140 lines
18 KiB
XML
140 lines
18 KiB
XML
<document id="68CCF61D24519D8C7C489042FCAA421C" ID-DOI="10.5281/zenodo.184695" ID-GBIF-Dataset="d43a5b94-5394-443c-afc8-1baf086959b5" ID-ISSN="1175-5326" ID-Zenodo-Dep="184695" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459997035449" checkinUser="plazi" docAuthor="Amorim, Dalton De Souza & Oliveira, Sarah Siqueira" docDate="2008" docId="0390F003C6183C06FF4DFA50FE9DFB0F" docLanguage="en" docName="zt01920p028.pdf" docOrigin="Zootaxa 1920" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cluzobra elpidia Amorim & Oliveira, 2008, sp.n." docType="treatment" docVersion="10" lastPageNumber="24" masterDocId="FFA9887BC60E3C11FFDAFFF0FFB2FFC2" masterDocTitle="Eleven new species of the genus Cluzobra Edwards (Diptera, Mycetophilidae, Sciophilinae) from the Atlantic Forest of Brazil" masterLastPageNumber="28" masterPageNumber="1" pageNumber="23" updateTime="1698232936749" updateUser="plazi">
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<mods:mods id="52A2226AC54D18F4ED9BED364D29D0A7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9B3394EBF65A9158EB5D70592CBC5BAF">
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<mods:title id="B02E114AD6C3A328A06A78D29383B8ED">Eleven new species of the genus Cluzobra Edwards (Diptera, Mycetophilidae, Sciophilinae) from the Atlantic Forest of Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="B23696A6FA60739A09D1ECC73074706D" type="personal">
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<mods:roleTerm id="86E98BFA8F7B3E9241152AE48B3969BF">Author</mods:roleTerm>
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<mods:namePart id="4C927D93DA9116A47809292E3EAE4951">Amorim, Dalton De Souza</mods:namePart>
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<mods:name id="0C688570725930DF6BBCFD935F1E7B59" type="personal">
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<mods:roleTerm id="CDB3670C26940CB75E96CFD8D4360895">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1057AFFBC153ED3D271C2D0923A800B">Oliveira, Sarah Siqueira</mods:namePart>
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<mods:typeOfResource id="E56893229FF9C9F834E196EA4B2CC6E5">text</mods:typeOfResource>
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<mods:title id="1F766A557B6090D326DEEEEFA19EB1B7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9B9337C901E16D521FAFF47681EF89DD">
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<mods:date id="5A61DFCD9997B336889C22DC1FB72C38">2008</mods:date>
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<mods:detail id="A03E682619EEAEC867B8085FE5C33621" type="volume">
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<mods:number id="54F6C168E3E02B74502D70A60B8D2231">1920</mods:number>
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<mods:start id="031A22C6B679C2D19F45BEB51C20FA55">1</mods:start>
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<mods:end id="D0EB7211CA01B221FD260E0E8DB9E936">28</mods:end>
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<mods:classification id="B3D0D42A5504039A7E902D22633D551A">journal article</mods:classification>
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<mods:identifier id="359D2E1E79E9D8668AF375786CF7FABF" type="DOI">10.5281/zenodo.184695</mods:identifier>
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<mods:identifier id="6D3B7A6AA7828281ABE53B45900FD354" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C2A78EC027E9E9A18E4F34D706AEC757" type="Zenodo-Dep">184695</mods:identifier>
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<treatment id="0390F003C6183C06FF4DFA50FE9DFB0F" ID-DOI="http://doi.org/10.5281/zenodo.5685415" ID-GBIF-Taxon="119380411" ID-Zenodo-Dep="5685415" LSID="urn:lsid:plazi:treatment:0390F003C6183C06FF4DFA50FE9DFB0F" httpUri="http://treatment.plazi.org/id/0390F003C6183C06FF4DFA50FE9DFB0F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection id="C323129EC6183C07FF4DFA50FE87FA1F" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8B864115C6183C07FF4DFA50FE1AFA78" blockId="22.[151,424,1440,1501]" box="[151,424,1440,1466]" pageId="22" pageNumber="23">
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<heading id="D0CEF679C6183C07FF4DFA50FE1AFA78" bold="true" box="[151,424,1440,1466]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="B94D9D07C6183C07FF4DFA50FE1AFA78" bold="true" box="[151,424,1440,1466]" pageId="22" pageNumber="23">
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<taxonomicName id="4C393A96C6183C07FF4DFA50FEECFA78" ID-CoL="WDNM" box="[151,350,1440,1466]" class="Insecta" family="Mycetophilidae" genus="Cluzobra" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="elpidia" status="sp. nov.">
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<emphasis id="B94D9D07C6183C07FF4DFA50FEECFA78" bold="true" box="[151,350,1440,1466]" italics="true" pageId="22" pageNumber="23">Cluzobra elpidia</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27E207CC6183C07FEB7FA50FE1AFA78" box="[365,424,1440,1466]" pageId="22" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B864115C6183C07FF4DFA33FE87FA1F" blockId="22.[151,424,1440,1501]" box="[151,309,1475,1501]" pageId="22" pageNumber="23">
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(
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<figureCitation id="13025D90C6183C07FF45FA33FEB0FA1F" box="[159,258,1475,1501]" captionStart="FIGURES 11 – 19" captionStartId="6.[151,269,1898,1922]" captionTargetBox="[160,1420,189,1875]" captionTargetId="figure@6.[160,1425,182,1876]" captionTargetPageId="6" captionText="FIGURES 11 – 19. Wings of Cluzobra species. 11. Cluzobra vockerothi, sp. n. 12. Cluzobra sapiranga, sp. n. 13. Cluzobra fluminense, sp. n. 14. Cluzobra binocellaris (Edwards). 15. Cluzobra fuscipennis Edwards, specimen from Embu, State of São Paulo. 16. Cluzobra plaumanni Edwards, specimen from Nova Teutônia, State of Santa Catarina. 17. Cluzobra fritzmuelleri, sp. n. 18. Cluzobra spinata, sp. n. 19. Cluzobra elpidia, sp. n." httpUri="https://zenodo.org/record/184699/files/figure.png" pageId="22" pageNumber="23">Figs. 19</figureCitation>
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,
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<figureCitation id="13025D90C6183C07FED4FA33FE9EFA1F" box="[270,300,1475,1501]" captionStart="FIGURES 35 – 37" captionStartId="21.[151,269,1823,1847]" captionTargetBox="[185,1392,178,1814]" captionTargetId="figure@21.[185,1394,176,1815]" captionTargetPageId="21" captionText="FIGURES 35 – 37. Male terminalia of Cluzobra species. 35. Cluzobra spinata, sp. n., ventral view. 36. Same, dorsal view. 37. Cluzobra elpidia, sp. n., dorsal view. Abbreviations: ae, aedeagus; ce, cercus; dc, dorsal crypt of syngonocoxite; gp, gonocoxite projection ventrad to gonostyle insertion; gs, gonostyle; s 8, sternite 8; sg, syngonocoxite; t 8, tergite 8; t 9, tergite 9." httpUri="https://zenodo.org/record/184704/files/figure.png" pageId="22" pageNumber="23">37</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C323129EC6183C07FF4DF9E3FC5BF9BF" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="8B864115C6183C07FF4DF9E3FC5BF9BF" blockId="22.[151,1436,1555,2021]" pageId="22" pageNumber="23">
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<emphasis id="B94D9D07C6183C07FF4DF9E3FEA5F9EF" bold="true" box="[151,279,1555,1581]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
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Wing brownish, nearly devoid of maculae, Sc long reaching C at level of origin of Rs; syngonocoxite without mesal notch at distal margin, a conical projection ventrad to insertion of gonostyle, bearing a pair of strong, short spines at apex; gonostyle strongly sclerotized, short.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323129EC6183C07FF1FF97BFE0FF90F" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="8B864115C6183C07FF1FF97BFE0FF90F" blockId="22.[151,1436,1555,2021]" pageId="22" pageNumber="23">
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<emphasis id="B94D9D07C6183C07FF1FF97BFE09F967" bold="true" box="[197,443,1675,1701]" pageId="22" pageNumber="23">Material examined.</emphasis>
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<typeStatus id="5482FFB7C6183C07FE1EF97BFD83F967" box="[452,561,1675,1701]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
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ɗ,
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<collectingCountry id="F32E0185C6183C07FD84F97BFD79F967" box="[606,715,1675,1701]" name="Brazil" pageId="22" pageNumber="23">BRAZIL</collectingCountry>
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, State of Bahia, Ilhéus,
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<collectingCountry id="F32E0185C6183C07FC30F97BFB94F967" box="[1002,1062,1675,1701]" name="Malta" pageId="22" pageNumber="23">Mata</collectingCountry>
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Esperança,
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<geoCoordinate id="EE0D27D2C6183C07FB62F97BFAD8F967" box="[1208,1386,1675,1701]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="-39.066666">39° 03’ 60” W</geoCoordinate>
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<geoCoordinate id="EE0D27D2C6183C07FAA9F97BFEBBF90F" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="15" value="-14.783333">14° 46’ 60” S</geoCoordinate>
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,
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<date id="FF8767D5C6183C07FECCF943FE0AF90F" box="[278,440,1715,1741]" pageId="22" pageNumber="23" value="2002-05-15" valueMax="2002-05-18" valueMin="2002-05-15">15–18.v.2002</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323129EC6183C06FF1FF92BFEDEFCD7" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
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<paragraph id="8B864115C6183C06FF1FF92BFEDEFCD7" blockId="22.[151,1436,1555,2021]" lastBlockId="23.[151,1437,152,1229]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<emphasis id="B94D9D07C6183C07FF1FF92BFE49F937" bold="true" box="[197,507,1755,1781]" pageId="22" pageNumber="23">Description. Male. Head.</emphasis>
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Occiput light brown, darker at apex; mid ocellus absent, lateral ocelli separated from eye margins by a distance larger than ocellus diameter, ocellar area brown; front and clypeus yellowish, setose; maxillary palpus brown, 1+4 palpomeres, setose, palpomeres increasing in length to apex; labella yellowish, except for the brownish sclerite of basal article. Scape and pedicel rounded, yellowish, setose; flagellomeres brownish, setose.
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<emphasis id="B94D9D07C6183C07FDC3F88BFDCFF857" bold="true" box="[537,637,1915,1941]" pageId="22" pageNumber="23">Thorax.</emphasis>
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Pronotum light brown, with some longer and other smaller setae. Scutum brown, with four yellowish brown bands intertwined with brown bands, smaller setae scattered and some stronger acrostical and dorsocentral bristles; scutellum yellowish with a brown mid band, six stronger scutellar bristles and some smaller setae. Mediotergite yellowish brown, with a mesal and a pair of more lateral tufts of setae. Pleural membrane yellowish. Proepisternum light yellowish brown. Pleural sclerites light brown. Laterotergite setose; anepisternum and katepisternum bare. Halter yellowish, setose.
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<emphasis id="B94D9D07C6193C06FB16FF17FABCFEC3" bold="true" box="[1228,1294,231,257]" pageId="23" pageNumber="24">Legs.</emphasis>
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Coxae light brown, hind coxa lighter at base, femora brown, mid femur darker, tibiae and tarsi light brown; tibial spurs 1:2:2, yellowish, front spur more than twice front tibial width at apex.
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<emphasis id="B94D9D07C6193C06FC34FEC7FB80FE93" bold="true" box="[1006,1074,311,337]" pageId="23" pageNumber="24">Wing</emphasis>
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(
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<figureCitation id="13025D90C6193C06FB9EFEC8FB08FE90" box="[1092,1210,312,338]" captionStart="FIGURES 11 – 19" captionStartId="6.[151,269,1898,1922]" captionTargetBox="[160,1420,189,1875]" captionTargetId="figure@6.[160,1425,182,1876]" captionTargetPageId="6" captionText="FIGURES 11 – 19. Wings of Cluzobra species. 11. Cluzobra vockerothi, sp. n. 12. Cluzobra sapiranga, sp. n. 13. Cluzobra fluminense, sp. n. 14. Cluzobra binocellaris (Edwards). 15. Cluzobra fuscipennis Edwards, specimen from Embu, State of São Paulo. 16. Cluzobra plaumanni Edwards, specimen from Nova Teutônia, State of Santa Catarina. 17. Cluzobra fritzmuelleri, sp. n. 18. Cluzobra spinata, sp. n. 19. Cluzobra elpidia, sp. n." httpUri="https://zenodo.org/record/184699/files/figure.png" pageId="23" pageNumber="24">Figure 19</figureCitation>
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). Length, 2.6 mm, with no transverse maculae, membrane brown, slightly lighter at base of cell cuA. Sc complete, well sclerotized, short, reaching C at level of origin of Rs; R1 long, more than five times r–m length, reaching C at distal third of wing; r–m short, slightly longer than base of Rs; R5 long, reaching C close to wing apex; C extending two fifth the distance between R5 and M1; r–m oblique; M1+2 short, slightly longer than r–m; medial fork complete; M4 entirely absent; A1 very short, scarcely visible.
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<emphasis id="B94D9D07C6193C06FCF1FDF8FC1FFDE0" bold="true" box="[811,941,520,546]" pageId="23" pageNumber="24">Abdomen.</emphasis>
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Abdominal tergites and sternites 1–7 light brown. S8 U–shape, T8 wide and short.
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<emphasis id="B94D9D07C6193C06FDAFFDC3FD4CFD8F" bold="true" box="[629,766,563,589]" pageId="23" pageNumber="24">Terminalia</emphasis>
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(
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<figureCitation id="13025D90C6193C06FCD5FDC3FC31FD8F" box="[783,899,563,589]" captionStart="FIGURES 35 – 37" captionStartId="21.[151,269,1823,1847]" captionTargetBox="[185,1392,178,1814]" captionTargetId="figure@21.[185,1394,176,1815]" captionTargetPageId="21" captionText="FIGURES 35 – 37. Male terminalia of Cluzobra species. 35. Cluzobra spinata, sp. n., ventral view. 36. Same, dorsal view. 37. Cluzobra elpidia, sp. n., dorsal view. Abbreviations: ae, aedeagus; ce, cercus; dc, dorsal crypt of syngonocoxite; gp, gonocoxite projection ventrad to gonostyle insertion; gs, gonostyle; s 8, sternite 8; sg, syngonocoxite; t 8, tergite 8; t 9, tergite 9." httpUri="https://zenodo.org/record/184704/files/figure.png" pageId="23" pageNumber="24">Figure 37</figureCitation>
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). Terminalia yellowish. Syngonocoxites covering the entire ventral face of the terminalia, with a conical projection laterally just before the insertion of the gonostyle bearing two short and strong spines. Gonostyle with the shape of a short, wide spoon, relatively well sclerotized. Aedeagus weakly sclerotized, with apodemes laterally projected, expanded at apex. T9 wider than long, with no projections or spines laterally at distal margin. Cercus simple, large, but weakly sclerotized and without setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323129EC6193C06FF1FFCD3FAAFFCA7" pageId="23" pageNumber="24" type="etymology">
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<paragraph id="8B864115C6193C06FF1FFCD3FAAFFCA7" blockId="23.[151,1437,152,1229]" pageId="23" pageNumber="24">
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<emphasis id="B94D9D07C6193C06FF1FFCD3FEE0FCFF" bold="true" box="[197,338,803,829]" pageId="23" pageNumber="24">Etymology.</emphasis>
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The name of this species is feminine and comes from the Greek name
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<emphasis id="B94D9D07C6193C06FB4AFCD4FB75FCFF" box="[1168,1223,804,829]" italics="true" pageId="23" pageNumber="24">elpis</emphasis>
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, that means hope, as a reference to the biological reserve of the
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<typeStatus id="5482FFB7C6193C06FD71FCBBFD6CFCA7" box="[683,734,843,869]" pageId="23" pageNumber="24">type</typeStatus>
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locality,
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<collectingCountry id="F32E0185C6193C06FC93FCBBFC37FCA7" box="[841,901,843,869]" name="Malta" pageId="23" pageNumber="24">Mata</collectingCountry>
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Esperança, meaning “good hope”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323129EC6193C06FF1FFC83FE9DFB0F" pageId="23" pageNumber="24" type="discussion">
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<paragraph id="8B864115C6193C06FF1FFC83FE9DFB0F" blockId="23.[151,1437,152,1229]" pageId="23" pageNumber="24">
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<emphasis id="B94D9D07C6193C06FF1FFC83FEE6FC4F" bold="true" box="[197,340,883,909]" pageId="23" pageNumber="24">Comments.</emphasis>
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This species does not properly fits into any of the groups of species described by
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<bibRefCitation id="EFA83CE4C6193C06FA95FC83FF5BFC77" author="Matile" pageId="23" pageNumber="28" refString="Matile, L. (1996) Revision des Cluzobra Neotropicaux (Diptera: Mycetophilidae). Annales de la Societe Entomolique de France (N. S.), 32, 3 - 57." type="journal article" year="1996">Matile (1996)</bibRefCitation>
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. The species has an apparently unique combination of features, not seen in any other of the groups of species of
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<taxonomicName id="4C393A96C6193C06FEC8FC34FECDFC1F" box="[274,383,964,989]" class="Insecta" family="Mycetophilidae" genus="Cluzobra" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
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<emphasis id="B94D9D07C6193C06FEC8FC34FECDFC1F" box="[274,383,964,989]" italics="true" pageId="23" pageNumber="24">Cluzobra</emphasis>
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</taxonomicName>
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– a quite homogeneous brownish wing membrane, loss of mid ocellus, long Sc, brownish legs and abdomen, and the shape of the male terminalia, with a simple syngonocoxite covering the entire ventral face and a projection just ventrad to the insertion of the gonostyle, bearing a pair of short and strong spines, and the unique shape of the gonostyle. The gonocoxal projection ventrad to the gonostyle insertion is also seen in species of the
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<emphasis id="B94D9D07C6193C06FE10FB94FDEEFBBF" box="[458,604,1124,1149]" italics="true" pageId="23" pageNumber="24">
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<taxonomicName id="4C393A96C6193C06FE10FB94FDFFFBBF" box="[458,589,1124,1149]" class="Insecta" family="Mycetophilidae" genus="Cluzobra" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fascipennis">fascipennis</taxonomicName>
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–
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</emphasis>
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group, such as
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<taxonomicName id="4C393A96C6193C06FCD0FB94FC15FBBF" box="[778,935,1124,1149]" class="Insecta" family="Mycetophilidae" genus="Cluzobra" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="vockerothi" status="sp. nov.">
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<emphasis id="B94D9D07C6193C06FCD0FB94FC15FBBF" box="[778,935,1124,1149]" italics="true" pageId="23" pageNumber="24">C. vockerothi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27E207CC6193C06FC77FB93FC5AFBBF" box="[941,1000,1123,1149]" pageId="23" pageNumber="24" rank="species">
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<emphasis id="B94D9D07C6193C06FC77FB93FC5AFBBF" bold="true" box="[941,1000,1123,1149]" pageId="23" pageNumber="24">sp.n.</emphasis>
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</taxonomicNameLabel>
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, but in many other details the species does not fit in that group. In Matile’s (1996) key, it would run into couplet 28, but it differs significantly from both species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |