifications of 524 species from FRV Southern Surveyor voyage SS 10 - 2005. Museum Victoria Science Reports, 11, 1 - 106." type="journal article" year="2008">
Poore
<emphasis id="B94CF9548B78FF9A1D1E06E5FDF1F963" box="[512,564,1662,1685]" italics="true" pageId="6" pageNumber="207">et al.</emphasis>
2008
</bibRefCitation>
: 18
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(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32276CD8B78FF9A1F8906F8FC96F944" box="[151,851,1693,1715]" pageId="6" pageNumber="207" type="reference_group">
<paragraph id="8B8725468B78FF9A1F8906F8FC96F944" blockId="6.[151,851,1662,1715]" box="[151,851,1693,1715]" pageId="6" pageNumber="207">
<treatmentCitationGroup id="AB2802688B78FF9A1F8906F8FC96F944" box="[151,851,1693,1715]" pageId="6" pageNumber="207">
<taxonomicName id="4C385EC58B78FF9A1F8906F8FD56F944" authority="Poore &amp; Andreakis, 2012: 16" authorityName="Poore &amp; Andreakis" authorityPageNumber="16" authorityYear="2012" box="[151,659,1693,1715]" class="Malacostraca" family="Galatheidae" genus="Agononida" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="auscerta">
<emphasis id="B94CF9548B78FF9A1F8906F8FEA6F944" box="[151,355,1693,1715]" italics="true" pageId="6" pageNumber="207">Agononida auscerta</emphasis>
<treatmentCitation id="0A9903578B78FF9A1E7706FBFD56F944" ID-DOI="http://doi.org/10.5281/zenodo.3501407" ID-Zenodo-Dep="3501407" author="Poore" box="[361,659,1693,1715]" httpUri="http://treatment.plazi.org/id/743287A5B57F120BFF6D0663503A45EB" page="16" pageId="6" pageNumber="207" year="2012">
<bibRefCitation id="EFA958B78B78FF9A1E7706FBFDB4F944" author="Poore" box="[361,625,1693,1715]" pageId="6" pageNumber="207" refString="Poore, G. C. B. &amp; Andreakis, N. (2012) The Agononida incerta species complex unravelled (Crustacea: Decapoda: Anomura: Munididae). Zootaxa, 3429, 1 - 29." type="journal article" year="2012">Poore &amp; Andreakis, 2012</bibRefCitation>
: 16
</treatmentCitation>
</taxonomicName>
18, figs 2F, 2G, 9.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32276CD8B78FF9A1F8906BAFC05F8EA" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="8B8725468B78FF9A1F8906BAFC05F8EA" blockId="6.[151,1436,1759,2001]" pageId="6" pageNumber="207">
<emphasis id="B94CF9548B78FF9A1F8906BAFE4EF90F" bold="true" box="[151,395,1759,1784]" pageId="6" pageNumber="207">Material examined</emphasis>
.
<collectingCountry id="F32F65D68B78FF9A1EB50685FD26F90F" box="[427,739,1760,1784]" name="Papua New Guinea" pageId="6" pageNumber="207">
<collectionCode id="ED29BD838B78FF9A1EB50685FDC1F90F" box="[427,516,1760,1784]" pageId="6" pageNumber="207">PAPUA</collectionCode>
<collectionCode id="ED29BD838B78FF9A1D030685FD98F900" box="[541,605,1760,1783]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="207">NEW</collectionCode>
<collectionCode id="ED29BD838B78FF9A1D6B0685FD26F90F" box="[629,739,1760,1784]" pageId="6" pageNumber="207">GUINEA</collectionCode>
</collectingCountry>
, W of New Hanover,
<geoCoordinate id="EE0C43818B78FF9A1B2F06BAFB4EF90F" box="[1073,1163,1759,1784]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="925" value="-2.25">02°15'S</geoCoordinate>
,
<geoCoordinate id="EE0C43818B78FF9A1B8806BAFAC5F90F" box="[1174,1280,1759,1784]" direction="east" orientation="longitude" pageId="6" pageNumber="207" precision="925" value="150.26666">150°16'E</geoCoordinate>
,
<quantity id="4CC088A38B78FF9A1A030685FA59F900" box="[1309,1436,1760,1784]" metricMagnitude="2" metricUnit="m" metricValue="4.21" metricValueMax="4.4" metricValueMin="4.02" pageId="6" pageNumber="207" unit="m" value="421.0" valueMax="440.0" valueMin="402.0">402440 m</quantity>
(BIOPAPUA stn CP3655),
<collectionCode id="ED29BD838B78FF9A1ED70760FDDAF8EB" LSID="urn:lsid:biocol.org:col:34988" box="[457,543,1797,1820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="207">MNHN</collectionCode>
<collectionCode id="ED29BD838B78FF9A1D380761FD88F8EA" box="[550,589,1796,1821]" pageId="6" pageNumber="207">IU-</collectionCode>
<date id="FF8603868B78FF9A1D530761FD00F8EA" box="[589,709,1796,1821]" pageId="6" pageNumber="207" value="2011" valueMax="2426">2011-2426</date>
(det. E. Macpherson).
</paragraph>
</subSubSection>
<subSubSection id="C32276CD8B78FF9A1FD90742FAEDF85B" pageId="6" pageNumber="207" type="diagnosis">
<paragraph id="8B8725468B78FF9A1FD90742FAEDF85B" blockId="6.[151,1436,1759,2001]" pageId="6" pageNumber="207">
<emphasis id="B94CF9548B78FF9A1FD90742FEFFF8B7" bold="true" box="[199,314,1831,1856]" pageId="6" pageNumber="207">Diagnosis</emphasis>
. Telson anterolateral margin almost straight over anterior three-quarters, posterior quarter separated by shallow hiatus. Pereopods 24 dactyli with 89 times as long as greatest basal width; robust setae on flexor margin of dactylus of pereopod 2 number 26 (median 4; rarely 8), of pereopod 3, 14 (median 2) and of pereopod 4, 12 (rarely 3). Pereopod 4 merus without spines proximally on upper face. (
<bibRefCitation id="EFA958B78B78FF9A1B1B07F1FAE5F85B" author="Poore" box="[1029,1312,1940,1965]" pageId="6" pageNumber="207" refString="Poore, G. C. B. &amp; Andreakis, N. (2012) The Agononida incerta species complex unravelled (Crustacea: Decapoda: Anomura: Munididae). Zootaxa, 3429, 1 - 29." type="journal article" year="2012">Poore &amp; Andreakis 2012</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32276CD8B78FF9A1FD907DDFB47F826" box="[199,1154,1976,2001]" pageId="6" pageNumber="207" type="distribution">
<paragraph id="8B8725468B78FF9A1FD907DDFB47F826" blockId="6.[151,1436,1759,2001]" box="[199,1154,1976,2001]" pageId="6" pageNumber="207">
<emphasis id="B94CF9548B78FF9A1FD907DDFE9CF826" bold="true" box="[199,345,1976,2001]" pageId="6" pageNumber="207">Distribution</emphasis>
. Western
<collectingCountry id="F32F65D68B78FF9A1ED707DDFDF6F826" box="[457,563,1976,2001]" name="Australia" pageId="6" pageNumber="207">Australia</collectingCountry>
,
<collectingCountry id="F32F65D68B78FF9A1D2007DDFDB3F827" box="[574,630,1976,2000]" name="Papua New Guinea" pageId="6" pageNumber="207">PNG</collectingCountry>
, 2°S22°S;
<quantity id="4CC088A38B78FF9A1DE307DDFCB9F827" box="[765,892,1976,2001]" metricMagnitude="2" metricUnit="m" metricValue="4.095" metricValueMax="4.46" metricValueMin="3.7299999999999995" pageId="6" pageNumber="207" unit="m" value="409.5" valueMax="446.0" valueMin="373.0">373446 m</quantity>
(median
<quantity id="4CC088A38B78FF9A1CFB07DDFBE8F827" box="[997,1069,1976,2000]" metricMagnitude="2" metricUnit="m" metricValue="4.04" pageId="6" pageNumber="207" unit="m" value="404.0">404 m</quantity>
) depth.
</paragraph>
</subSubSection>
<caption id="DF4775CE8B79FF9B1F890735FC2EF835" httpUri="https://zenodo.org/record/228958/files/figure.png" pageId="7" pageNumber="208" targetBox="[158,1434,193,1843]" targetPageId="7">
<paragraph id="8B8725468B79FF9B1F890735FC2EF835" blockId="7.[151,1436,1872,1986]" pageId="7" pageNumber="208">
<emphasis id="B94CF9548B79FF9B1F890735FEC9F892" bold="true" box="[151,268,1872,1894]" pageId="7" pageNumber="208">FIGURE 3</emphasis>
.
<taxonomicName id="4C385EC58B79FF9B1E060735FDFCF891" authority="Telsons" authorityName="Telsons" box="[280,569,1872,1894]" class="Malacostraca" family="Galatheidae" genus="Agononida" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="africerta">
<emphasis id="B94CF9548B79FF9B1E060735FE23F891" box="[280,486,1872,1894]" italics="true" pageId="7" pageNumber="208">Agononida africerta</emphasis>
Telsons
</taxonomicName>
and detail of anterolateral margins: a (top row), male, 21 mm, MNHN IU-2010-1158; b, male, 17 mm, MNHN IU-2008-10440; c, ovigerous female, 26 mm, MNHN IU-2008-10441; d, supermale, 31 mm, MNHN- IU-2008-10192. Pereopod 1, upper and lateral views: e, ovigerous female, 26 mm, MNHN IU-2008-10441; f, supermale, 31 mm, MNHN-IU-2008-10192. Line drawings to same scale; chelipeds to same scale.
</paragraph>
</caption>
<subSubSection id="C32276CD8B76FF941FD900F2FDF9FE02" pageId="8" pageNumber="209" type="discussion">
<paragraph id="8B8725468B76FF941FD900F2FDF9FE02" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="209">
<emphasis id="B94CF9548B76FF941FD900F2FEF1FF47" bold="true" box="[199,308,151,176]" pageId="8" pageNumber="209">Remarks</emphasis>
.
<taxonomicName id="4C385EC58B76FF941E5000F2FDFBFF47" box="[334,574,151,176]" class="Malacostraca" family="Galatheidae" genus="Agononida" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="auscerta">
<emphasis id="B94CF9548B76FF941E5000F2FDFBFF47" box="[334,574,151,176]" italics="true" pageId="8" pageNumber="209">Agononida auscerta</emphasis>
</taxonomicName>
was recovered sister species to
<taxonomicName id="4C385EC58B76FF941CF100FDFB5AFF47" box="[1007,1183,151,176]" class="Malacostraca" family="Galatheidae" genus="Agononida" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="rubrizonata">
<emphasis id="B94CF9548B76FF941CF100FDFB5AFF47" box="[1007,1183,151,176]" italics="true" pageId="8" pageNumber="209">A. rubrizonata</emphasis>
</taxonomicName>
in clade B but is morphologically most similar to
<taxonomicName id="4C385EC58B76FF941D1500D9FDBFFF23" box="[523,634,188,212]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B94CF9548B76FF941D1500D9FDBFFF23" box="[523,634,188,212]" italics="true" pageId="8" pageNumber="209">A. incerta</emphasis>
</taxonomicName>
. The anterolateral margin of the telson has only a slight indentation in the male (almost none in the female), the anterior three-quarters being only slightly concave if at all and the posterior quarter barely convex (
<taxonomicName id="4C385EC58B76FF941D1A0161FDB6FEEB" box="[516,627,260,284]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B94CF9548B76FF941D1A0161FDB6FEEB" box="[516,627,260,284]" italics="true" pageId="8" pageNumber="209">A. incerta</emphasis>
</taxonomicName>
always has a clear notch separating these two sections). The numbers of robust setae on pereopodal dactyli are fewer, usually 26 on pereopod 2 (1230, median 17, in
<taxonomicName id="4C385EC58B76FF941BDD014DFAF7FEB7" box="[1219,1330,296,320]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B94CF9548B76FF941BDD014DFAF7FEB7" box="[1219,1330,296,320]" italics="true" pageId="8" pageNumber="209">A. incerta</emphasis>
</taxonomicName>
) and 14 on pereopod 3 (210, median 6, in
<taxonomicName id="4C385EC58B76FF941D050129FD4DFE93" box="[539,648,332,356]" class="Malacostraca" family="Galatheidae" genus="Munida" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="B94CF9548B76FF941D050129FD4DFE93" box="[539,648,332,356]" italics="true" pageId="8" pageNumber="209">A. incerta</emphasis>
</taxonomicName>
). The cheliped of an ovigerous female (
<figureCitation id="130339C38B76FF941B5B0129FB42FE93" box="[1093,1159,332,357]" captionStart="FIGURE 4" captionStartId="8.[151,250,1142,1164]" captionTargetBox="[160,1427,555,1118]" captionTargetId="figure@8.[606,1030,801,1119]" captionTargetPageId="8" captionText="FIGURE 4. Agononida auscerta. Pereopods 1, upper and lateral views: a, ovigerous female, 24 mm, NMV J 55995. Agononida incerta. Anterolateral margin of telson: b, ovigerous female, 24 mm, NMV J 5521; c, male, 28 mm, NMV J 55025." httpUri="https://zenodo.org/record/228959/files/figure.png" pageId="8" pageNumber="209">Fig. 4</figureCitation>
a) is similar to that of the small male figured by
<bibRefCitation id="EFA958B78B76FF941E890115FCEEFE7F" author="Poore" box="[407,811,367,392]" pageId="8" pageNumber="209" refString="Poore, G. C. B. &amp; Andreakis, N. (2012) The Agononida incerta species complex unravelled (Crustacea: Decapoda: Anomura: Munididae). Zootaxa, 3429, 1 - 29." type="journal article" year="2012">
Poore &amp; Andreakis (2012:
<figureCitation id="130339C38B76FF941DD5010AFCCBFE7F" box="[715,782,367,392]" captionStart="FIGURE 9" captionStartId="17.[151,250,865,887]" captionText="FIGURE 9. Agononida polycerta n. sp. Paratype, male, 28 mm, MNHN IU- 2010 - 6091: a, anterolateral margin of telson; b d, dactyli of pereopods 2 4. Paratype, female, 29 mm, MNHN IU- 2010 - 6090: e, anterolateral margin of telson; f h, dactyli of pereopods 2 4." pageId="8" pageNumber="209">Fig. 9</figureCitation>
A)
</bibRefCitation>
and is of a delicate form; it is uncertain if more robust forms exist as in larger females of other species. Supermales are unknown. The specimen identified by Enrique Macpherson and photographed by T.-Y. Chan extends the range of this species further east and north from its original records in Western
<collectingCountry id="F32F65D68B76FF941ED301B9FDF2FE02" box="[461,567,476,501]" name="Australia" pageId="8" pageNumber="209">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>