379 lines
53 KiB
XML
379 lines
53 KiB
XML
<document id="55D5B853D08469F7FEB863E17404AAAC" ID-DOI="10.5281/zenodo.5358925" ID-ISSN="2345-7600" ID-Zenodo-Dep="5358925" ID-ZooBank="6FECC405-FF06-41AF-8249-97B1F3528085" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630225870716" checkinUser="valdenar" docAuthor="Golovatch, Sergei I." docDate="2018" docId="03916D22C543496AFC5D7497FC96DC37" docLanguage="en" docName="66rbz361-370.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Circulocryptus kompantsevi Golovatch 2018, new species" docType="treatment" docUuid="1339E07B-6D35-4EAB-A2D5-06ED04CF6B66" docUuidSource="ZooBank" docVersion="4" lastPageNumber="364" masterDocId="FFA8155AC5434969FFC37312FE78D415" masterDocTitle="Further notes on the millipede family Cryptodesmidae in Southeast Asia, with descriptions of a new genus and two new species from Indochina (Diplopoda: Polydesmida)" masterLastPageNumber="370" masterPageNumber="361" pageNumber="361" updateTime="1699192007770" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="009ADFEB24F0E2A74CCA7767A6481FF0">
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<mods:title id="180EF13ABA67EB2C79A15A247F2BA6BF">Further notes on the millipede family Cryptodesmidae in Southeast Asia, with descriptions of a new genus and two new species from Indochina (Diplopoda: Polydesmida)</mods:title>
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<mods:name id="E2E45034988A616CC118004880F5E2A7" type="personal">
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<mods:namePart id="981354ABF03FAB22F1D07CF503AF16E8">Golovatch, Sergei I.</mods:namePart>
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<mods:title id="F99F3578FB4CEBF935A175904C9B0236">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="183109370EE5043C0650277D81BA3C12">
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<mods:date id="B805E73865CA7487FF389387E84CDA1D">2018</mods:date>
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<mods:number id="E47456383998C6BCFFC63B5A4296FCD2">2018-06-26</mods:number>
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<mods:detail id="2ED5E70600F2CB0D3AC18A322C464A15" type="volume">
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<mods:number id="254ABC901458B31D323217679E7C2201">66</mods:number>
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<mods:classification id="AACF39C827613DA49A2C717577DFEA61">journal article</mods:classification>
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<mods:identifier id="261B652484D044C74098A836DAF84FC3" type="DOI">10.5281/zenodo.5358925</mods:identifier>
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<treatment id="03916D22C543496AFC5D7497FC96DC37" LSID="urn:lsid:zoobank.org:act:1339E07B-6D35-4EAB-A2D5-06ED04CF6B66" httpUri="http://treatment.plazi.org/id/03916D22C543496AFC5D7497FC96DC37" lastPageId="3" lastPageNumber="364" pageId="0" pageNumber="361">
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<subSubSection id="C3228FBFC5434969FC5D7497FB10D38B" box="[926,1384,1925,1950]" pageId="0" pageNumber="361" type="nomenclature">
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<paragraph id="8B87DC34C5434969FC5D7497FB10D38B" blockId="0.[926,1384,1925,1982]" box="[926,1384,1925,1950]" pageId="0" pageNumber="361">
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<heading id="D0CF6B58C5434969FC5D7497FB10D38B" bold="true" box="[926,1384,1925,1950]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="361" reason="2">
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<emphasis id="B94C0026C5434969FC5D7497FB10D38B" bold="true" box="[926,1384,1925,1950]" pageId="0" pageNumber="361">
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<taxonomicName id="4C38A7B7C5434969FC5D7497FAABD388" ID-CoL="VG5Q" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[926,1235,1925,1949]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
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<emphasis id="B94C0026C5434969FC5D7497FAABD388" bold="true" box="[926,1235,1925,1949]" italics="true" pageId="0" pageNumber="361">Circulocryptus kompantsevi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27FBD5DC5434969FB207494FB10D38B" box="[1251,1384,1926,1950]" pageId="0" pageNumber="361" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3228FBFC5434969FB8574B4FAC7D3AB" box="[1094,1215,1958,1982]" pageId="0" pageNumber="361" type="description">
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<paragraph id="8B87DC34C5434969FB8574B4FAC7D3AB" blockId="0.[926,1384,1925,1982]" box="[1094,1215,1958,1982]" pageId="0" pageNumber="361">
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(
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<figureCitation id="1303C0B1C5434969FB8D74B4FAE1D3AB" box="[1102,1177,1958,1982]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="0" pageNumber="361">Figs. 1</figureCitation>
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,
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<figureCitation id="1303C0B1C5434969FB6A74B4FACFD3AB" box="[1193,1207,1958,1982]" captionStart="Fig" captionStartId="3.[113,148,929,951]" captionTargetBox="[114,1450,190,916]" captionTargetId="figure-8@3.[97,1464,184,919]" captionTargetPageId="3" captionText="Fig. 2. Circulocryptus kompantsevi, new species, holotype. A, Leg 9, lateral view; B, right gonopod, lateral view; C, right gonopod, mesal view. Scale bar = 0.1 mm (A) and 0.2 mm (B, C). Designations: a = accessory seminal chamber; b = basal branch; l = hyaline fringed lobe at base of solenomere; p = prefemoral process; sl = solenomere; sph = solenophorous branch." figureDoi="http://doi.org/10.5281/zenodo.5358929" httpUri="https://zenodo.org/record/5358929/files/figure.png" pageId="0" pageNumber="361">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3228FBFC5434968FC8074F4FF79D4E0" lastPageId="1" lastPageNumber="362" pageId="0" pageNumber="361" type="materials_examined">
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<paragraph id="8B87DC34C5434969FC8074F4FBB9DC2B" blockId="0.[835,1474,2022,2110]" pageId="0" pageNumber="361">
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<emphasis id="B94C0026C5434969FC8074F4FA5DD3EB" bold="true" box="[835,1061,2022,2046]" pageId="0" pageNumber="361">Material examined.</emphasis>
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<materialsCitation id="3B50D669C5434969FBED74F4FBB9DC2B" collectionCode="ZMUM" country="Cambodia" county="Elephant Mountains" location="Gonoderma" municipality="Phnom Pokor National Park" pageId="0" pageNumber="361" specimenCount="1" stateProvince="Kampot" typeStatus="holotype">
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<typeStatus id="54836296C5434969FBED74F4FAE9D3EB" box="[1070,1169,2022,2046]" pageId="0" pageNumber="361" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED2944F1C5434969FB2374F4FB41D3EB" box="[1248,1337,2022,2046]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="0" pageNumber="361">ZMUM</collectionCode>
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),
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<collectingCountry id="F32F9CA4C5434969FA8D74F4FBC6D3EB" box="[1358,1470,2022,2046]" name="Cambodia" pageId="0" pageNumber="361">Cambodia</collectingCountry>
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,
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<collectingRegion id="49FC12D6C5434969FC807B14FDA4DC08" box="[835,988,2054,2077]" country="Cambodia" name="Kampot" pageId="0" pageNumber="361">Kampot Prov.</collectingRegion>
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,
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<collectingCounty id="62E6A4B8C5434969FC247B14FABADC0B" box="[999,1218,2054,2078]" pageId="0" pageNumber="361">Elephant Mountains</collectingCounty>
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,
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<collectingMunicipality id="6BE3464EC5434969FB0E7B14FD02DC2B" pageId="0" pageNumber="361">Phnom Pokor National Park</collectingMunicipality>
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, inside a rotten fruit body of
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<emphasis id="B94C0026C5434969FB1D7B34FB1BDC2B" box="[1246,1379,2086,2110]" italics="true" pageId="0" pageNumber="361">
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<location id="8EE78AEFC5434969FB1D7B34FB1BDC2B" LSID="urn:lsid:plazi:treatment:03916D22C543496AFC5D7497FC96DC37:8EE78AEFC5434969FB1D7B34FB1BDC2B" box="[1246,1379,2086,2110]" country="Cambodia" county="Elephant Mountains" municipality="Phnom Pokor National Park" name="Gonoderma" pageId="0" pageNumber="361" stateProvince="Kampot">Gonoderma</location>
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</emphasis>
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fungus,
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</materialsCitation>
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</paragraph>
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<paragraph id="8B87DC34C5424968FFB073AEFF79D4E0" blockId="1.[115,753,188,245]" pageId="1" pageNumber="362">
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<geoCoordinate id="EE0CBAF3C5424968FFB073AEFF5FD4C0" box="[115,295,188,213]" degrees="10" direction="north" minutes="37" orientation="latitude" pageId="1" pageNumber="362" precision="1" seconds="08.89" value="10.619136">10°37′08.89″N</geoCoordinate>
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,
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<geoCoordinate id="EE0CBAF3C5424968FEFB73AEFF80D4C0" box="[312,504,188,213]" degrees="104" direction="east" minutes="04" orientation="longitude" pageId="1" pageNumber="362" precision="1" seconds="40.87" value="104.07802">104°04′40.87″E</geoCoordinate>
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,
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<date id="FF86FAF4C5424968FDCA73AFFCF5D4C0" box="[521,653,189,213]" pageId="1" pageNumber="362" value="2017-12-08">8.XII.2017</date>
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, leg. A. Kompantsev.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3228FBFC5424968FFB2720FFC96D760" pageId="1" pageNumber="362" type="diagnosis">
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<paragraph id="8B87DC34C5424968FFB2720FFE85D720" blockId="1.[112,754,285,821]" pageId="1" pageNumber="362">
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<emphasis id="B94C0026C5424968FFB2720FFE90D520" bold="true" box="[113,232,285,309]" pageId="1" pageNumber="362">Diagnosis.</emphasis>
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Differs from both known congeners,
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<taxonomicName id="4C38A7B7C5424968FD56720CFF41D540" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="fallei">
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<emphasis id="B94C0026C5424968FD56720CFC88D520" box="[661,752,285,309]" italics="true" pageId="1" pageNumber="362">C. fallei</emphasis>
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<bibRefCitation id="EFA9A1C5C5424968FFB2722FFF41D540" author="Golovatch SI" box="[113,313,317,341]" pageId="1" pageNumber="362" pagination="33 - 43" refId="ref6834" refString="Golovatch SI (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda: Polydesmida). ZooKeys, 578: 33 - 43. http: // dx. doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
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</taxonomicName>
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, the type-species from southern and central-southern
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<collectingCountry id="F32F9CA4C5424968FEE9724FFFFED560" box="[298,390,349,373]" name="Vietnam" pageId="1" pageNumber="362">Vietnam</collectingCountry>
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||
(
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<bibRefCitation id="EFA9A1C5C5424968FE5B724FFC2BD560" author="Golovatch SI" box="[408,595,349,373]" pageId="1" pageNumber="362" pagination="33 - 43" refId="ref6834" refString="Golovatch SI (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda: Polydesmida). ZooKeys, 578: 33 - 43. http: // dx. doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
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;
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<bibRefCitation id="EFA9A1C5C5424968FDA2724FFF26D580" author="Golovatch SI & VandenSpiegel D" pageId="1" pageNumber="362" pagination="757 - 767" refId="ref6912" refString="Golovatch SI & VandenSpiegel D (2017) One new and two littleknown species of the millipede family Cryptodesmidae from Indochina (Diplopoda, Polydesmida). Zoologicheskii zhurnal, 96 (7): 757 - 767." type="journal article" year="2017">Golovatch & VandenSpiegel, 2017</bibRefCitation>
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), and
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<taxonomicName id="4C38A7B7C5424968FE60726CFC94D580" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[419,748,381,405]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="javanicus">
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<emphasis id="B94C0026C5424968FE60726CFC53D580" box="[419,555,382,405]" italics="true" pageId="1" pageNumber="362">C. javanicus</emphasis>
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<bibRefCitation id="EFA9A1C5C5424968FDF6726FFC94D580" author="Golovatch SI" box="[565,748,381,405]" pageId="1" pageNumber="362" pagination="1 - 10" refId="ref6873" refString="Golovatch SI (2018) Two new species of polydesmoid millipedes from western Java, Indonesia (Diplopoda: Polydesmida: Cryptodesmidae, Haplodesmidae). Tropical Natural History, 18 (1): 1 - 10." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
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</taxonomicName>
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, from western
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<collectingRegion id="49FC12D6C5424968FED2728CFF3CD5A0" box="[273,324,414,437]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="362">Java</collectingRegion>
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,
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<collectingCountry id="F32F9CA4C5424968FE97728FFFC7D5A0" box="[340,447,413,437]" name="Indonesia" pageId="1" pageNumber="362">Indonesia</collectingCountry>
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(
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<bibRefCitation id="EFA9A1C5C5424968FE12728FFCEAD5A0" author="Golovatch SI" box="[465,658,413,437]" pageId="1" pageNumber="362" pagination="1 - 10" refId="ref6873" refString="Golovatch SI (2018) Two new species of polydesmoid millipedes from western Java, Indonesia (Diplopoda: Polydesmida: Cryptodesmidae, Haplodesmidae). Tropical Natural History, 18 (1): 1 - 10." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
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), by the conspicuously swollen anterior half of the collum and certain minor details of the gonopodal structure, i.e., the shapes of the prefemoral process and basal branch. In showing 25 radii at the anterior margin of the collum, only four transverse rows of tubercles and four lateral lobulations on each of postcollum metaterga,
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<taxonomicName id="4C38A7B7C5424968FEA9714CFC77D660" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[362,527,605,629]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
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<emphasis id="B94C0026C5424968FEA9714CFC77D660" box="[362,527,605,629]" italics="true" pageId="1" pageNumber="362">C. kompantsevi</emphasis>
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A27FBD5DC5424968FDDD714CFCDBD660" box="[542,675,605,629]" pageId="1" pageNumber="362" rank="species">new species</taxonomicNameLabel>
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, seems to be more similar to the geographically closer
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<taxonomicName id="4C38A7B7C5424968FD4D716CFC91D680" authorityName="Golovatch" authorityYear="2016" box="[654,745,637,661]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="fallei">
|
||
<emphasis id="B94C0026C5424968FD4D716CFC91D680" box="[654,745,637,661]" italics="true" pageId="1" pageNumber="362">C. fallei</emphasis>
|
||
</taxonomicName>
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||
, but differs clearly in the light body (vs. sometimes dark), the presence of a fimbriate prefemoral process
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||
<emphasis id="B94C0026C5424968FD5471AFFCDED6C0" bold="true" box="[663,678,701,725]" pageId="1" pageNumber="362">p</emphasis>
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||
(vs. a finger-shaped trichostele) on the gonopodal telopodite, the latter’s branch
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<emphasis id="B94C0026C5424968FEDA71EFFF50D700" bold="true" box="[281,296,765,789]" pageId="1" pageNumber="362">b</emphasis>
|
||
being somewhat longer than branch
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||
<emphasis id="B94C0026C5424968FD0B71EFFC88D700" bold="true" box="[712,752,765,789]" pageId="1" pageNumber="362">sph</emphasis>
|
||
(
|
||
<figureCitation id="1303C0B1C5424968FFBA700FFE89D720" box="[121,241,797,821]" captionStart="Fig" captionStartId="3.[113,148,929,951]" captionTargetBox="[114,1450,190,916]" captionTargetId="figure-8@3.[97,1464,184,919]" captionTargetPageId="3" captionText="Fig. 2. Circulocryptus kompantsevi, new species, holotype. A, Leg 9, lateral view; B, right gonopod, lateral view; C, right gonopod, mesal view. Scale bar = 0.1 mm (A) and 0.2 mm (B, C). Designations: a = accessory seminal chamber; b = basal branch; l = hyaline fringed lobe at base of solenomere; p = prefemoral process; sl = solenomere; sph = solenophorous branch." figureDoi="http://doi.org/10.5281/zenodo.5358929" httpUri="https://zenodo.org/record/5358929/files/figure.png" pageId="1" pageNumber="362">Fig. 2B, C</figureCitation>
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).
|
||
</paragraph>
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<paragraph id="8B87DC34C5424968FFB2704FFC96D760" blockId="1.[113,750,861,885]" box="[113,750,861,885]" pageId="1" pageNumber="362">
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||
<emphasis id="B94C0026C5424968FFB2704FFEC3D760" bold="true" box="[113,187,861,885]" pageId="1" pageNumber="362">Name.</emphasis>
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||
Honours Dr. Aleksandr Kompantsev, the collector.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C3228FBFC5424968FFB2708FFB2AD2A0" pageId="1" pageNumber="362" type="description">
|
||
<paragraph id="8B87DC34C5424968FFB2708FFF88D063" blockId="1.[112,753,925,1142]" pageId="1" pageNumber="362">
|
||
<emphasis id="B94C0026C5424968FFB2708FFF79D7A0" bold="true" box="[113,257,925,949]" pageId="1" pageNumber="362">Description.</emphasis>
|
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Length ca
|
||
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||
, width of midbody pro- and metazonae 1.6 and
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||
<quantity id="4CC071D1C5424968FE4E70AFFF91D7C0" box="[397,489,957,981]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="362" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, respectively. General colouration in alcohol rather uniformly light pinkish (
|
||
<figureCitation id="1303C0B1C5424968FD0670CFFE05D000" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1</figureCitation>
|
||
), only antennae brown to dark purplish brown, increasingly infuscate distad; head, anterior half of collum, distal halves of postcollum paraterga, mid-dorsal parts of prozonae, sterna and legs light yellowish to creamy.
|
||
</paragraph>
|
||
<paragraph id="8B87DC34C5424968FFB2778FFF5AD1E0" blockId="1.[112,754,1181,1525]" pageId="1" pageNumber="362">
|
||
Body with 20 segments. In width, head << collum <segment 2 <3 <4 = 15; thereafter body gradually tapering towards telson. Head densely setose, between and behind antennae densely and clearly microgranulate; epicranial suture superficial, more distinct in vertigial region. Antennae very short and densely setose, clearly clavate, C-shaped (
|
||
<figureCitation id="1303C0B1C5424968FD00762FFECDD160" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1A–C</figureCitation>
|
||
); in length, antennomere 1 = 2 = 4 = 7 <3 <6 <5; antennomeres 5 and 6 subequally high, each with a small, compact, apicodorsal group of minute bacilliform sensilla. Interantennal isthmus ca 1.3 times as broad as diameter of antennal socket.
|
||
</paragraph>
|
||
<paragraph id="8B87DC34C5424968FFB2750CFDE0D7C0" blockId="1.[112,754,1565,2101]" lastBlockId="1.[811,1453,189,981]" pageId="1" pageNumber="362">
|
||
Tegument mostly dull and clearly microgranulate almost throughout, only prozonae and strictures between pro- and metazonae shining, very finely shagreened (prozonae) or micro-alveolate (strictures). Dorsum strongly and regularly convex; paraterga mostly set at about upper 1/3 of body, very strongly developed, beginning with collum, regularly declined ventrad much like dorsum (
|
||
<figureCitation id="1303C0B1C5424968FE2475CFFC5AD2E0" box="[487,546,1757,1781]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1</figureCitation>
|
||
). Collum flabellate, completely covering the head from above, tauriform, anterior half distinctly swollen, posterior half clearly impressed and with 5–6 transverse, irregular rows of small, round, setigerous tuberculations; anterior margin regularly rounded, with 25 indistinct radii, central setigerous lobulations especially inconspicuous compared to more lateral ones; paraterga narrowly rounded caudally, indistinctly lobulate laterally and caudally as well. Postcollum paraterga very broad, each much broader than prozona; anterior margin/shoulder always entire, mostly nearly straight and narrowly bordered, increasingly, but faintly curved caudad towards telson, with neither radii nor crenulations, only anterolateral corner slightly rounded and bearing a particularly small lobulation, followed by three radii and larger lobulations along a subquadrate lateral edge, caudal corner drawn increasingly behind rear tergal margin only in segments 16–19; paraterga 19 produced nearly as far behind as tip of epiproct. Caudal margins of paraterga both radiate and increasingly strongly lobulate laterad, typically with 12+12 setigerous lobulations on each side, regardless of caudolateral ones (
|
||
<figureCitation id="1303C0B1C5424968FBD772CFFA2CD5E0" box="[1044,1108,477,501]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1</figureCitation>
|
||
). Mid-dorsal parts of metaterga with four transverse, rather regular rows of small, setigerous, round tubercles/knobs, caudal row being especially regular; 1–3 irregular rows of dorsal knobs increasingly obliterated when extended onto distal parts of paraterga. Tergal setae short, light, pointed, usually well visible, 1/4–1/5 as long as metatergite. Neither an axial line nor pleurosternal carinae. Ozopores present anteriorly at least at the very base of paraterga 7 (in a broken body), invisible, pore formula untraceable. Stricture dividing pro- and metazonae broad, shallow and more shining than adjacent metazonae. Limbus brush-like, very densely microspiculate (
|
||
<figureCitation id="1303C0B1C5424968FB12702FFB38D740" box="[1233,1344,829,853]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1C, D</figureCitation>
|
||
). Epiproct (
|
||
<figureCitation id="1303C0B1C5424968FCF0704FFDB2D760" box="[819,970,861,885]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1A, B, D</figureCitation>
|
||
) conical, slightly emarginate at apex, dorsal and lateral tubercles evident. Hypoproct semi-circular, 1+1 caudal setae clearly separated, borne on evident tubercles (
|
||
<figureCitation id="1303C0B1C5424968FCF670AFFDF3D7C0" box="[821,907,957,981]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87DC34C5424968FCEE70EFFA29D120" blockId="1.[813,1453,1021,1333]" pageId="1" pageNumber="362">
|
||
Sternites narrow (
|
||
<figureCitation id="1303C0B1C5424968FBC170EFFAFFD000" box="[1026,1159,1021,1045]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1C, D</figureCitation>
|
||
), densely setose, clearly impressed along main axis, distinctly broadened only between coxae 7 and 9 (male). Legs long and slender, without modifications (
|
||
<figureCitation id="1303C0B1C5424968FBE4774FFAB2D060" box="[1063,1226,1117,1141]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Figs. 1A, C, D</figureCitation>
|
||
,
|
||
<figureCitation id="1303C0B1C5424968FB15774FFA82D060" box="[1238,1274,1117,1141]" captionStart="Fig" captionStartId="3.[113,148,929,951]" captionTargetBox="[114,1450,190,916]" captionTargetId="figure-8@3.[97,1464,184,919]" captionTargetPageId="3" captionText="Fig. 2. Circulocryptus kompantsevi, new species, holotype. A, Leg 9, lateral view; B, right gonopod, lateral view; C, right gonopod, mesal view. Scale bar = 0.1 mm (A) and 0.2 mm (B, C). Designations: a = accessory seminal chamber; b = basal branch; l = hyaline fringed lobe at base of solenomere; p = prefemoral process; sl = solenomere; sph = solenophorous branch." figureDoi="http://doi.org/10.5281/zenodo.5358929" httpUri="https://zenodo.org/record/5358929/files/figure.png" pageId="1" pageNumber="362">2A</figureCitation>
|
||
), almost as long as paratergal width; in length, tarsi = femora >> coxae = prefemora = postfemora = tibiae; gonapophyses on coxae 2 small cones (male); neither adenostyles nor tarsal brushes. Claws simple, slightly curved ventrad. Gonopod aperture transversely ovoid, caudal and lateral margins thin and slightly elevated (
