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<document id="9BF9365EA0BBB3355F133FAEDE6E1788" ID-CLB-Dataset="68941" ID-DOI="10.5281/zenodo.5184182" ID-GBIF-Dataset="dcb37aba-9f9f-402a-91db-c1c6e310d61c" ID-ISSN="0253-116X" ID-Zenodo-Dep="5184182" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" checkinTime="1628735848729" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2015" docId="03915A153A1CFF87FF0AEC43FE4AFCAF" docLanguage="en" docName="LinzerbiolBeitr.47.2.1133-1178.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Lithocharis parvincisa Assing 2015, nov.sp." docType="treatment" docVersion="10" lastPageNumber="1163" masterDocId="FFA8226D3A00FF99FFB8EF41FFB6FFD8" masterDocTitle="A revison of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina)" masterLastPageNumber="1178" masterPageNumber="1133" pageNumber="1161" updateTime="1699050958296" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="56E93868260E369DE7DF6F118E6D7633" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0B78176A0F0BC16CC1181516B9213274">
<mods:title id="87FDA5558EC2D696993E3D70C97D1542">A revison of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina)</mods:title>
</mods:titleInfo>
<mods:name id="144EA97F4ED517459EC89C687175290E" type="personal">
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</mods:role>
<mods:namePart id="2DA4F2190BDEB2900A0E2286DF58FD1F">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="E79460FA3F056271FE8D406336986C71">text</mods:typeOfResource>
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<mods:title id="34C11CE7AB1E3DEAE8004E8A5D176B29">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="9D03AA893DD9F151B34DDD8B07B266A0">
<mods:date id="D25FAD6860D8510624601CB025DEDE7E">2015</mods:date>
<mods:detail id="B18D281DD29DD3480C67B9379A46E4E2" type="pubDate">
<mods:number id="C064D583293D930534FB71DAC04DF4C2">2015-12-30</mods:number>
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<mods:number id="1D6FD961EE4D17CD85464F5A5DEE18F0">47</mods:number>
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<mods:classification id="B66509B42F5768C7650BC30B678AA6E3">journal article</mods:classification>
<mods:identifier id="8C6EDC60E83EF7B12CAB50ADA1C3EFEB" type="CLB-Dataset">68941</mods:identifier>
<mods:identifier id="8F0BAE0F535ED76C0579CC02AEFC731D" type="DOI">10.5281/zenodo.5184182</mods:identifier>
<mods:identifier id="EF023AEB0D18D6BD6796D479ED034766" type="GBIF-Dataset">dcb37aba-9f9f-402a-91db-c1c6e310d61c</mods:identifier>
<mods:identifier id="B0ECE1127070992B8650ED00ED0F7488" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="3CA3A74A55A25BC58DB39DBE326028E2" type="Zenodo-Dep">5184182</mods:identifier>
<mods:identifier id="F251E448CFDEA2F1639A5B7F31A50983" type="ZooBank">7F05EEC9-77C2-4592-A593-EE0139FE94FE</mods:identifier>
</mods:mods>
<treatment id="03915A153A1CFF87FF0AEC43FE4AFCAF" ID-DOI="http://doi.org/10.5281/zenodo.6631528" ID-GBIF-Taxon="195832409" ID-Zenodo-Dep="6631528" LSID="urn:lsid:plazi:treatment:03915A153A1CFF87FF0AEC43FE4AFCAF" httpUri="http://treatment.plazi.org/id/03915A153A1CFF87FF0AEC43FE4AFCAF" lastPageId="30" lastPageNumber="1163" pageId="28" pageNumber="1161">
<subSubSection id="C322B8883A1CFF85FF0AEC43FDC3FCC0" box="[178,629,770,792]" pageId="28" pageNumber="1161" type="nomenclature">
<paragraph id="8B87EB033A1CFF85FF0AEC43FDC3FCC0" blockId="28.[178,629,770,792]" box="[178,629,770,792]" pageId="28" pageNumber="1161">
<heading id="D0CF5C6F3A1CFF85FF0AEC43FDC3FCC0" box="[178,629,770,792]" fontSize="8" level="2" pageId="28" pageNumber="1161" reason="2">
<emphasis id="B94C37113A1CFF85FF0AEC43FDC3FCC0" bold="true" box="[178,629,770,792]" pageId="28" pageNumber="1161">
<taxonomicName id="4C3890803A1CFF85FF0AEC43FE2BFCCF" ID-CoL="3VKD3" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[178,413,770,792]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1161" phylum="Arthropoda" rank="species" species="parvincisa" status="sp. nov.">
<emphasis id="B94C37113A1CFF85FF0AEC43FE2BFCCF" bold="true" box="[178,413,770,792]" italics="true" pageId="28" pageNumber="1161">Lithocharis parvincisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F8A6A3A1CFF85FE1CEC42FE46FCC0" box="[420,496,771,792]" pageId="28" pageNumber="1161" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1303F7863A1CFF85FE45EC43FDD9FCC0" box="[509,623,770,792]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="28" pageNumber="1161">Figs 77-84</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A1CFF85FF0AEC69FBB2FBEF" pageId="28" pageNumber="1161" type="materials_examined">
<paragraph id="8B87EB033A1CFF85FF0AEC69FBB2FBEF" blockId="28.[178,1086,807,1079]" pageId="28" pageNumber="1161">
Type material:
<materialsCitation id="3B50E15E3A1CFF85FE1DEC66FEBEFC5A" ID-GBIF-Occurrence="3805307489" collectingDate="1985-02" collectingMethod="human dung baited trap" collectionCode="BMNH" collectorName="I. Hanski" country="Indonesia" determinerName="V. Assing" elevation="920" location="Dumoga-Bone N. P." pageId="28" pageNumber="1161" specimenCount="1" specimenCount-male="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<typeStatus id="548355A13A1CFF85FE1DEC66FE4FFCE5" box="[421,505,807,829]" pageId="28" pageNumber="1161" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3E208A3A1CFF85FE47EC66FDA4FCE4" box="[511,530,807,828]" pageId="28" pageNumber="1161" type="male">♂</specimenCount>
: &quot;
<collectingCountry id="F32FAB933A1CFF85FD9EEC66FD11FCE4" box="[550,679,807,828]" name="Indonesia" pageId="28" pageNumber="1161">INDONESIA</collectingCountry>
:
<collectingRegion id="49FC25E13A1CFF85FD08EC66FCCCFCE4" box="[688,890,807,828]" country="Indonesia" name="Sulawesi Utara" pageId="28" pageNumber="1161">SULAWESI UTARA</collectingRegion>
,
<location id="8EE7BDD83A1CFF85FC3BEC69FB8FFCE5" LSID="urn:lsid:plazi:treatment:03915A153A1CFF87FF0AEC43FE4AFCAF:8EE7BDD83A1CFF85FC3BEC69FB8FFCE5" box="[899,1081,808,829]" country="Indonesia" name="Dumoga-Bone N. P." pageId="28" pageNumber="1161" stateProvince="SULAWESI UTARA">Dumoga-Bone N.P.</location>
,
<date id="FF86CDC33A1CFF85FF0AEC7FFE8DFC8C" box="[178,315,830,852]" pageId="28" pageNumber="1161" value="1985-02">
<collectingDate id="EFC2342B3A1CFF85FF0AEC7FFE8DFC8C" box="[178,315,830,852]" pageId="28" pageNumber="1161" value="1985-02">February 1985</collectingDate>
</date>
/
<collectingMethod id="527993143A1CFF85FEEFEC7FFD89FC8B" box="[343,575,830,852]" pageId="28" pageNumber="1161">human dung baited trap</collectingMethod>
,
<collectorName id="26CD8ED53A1CFF85FDF4EC7FFD12FC8C" box="[588,676,830,852]" pageId="28" pageNumber="1161">I. Hanski</collectorName>
<quantity id="4CC046E63A1CFF85FD16EC7FFD5BFC8B" box="[686,749,830,851]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="28" pageNumber="1161" unit="m" value="920.0">
<elevation id="00150C303A1CFF85FD16EC7FFD5BFC8B" box="[686,749,830,851]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="28" pageNumber="1161" unit="m" value="920.0">920 m</elevation>
</quantity>
/ 26.29 / R.Ent.Soc.Lond., Project Wallace, B.M. 1985-10 / Holotypus ♂
<originalDetermination id="A82653FF3A1CFF85FDFAEC14FCEBFCB2" box="[578,861,853,874]" pageId="28" pageNumber="1161">
<taxonomicName id="4C3890803A1CFF85FDFAEC14FCA1FCB2" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[578,791,853,874]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1161" phylum="Arthropoda" rank="species" species="parvincisa" status="sp. nov.">
<emphasis id="B94C37113A1CFF85FDFAEC14FCA1FCB2" box="[578,791,853,874]" italics="true" pageId="28" pageNumber="1161">Lithocharis parvincisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F8A6A3A1CFF85FC9CEC14FCEBFCB2" box="[804,861,853,874]" pageId="28" pageNumber="1161" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
det.
<determinerName id="7E5B07C73A1CFF85FC2FEC14FC4FFCB3" box="[919,1017,853,875]" pageId="28" pageNumber="1161">
V.
<bibRefCitation id="EFA996F23A1CFF85FC02EC14FC4FFCB3" author="ASSING V" box="[954,1017,853,875]" pageId="28" pageNumber="1161" pagination="191 - 206" refId="ref27538" refString="ASSING V. (2015): Two new genera of Medonina from China and Japan (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 47 (1): 191 - 206." type="journal article" year="2015">Assing</bibRefCitation>
</determinerName>
2015&quot; (
<collectionCode id="ED2973C63A1CFF85FF00EC2DFEB7FC5A" box="[184,257,876,898]" pageId="28" pageNumber="1161">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B50E15E3A1CFF85FEA9EC2DFD70FC5A" ID-GBIF-Occurrence="3805307330" box="[273,710,875,902]" collectingDate="1985-02" collectionCode="BMNH" country="Indonesia" elevation="920" location="SULAWESI UTARA" pageId="28" pageNumber="1161" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="548355A13A1CFF85FEA9EC2DFEDFFC59" box="[273,361,876,897]" pageId="28" pageNumber="1161" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3E208A3A1CFF85FECEEC2DFE23FC5E" box="[374,405,876,902]" pageId="28" pageNumber="1161" type="female">1♀</specimenCount>
: same data as holotype (
<collectionCode id="ED2973C63A1CFF85FDCFEC2DFD76FC5A" box="[631,704,876,898]" pageId="28" pageNumber="1161">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B50E15E3A1CFF85FD68EC2DFCD4FC77" ID-GBIF-Occurrence="3805307323" collectingDate="1985-04" collectionCode="BMNH" country="Indonesia" elevation="1200" location="SULAWESI UTARA" pageId="28" pageNumber="1161" specimenCount="201" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
2♂♂,
<specimenCount id="9D3E208A3A1CFF85FCB5EC2DFC9AFC5E" box="[781,812,876,902]" pageId="28" pageNumber="1161" type="female">1♀</specimenCount>
: &quot;
<collectingCountry id="F32FAB933A1CFF85FC87EC2DFC76FC59" box="[831,960,876,897]" name="Indonesia" pageId="28" pageNumber="1161">INDONESIA</collectingCountry>
:
<collectingRegion id="49FC25E13A1CFF85FC72EC2DFEB5FC4F" country="Indonesia" name="Sulawesi Utara" pageId="28" pageNumber="1161">SULAWESI UTARA</collectingRegion>
, Dumoga-Bone N.P.,
<date id="FF86CDC33A1CFF85FE60ECC3FD89FC40" box="[472,575,898,920]" pageId="28" pageNumber="1161" value="1985-04">
<collectingDate id="EFC2342B3A1CFF85FE60ECC3FD89FC40" box="[472,575,898,920]" pageId="28" pageNumber="1161" value="1985-04">April 1985</collectingDate>
</date>
/
<collectingMethod id="527993143A1CFF85FDE2ECC3FC9DFC40" box="[602,811,898,920]" pageId="28" pageNumber="1161">Rothamsted light trap</collectingMethod>
, site
<quantity id="4CC046E63A1CFF85FCD9ECC3FC0AFC40" box="[865,956,898,920]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="1161" unit="m" value="1200.0">
<elevation id="00150C303A1CFF85FCD9ECC3FC0AFC40" box="[865,956,898,920]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="1161" unit="m" value="1200.0">
1,
<specimenCount id="9D3E208A3A1CFF85FCC4ECC3FC0AFC40" box="[892,956,898,920]" pageId="28" pageNumber="1161" type="generic">200 m</specimenCount>
</elevation>
</quantity>
. H. Barlow / R.Ent.Soc.Lond., Project Wallace, B.M. 1985-10&quot; (
<collectionCode id="ED2973C63A1CFF85FD75ECD8FCA3FC76" box="[717,789,921,942]" pageId="28" pageNumber="1161">BMNH</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B50E15E3A1CFF85FCC1ECD8FCEFFC04" ID-GBIF-Occurrence="3805307389" collectingDate="1985-04" collectionCode="BMNH" country="Indonesia" elevation="400" isTailEnumeration="true" location="Lowland forest" pageId="28" pageNumber="1161" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D3E208A3A1CFF85FCC1ECD8FC2EFC6B" box="[889,920,921,947]" pageId="28" pageNumber="1161" type="female">1♀</specimenCount>
: &quot;
<collectingCountry id="F32FAB933A1CFF85FC01ECD8FB8CFC76" box="[953,1082,921,942]" name="Indonesia" pageId="28" pageNumber="1161">INDONESIA</collectingCountry>
:
<collectingRegion id="49FC25E13A1CFF85FF0AECF1FECBFC1D" box="[178,381,944,965]" country="Indonesia" name="Sulawesi Utara" pageId="28" pageNumber="1161">SULAWESI UTARA</collectingRegion>
, Dumoga-Bone N.P.,
<date id="FF86CDC33A1CFF85FDF2ECF1FD18FC1D" box="[586,686,943,966]" pageId="28" pageNumber="1161" value="1985-04">
<collectingDate id="EFC2342B3A1CFF85FDF2ECF1FD18FC1D" box="[586,686,943,966]" pageId="28" pageNumber="1161" value="1985-04">April 1985</collectingDate>
</date>
/
<collectingMethod id="527993143A1CFF85FD7CECF1FC9AFC1D" box="[708,812,944,966]" pageId="28" pageNumber="1161">Pit fall trap</collectingMethod>
/ Plot C, ca
<quantity id="4CC046E63A1CFF85FC1CECF1FC54FC1D" box="[932,994,944,965]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="1161" unit="m" value="400.0">
<elevation id="00150C303A1CFF85FC1CECF1FC54FC1D" box="[932,994,944,965]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="1161" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<location id="8EE7BDD83A1CFF85FC54ECF1FF50FC04" LSID="urn:lsid:plazi:treatment:03915A153A1CFF87FF0AEC43FE4AFCAF:8EE7BDD83A1CFF85FC54ECF1FF50FC04" country="Indonesia" name="Lowland forest" pageId="28" pageNumber="1161" stateProvince="Sulawesi Utara">Lowland forest</location>
/ R.Ent.Soc.Lond., Project Wallace, B.M. 1985-10&quot; (
<collectionCode id="ED2973C63A1CFF85FCB2EC87FCE5FC04" box="[778,851,966,988]" pageId="28" pageNumber="1161">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B50E15E3A1CFF85FCD4EC87FBBFFBD2" ID-GBIF-Occurrence="3805307379" collectionCode="BMNH" country="Indonesia" elevation="664" location="Lowland forest" municipality="Edwards' Camp" pageId="28" pageNumber="1161" specimenCount="3" specimenCount-female="3" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D3E208A3A1CFF85FCD4EC87FC2BFC39" box="[876,925,966,993]" pageId="28" pageNumber="1161" type="female">3♀♀</specimenCount>
: &quot;
<collectingCountry id="F32FAB933A1CFF85FC01EC87FB8CFC03" box="[953,1082,966,987]" name="Indonesia" pageId="28" pageNumber="1161">INDONESIA</collectingCountry>
:
<collectingRegion id="49FC25E13A1CFF85FF0AEC9CFE34FC2A" box="[178,386,989,1010]" country="Indonesia" name="Sulawesi Utara" pageId="28" pageNumber="1161">SULAWESI UTARA</collectingRegion>
, Dumoga-Bone N.P., 1985. / '
<collectingMunicipality id="6BE371793A1CFF85FD16EC9CFCFEFC2A" box="[686,840,989,1011]" pageId="28" pageNumber="1161">Edwards' Camp</collectingMunicipality>
,
<location id="8EE7BDD83A1CFF85FCEFEC9CFC5FFC2B" LSID="urn:lsid:plazi:treatment:03915A153A1CFF87FF0AEC43FE4AFCAF:8EE7BDD83A1CFF85FCEFEC9CFC5FFC2B" box="[855,1001,989,1011]" country="Indonesia" municipality="Edwards' Camp" name="Lowland forest" pageId="28" pageNumber="1161" stateProvince="Sulawesi Utara">Lowland forest</location>
,
<quantity id="4CC046E63A1CFF85FC40EC9CFB8CFC2A" box="[1016,1082,989,1011]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="28" pageNumber="1161" unit="m" value="664.0">
<elevation id="00150C303A1CFF85FC40EC9CFB8CFC2A" box="[1016,1082,989,1011]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="28" pageNumber="1161" unit="m" value="664.0">664 m</elevation>
</quantity>
, 26.iv.-4.vi. /
<collectingMethod id="527993143A1CFF85FE8BECB5FE1CFBD1" box="[307,426,1012,1034]" pageId="28" pageNumber="1161">Malaise trap</collectingMethod>
/ R.Ent.Soc.Lond., Project Wallace, B.M. 1985-10&quot; (
<collectionCode id="ED2973C63A1CFF85FC02ECB5FBB5FBD2" box="[954,1027,1012,1034]" pageId="28" pageNumber="1161">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B50E15E3A1CFF85FBA0ECB5FBB6FBEF" ID-GBIF-Occurrence="3805307382" collectingDate="1985-05" country="Indonesia" elevation="1140" location="INDONESIA" pageId="28" pageNumber="1161" specimenCount="1" typeStatus="paratype">
1♂: &quot;
<collectingCountry id="F32FAB933A1CFF85FF01EB4AFE8CFBF8" box="[185,314,1035,1056]" name="Indonesia" pageId="28" pageNumber="1161">INDONESIA</collectingCountry>
: SULAWESE UTARA, Dumoga-Bone N.P.,
<date id="FF86CDC33A1CFF85FD56EB4AFCE4FBF8" box="[750,850,1034,1056]" pageId="28" pageNumber="1161" value="1985-05">
<collectingDate id="EFC2342B3A1CFF85FD56EB4AFCE4FBF8" box="[750,850,1034,1056]" pageId="28" pageNumber="1161" value="1985-05">May 1985</collectingDate>
</date>
. / 'Clarke' Camp, Lower montane forest,
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<elevation id="00150C303A1CFF85FEFDEB60FE3CFBEF" box="[325,394,1057,1079]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="28" pageNumber="1161" unit="m" value="1140.0">1140 m</elevation>
</quantity>
/ At light / R.Ent.Soc.Lond., Project Wallace, B.M. 1985-10&quot; (cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A1CFF85FF0AEB04FEDBFB4D" pageId="28" pageNumber="1161" type="etymology">
<paragraph id="8B87EB033A1CFF85FF0AEB04FEDBFB4D" blockId="28.[178,1085,1093,1174]" pageId="28" pageNumber="1161">Etymology: The specific epithet (adjective) is composed of the adjectives parvus (small) and incisus (incised). It alludes to the relatively small posterior incision of the male sternite VIII.</paragraph>
</subSubSection>
<subSubSection id="C322B8883A1CFF87FF0AEBE4FEF9FDFD" lastPageId="30" lastPageNumber="1163" pageId="28" pageNumber="1161" type="description">
<paragraph id="8B87EB033A1CFF85FF0AEBE4FDE7FACB" blockId="28.[178,1085,1189,1299]" pageId="28" pageNumber="1161">
Description: Body length 4.0-
<quantity id="4CC046E63A1CFF85FD87EBE4FD24FB62" box="[575,658,1189,1210]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="28" pageNumber="1161" unit="mm" value="4.7">4.7 mm</quantity>
; length of forebody
<quantity id="4CC046E63A1CFF85FCD1EBE4FC52FB62" box="[873,996,1189,1211]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="28" pageNumber="1161" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3-2.5 mm</quantity>
. Coloration: head dark-brown to black; pronotum brown to dark-brown; elytra brown; abdomen brown to dark-brown, with the posterior margins of the segments yellowish to yellowishbrown; legs yellowish; antennae reddish.
</paragraph>
<paragraph id="8B87EB033A1CFF85FF0AEA63FED2FA75" blockId="28.[178,1085,1314,1453]" pageId="28" pageNumber="1161">
Head (
<figureCitation id="1303F7863A1CFF85FF43EA63FEFCFAE0" box="[251,330,1314,1336]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="28" pageNumber="1161">Fig. 77</figureCitation>
) weakly transverse, 1.06-1.09 times as broad as long, lateral margins behind eyes subparallel in dorsal view; punctation dense and fine; interstices with shallow microreticulation, glossy (
<figureCitation id="1303F7863A1CFF85FE4EEA1CFDF1FAAB" box="[502,583,1373,1395]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="28" pageNumber="1161">Fig. 78</figureCitation>
). Eyes large, noticeably longer than postocular region in dorsal view. Antennae approximately
<quantity id="4CC046E63A1CFF85FD17EA3BFCB6FA57" box="[687,768,1402,1423]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="28" pageNumber="1161" unit="mm" value="1.7">1.7 mm</quantity>
long; preapical antennomeres distinctly oblong.
</paragraph>
<paragraph id="8B87EB033A1CFF87FF0AEAFCFBB0FEF3" blockId="28.[178,1085,1468,1578]" lastBlockId="30.[178,1086,219,299]" lastPageId="30" lastPageNumber="1163" pageId="28" pageNumber="1161">
Pronotum (
<figureCitation id="1303F7863A1CFF85FE9CEAFDFED8FA0A" box="[292,366,1468,1490]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="28" pageNumber="1161">Fig. 77</figureCitation>
) approximately as long as broad and as broad as head; anterior angles sharply, posterior angles weakly marked; lateral margins weakly converging posteriad in dorsal view; midline with pronounced impunctate band; punctation dense and fine, slightly more distinct than that of head; interstices without microsculpture (
<figureCitation id="1303F7863A1CFF85FC14E955FC4FF9F2" box="[940,1017,1556,1578]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="28" pageNumber="1161">Fig. 79</figureCitation>
). Elytra (
<figureCitation id="1303F7863A1EFF87FF47EF9AFEFFFF28" box="[255,329,219,241]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="30" pageNumber="1163">Fig. 77</figureCitation>
) approximately 1.1 times as long as pronotum; punctation very dense and fine; interstices without microsculpture. Hind wings present. Protarsomeres I-IV weakly dilated. Metatarsomere I approximately as long as the combined length of II and III.
</paragraph>
<caption id="DF47BB8B3A1DFF84FF0AEA9DFC31F9EE" ID-DOI="http://doi.org/10.5281/zenodo.5184208" ID-Zenodo-Dep="5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="29" pageNumber="1162" startId="29.[178,218,1500,1522]" targetBox="[182,1091,212,1480]" targetPageId="29">
<paragraph id="8B87EB033A1DFF84FF0AEA9DFC31F9EE" blockId="29.[178,1085,1500,1590]" pageId="29" pageNumber="1162">
<emphasis id="B94C37113A1DFF84FF0AEA9DFEA0FA29" bold="true" box="[178,278,1500,1522]" pageId="29" pageNumber="1162">Figs 76-84</emphasis>
:
<taxonomicName id="4C3890803A1DFF84FE9BEA9DFDD9FA29" authority="CAMERON" authorityName="Cameron" authorityYear="1936" box="[291,623,1500,1522]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="1162" phylum="Arthropoda" rank="species" species="preangerana">
<emphasis id="B94C37113A1DFF84FE9BEA9DFDB1FA2A" box="[291,519,1500,1522]" italics="true" pageId="29" pageNumber="1162">Lithocharis preangerana</emphasis>
CAMERON
</taxonomicName>
, holotype (
<emphasis id="B94C37113A1DFF84FD60EA9DFD59FA29" bold="true" box="[728,751,1500,1521]" pageId="29" pageNumber="1162">76</emphasis>
), and
<taxonomicName id="4C3890803A1DFF84FC93EA9CFC12FA2A" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[811,932,1501,1522]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="1162" phylum="Arthropoda" rank="species" species="parvincisa" status="sp. nov.">
<emphasis id="B94C37113A1DFF84FC93EA9CFC12FA2A" box="[811,932,1501,1522]" italics="true" pageId="29" pageNumber="1162">L. parvincisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F8A6A3A1DFF84FC12EA9CFC5BFA2A" box="[938,1005,1501,1522]" pageId="29" pageNumber="1162" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B94C37113A1DFF84FC42EA9DFB99FA29" bold="true" box="[1018,1071,1500,1521]" pageId="29" pageNumber="1162">77-84</emphasis>
): (
<emphasis id="B94C37113A1DFF84FF02EAB2FF59F9D0" bold="true" box="[186,239,1523,1544]" pageId="29" pageNumber="1162">76-77</emphasis>
) forebody; (
<emphasis id="B94C37113A1DFF84FEDBEAB2FECCF9D0" bold="true" box="[355,378,1523,1544]" pageId="29" pageNumber="1162">78</emphasis>
) median dorsal portion of head; (
<emphasis id="B94C37113A1DFF84FD08EAB2FD71F9D0" bold="true" box="[688,711,1523,1544]" pageId="29" pageNumber="1162">79</emphasis>
) median portion of pronotum; (
<emphasis id="B94C37113A1DFF84FC54EAB2FBB5F9D0" bold="true" box="[1004,1027,1523,1544]" pageId="29" pageNumber="1162">80</emphasis>
) male sternite VII; (
<emphasis id="B94C37113A1DFF84FE8BE948FEFCF9C6" bold="true" box="[307,330,1545,1566]" pageId="29" pageNumber="1162">81</emphasis>
) male sternite VIII; (
<emphasis id="B94C37113A1DFF84FDAFE948FDFAF9C6" bold="true" box="[535,588,1545,1566]" pageId="29" pageNumber="1162">82-83</emphasis>
) aedeagus in lateral and in ventral view; (
<emphasis id="B94C37113A1DFF84FC58E948FC41F9C6" bold="true" box="[992,1015,1545,1566]" pageId="29" pageNumber="1162">84</emphasis>
) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B87EB033A1EFF87FF0AEE7AFCCEFEB6" blockId="30.[178,1085,315,366]" pageId="30" pageNumber="1163">Abdomen narrower than elytra; punctation fine and very dense; interstices with shallow microreticulation; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="8B87EB033A1EFF87FF0AEE3CFEF9FDFD" blockId="30.[178,1086,381,549]" pageId="30" pageNumber="1163">
♂: sternite VII (
<figureCitation id="1303F7863A1EFF87FED8EE3CFE1BFE4B" box="[352,429,381,403]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="30" pageNumber="1163">Fig. 80</figureCitation>
) moderately transverse, posterior margin with shallow median excision, this excision with a comb of 20-25 moderately long palisade setae; sternite VIII (
<figureCitation id="1303F7863A1EFF87FF02EEF9FEB1FE16" box="[186,263,440,462]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="30" pageNumber="1163">Fig. 81</figureCitation>
) weakly transverse and with unmodified pubescence, posterior incision small and broadly V-shaped; aedeagus (
<figureCitation id="1303F7863A1EFF87FD9CEE94FD2CFE33" box="[548,666,469,491]" captionStart="Figs 76-84" captionStartId="29.[178,218,1500,1522]" captionTargetBox="[182,1091,212,1480]" captionTargetId="figure-1@29.[178,1091,212,1480]" captionTargetPageId="29" captionText="Figs 76-84: Lithocharis preangerana CAMERON, holotype (76), and L. parvincisa nov.sp. (77-84): (76-77) forebody; (78) median dorsal portion of head; (79) median portion of pronotum; (80) male sternite VII; (81) male sternite VIII; (82-83) aedeagus in lateral and in ventral view; (84) apical portion of aedeagus in ventral view. Scale bars: 76-77: 1.0 mm; 78-84: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5184208" httpUri="https://zenodo.org/record/5184208/files/figure.png" pageId="30" pageNumber="1163">Figs 82-84</figureCitation>
) approximately
<quantity id="4CC046E63A1EFF87FCF7EE94FC14FE32" box="[847,930,469,490]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="1163" unit="mm" value="0.8">0.8 mm</quantity>
long, without dorso-lateral apophyses; ventral process apically deeply bifid; internal structures of distinctive shapes.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A1EFF87FF0AED74FCE3FD22" pageId="30" pageNumber="1163" type="discussion">
<paragraph id="8B87EB033A1EFF87FF0AED74FCE3FD22" blockId="30.[178,1086,565,762]" pageId="30" pageNumber="1163">
Comparative notes: Regarding its external and male sexual characters,
<taxonomicName id="4C3890803A1EFF87FB90ED74FEABFDB0" authorityName="Assing" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1163" phylum="Arthropoda" rank="species" species="parvincisa">
<emphasis id="B94C37113A1EFF87FB90ED74FEABFDB0" italics="true" pageId="30" pageNumber="1163">L. parvincisa</emphasis>
</taxonomicName>
is similar to the sympatric
<taxonomicName id="4C3890803A1EFF87FD8CED12FDCFFDBF" authorityName="Kraatz" authorityYear="1859" box="[564,633,594,616]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1163" phylum="Arthropoda" rank="species" species="vilis">
<emphasis id="B94C37113A1EFF87FD8CED12FDCFFDBF" box="[564,633,594,616]" italics="true" pageId="30" pageNumber="1163">L. vilis</emphasis>
</taxonomicName>
, from which it is distinguished by the shape of the pronotum, the more slender antennae, the less distinctly dilated protarsomeres I-IV, the shorter comb of palisade setae at the posterior margin of the male sternite VII, the less transverse male sternite VIII with a much smaller posterior excision and with unmodified pubescence, and by the smaller aedeagus without dorso-lateral apophyses and with smaller sclerotized internal structures of different shapes.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A1EFF87FF0AEC4BFE4AFCAF" pageId="30" pageNumber="1163" type="distribution">
<paragraph id="8B87EB033A1EFF87FF0AEC4BFE4AFCAF" blockId="30.[178,1085,778,888]" pageId="30" pageNumber="1163">
Distribution and natural history: The
<typeStatus id="548355A13A1EFF87FCEEEC4BFCCCFCC7" box="[854,890,778,799]" pageId="30" pageNumber="1163">type</typeStatus>
specimens were collected in Dumoga-Bone National Park in
<collectingRegion id="49FC25E13A1EFF87FD09EC66FCE3FCE4" box="[689,853,807,829]" country="Indonesia" name="Sulawesi Utara" pageId="30" pageNumber="1163">Sulawesi Utara</collectingRegion>
(
<collectingCountry id="F32FAB933A1EFF87FCD4EC66FC62FCE5" box="[876,980,807,829]" name="Indonesia" pageId="30" pageNumber="1163">Indonesia</collectingCountry>
: northern
<collectingRegion id="49FC25E13A1EFF87FF0AEC05FEB9FC82" box="[178,271,836,858]" country="Indonesia" name="Sulawesi Utara" pageId="30" pageNumber="1163">Sulawesi</collectingRegion>
) with traps baited with human faeces, with light traps, Malaise traps, and pitfall traps at altitudes of
<quantity id="4CC046E63A1EFF87FEC2EC23FE41FCAF" box="[378,503,866,887]" metricMagnitude="2" metricUnit="m" metricValue="6.7" metricValueMax="11.4" metricValueMin="2.0" pageId="30" pageNumber="1163" unit="m" value="670.0" valueMax="1140.0" valueMin="200.0">200-1140 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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