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<document id="585B190C552719FF44CC5D2A7F48D998" ID-CLB-Dataset="51205" ID-DOI="10.11646/zootaxa.4061.5.3" ID-GBIF-Dataset="8e4c370b-2a1d-4bc4-a714-16f9f3a1bd65" ID-ISSN="1175-5326" ID-Zenodo-Dep="264294" ID-ZooBank="6EEE4C42-D292-490A-9549-CB3428430920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459680617424" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2016" docId="038F87EE01523005FF0BD57FFD2CF992" docLanguage="en" docName="zootaxa.4061.5.3.pdf" docOrigin="Zootaxa 4061 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus arnosti Häckel, 2016, sp. n." docType="treatment" docVersion="10" lastPageNumber="507" masterDocId="FFB6FF9601513006FF9CD433FFDDFFEB" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)" masterLastPageNumber="526" masterPageNumber="504" pageNumber="507" updateTime="1698207076058" updateUser="ExternalLinkService">
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<mods:title id="4F877D3C713C3BA991990F4797CCAFB7">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="F615B5B194F54F34EE10742A648D3EF7">Häckel, Martin</mods:namePart>
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<mods:date id="C7DE9AFB6EF8E5C4BC48A5E4306FE366">2016</mods:date>
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<treatment id="038F87EE01523005FF0BD57FFD2CF992" ID-DOI="http://doi.org/10.5281/zenodo.6088121" ID-GBIF-Taxon="119333004" ID-Zenodo-Dep="6088121" LSID="urn:lsid:plazi:treatment:038F87EE01523005FF0BD57FFD2CF992" httpUri="http://treatment.plazi.org/id/038F87EE01523005FF0BD57FFD2CF992" lastPageNumber="507" pageId="3" pageNumber="507">
<subSubSection id="C33C657301523005FF0BD57FFDFDFE8D" box="[151,544,332,358]" pageId="3" pageNumber="507" type="nomenclature">
<paragraph id="8B9936F801523005FF0BD57FFDFDFE8D" blockId="3.[151,699,332,391]" box="[151,544,332,358]" pageId="3" pageNumber="507">
<heading id="D0D1819401523005FF0BD57FFDFDFE8D" bold="true" box="[151,544,332,358]" fontSize="11" level="1" pageId="3" pageNumber="507" reason="1">
<emphasis id="B952EAEA01523005FF0BD57FFDFDFE8D" bold="true" box="[151,544,332,358]" pageId="3" pageNumber="507">
1.
<taxonomicName id="4C264D7B01523005FF28D57FFE0AFE8D" ID-CoL="Z5SW" box="[180,471,332,358]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="arnosti" status="sp. nov.">
<emphasis id="B952EAEA01523005FF28D57FFE0AFE8D" bold="true" box="[180,471,332,358]" italics="true" pageId="3" pageNumber="507">Craspedophorus arnosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101523005FE42D57FFDFDFE8D" box="[478,544,332,358]" pageId="3" pageNumber="507" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C657301523005FF0BD55DFD66FE6C" box="[151,699,366,391]" pageId="3" pageNumber="507" type="description">
<paragraph id="8B9936F801523005FF0BD55DFD66FE6C" blockId="3.[151,699,332,391]" box="[151,699,366,391]" pageId="3" pageNumber="507">
(Plate 2, Figs 13, 14,
<figureCitation id="131D2A7D01523005FE1AD55DFE0BFE6C" box="[390,470,366,391]" captionStart="FIGURE 25. C" captionStartId="13.[151,250,1927,1949]" captionTargetBox="[362,1246,1183,1901]" captionTargetId="figure@13.[340,1247,1183,1906]" captionTargetPageId="13" captionText="FIGURE 25. C. arnosti sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. pronotum, dorsal view; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264295/files/figure.png" pageId="3" pageNumber="507">Fig. 25</figureCitation>
af, Plate 4, Fig. 29)
</paragraph>
</subSubSection>
<subSubSection id="C33C657301523005FF0BD585FDD1FDFC" pageId="3" pageNumber="507" type="materials_examined">
<paragraph id="8B9936F801523005FF0BD585FDD1FDFC" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">
<emphasis id="B952EAEA01523005FF0BD585FE98FE24" bold="true" box="[151,325,438,463]" pageId="3" pageNumber="507">
<typeStatus id="549D885A01523005FF0BD585FF0FFE24" box="[151,210,438,463]" pageId="3" pageNumber="507">Type</typeStatus>
material.
</emphasis>
<typeStatus id="549D885A01523005FED0D585FE69FE24" box="[332,436,438,463]" pageId="3" pageNumber="507" type="holotype">Holotype</typeStatus>
(♂): “W Centr. Africa Rep. of, Nana-Mambéré prov.
<quantity id="4CDE9B1D01523005FB87D585FBBFFE24" box="[1051,1122,438,463]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="507" unit="km" value="60.0">60 km</quantity>
SE Bouar,
<quantity id="4CDE9B1D01523005FB7FD585FAC0FE24" box="[1251,1309,438,463]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="507" unit="km" value="5.0">5 km</quantity>
NE Baoro,
<quantity id="4CDE9B1D01523005FF0BD5E9FF3DFE19" box="[151,224,474,499]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="507" unit="m" value="600.0">600 m</quantity>
<date id="FF98103801523005FF75D5E8FEF1FE18" box="[233,300,475,499]" pageId="3" pageNumber="507" value="2009-06">VI-09</date>
lgt. A. Kudrna jr.”, (Plate 2, Figs 13, 14,
<figureCitation id="131D2A7D01523005FC8DD5E9FCBEFE18" box="[785,867,474,499]" captionStart="FIGURE 25. C" captionStartId="13.[151,250,1927,1949]" captionTargetBox="[362,1246,1183,1901]" captionTargetId="figure@13.[340,1247,1183,1906]" captionTargetPageId="13" captionText="FIGURE 25. C. arnosti sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. pronotum, dorsal view; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264295/files/figure.png" pageId="3" pageNumber="507">Fig. 25</figureCitation>
af, Plate 4, Fig. 29).
<typeStatus id="549D885A01523005FBFDD5E8FB0EFE18" box="[1121,1235,475,499]" pageId="3" pageNumber="507" type="paratype">Paratypes</typeStatus>
: 2♀ same data as
<typeStatus id="549D885A01523005FF0BD5CDFF24FDFC" box="[151,249,510,535]" pageId="3" pageNumber="507" type="holotype">holotype</typeStatus>
(Plate 2, Fig. 14, cMH).
</paragraph>
</subSubSection>
<subSubSection id="C33C657301523005FF0BD611FF31F9E7" pageId="3" pageNumber="507" type="description">
<paragraph id="8B9936F801523005FF0BD611FC50FDB5" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">
<emphasis id="B952EAEA01523005FF0BD611FEF4FDD0" bold="true" box="[151,297,546,571]" pageId="3" pageNumber="507">Description.</emphasis>
Length
<quantity id="4CDE9B1D01523005FE11D610FDEBFDD1" box="[397,566,547,571]" metricMagnitude="-2" metricUnit="m" metricValue="2.565" metricValueMax="2.7199999999999998" metricValueMin="2.41" pageId="3" pageNumber="507" unit="mm" value="25.65" valueMax="27.2" valueMin="24.1">24.127.2 mm</quantity>
, width
<quantity id="4CDE9B1D01523005FD0CD610FCE5FDD1" box="[656,824,547,571]" metricMagnitude="-2" metricUnit="m" metricValue="1.11" metricValueMax="1.19" metricValueMin="1.03" pageId="3" pageNumber="507" unit="mm" value="11.1" valueMax="11.9" valueMin="10.3">10.311.9 mm</quantity>
. Proportions: Pronotum 1.361.45x wider than long, 1.792.2x wider than head, elytra 1.271.51x wider than pronotum.
</paragraph>
<paragraph id="8B9936F801523005FF5BD659FC4BFD19" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">Coloration. Body throughout piceous black, opaque, densely covered by black setae; each elytron with a large compound yellowish-orange humeral macula extending from 2nd to 8th interval and consisting of seven about equally elongated spots with margins moderately serrate; elytral margin black. Underside piceous black, opaque, antennae and palps black. Head long, eyes convex, neck constricted.</paragraph>
<paragraph id="8B9936F801523005FF5BD6CDFA40FC05" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">Pronotum transverse, 1.36x (females)1.45x (male HT) wider than long, almost hexagonal; maximum width immediately behind midlength; anterior margin flat, front angles distinctly projecting anteriorly and strongly rounded; lateral margins evenly rounded (in male front part of lateral margin more rounded and lateral rim wider; Plate 2, Figs 13, 14), widening gradually to maximum width, then narrowing in straight line toward hind angles; each hind angle with a small indentation preceded by an indistinct incision (male with incision more distinct); base medially parallel with elytral base; laterally base slanted forward, causing hind angle to be wide open; sagittal line indistinct, basal pits strongly impressed, elongated to midlength; surface strongly coarsely and unevenly punctured.</paragraph>
<paragraph id="8B9936F801523005FF5BD7CAFCB0FBB7" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">Elytra fused, almost parallel-sided, weekly convex, flattened on disc, wings reduced. Striae deeply impressed, punctured, 1st stria strongly oblique to 2nd stria in the first basal quarter, joining to it near base; intervals fairly convex, finely and densely punctured, sparsely covered by setae.</paragraph>
<paragraph id="8B9936F801523005FF5BD05BFAC5FB4F" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">
Underside covered by short setae, finely punctured medially, coarsely punctured near margins; metepisterna longer than wide, wider anteriorly than posteriorly (
<figureCitation id="131D2A7D01523005FD40D0BFFCF2FB4E" box="[732,815,1164,1189]" captionStart="FIGURE 25. C" captionStartId="13.[151,250,1927,1949]" captionTargetBox="[362,1246,1183,1901]" captionTargetId="figure@13.[340,1247,1183,1906]" captionTargetPageId="13" captionText="FIGURE 25. C. arnosti sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. pronotum, dorsal view; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264295/files/figure.png" pageId="3" pageNumber="507">Fig. 25</figureCitation>
e). Legs moderately covered by black setae.
</paragraph>
<paragraph id="8B9936F801523005FF5BD083FE83FB06" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">
Aedeagus similar to that in
<taxonomicName id="4C264D7B01523005FE66D082FCE1FB23" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[506,828,1200,1225]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01523005FE66D082FDAFFB22" box="[506,626,1200,1225]" italics="true" pageId="3" pageNumber="507">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901523005FD1CD083FCEEFB23" author="Fabricius" box="[640,819,1200,1225]" pageId="3" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
, but in dorsal view differing by apical curvature (Plate 4 cf. Figs 2829).
</paragraph>
<paragraph id="8B9936F801523005FF5BD0CBFF31F9E7" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">
<emphasis id="B952EAEA01523005FF5BD0CBFE0EFAFA" bold="true" box="[199,467,1272,1297]" pageId="3" pageNumber="507">Differential diagnosis.</emphasis>
<taxonomicName id="4C264D7B01523005FE7CD0CBFD2AFAFB" box="[480,759,1272,1297]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="arnosti" status="sp. nov.">
<emphasis id="B952EAEA01523005FE7CD0CBFD2AFAFB" box="[480,759,1272,1297]" italics="true" pageId="3" pageNumber="507">Craspedophorus arnosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101523005FC98D0CAFCBCFAFB" box="[772,865,1273,1296]" pageId="3" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
mostly resembles
<taxonomicName id="4C264D7B01523005FBDFD0CAFA45FAFB" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[1091,1432,1272,1297]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01523005FBDFD0CAFB1FFAFA" box="[1091,1218,1272,1297]" italics="true" pageId="3" pageNumber="507">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901523005FB4AD0CBFA52FAFB" author="Fabricius" box="[1238,1423,1272,1297]" pageId="3" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
, differing from it by elytral coloration. In contrast to
<taxonomicName id="4C264D7B01523005FC9CD12EFCA1FADE" box="[768,892,1308,1333]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01523005FC9CD12EFCA1FADE" box="[768,892,1308,1333]" italics="true" pageId="3" pageNumber="507">C. reflexus</emphasis>
</taxonomicName>
, which has both elytral maculae (humeral and preapical), in
<taxonomicName id="4C264D7B01523005FEA8D172FE78FAB3" box="[308,421,1345,1368]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="arnosti" status="sp. nov.">
<emphasis id="B952EAEA01523005FEA8D172FE78FAB3" box="[308,421,1345,1368]" italics="true" pageId="3" pageNumber="507">C. arnosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101523005FE32D172FDDBFAB3" box="[430,518,1345,1368]" pageId="3" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
the preapical macula is absent in both sexes. The new species also resembles
<taxonomicName id="4C264D7B01523005FA1ED172FE62FA96" authority="Alluaud, 1915" authorityName="Alluaud" authorityYear="1915" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="chevalieri">
<emphasis id="B952EAEA01523005FA1ED172FEDAFA96" italics="true" pageId="3" pageNumber="507">C. chevalieri</emphasis>
(
<bibRefCitation id="EFB74B0901523005FE89D157FE6AFA96" author="Alluaud" box="[277,439,1380,1405]" pageId="3" pageNumber="525" refString="Alluaud, C. (1915) Contributions a l'etude des carabiques d'Afrique et de Madagascar (Col. Carabidae). V. Les grands Panageides du bassin du Chari. Bulletin de la Societe Entomologique de France, 1915, 152 - 156." type="journal article" year="1915">Alluaud, 1915</bibRefCitation>
)
</taxonomicName>
of the
<taxonomicName id="4C264D7B01523005FD90D156FD4BFA96" box="[524,662,1380,1405]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="strachani">
<emphasis id="B952EAEA01523005FD90D156FD4BFA96" box="[524,662,1380,1405]" italics="true" pageId="3" pageNumber="507">C. strachani</emphasis>
</taxonomicName>
species group in elytral coloration presenting only a single (humeral) macula on each elytron, but exhibiting a more flattened body, differently shaped metepisterna (narrowing posteriorly and longer than wide), and anterior margins of ventrites not crenulated. The last two characters place the new species in the
<taxonomicName id="4C264D7B01523005FE03D1E2FDC7FA02" box="[415,538,1488,1513]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01523005FE03D1E2FDC7FA02" box="[415,538,1488,1513]" italics="true" pageId="3" pageNumber="507">C. reflexus</emphasis>
</taxonomicName>
group, within which it can be easily distinguished by the unique elytral color pattern.
</paragraph>
</subSubSection>
<subSubSection id="C33C657301523005FF5BD22BFEF1F9BE" pageId="3" pageNumber="507" type="etymology">
<paragraph id="8B9936F801523005FF5BD22BFEF1F9BE" blockId="3.[151,1437,438,1657]" pageId="3" pageNumber="507">
<emphasis id="B952EAEA01523005FF5BD22BFE93F9DA" bold="true" box="[199,334,1560,1585]" pageId="3" pageNumber="507">Etymology.</emphasis>
Named after my friend Arnošt Kudrna (České Budějovice,
<collectingCountry id="F331766801523005FBBED22BFB02F9DA" box="[1058,1247,1560,1585]" name="Czech Republic" pageId="3" pageNumber="507">Czech Republic</collectingCountry>
), a specialist in
<taxonomicName id="4C264D7B01523005FF0BD20FFEF5F9BE" box="[151,296,1596,1621]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="subFamily" subFamily="Cicindelinae">Cicindelinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C657301523005FF5BD253FD2CF992" box="[199,753,1632,1657]" pageId="3" pageNumber="507" type="distribution">
<paragraph id="8B9936F801523005FF5BD253FD2CF992" blockId="3.[151,1437,438,1657]" box="[199,753,1632,1657]" pageId="3" pageNumber="507">
<emphasis id="B952EAEA01523005FF5BD253FEBDF992" bold="true" box="[199,352,1632,1657]" pageId="3" pageNumber="507">Distribution.</emphasis>
Known only from the
<typeStatus id="549D885A01523005FDFCD252FD4DF992" box="[608,656,1633,1657]" pageId="3" pageNumber="507">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>