|
||
<figureCitation id="1303C0B1C5424968FC2C760FFA3CD120" box="[1007,1092,1309,1333]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87DC34C5424968FCEE764FFB2AD2A0" blockId="1.[811,1452,1373,1717]" pageId="1" pageNumber="362">
|
||
Gonopods (
|
||
<figureCitation id="1303C0B1C5424968FC65764FFA78D160" box="[934,1024,1373,1397]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="1" pageNumber="362">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="1303C0B1C5424968FBC9764FFA30D160" box="[1034,1096,1373,1397]" captionStart="Fig" captionStartId="3.[113,148,929,951]" captionTargetBox="[114,1450,190,916]" captionTargetId="figure-8@3.[97,1464,184,919]" captionTargetPageId="3" captionText="Fig. 2. Circulocryptus kompantsevi, new species, holotype. A, Leg 9, lateral view; B, right gonopod, lateral view; C, right gonopod, mesal view. Scale bar = 0.1 mm (A) and 0.2 mm (B, C). Designations: a = accessory seminal chamber; b = basal branch; l = hyaline fringed lobe at base of solenomere; p = prefemoral process; sl = solenomere; sph = solenophorous branch." figureDoi="http://doi.org/10.5281/zenodo.5358929" httpUri="https://zenodo.org/record/5358929/files/figure.png" pageId="1" pageNumber="362">2B, C</figureCitation>
|
||
) in situ held parallel to each other. Telopodite typically subcircular, very strongly curved caudad, bipartite, with a short, “prefemoral”, fimbriate, basal process (
|
||
<emphasis id="B94C0026C5424968FCF676AFFD3CD1C0" bold="true" box="[821,836,1469,1493]" pageId="1" pageNumber="362">p</emphasis>
|
||
) on lateral face. Main, basal part (
|
||
<emphasis id="B94C0026C5424968FB7276AFFAB8D1C0" bold="true" box="[1201,1216,1469,1493]" pageId="1" pageNumber="362">b</emphasis>
|
||
) of telopodite ribbonshaped, slightly curved anteriad and subtruncate apically, extended in distal third into a long, solid, subunciform, bare, lateral solenomere (
|
||
<emphasis id="B94C0026C5424968FC3E750FFA77D220" bold="true" box="[1021,1039,1565,1589]" pageId="1" pageNumber="362">sl</emphasis>
|
||
), the latter at its base bearing a fringed, membranous, hyaline lobe (
|
||
<emphasis id="B94C0026C5424968FB9A752FFA18D240" bold="true" box="[1113,1120,1597,1621]" pageId="1" pageNumber="362">l</emphasis>
|
||
). A distinct accessory seminal chamber (
|
||
<emphasis id="B94C0026C5424968FC5E754FFDD2D260" bold="true" box="[925,938,1629,1653]" pageId="1" pageNumber="362">a</emphasis>
|
||
) and a short duct devoid of a hairy pulvillus lying in distal part of solenophore (
|
||
<emphasis id="B94C0026C5424968FB7C756FFA9FD280" bold="true" box="[1215,1255,1661,1685]" pageId="1" pageNumber="362">sph</emphasis>
|
||
) just opposite
|
||
<emphasis id="B94C0026C5424968FA50756FFBD4D280" bold="true" box="[1427,1452,1661,1685]" pageId="1" pageNumber="362">sl,</emphasis>
|
||
branch
|
||
<emphasis id="B94C0026C5424968FCBD758FFDF5D2A0" bold="true" box="[894,909,1693,1717]" pageId="1" pageNumber="362">b</emphasis>
|
||
being somewhat longer than branch
|
||
<emphasis id="B94C0026C5424968FAE6758FFB33D2A0" bold="true" box="[1317,1355,1693,1717]" pageId="1" pageNumber="362">sph</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3228FBFC542496AFCEE75CFFC4AD1CA" lastPageId="3" lastPageNumber="364" pageId="1" pageNumber="362" type="discussion">
|
||
<paragraph id="8B87DC34C542496AFCEE75CFFFB6D16A" blockId="1.[812,1452,1757,2101]" lastBlockId="3.[112,753,1063,1408]" lastPageId="3" lastPageNumber="364" pageId="1" pageNumber="362">
|
||
<emphasis id="B94C0026C5424968FCEE75CFFDDBD2E0" bold="true" box="[813,931,1757,1781]" pageId="1" pageNumber="362">Remarks.</emphasis>
|
||
As noted above, the genus
|
||
<taxonomicName id="4C38A7B7C5424968FB3E75CFFD99D300" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C0026C5424968FB3E75CFFBD3D2E0" box="[1277,1451,1757,1781]" italics="true" pageId="1" pageNumber="362">Circulocryptus</emphasis>
|
||
<bibRefCitation id="EFA9A1C5C5424968FCEE75EFFD99D300" author="Golovatch SI" box="[813,993,1789,1813]" pageId="1" pageNumber="362" pagination="33 - 43" refId="ref6834" refString="Golovatch SI (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda: Polydesmida). ZooKeys, 578: 33 - 43. http: // dx. doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
has hitherto been known to contain only two species:
|
||
<taxonomicName id="4C38A7B7C5424968FC05740CFA5CD320" authorityName="Golovatch" authorityYear="2016" box="[966,1060,1821,1845]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="fallei">
|
||
<emphasis id="B94C0026C5424968FC05740CFA5CD320" box="[966,1060,1821,1845]" italics="true" pageId="1" pageNumber="362">C. fallei</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F32F9CA4C5424968FBB3740FFAA8D320" box="[1136,1232,1821,1845]" name="Vietnam" pageId="1" pageNumber="362">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFA9A1C5C5424968FB20740FFBDFD320" author="Golovatch SI" box="[1251,1447,1821,1845]" pageId="1" pageNumber="362" pagination="33 - 43" refId="ref6834" refString="Golovatch SI (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda: Polydesmida). ZooKeys, 578: 33 - 43. http: // dx. doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA9A1C5C5424968FCEE742FFAB5D340" author="Golovatch SI & VandenSpiegel D" box="[813,1229,1853,1877]" pageId="1" pageNumber="362" pagination="757 - 767" refId="ref6912" refString="Golovatch SI & VandenSpiegel D (2017) One new and two littleknown species of the millipede family Cryptodesmidae from Indochina (Diplopoda, Polydesmida). Zoologicheskii zhurnal, 96 (7): 757 - 767." type="journal article" year="2017">Golovatch & VandenSpiegel, 2017</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="4C38A7B7C5424968FADA742CFBD3D340" authorityName="Golovatch" authorityYear="2018" box="[1305,1451,1854,1877]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="javanicus">
|
||
<emphasis id="B94C0026C5424968FADA742CFBD3D340" box="[1305,1451,1854,1877]" italics="true" pageId="1" pageNumber="362">C. javanicus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49FC12D6C5424968FCAF744CFDE7D360" box="[876,927,1886,1909]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="362">Java</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32F9CA4C5424968FC6C744FFA62D360" box="[943,1050,1885,1909]" name="Indonesia" pageId="1" pageNumber="362">Indonesia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFA9A1C5C5424968FBEF744FFA95D360" author="Golovatch SI" box="[1068,1261,1885,1909]" pageId="1" pageNumber="362" pagination="1 - 10" refId="ref6873" refString="Golovatch SI (2018) Two new species of polydesmoid millipedes from western Java, Indonesia (Diplopoda: Polydesmida: Cryptodesmidae, Haplodesmidae). Tropical Natural History, 18 (1): 1 - 10." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
|
||
). The third, new species fails to alter the generic diagnosis as reformulated by
|
||
<bibRefCitation id="EFA9A1C5C5424968FC8C748FFA68D3A0" author="Golovatch SI" box="[847,1040,1949,1973]" pageId="1" pageNumber="362" pagination="1 - 10" refId="ref6873" refString="Golovatch SI (2018) Two new species of polydesmoid millipedes from western Java, Indonesia (Diplopoda: Polydesmida: Cryptodesmidae, Haplodesmidae). Tropical Natural History, 18 (1): 1 - 10." type="journal article" year="2018">Golovatch (2018)</bibRefCitation>
|
||
. This primarily concerns an obviously subcircular, biramous, highly complex gonopodal telopodite that shows an accessory seminal chamber and its duct on a separate solenophorous branch (
|
||
<emphasis id="B94C0026C5424968FB6374EFFAB0DC00" bold="true" box="[1184,1224,2045,2069]" pageId="1" pageNumber="362">sph</emphasis>
|
||
) which lies opposite the basal branch (
|
||
<emphasis id="B94C0026C5424968FC2C7B0FFD86DC20" bold="true" box="[1007,1022,2077,2101]" pageId="1" pageNumber="362">b</emphasis>
|
||
) with a spiniform solenomere (
|
||
<emphasis id="B94C0026C5424968FA9B7B0FFB12DC20" bold="true" box="[1368,1386,2077,2101]" pageId="1" pageNumber="362">sl</emphasis>
|
||
). The presence in
|
||
<taxonomicName id="4C38A7B7C540496AFF39773AFFDBD02A" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[250,419,1063,1087]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
|
||
<emphasis id="B94C0026C540496AFF39773AFFDBD02A" box="[250,419,1063,1087]" italics="true" pageId="3" pageNumber="364">C. kompantsevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27FBD5DC540496AFE76773AFC45D02A" box="[437,573,1063,1087]" pageId="3" pageNumber="364" rank="species">new species</taxonomicNameLabel>
|
||
, of a small, but evident gonoprefemoral process (
|
||
<emphasis id="B94C0026C540496AFE27775AFF8BD075" bold="true" box="[484,499,1096,1120]" pageId="3" pageNumber="364">p</emphasis>
|
||
) which is absent from
|
||
<taxonomicName id="4C38A7B7C540496AFFB2777AFE86D06A" authorityName="Golovatch" authorityYear="2018" box="[113,254,1128,1151]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="javanicus">
|
||
<emphasis id="B94C0026C540496AFFB2777AFE86D06A" box="[113,254,1128,1151]" italics="true" pageId="3" pageNumber="364">C. javanicus</emphasis>
|
||
</taxonomicName>
|
||
and often absent from
|
||
<taxonomicName id="4C38A7B7C540496AFDCC777AFC14D06A" authorityName="Golovatch" authorityYear="2016" box="[527,620,1127,1151]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="fallei">
|
||
<emphasis id="B94C0026C540496AFDCC777AFC14D06A" box="[527,620,1127,1151]" italics="true" pageId="3" pageNumber="364">C. fallei</emphasis>
|
||
</taxonomicName>
|
||
also seems characteristic of the genus. Whereas the latter trait is shared with a few other genera of
|
||
<taxonomicName id="4C38A7B7C540496AFE4477B5FC4DD0AA" authorityName="Karsch" authorityYear="1880" box="[391,565,1191,1215]" class="Diplopoda" family="Cryptodesmidae" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="family">Cryptodesmidae</taxonomicName>
|
||
in Southeast Asia (
|
||
<bibRefCitation id="EFA9A1C5C540496AFFB477D5FF4DD0CA" author="Golovatch SI" box="[119,309,1223,1247]" pageId="3" pageNumber="364" pagination="33 - 43" refId="ref6834" refString="Golovatch SI (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda: Polydesmida). ZooKeys, 578: 33 - 43. http: // dx. doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
|
||
), the former feature is unique not only within the family, but also amongst all known genera of the order
|
||
<taxonomicName id="4C38A7B7C540496AFF707615FC76D135" authority="(Golovatch, 2018)" baseAuthorityName="Golovatch" baseAuthorityYear="2018" box="[179,526,1287,1312]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="order">
|
||
Polydesmida (
|
||
<bibRefCitation id="EFA9A1C5C540496AFE8F7615FC7ED135" author="Golovatch SI" box="[332,518,1287,1312]" pageId="3" pageNumber="364" pagination="1 - 10" refId="ref6873" refString="Golovatch SI (2018) Two new species of polydesmoid millipedes from western Java, Indonesia (Diplopoda: Polydesmida: Cryptodesmidae, Haplodesmidae). Tropical Natural History, 18 (1): 1 - 10." type="journal article" year="2018">Golovatch, 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. This alone warrants the promotion of
|
||
<taxonomicName id="4C38A7B7C540496AFEF27635FFA9D12A" authorityName="Golovatch" authorityYear="2016" box="[305,465,1319,1343]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C0026C540496AFEF27635FFA9D12A" box="[305,465,1319,1343]" italics="true" pageId="3" pageNumber="364">Circulocryptus</emphasis>
|
||
</taxonomicName>
|
||
to a cryptodesmid tribe of its own,
|
||
<taxonomicName id="4C38A7B7C540496AFF167655FF04D14A" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[213,380,1351,1375]" class="Diplopoda" family="Cryptodesmidae" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Circulocryptini">Circulocryptini</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27FBD5DC540496AFE497655FC71D14A" box="[394,521,1351,1375]" pageId="3" pageNumber="364" rank="tribe">tribus nova</taxonomicNameLabel>
|
||
, which also includes the new genus described below.
|
||
</paragraph>
|
||
<caption id="DF478CBCC541496BFF447B02FFB4DC56" ID-DOI="http://doi.org/10.5281/zenodo.5358927" ID-Zenodo-Dep="5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="2" pageNumber="363" startId="2.[135,170,2064,2086]" targetBox="[310,1291,203,2049]" targetPageId="2">
|
||
<paragraph id="8B87DC34C541496BFF447B02FFB4DC56" blockId="2.[134,1473,2064,2115]" pageId="2" pageNumber="363">
|
||
Fig. 1.
|
||
<taxonomicName id="4C38A7B7C541496BFF0A7B02FFAEDC30" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[201,470,2064,2085]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
|
||
<emphasis id="B94C0026C541496BFF0A7B02FFAEDC30" box="[201,470,2064,2085]" italics="true" pageId="2" pageNumber="363">Circulocryptus kompantsevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27FBD5DC541496BFE217B02FC2FDC33" box="[482,599,2064,2086]" pageId="2" pageNumber="363" rank="species">new species</taxonomicNameLabel>
|
||
, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF478CBCC540496AFFB270B3FA44D7E5" ID-DOI="http://doi.org/10.5281/zenodo.5358929" ID-Zenodo-Dep="5358929" httpUri="https://zenodo.org/record/5358929/files/figure.png" pageId="3" pageNumber="364" startId="3.[113,148,929,951]" targetBox="[114,1450,190,916]" targetPageId="3">
|
||
<paragraph id="8B87DC34C540496AFFB270B3FA44D7E5" blockId="3.[113,1452,929,1008]" pageId="3" pageNumber="364">
|
||
Fig. 2.
|
||
<taxonomicName id="4C38A7B7C540496AFF7470B3FFBCD7A3" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[183,452,929,950]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
|
||
<emphasis id="B94C0026C540496AFF7470B3FFBCD7A3" box="[183,452,929,950]" italics="true" pageId="3" pageNumber="364">Circulocryptus kompantsevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27FBD5DC540496AFE1270B3FC30D7A2" box="[465,584,929,951]" pageId="3" pageNumber="364" rank="species">new species</taxonomicNameLabel>
|
||
, holotype. A, Leg 9, lateral view; B, right gonopod, lateral view; C, right gonopod, mesal view. Scale bar = 0.1 mm (A) and 0.2 mm (B, C). Designations: a = accessory seminal chamber; b = basal branch; l = hyaline fringed lobe at base of solenomere; p = prefemoral process; sl = solenomere; sph = solenophorous branch.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87DC34C540496AFFB276B5FC4AD1CA" blockId="3.[113,752,1447,1503]" pageId="3" pageNumber="364">
|
||
The following key can be proposed to separate all three species of
|
||
<taxonomicName id="4C38A7B7C540496AFF2B76D5FFF0D1CA" authorityName="Golovatch" authorityYear="2016" box="[232,392,1479,1503]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C0026C540496AFF2B76D5FFF0D1CA" box="[232,392,1479,1503]" italics="true" pageId="3" pageNumber="364">Circulocryptus</emphasis>
|
||
</taxonomicName>
|
||
known to date:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3228FBFC540496AFFB27514FC96DC37" pageId="3" pageNumber="364" type="key">
|
||
<key id="FFA98B05C540496AFFB27514FC96DC37" pageId="3" pageNumber="364">
|
||
<keyStep id="30CCC491C540496AFFB27514FC97D32E" pageId="3" pageNumber="364">
|
||
<paragraph id="8B87DC34C540496AFFB27514FC96D2F3" blockId="3.[113,755,1542,2083]" pageId="3" pageNumber="364">
|
||
<keyLead id="30C97F01C540496AFFB27514FC96D2F3" pageId="3" pageNumber="364">
|
||
1. General colouration light, mostly light brown to yellow (
|
||
<figureCitation id="1303C0B1C540496AFD687514FC99D209" box="[683,737,1542,1564]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="3" pageNumber="364">Fig. 1</figureCitation>
|
||
). Collum conspicuously swollen in anterior half and impressed in posterior one (
|
||
<figureCitation id="1303C0B1C540496AFE857552FFCDD240" box="[326,437,1600,1622]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="3" pageNumber="364">Fig. 1A, B</figureCitation>
|
||
), anterior margin with 25 radii. Postcollum paraterga each with four lateral lobulations, anteriormost of which especially small (
|
||
<figureCitation id="1303C0B1C540496AFDFF756BFC03D29A" box="[572,635,1657,1679]" captionStart="Fig" captionStartId="2.[135,170,2064,2086]" captionTargetBox="[310,1291,203,2049]" captionTargetId="figure-7@2.[310,1298,189,2049]" captionTargetPageId="2" captionText="Fig. 1. Circulocryptus kompantsevi, new species, holotype. Habitus, A, lateral view; B, dorsal view; C, anteroventral view; D, posteroventral view. Pictures taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.5358927" httpUri="https://zenodo.org/record/5358927/files/figure.png" pageId="3" pageNumber="364">Fig. 1</figureCitation>
|
||
). Gonopod with a fimbriate, basal, prefemoral process (
|
||
<emphasis id="B94C0026C540496AFD877584FC29D2BE" bold="true" box="[580,593,1686,1707]" pageId="3" pageNumber="364">p</emphasis>
|
||
) on lateral face, telopodite with branch
|
||
<emphasis id="B94C0026C540496AFEB875A1FFF0D2DD" bold="true" box="[379,392,1715,1736]" pageId="3" pageNumber="364">b</emphasis>
|
||
being somewhat shorter than branch
|
||
<emphasis id="B94C0026C540496AFF5A75C2FEC5D2F0" bold="true" box="[153,189,1744,1765]" pageId="3" pageNumber="364">sph</emphasis>
|
||
(
|
||
<figureCitation id="1303C0B1C540496AFF0975C2FF7FD2F3" box="[202,263,1744,1766]" captionStart="Fig" captionStartId="3.[113,148,929,951]" captionTargetBox="[114,1450,190,916]" captionTargetId="figure-8@3.[97,1464,184,919]" captionTargetPageId="3" captionText="Fig. 2. Circulocryptus kompantsevi, new species, holotype. A, Leg 9, lateral view; B, right gonopod, lateral view; C, right gonopod, mesal view. Scale bar = 0.1 mm (A) and 0.2 mm (B, C). Designations: a = accessory seminal chamber; b = basal branch; l = hyaline fringed lobe at base of solenomere; p = prefemoral process; sl = solenomere; sph = solenophorous branch." figureDoi="http://doi.org/10.5281/zenodo.5358929" httpUri="https://zenodo.org/record/5358929/files/figure.png" pageId="3" pageNumber="364">Fig. 2</figureCitation>
|
||
).
|
||
<collectingCountry id="F32F9CA4C540496AFED875C2FFF8D2F3" box="[283,384,1744,1766]" name="Cambodia" pageId="3" pageNumber="364">Cambodia</collectingCountry>
|
||
..............
|
||
<taxonomicName id="4C38A7B7C540496AFE1575C2FC14D2F0" ID-CoL="VG5Q" authority="Golovatch, 2018" authorityName="Golovatch" authorityYear="2018" box="[470,620,1744,1765]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
|
||
<emphasis id="B94C0026C540496AFE1575C2FC14D2F0" box="[470,620,1744,1765]" italics="true" pageId="3" pageNumber="364">C. kompantsevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27FBD5DC540496AFDB975C2FC96D2F3" box="[634,750,1744,1766]" pageId="3" pageNumber="364" rank="species">new species</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B87DC34C540496AFFB275FFFC97D32E" blockId="3.[113,755,1542,2083]" pageId="3" pageNumber="364">
|
||
<keyLead id="30C97F01C540496AFFB275FFFC97D32E" pageId="3" pageNumber="364">
|
||
– General colouration sometimes dark, red-brown to blackish. Collum not swollen in anterior half. Gonopodal telopodite with branch
|
||
<emphasis id="B94C0026C540496AFF237434FE95D32E" bold="true" box="[224,237,1830,1851]" pageId="3" pageNumber="364">b</emphasis>
|
||
being somewhat longer than branch
|
||
<emphasis id="B94C0026C540496AFD9E7434FC07D32E" bold="true" box="[605,639,1830,1851]" pageId="3" pageNumber="364">sph</emphasis>
|
||
................2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30CCC491C540496AFFB27451FC96DC37" pageId="3" pageNumber="364">
|
||
<paragraph id="8B87DC34C540496AFFB27451FC97D3BA" blockId="3.[113,755,1542,2083]" pageId="3" pageNumber="364">
|
||
<keyLead id="30C97F01C540496AFFB27451FC97D3BA" pageId="3" pageNumber="364">
|
||
2. Collum with 29 radii at anterior margin, postcollum metaterga each with 5–6 transverse rows of setigerous tubercles and five lateral lobulations. Tip of branch
|
||
<emphasis id="B94C0026C540496AFE2F746FFF81D387" bold="true" box="[492,505,1917,1938]" pageId="3" pageNumber="364">b</emphasis>
|
||
of gonopodal telopodite retrorse.
|
||
<collectingRegion id="49FC12D6C540496AFF2D7488FF63D3BA" box="[238,283,1946,1967]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="364">Java</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32F9CA4C540496AFEE4748BFFFCD3BA" box="[295,388,1945,1967]" name="Indonesia" pageId="3" pageNumber="364">Indonesia</collectingCountry>
|
||
.......................................
|
||
<taxonomicName id="4C38A7B7C540496AFDB67488FC97D3BA" ID-CoL="VG5P" authorityName="Golovatch" authorityYear="2018" box="[629,751,1946,1967]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="javanicus">
|
||
<emphasis id="B94C0026C540496AFDB67488FC97D3BA" box="[629,751,1946,1967]" italics="true" pageId="3" pageNumber="364">C. javanicus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B87DC34C540496AFFB274A5FC96DC37" blockId="3.[113,755,1542,2083]" pageId="3" pageNumber="364">
|
||
<keyLead id="30C97F01C540496AFFB274A5FC96DC37" pageId="3" pageNumber="364">
|
||
– Collum with 25 radii at anterior margin, postcollum metaterga each with four transverse rows of setigerous tubercles and four lateral lobulations. Tip of branch
|
||
<emphasis id="B94C0026C540496AFE2F74E2FF81DC10" bold="true" box="[492,505,2032,2053]" pageId="3" pageNumber="364">b</emphasis>
|
||
of gonopodal telopodite curved forward.
|
||
<collectingCountry id="F32F9CA4C540496AFEF97B1FFFE8DC36" box="[314,400,2061,2083]" name="Vietnam" pageId="3" pageNumber="364">Vietnam</collectingCountry>
|
||
. ............................................
|
||
<taxonomicName id="4C38A7B7C540496AFD637B1FFC96DC37" authorityName="Golovatch" authorityYear="2016" box="[672,750,2061,2082]" class="Diplopoda" family="Cryptodesmidae" genus="Circulocryptus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="fallei">
|
||
<emphasis id="B94C0026C540496AFD637B1FFC96DC37" box="[672,750,2061,2082]" italics="true" pageId="3" pageNumber="364">C. fallei</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |