175 lines
22 KiB
XML
175 lines
22 KiB
XML
<document id="2CFB2C44BCC0EEF8D94EEAFFC231AE45" ID-DOI="10.11646/zootaxa.5047.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5540953" ID-ZooBank="2FDA4955-4718-4A34-8053-09ACA9BB0241" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632986400249" checkinUser="plazi" docAuthor="Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng" docDate="2021" docId="038FEA2D9A4BFFA819AA4B72FDB13149" docLanguage="en" docName="zootaxa.5047.3.7.pdf" docOrigin="Zootaxa 5047 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ricanula Melichar 1898" docType="treatment" docVersion="3" lastPageNumber="355" masterDocId="FFB692559A4AFFAA193D486DFF9B3270" masterDocTitle="Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha)" masterLastPageNumber="369" masterPageNumber="353" pageNumber="354" updateTime="1699416654639" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="10B78CC6A0B2E8FF08779071649052D7">Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha)</mods:title>
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<mods:name id="5D3545CFC380C972A3A196CADFC98242" type="personal">
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<mods:namePart id="DAB9843D2979CDB5C1BAACF8D7E9B9B0">Zhang, Huan</mods:namePart>
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<mods:nameIdentifier id="424545F9232436E8C60930852FC6DA07" type="ORCID">0000-0002-4372-8419</mods:nameIdentifier>
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<mods:affiliation id="656E120F6233419FCCAA692430075E38">Key Laboratory of Plant Protection Resources and Pest Management of the Ministry of Education; Entomological Museum, Northwest A & F University, Yangling, Shaanxi 712100, China & 18335481296 @ 163. com; https: // orcid. org / 0000 - 0002 - 4372 - 8419</mods:affiliation>
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<mods:namePart id="CE9BD27A8DF3077EB7E7FBB6CBAD4E89">Wang, Wen-Qian</mods:namePart>
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<mods:affiliation id="BF88FD2B2D017729C0E0A47D20173757">Key Laboratory of Plant Protection Resources and Pest Management of the Ministry of Education; Entomological Museum, Northwest A & F University, Yangling, Shaanxi 712100, China & wanghan 19931112 @ 163. com; https: // orcid. org / 0000 - 0003 - 0446 - 4948</mods:affiliation>
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<mods:namePart id="B26AAFE181CD6052AEECB1605AEC08C2">Stroiński, Adam</mods:namePart>
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<mods:affiliation id="C86E6096FA809A26EC28B18DAFEEA6F5">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland.</mods:affiliation>
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<mods:namePart id="AF5EA7577493277285F2B4ACFB831657">Qin, Dao-Zheng</mods:namePart>
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<mods:affiliation id="8D4163C0A944DCD81E595ACE2032819E">Key Laboratory of Plant Protection Resources and Pest Management of the Ministry of Education; Entomological Museum, Northwest A & F University, Yangling, Shaanxi 712100, China & qindaozh 0426 @ aliyun. com, qindaozh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0001 - 8219 - 6753</mods:affiliation>
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<mods:nameIdentifier id="ADD0EFD44B08018DF5AA0F3943F3DD85" type="email">qindaozh@nwsuaf.edu.cn</mods:nameIdentifier>
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<mods:title id="12D1EDCEFBBECF2F9A1BF8F24B0A6DCD">Zootaxa</mods:title>
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<mods:date id="8E49F7E58FC5C39FEDB6426358000C27">2021</mods:date>
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<mods:number id="D928F801FCA3EB9692ABF547FB7FD26C">2021-09-30</mods:number>
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<mods:number id="90D350E604DF6F3250317BEF86548E57">5047</mods:number>
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<taxonomicName id="4C2620B89A4BFFAB19AA4B72FE5D3149" ID-CoL="79VS" ID-ENA="130628" authority="Melichar, 1898" authorityName="Melichar" authorityYear="1898" box="[151,454,799,826]" class="Insecta" family="Ricaniidae" genus="Ricanula" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="354" phylum="Arthropoda" rank="genus">
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<emphasis id="B95287299A4BFFAB19AA4B72FE9D314A" bold="true" box="[151,262,799,826]" italics="true" pageId="1" pageNumber="354">Ricanula</emphasis>
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<bibRefCitation id="EFB726CA9A4BFFAB18304B72FE5D3149" author="Melichar, L." box="[269,454,799,825]" pageId="1" pageNumber="354" pagination="197 - 359" refId="ref8707" refString="Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359." type="journal article" year="1898">Melichar, 1898</bibRefCitation>
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</emphasis>
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</taxonomicName>
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<subSubSection id="C33C08B09A4BFFAB19AA4B09FE2131EA" pageId="1" pageNumber="354" type="reference_group">
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<paragraph id="8B995B3B9A4BFFAB19AA4B09FDA8310C" blockId="1.[151,973,868,953]" box="[151,563,868,892]" pageId="1" pageNumber="354">
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<treatmentCitation id="0A877D2A9A4BFFAB19AA4B09FDB4310C" author="Melichar, L." box="[151,559,868,892]" page="218" pageId="1" pageNumber="354" year="1898">
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<taxonomicName id="4C2620B89A4BFFAB19AA4B09FDB4310C" authority="Melichar, 1898: 218" authorityName="Melichar" authorityPageNumber="218" authorityYear="1898" box="[151,559,868,892]" class="Insecta" family="Ricaniidae" genus="Ricania" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="354" phylum="Arthropoda" rank="subGenus" subGenus="Ricanula">
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<emphasis id="B95287299A4BFFAB19AA4B09FF7D310B" box="[151,230,868,891]" italics="true" pageId="1" pageNumber="354">Ricania</emphasis>
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(
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<emphasis id="B95287299A4BFFAB19C84B09FECB310C" box="[245,336,868,892]" italics="true" pageId="1" pageNumber="354">Ricanula</emphasis>
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)
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<bibRefCitation id="EFB726CA9A4BFFAB18624B09FDB4310C" author="Melichar, L." box="[351,559,868,892]" pageId="1" pageNumber="354" pagination="197 - 359" refId="ref8707" refString="Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359." type="journal article" year="1898">Melichar, 1898: 218</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19AA4BEFFE2131EA" blockId="1.[151,973,868,953]" box="[151,442,898,922]" pageId="1" pageNumber="354">
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<treatmentCitationGroup id="AB367C159A4BFFAB19AA4BEFFE2131EA" box="[151,442,898,922]" pageId="1" pageNumber="354">
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<treatmentCitation id="0A877D2A9A4BFFAB19AA4BEFFE2D31EA" author="Schmidt, E." box="[151,438,898,922]" page="75" pageId="1" pageNumber="354" year="1912">
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<taxonomicName id="4C2620B89A4BFFAB19AA4BEFFE2D31EA" ID-CoL="79VS" authority="Schmidt, 1912: 75" authorityName="Schmidt" authorityPageNumber="75" authorityYear="1912" box="[151,438,898,922]" class="Insecta" family="Ricaniidae" genus="Ricanula" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="354" phylum="Arthropoda" rank="genus">
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<emphasis id="B95287299A4BFFAB19AA4BEFFF6931EA" box="[151,242,898,922]" italics="true" pageId="1" pageNumber="354">Ricanula</emphasis>
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<bibRefCitation id="EFB726CA9A4BFFAB19C44BEFFE2D31EA" author="Schmidt, E." box="[249,438,898,922]" pageId="1" pageNumber="354" pagination="67 - 102" refId="ref9019" refString="Schmidt, E. (1912) Diagnosen neuer Fulgoriden-Gattungen und Arten nebst einigen Bemerkungen. Stettiner Entomologische Zeitung, 73, 67 - 102." type="journal article" year="1912">Schmidt, 1912: 75</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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<subSubSection id="C33C08B09A4BFFAB19AA4BCCFC5631C9" box="[151,973,929,953]" pageId="1" pageNumber="354" type="type_taxon">
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<paragraph id="8B995B3B9A4BFFAB19AA4BCCFC5631C9" blockId="1.[151,973,868,953]" box="[151,973,929,953]" pageId="1" pageNumber="354">
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<emphasis id="B95287299A4BFFAB19AA4BCCFEB831C9" bold="true" box="[151,291,929,953]" inLineHeading="true" pageId="1" pageNumber="354" reason="1">
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<typeStatus id="549DE5999A4BFFAB19AA4BCCFF5731C9" box="[151,204,929,953]" pageId="1" pageNumber="354">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C2620B89A4BFFAB18174BCFFD1031C9" ID-CoL="4SYSH" authority="Melichar, 1898" authorityName="Melichar" authorityYear="1898" box="[298,651,929,953]" class="Insecta" family="Ricaniidae" genus="Ricania" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="354" phylum="Arthropoda" rank="species" species="noualhieri">
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<emphasis id="B95287299A4BFFAB18174BCFFE7231C9" box="[298,489,929,953]" italics="true" pageId="1" pageNumber="354">Ricania noualhieri</emphasis>
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<bibRefCitation id="EFB726CA9A4BFFAB18D24BCCFD1031C9" author="Melichar, L." box="[495,651,929,953]" pageId="1" pageNumber="354" pagination="197 - 359" refId="ref8707" refString="Melichar, L. (1898) Monographie der Ricaniiden (Homoptera). Annalen des K. K. Naturhistorischen Hofmuseums, 8 (2 - 3), 197 - 359." type="journal article" year="1898">Melichar, 1898</bibRefCitation>
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</taxonomicName>
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, designated by
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<bibRefCitation id="EFB726CA9A4BFFAB1A154BCCFC5231C9" author="Schmidt, E." box="[808,969,929,953]" pageId="1" pageNumber="354" pagination="67 - 102" refId="ref9019" refString="Schmidt, E. (1912) Diagnosen neuer Fulgoriden-Gattungen und Arten nebst einigen Bemerkungen. Stettiner Entomologische Zeitung, 73, 67 - 102." type="journal article" year="1912">Schmidt (1912)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C08B09A4BFFA819AA4B8AFDB13149" lastPageId="2" lastPageNumber="355" pageId="1" pageNumber="354" type="discussion">
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<paragraph id="8B995B3B9A4BFFAB19AA4B8AFE5E361D" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">
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<emphasis id="B95287299A4BFFAB19AA4B8AFE903671" bold="true" box="[151,267,999,1025]" pageId="1" pageNumber="354">Remarks.</emphasis>
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<taxonomicName id="4C2620B89A4BFFAB182F4B8AFEEC3671" authorityName="Schmidt" authorityYear="1912" box="[274,375,999,1025]" class="Insecta" family="Ricaniidae" genus="Ricanula" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="354" phylum="Arthropoda" rank="genus">
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<emphasis id="B95287299A4BFFAB182F4B8AFEEC3671" box="[274,375,999,1025]" italics="true" pageId="1" pageNumber="354">Ricanula</emphasis>
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</taxonomicName>
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can be distinguished from other genera in
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<taxonomicName id="4C2620B89A4BFFAB1A654B8AFC4B3671" authorityName="Amyot & Audinet-Serville" authorityYear="1843" box="[856,976,999,1025]" class="Insecta" family="Ricaniidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="354" phylum="Arthropoda" rank="family">Ricaniidae</taxonomicName>
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by the combination of following characters: costal area of tegmen with sparse and curved transverse veinlets; postero-apical part of tegmen with eyes-spot black cells; tegmen with one line of transverse veinlets; median and posterior parts of tegmen with numerous irregular transverse veinlets.
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</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4C1AFAF436C5" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">
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<emphasis id="B95287299A4BFFAB19FA4C1AFE4836E1" bold="true" box="[199,467,1143,1169]" pageId="1" pageNumber="354">Diagnostic characters.</emphasis>
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<emphasis id="B95287299A4BFFAB18E64C1AFDBB36E1" box="[475,544,1143,1169]" italics="true" pageId="1" pageNumber="354">Head.</emphasis>
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Head including eyes narrower than mesonotum/thorax. Vertex transverse, distinctly wider than long at midline, with all margins well carinate; disc of vertex with or without median carina.
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</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4CADFD753735" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">Frons with all margins well carinated, at upper margin longer than high at midline, widest at the level of lower margin of compound eyes; lateral margins covering base of pedicel, not incised near the level of ocelli. Frontal disc tricarinate, carinae distinctly separated basally, median carina straight; lateral carinae arcuate, almost parallel to lateral margins. Fronto-clypeal suture arched/arcuate.</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4D22FC1D3719" blockId="1.[151,1437,999,2033]" box="[199,902,1359,1386]" pageId="1" pageNumber="354">Clypeus distinctly narrower than frons, without median carina.</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4D1EFE5F37C2" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">Compound eyes, with small callus in lower part of posterior margin. Ocelli present. Rostrum-apical segment shorter than subapical one.</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4DD6FC5537A6" blockId="1.[151,1437,999,2033]" box="[199,974,1467,1494]" pageId="1" pageNumber="354">Antenna pedicel elongate, cylindrical, with functional area at the top.</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4DB2FEBE346D" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">
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<emphasis id="B95287299A4BFFAB19FA4DB2FE833789" box="[199,280,1503,1529]" italics="true" pageId="1" pageNumber="354">Thorax</emphasis>
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. Pronotum distinctly longer than vertex at midline; disc of pronotum with median carina and two lateral impressions.
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</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4E45FD4434A2" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">Mesonotum elongate, diamond shape, longer in midlength than wide at lateral angles and longer in midlength longer than combined length of vertex and pronotum at midlength; lateral angles placed before midlength; median carina, lateral and antero-lateral carinae present; median carina and lateral carinae connected basally; median carina reaching scutellum, lateral carinae reaching posterior margin, anterolateral carinae not connected with lateral and not surpassing level of lateral angles of mesonotum.</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4EB6FBB935D9" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">
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Tegmina membranous, elongately-triangular, flattened, with distinct venation and transverse veinlets. Costal margin weakly arcuate, apical angle broadly rounded, placed distad to claval angle, posterior margin weakly sinuate, postclaval margin (tornus) absent. Costal area with transverse veinlets and oblique shallowly incisions; postero-apical part with eyes-spot black cells. Longitudinal veins ScP+RA, MP and CuA leaving basal cell separated. CuA vein first fork placed before the connection of claval veins Pcu+A
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<subScript id="17A2597E9A4BFFAB1A064F13FCD835FC" attach="right" box="[827,835,1918,1932]" fontSize="6" pageId="1" pageNumber="354">1</subScript>
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. Tegmen with single apical line of transverse veinlets, median and posterior part of tegmen with numerous irregular transverse veinlets.
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</paragraph>
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<paragraph id="8B995B3B9A4BFFAB19FA4FDEFE7E3581" blockId="1.[151,1437,999,2033]" pageId="1" pageNumber="354">Hind wing with precostal cell present; ScRA and MP forking distinctly after midlength of wing, CuA forking distinctly before half of wing.</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA48FAFE833289" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">Hing legs. Metatibia with 2 lateral spines; apically with 6 well developed spines, external lateral spines bigger than internal lateral spines; basitarsomere of metatarsus a little longer than cumulative length of second and apical tarsomeres.</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA496EFB1A3315" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">
|
||
<emphasis id="B95287299A48FFA819FA496EFEE5336D" box="[199,382,259,285]" italics="true" pageId="2" pageNumber="355">Male terminalia</emphasis>
|
||
. Anal tube with ventral margin strongly concaved in lateral view. Pygofer higher than wide; dorsal part narrower than ventral one. Genital styles broadly triangular (dorsal and ventral margins nearly parallel in
|
||
<taxonomicName id="4C2620B89A48FFA819AA4926FEB43315" box="[151,303,331,357]" class="Insecta" family="Ricaniidae" genus="Ricanula" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="355" phylum="Arthropoda" rank="species" species="fujianensis">
|
||
<emphasis id="B95287299A48FFA819AA4926FEB43315" box="[151,303,331,357]" italics="true" pageId="2" pageNumber="355">R. fujianensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2620B89A48FFA8185B4926FE723314" authorityName="Chou et Lu" authorityYear="1977" box="[358,489,331,356]" class="Insecta" family="Ricaniidae" genus="Ricania" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="355" phylum="Arthropoda" rank="species" species="cacaonis">
|
||
<emphasis id="B95287299A48FFA8185B4926FE723314" box="[358,489,331,356]" italics="true" pageId="2" pageNumber="355">R. cacaonis</emphasis>
|
||
</taxonomicName>
|
||
), with sharp spine-like process at the end of dorsal margin.
|
||
</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA4902FE3133A1" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">Periandrium with or without processes, with long lateral split surpassing the half of its length; dorsal periandrium a bit shorter than ventral one. Aedeagus long and narrow, apically with 1–2 pairs of symmetrical, well sclerotized, spinose processes.</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA49B6FB193069" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">
|
||
<emphasis id="B95287299A48FFA819FA49B6FE0D3385" box="[199,406,475,501]" italics="true" pageId="2" pageNumber="355">Female terminalia</emphasis>
|
||
. Pregenital sternite with well-developed lateral lobes, median portion of the posterior margin with or without processes. Anal tube not surpassing half of upper margin of the gonoplac.
|
||
</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA4A4EFBA33011" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">Gonoplac well developed, laterally flattened; posterior margin of the gonoplac with 2–3 rows of small teeth; membranous parts of gonoplac well developed, placed medially on ventral margin.</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA4A06FF6830BD" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">Gonapophysis VIII sabre-like, v-shape in cross section, with teeth at dorsal margin; endogonocoxal process with spiniferous microsculptures, well sclerotized medially; lateral parts membranous, reaching apex of gonapophysis VIII.</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA4ABAFCE03081" blockId="2.[151,1437,151,825]" box="[199,891,727,753]" pageId="2" pageNumber="355">Gonaphophyses IX and gonospiculum bridge well developed.</paragraph>
|
||
<paragraph id="8B995B3B9A48FFA819FA4A96FDB13149" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="355">Bursa copulatrix with two isometric pouches: first pouch with well visible cell and sclerotized ornamentation; second pouch with numerous pores.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |