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<document id="59CE01AC8E9E9FEFD60C3588CA6172D6" ID-CLB-Dataset="3791" ID-DOI="10.11646/zootaxa.5060.1.1" ID-GBIF-Dataset="40f89bbe-7f7b-4abd-aa02-a45e410da232" ID-ISSN="1175-5326" ID-Zenodo-Dep="5606671" ID-ZooBank="14126B35-AEEA-40F5-A892-FE6A8D64E5D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1635406161957" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2021" docId="038F87D5FF94FF86FF6D21B4FBB39D8E" docLanguage="en" docName="zootaxa.5060.1.1.pdf" docOrigin="Zootaxa 5060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pacifiosoma cristofer" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFB6FFADFF8FFF9DFFFA2217FF8C9A76" masterDocTitle="The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="28" updateTime="1699419728057" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AA6A673D0AAE56B1DE2AA24ABA9FE121">The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)</mods:title>
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<mods:namePart id="4A7039666D3EB09755FBE79DF032A6F9">Mikhaljova, Elena V.</mods:namePart>
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<mods:date id="3CDFF6517379C992E4DD24A8DCC84909">2021</mods:date>
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<treatment id="038F87D5FF94FF86FF6D21B4FBB39D8E" ID-DOI="http://doi.org/10.5281/zenodo.5608271" ID-GBIF-Taxon="189548890" ID-Zenodo-Dep="5608271" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E" httpUri="http://treatment.plazi.org/id/038F87D5FF94FF86FF6D21B4FBB39D8E" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="C33C6548FF94FF86FF6D21B4FD0299C8" box="[151,654,931,958]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B9936C3FF94FF86FF6D21B4FD0299C8" blockId="27.[151,654,931,994]" box="[151,654,931,958]" pageId="27" pageNumber="28">
<heading id="D0D181AFFF94FF86FF6D21B4FD0299C8" bold="true" box="[151,654,931,958]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<taxonomicName id="4C264D40FF94FF86FF6D21B4FD0299C8" ID-CoL="4BW6F" authority="(Mikhaljova, 1993)" baseAuthorityName="Mikhaljova" baseAuthorityYear="1993" box="[151,654,931,958]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cristofer">
<emphasis id="B952EAD1FF94FF86FF6D21B4FD0299C8" bold="true" box="[151,654,931,958]" pageId="27" pageNumber="28">
<emphasis id="B952EAD1FF94FF86FF6D21B4FE1599CB" bold="true" box="[151,409,931,957]" italics="true" pageId="27" pageNumber="28">Pacifiosoma cristofer</emphasis>
(
<bibRefCitation id="EFB74B32FF94FF86FE5321B3FD0A99C8" author="Mikhaljova, E. V." box="[425,646,932,958]" pageId="27" pageNumber="28" pagination="3 - 36" refId="ref29575" refString="Mikhaljova, E. V. (1993) The millipedes (Diplopoda) of Siberia and the Far East of Russia. Arthropoda Selecta, 2 (2), 3 - 36." type="journal article" year="1993">Mikhaljova, 1993</bibRefCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="C33C6548FF94FF86FF6D21DFFE989994" box="[151,276,968,994]" pageId="27" pageNumber="28" type="description">
<paragraph id="8B9936C3FF94FF86FF6D21DFFE989994" blockId="27.[151,654,931,994]" box="[151,276,968,994]" pageId="27" pageNumber="28">
<figureCitation id="131D2A46FF94FF86FF6D21DFFE989994" box="[151,276,968,994]" captionStart-0="FIGURES 40–44" captionStart-1="FIGURES 45–48" captionStartId-0="28.[151,264,1476,1500]" captionStartId-1="29.[151,264,1004,1028]" captionTargetBox-0="[151,1435,181,1452]" captionTargetBox-1="[151,1435,181,980]" captionTargetId-0="figure-24@28.[151,1435,181,1453]" captionTargetId-1="figure-16@29.[151,1435,181,980]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURES 40–44. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 40–41, 44) and Bolshekhekhtsirsky Nature Reserve (Figs 42–43). 40, 42, gonopods, caudal view, telopodites of posterior gonopods broken on fig. 42; 41, central part of gonopods; 43, distal parts of gonopods; 44, apex of colpocoxite and anterior gonopod telopodite; pp, posterior angicoxal process; aat, apex of anterior gonopod telopodite. Scales: 10 μm (Fig. 44), 20 μm (Figs 41, 43), 100 μm (Figs 40, 42)." captionText-1="FIGURES 45–48. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 45–46) and Bolshekhekhtsirsky Nature Reserve (Figs 47–48). 45, 47, gonopods, front view; 46, 48, distal parts of colpocoxites; ap, anterior angiocoxal process; cl, colpocoxites. Scales: 20 μm (Figs 46, 48), 100 μm (Figs 45, 47)." figureDoi-0="http://doi.org/10.5281/zenodo.5606705" figureDoi-1="http://doi.org/10.5281/zenodo.5606709" httpUri-0="https://zenodo.org/record/5606705/files/figure.png" httpUri-1="https://zenodo.org/record/5606709/files/figure.png" pageId="27" pageNumber="28">Figs 40–48</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF94FF86FF6D2611FEBF9E0C" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8B9936C3FF94FF86FF6D2611FD119E68" blockId="27.[151,1436,1030,1146]" box="[151,669,1030,1054]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB3611EDFF94FF86FF6D2611FD119E68" box="[151,669,1030,1054]" pageId="27" pageNumber="28">
<treatmentCitation id="0A8710D2FF94FF86FF6D2611FDE99E68" author="Mikhaljova, E. V." box="[151,613,1030,1054]" page="18" pageId="27" pageNumber="28" year="1993">
<taxonomicName id="4C264D40FF94FF86FF6D2611FDE99E68" ID-CoL="36LVJ" authority="Mikhaljova 1993: 18" authorityName="Mikhaljova" authorityPageNumber="18" authorityYear="1993" box="[151,613,1030,1054]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cristofer">
<emphasis id="B952EAD1FF94FF86FF6D2611FE059E68" box="[151,393,1030,1054]" italics="true" pageId="27" pageNumber="28">Diplomaragna cristofer</emphasis>
<bibRefCitation id="EFB74B32FF94FF86FE752611FDE99E68" author="Mikhaljova, E. V." box="[399,613,1030,1054]" pageId="27" pageNumber="28" pagination="3 - 36" refId="ref29575" refString="Mikhaljova, E. V. (1993) The millipedes (Diplopoda) of Siberia and the Far East of Russia. Arthropoda Selecta, 2 (2), 3 - 36." type="journal article" year="1993">Mikhaljova 1993: 18</bibRefCitation>
</taxonomicName>
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, figs.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936C3FF94FF86FF6D2633FBB59E4B" blockId="27.[151,1436,1030,1146]" box="[151,1081,1060,1085]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB3611EDFF94FF86FF6D2633FBB59E4B" box="[151,1081,1060,1085]" pageId="27" pageNumber="28">
<taxonomicName id="4C264D40FF94FF86FF6D2633FE059E4B" ID-CoL="36LVJ" authorityName="Mikhaljova" authorityYear="1993" box="[151,393,1060,1085]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cristofer">
<emphasis id="B952EAD1FF94FF86FF6D2633FE059E4B" box="[151,393,1060,1085]" italics="true" pageId="27" pageNumber="28">Diplomaragna cristofer</emphasis>
</taxonomicName>
—
<treatmentCitation id="0A8710D2FF94FF86FE582633FD099E4B" author="Mikhaljova, E. V." box="[418,645,1060,1085]" page="128" pageId="27" pageNumber="28" year="1997">
<bibRefCitation id="EFB74B32FF94FF86FE582633FD099E4B" author="Mikhaljova, E. V." box="[418,645,1060,1085]" pageId="27" pageNumber="28" pagination="123 - 130" refId="ref29657" refString="Mikhaljova, E. V. (1997) New data on the millipede genus Diplomaragna Attems, 1907, in the Far East of Russia (Diplopoda, Choredumatida, Diplomaragnidae). Arthropoda Selecta, 6 (1 / 2), 123 - 130." type="journal article" year="1997">Mikhaljova 1997: 128</bibRefCitation>
</treatmentCitation>
, figs; 1998: 36, figs, map;
<treatmentCitation id="0A8710D2FF94FF86FC6F2633FBB99E4B" author="Ganin, G. N." box="[917,1077,1060,1085]" page="10" pageId="27" pageNumber="28" year="1997">
<bibRefCitation id="EFB74B32FF94FF86FC6F2633FBB99E4B" author="Ganin, G. N." box="[917,1077,1060,1085]" pageId="27" pageNumber="28" refId="ref28495" refString="Ganin, G. N. (1997) Soil animals of the Ussuri region. Dalnauka Publisher, Vladivostok &amp; Khabarovsk, 159 pp. [in Russian, English summary]" type="book" year="1997">Ganin 1997: 10</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9936C3FF94FF86FF6D2654FEBF9E0C" blockId="27.[151,1436,1030,1146]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB3611EDFF94FF86FF6D2654FEBF9E0C" pageId="27" pageNumber="28">
<taxonomicName id="4C264D40FF94FF86FF6D2654FEF89E2D" ID-CoL="4BW6F" baseAuthorityName="Mikhaljova" baseAuthorityYear="1993" box="[151,372,1091,1115]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cristofer">
<emphasis id="B952EAD1FF94FF86FF6D2654FEF89E2D" box="[151,372,1091,1115]" italics="true" pageId="27" pageNumber="28">Pacifiosoma cristofer</emphasis>
</taxonomicName>
—
<treatmentCitation id="0A8710D2FF94FF86FE772654FDFF9E2D" author="Mikhaljova, E. V." box="[397,627,1091,1115]" page="169" pageId="27" pageNumber="28" year="2000">
<bibRefCitation id="EFB74B32FF94FF86FE772654FDFF9E2D" author="Mikhaljova, E. V." box="[397,627,1091,1115]" pageId="27" pageNumber="28" pagination="153 - 181" refId="ref29738" refString="Mikhaljova, E. V. (2000 [for 1999]) Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida). Arthropoda Selecta, 8 (3), 153 - 181." type="journal article" year="2000">Mikhaljova 2000: 169</bibRefCitation>
</treatmentCitation>
, figs; 2004: 154, figs, map; 2009c: 4; 2017: 190, figs, map;
<treatmentCitation id="0A8710D2FF94FF86FB122654FA149E2D" author="Ganin, G. N." box="[1256,1432,1091,1115]" page="153" pageId="27" pageNumber="28" year="2009">
<bibRefCitation id="EFB74B32FF94FF86FB122654FA149E2D" author="Ganin, G. N." box="[1256,1432,1091,1115]" pageId="27" pageNumber="28" pagination="147 - 156" refId="ref28576" refString="Ganin, G. N. (2009) Hypothetical factors of origin and coexisting of closely related soil-dwelling species. In: Storozhenko, S. Yu. (Ed.), A. I. Kurentsov's Annual Memorial Meetings. Issue XX. Dalnauka Publisher, Vladivostok, pp. 147 - 156. [in Russian]" type="book chapter" year="2009">Ganin 2009: 153</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8710D2FF94FF86FF3D2675FEA39E0C" author="Ganin, G. N." box="[199,303,1122,1146]" page="236" pageId="27" pageNumber="28" year="2011">
<bibRefCitation id="EFB74B32FF94FF86FF3D2675FEA39E0C" author="Ganin, G. N." box="[199,303,1122,1146]" pageId="27" pageNumber="28" refId="ref28637" refString="Ganin, G. N. (2011) Structural and functional organization of mezopedobiont communities of the Southern Russia Far East. Dalnauka Publisher, Vladivostok, 380 pp." type="book" year="2011">2011: 236</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<emphasis id="B952EAD1FF94FF86FF6D26B6FE0D9ECD" bold="true" box="[151,385,1185,1211]" pageId="27" pageNumber="28">Material examined.</emphasis>
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<specimenCount id="9D20FD4AFF94FF86FE7326B6FE599ECD" box="[393,469,1185,1211]" pageId="27" pageNumber="28" type="male">1 male</specimenCount>
(
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),
<collectingCountry id="F3317653FF94FF86FDB926B6FD1C9ECD" box="[579,656,1185,1211]" name="Russia" pageId="27" pageNumber="28">Russia</collectingCountry>
, environs of
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, near
<collectingMunicipality id="6BFDACB9FF94FF86FC0B26B6FB3F9ECD" box="[1009,1203,1185,1211]" pageId="27" pageNumber="28">Bychikha village</collectingMunicipality>
,
<location id="8EF96018FF94FF86FB4526B6FEDC9E96" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E:8EF96018FF94FF86FB4526B6FEDC9E96" country="Russia" municipality="Bychikha village" name="Bolshekhekhtsirsky Nature Reserve" pageId="27" pageNumber="28" stateProvince="Khabarovsk">Bolshekhekhtsirsky Nature Reserve</location>
, 48˚17.5´N 134˚49.7´E,
<date id="FF981003FF94FF86FD8126D2FCAB9E96" box="[635,807,1221,1248]" pageId="27" pageNumber="28" value="2005-09-11" valueMax="2005-09-13" valueMin="2005-09-11">
<collectingDate id="EFDCE9EBFF94FF86FD8126D2FCAB9E96" box="[635,807,1221,1248]" pageId="27" pageNumber="28" value="2005-09-11" valueMax="2005-09-13" valueMin="2005-09-11">11–13.IX.2005</collectingDate>
</date>
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;
<materialsCitation id="3B4E3C9EFF94FF86FCC126D2FE959F5E" ID-GBIF-Occurrence="3395035328" collectingDate="2005-09-14" collectionCode="FSCB" country="Russia" county="Bychikha village" location="Klyuch Sosninskiy" municipality="Bolshekhekhtsirsky Nature Reserve" pageId="27" pageNumber="28" specimenCount="3" specimenCount-male="3" stateProvince="Khabarovsk">
<specimenCount id="9D20FD4AFF94FF86FCC126D2FC159EA9" box="[827,921,1221,1247]" pageId="27" pageNumber="28" type="male">3 males</specimenCount>
(
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),
<collectingCountry id="F3317653FF94FF86FBEC26D2FBEF9EA9" box="[1046,1123,1221,1247]" name="Russia" pageId="27" pageNumber="28">Russia</collectingCountry>
, environs of
<collectingRegion id="49E2F821FF94FF86FAF426D2FA149EA9" box="[1294,1432,1221,1247]" country="Russia" name="Khabarovsk" pageId="27" pageNumber="28">Khabarovsk</collectingRegion>
, near
<collectingCounty id="62F84E4FFF94FF86FF2926FEFE169F75" box="[211,410,1257,1283]" pageId="27" pageNumber="28">Bychikha village</collectingCounty>
,
<collectingMunicipality id="6BFDACB9FF94FF86FE5026FEFCC49F72" box="[426,840,1257,1284]" pageId="27" pageNumber="28">Bolshekhekhtsirsky Nature Reserve</collectingMunicipality>
, 48˚14.56´N 134˚47´E,
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cordon,
<date id="FF981003FF94FF86FF6D271AFE959F5E" box="[151,281,1293,1320]" pageId="27" pageNumber="28" value="2005-09-14">
<collectingDate id="EFDCE9EBFF94FF86FF6D271AFE959F5E" box="[151,281,1293,1320]" pageId="27" pageNumber="28" value="2005-09-14">14.IX.2005</collectingDate>
</date>
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, all leg. Yu.M. Marusik;
<materialsCitation id="3B4E3C9EFF94FF86FDBB271AFD7A9F3D" ID-GBIF-Occurrence="3395035330" collectingDate="2011-07-31" collectingDateMax="2011-08-11" collectingDateMin="2011-07-31" collectionCode="FSCB" country="Russia" county="Khabarovsky Krai" location="Gassi Lake" municipality="Nanaisky District" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1">
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(
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),
<collectingCountry id="F3317653FF94FF86FCF9271AFCDC9F51" box="[771,848,1293,1319]" name="Russia" pageId="27" pageNumber="28">Russia</collectingCountry>
,
<collectingCounty id="62F84E4FFF94FF86FCA5271AFBB89F51" box="[863,1076,1293,1319]" pageId="27" pageNumber="28">Khabarovsky Krai</collectingCounty>
,
<collectingMunicipality id="6BFDACB9FF94FF86FBB8271AFA819F51" box="[1090,1293,1293,1319]" pageId="27" pageNumber="28">Nanaisky District</collectingMunicipality>
,
<location id="8EF96018FF94FF86FAE6271AFA109F51" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E:8EF96018FF94FF86FAE6271AFA109F51" box="[1308,1436,1293,1319]" country="Russia" county="Khabarovsky Krai" municipality="Nanaisky District" name="Gassi Lake" pageId="27" pageNumber="28">Gassi Lake</location>
basin, forest along
<location id="8EF96018FF94FF86FE912726FE759F3D" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E:8EF96018FF94FF86FE912726FE759F3D" box="[363,505,1329,1355]" country="Russia" county="Khabarovsky Krai" municipality="Nanaisky District" name="Khaso River" pageId="27" pageNumber="28">Khaso River</location>
,
<date id="FF981003FF94FF86FDF92726FD7A9F3D" box="[515,758,1329,1355]" pageId="27" pageNumber="28" value="2011-07-31" valueMax="2011-08-11" valueMin="2011-07-31">
<collectingDate id="EFDCE9EBFF94FF86FDF92726FD7A9F3D" box="[515,758,1329,1355]" pageId="27" pageNumber="28" value="2011-07-31" valueMax="2011-08-11" valueMin="2011-07-31">31.VII.–11.VIII.2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B4E3C9EFF94FF86FCFA2726FACF9F06" ID-GBIF-Occurrence="3395035312" collectingDate="2011-08-05" collectingDateMax="2011-08-09" collectingDateMin="2011-08-05" collectionCode="FSCB" country="Russia" county="Khabarovsky Krai" location="Gassi Lake" municipality="Nanaisky District" pageId="27" pageNumber="28" specimenCount="3" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D20FD4AFF94FF86FCFA2726FCC29F3D" box="[768,846,1329,1355]" pageId="27" pageNumber="28" type="male">1 male</specimenCount>
,
<specimenCount id="9D20FD4AFF94FF86FCA32726FC309F3D" box="[857,956,1329,1355]" pageId="27" pageNumber="28" type="female">1 female</specimenCount>
,
<specimenCount id="9D20FD4AFF94FF86FC3D2726FB899F3D" box="[967,1029,1329,1355]" pageId="27" pageNumber="28" type="generic">1 juv.</specimenCount>
(
<collectionCode id="ED37AE06FF94FF86FBE92726FBD19F3D" box="[1043,1117,1329,1355]" pageId="27" pageNumber="28">FSCB</collectionCode>
),
<collectingCountry id="F3317653FF94FF86FB952726FB309F3D" box="[1135,1212,1329,1355]" name="Russia" pageId="27" pageNumber="28">Russia</collectingCountry>
,
<collectingCounty id="62F84E4FFF94FF86FB322726FA149F3D" box="[1224,1432,1329,1355]" pageId="27" pageNumber="28">Khabarovsky Krai</collectingCounty>
,
<collectingMunicipality id="6BFDACB9FF94FF86FF6D2742FED39F19" box="[151,351,1365,1391]" pageId="27" pageNumber="28">Nanaisky District</collectingMunicipality>
,
<location id="8EF96018FF94FF86FE912742FE659F19" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E:8EF96018FF94FF86FE912742FE659F19" box="[363,489,1365,1391]" country="Russia" county="Khabarovsky Krai" municipality="Nanaisky District" name="Gassi Lake" pageId="27" pageNumber="28">Gassi Lake</location>
basin,
<taxonomicName id="4C264D40FF94FF86FDC02742FD0F9F19" box="[570,643,1365,1391]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="B952EAD1FF94FF86FDC02742FD0F9F19" box="[570,643,1365,1391]" italics="true" pageId="27" pageNumber="28">Betula</emphasis>
</taxonomicName>
with
<taxonomicName id="4C264D40FF94FF86FD3E2741FC889F19" box="[708,772,1366,1391]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="B952EAD1FF94FF86FD3E2741FC889F19" box="[708,772,1366,1391]" italics="true" pageId="27" pageNumber="28">Picea</emphasis>
</taxonomicName>
forest near
<location id="8EF96018FF94FF86FC732742FBA09F19" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E:8EF96018FF94FF86FC732742FBA09F19" box="[905,1068,1365,1391]" country="Russia" county="Khabarovsky Krai" municipality="Nanaisky District" name="Tukhala River" pageId="27" pageNumber="28">Tukhala River</location>
, 337.7 m,
<date id="FF981003FF94FF86FB582742FACF9F06" box="[1186,1347,1365,1392]" pageId="27" pageNumber="28" value="2011-08-05" valueMax="2011-08-09" valueMin="2011-08-05">
<collectingDate id="EFDCE9EBFF94FF86FB582742FACF9F06" box="[1186,1347,1365,1392]" pageId="27" pageNumber="28" value="2011-08-05" valueMax="2011-08-09" valueMin="2011-08-05">5–9.VIII.2011</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4E3C9EFF94FF86FAAA2742FEA99FE5" ID-GBIF-Occurrence="3395035305" collectionCode="V" country="Russia" location="Budilov" pageId="27" pageNumber="28" specimenCount="1">
all leg. P.
<collectionCode id="ED37AE06FF94FF86FF50276DFF339FE2" box="[170,191,1402,1428]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="27" pageNumber="28" type="Museum">V</collectionCode>
.
<location id="8EF96018FF94FF86FF33276EFEA99FE5" LSID="urn:lsid:plazi:treatment:038F87D5FF94FF86FF6D21B4FBB39D8E:8EF96018FF94FF86FF33276EFEA99FE5" box="[201,293,1401,1427]" country="Russia" name="Budilov" pageId="27" pageNumber="28">Budilov</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF94FF86FF3D278AFC819FCE" box="[199,781,1437,1464]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8B9936C3FF94FF86FF3D278AFC819FCE" blockId="27.[151,1437,1185,2040]" box="[199,781,1437,1464]" pageId="27" pageNumber="28">
<emphasis id="B952EAD1FF94FF86FF3D278AFEED9FC1" bold="true" box="[199,353,1437,1463]" pageId="27" pageNumber="28">Distribution.</emphasis>
<collectingCountry id="F3317653FF94FF86FE9D278AFE3A9FC1" box="[359,438,1437,1463]" name="Russia" pageId="27" pageNumber="28">Russia</collectingCountry>
: Far East (Khabarovsky Krai).
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF94FF86FF3D27D6FBB39D8E" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8B9936C3FF94FF86FF3D27D6FEFD9C31" blockId="27.[151,1437,1185,2040]" pageId="27" pageNumber="28">
<emphasis id="B952EAD1FF94FF86FF3D27D6FEB79FAD" bold="true" box="[199,315,1473,1499]" pageId="27" pageNumber="28">Remarks.</emphasis>
This species was originally described in
<taxonomicName id="4C264D40FF94FF86FD0127D6FC2C9FAD" authorityName="Attems" authorityYear="1907" box="[763,928,1473,1499]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B952EAD1FF94FF86FD0127D6FC2C9FAD" box="[763,928,1473,1499]" italics="true" pageId="27" pageNumber="28">Diplomaragna</emphasis>
</taxonomicName>
from the Ulchsky District of the Khabarovsky Krai and the environs of
<collectingRegion id="49E2F821FF94FF86FE5327F2FDBC9F89" box="[425,560,1509,1535]" country="Russia" name="Khabarovsk" pageId="27" pageNumber="28">Khabarovsk</collectingRegion>
(
<bibRefCitation id="EFB74B32FF94FF86FDC727F2FC8D9F89" author="Mikhaljova, E. V." box="[573,769,1509,1535]" pageId="27" pageNumber="28" pagination="3 - 36" refId="ref29575" refString="Mikhaljova, E. V. (1993) The millipedes (Diplopoda) of Siberia and the Far East of Russia. Arthropoda Selecta, 2 (2), 3 - 36." type="journal article" year="1993">Mikhaljova 1993</bibRefCitation>
), but later transferred to the genus
<taxonomicName id="4C264D40FF94FF86FB7A27F2FF509C55" authority="(Mikhaljova 2000)" baseAuthorityName="Mikhaljova" baseAuthorityYear="2000" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B952EAD1FF94FF86FB7A27F2FA809F89" box="[1152,1292,1509,1535]" italics="true" pageId="27" pageNumber="28">Pacifiosoma</emphasis>
(
<bibRefCitation id="EFB74B32FF94FF86FAE327F2FF589C55" author="Mikhaljova, E. V." pageId="27" pageNumber="28" pagination="153 - 181" refId="ref29738" refString="Mikhaljova, E. V. (2000 [for 1999]) Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida). Arthropoda Selecta, 8 (3), 153 - 181." type="journal article" year="2000">Mikhaljova 2000</bibRefCitation>
)
</taxonomicName>
. Based on
<typeStatus id="549D8861FF94FF86FE9D241DFE1B9C52" box="[359,407,1546,1572]" pageId="27" pageNumber="28">type</typeStatus>
material, the gonopod structure of this species has been redescribed more adequately (
<bibRefCitation id="EFB74B32FF94FF86FF65243AFEE99C31" author="Mikhaljova, E. V." box="[159,357,1581,1607]" pageId="27" pageNumber="28" pagination="123 - 130" refId="ref29657" refString="Mikhaljova, E. V. (1997) New data on the millipede genus Diplomaragna Attems, 1907, in the Far East of Russia (Diplopoda, Choredumatida, Diplomaragnidae). Arthropoda Selecta, 6 (1 / 2), 123 - 130." type="journal article" year="1997">Mikhaljova 1997</bibRefCitation>
).
</paragraph>
<paragraph id="8B9936C3FF94FF86FF3D2446FAAC9CC5" blockId="27.[151,1437,1185,2040]" pageId="27" pageNumber="28">
The study of the above fresh material has revealed the morphological variability of the structure of the gonopod posterior angiocoxal processes. Thus, the processes in males from the environs of Gassi Lake (
<emphasis id="B952EAD1FF94FF86FB4A2462FB439CF9" bold="true" box="[1200,1231,1653,1679]" pageId="27" pageNumber="28">pp</emphasis>
,
<figureCitation id="131D2A46FF94FF86FB272462FAA09CF9" box="[1245,1324,1653,1679]" captionStart="FIGURES 40–44" captionStartId="28.[151,264,1476,1500]" captionTargetBox="[151,1435,181,1452]" captionTargetId="figure-24@28.[151,1435,181,1453]" captionTargetPageId="28" captionText="FIGURES 40–44. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 40–41, 44) and Bolshekhekhtsirsky Nature Reserve (Figs 42–43). 40, 42, gonopods, caudal view, telopodites of posterior gonopods broken on fig. 42; 41, central part of gonopods; 43, distal parts of gonopods; 44, apex of colpocoxite and anterior gonopod telopodite; pp, posterior angicoxal process; aat, apex of anterior gonopod telopodite. Scales: 10 μm (Fig. 44), 20 μm (Figs 41, 43), 100 μm (Figs 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5606705" httpUri="https://zenodo.org/record/5606705/files/figure.png" pageId="27" pageNumber="28">Fig. 41</figureCitation>
) are more long and slender as compared ones in males from the Bolshekhekhtsirsky Nature Reserve (
<emphasis id="B952EAD1FF94FF86FB69248EFB3E9CC5" bold="true" box="[1171,1202,1689,1715]" pageId="27" pageNumber="28">pp</emphasis>
,
<figureCitation id="131D2A46FF94FF86FB3A248EFA9F9CC5" box="[1216,1299,1689,1715]" captionStart="FIGURES 40–44" captionStartId="28.[151,264,1476,1500]" captionTargetBox="[151,1435,181,1452]" captionTargetId="figure-24@28.[151,1435,181,1453]" captionTargetPageId="28" captionText="FIGURES 40–44. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 40–41, 44) and Bolshekhekhtsirsky Nature Reserve (Figs 42–43). 40, 42, gonopods, caudal view, telopodites of posterior gonopods broken on fig. 42; 41, central part of gonopods; 43, distal parts of gonopods; 44, apex of colpocoxite and anterior gonopod telopodite; pp, posterior angicoxal process; aat, apex of anterior gonopod telopodite. Scales: 10 μm (Fig. 44), 20 μm (Figs 41, 43), 100 μm (Figs 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5606705" httpUri="https://zenodo.org/record/5606705/files/figure.png" pageId="27" pageNumber="28">Fig. 42</figureCitation>
).
</paragraph>
<paragraph id="8B9936C3FF94FF86FF3D24A9FBF99DC6" blockId="27.[151,1437,1185,2040]" pageId="27" pageNumber="28">
In addition, the study of the gonopods using a scanning electron microscope has allowed to clarify the detail structure of gonopods: the telopodite of the anterior gonopod is 1-segmented (joints 1 and 2 are only separated by a deep constriction, not by an intersegmental suture), subflagelliform, with a cristate apex (
<emphasis id="B952EAD1FF94FF86FB4C2512FB579D69" bold="true" box="[1206,1243,1797,1823]" pageId="27" pageNumber="28">aat</emphasis>
,
<figureCitation id="131D2A46FF94FF86FB162512FACC9D69" box="[1260,1344,1797,1823]" captionStart="FIGURES 40–44" captionStartId="28.[151,264,1476,1500]" captionTargetBox="[151,1435,181,1452]" captionTargetId="figure-24@28.[151,1435,181,1453]" captionTargetPageId="28" captionText="FIGURES 40–44. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 40–41, 44) and Bolshekhekhtsirsky Nature Reserve (Figs 42–43). 40, 42, gonopods, caudal view, telopodites of posterior gonopods broken on fig. 42; 41, central part of gonopods; 43, distal parts of gonopods; 44, apex of colpocoxite and anterior gonopod telopodite; pp, posterior angicoxal process; aat, apex of anterior gonopod telopodite. Scales: 10 μm (Fig. 44), 20 μm (Figs 41, 43), 100 μm (Figs 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5606705" httpUri="https://zenodo.org/record/5606705/files/figure.png" pageId="27" pageNumber="28">Fig. 44</figureCitation>
) visible as stretched beyond sheath groove; the sheath groove longitudinally splits the distal end of posterior gonopod colpocoxite (
<emphasis id="B952EAD1FF94FF86FED1255AFEB39D11" bold="true" box="[299,319,1869,1895]" pageId="27" pageNumber="28">cl</emphasis>
,
<figureCitation id="131D2A46FF94FF86FEB5255AFE2C9D11" box="[335,416,1869,1895]" captionStart="FIGURES 45–48" captionStartId="29.[151,264,1004,1028]" captionTargetBox="[151,1435,181,980]" captionTargetId="figure-16@29.[151,1435,181,980]" captionTargetPageId="29" captionText="FIGURES 45–48. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 45–46) and Bolshekhekhtsirsky Nature Reserve (Figs 47–48). 45, 47, gonopods, front view; 46, 48, distal parts of colpocoxites; ap, anterior angiocoxal process; cl, colpocoxites. Scales: 20 μm (Figs 46, 48), 100 μm (Figs 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5606709" httpUri="https://zenodo.org/record/5606709/files/figure.png" pageId="27" pageNumber="28">Fig. 47</figureCitation>
) into two tightly adjacent parts, so that the unciform apices of the colpocoxites look bifid (
<figureCitation id="131D2A46FF94FF86FF652566FEAE9DFD" box="[159,290,1905,1931]" captionStart="FIGURES 45–48" captionStartId="29.[151,264,1004,1028]" captionTargetBox="[151,1435,181,980]" captionTargetId="figure-16@29.[151,1435,181,980]" captionTargetPageId="29" captionText="FIGURES 45–48. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 45–46) and Bolshekhekhtsirsky Nature Reserve (Figs 47–48). 45, 47, gonopods, front view; 46, 48, distal parts of colpocoxites; ap, anterior angiocoxal process; cl, colpocoxites. Scales: 20 μm (Figs 46, 48), 100 μm (Figs 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5606709" httpUri="https://zenodo.org/record/5606709/files/figure.png" pageId="27" pageNumber="28">Figs 46, 48</figureCitation>
); the colpocoxite posterior surface is finely papillate/spinulate (
<figureCitation id="131D2A46FF94FF86FC102566FBE29DFD" box="[1002,1134,1905,1931]" captionStart="FIGURES 40–44" captionStartId="28.[151,264,1476,1500]" captionTargetBox="[151,1435,181,1452]" captionTargetId="figure-24@28.[151,1435,181,1453]" captionTargetPageId="28" captionText="FIGURES 40–44. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 40–41, 44) and Bolshekhekhtsirsky Nature Reserve (Figs 42–43). 40, 42, gonopods, caudal view, telopodites of posterior gonopods broken on fig. 42; 41, central part of gonopods; 43, distal parts of gonopods; 44, apex of colpocoxite and anterior gonopod telopodite; pp, posterior angicoxal process; aat, apex of anterior gonopod telopodite. Scales: 10 μm (Fig. 44), 20 μm (Figs 41, 43), 100 μm (Figs 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5606705" httpUri="https://zenodo.org/record/5606705/files/figure.png" pageId="27" pageNumber="28">Figs 42–43</figureCitation>
); the apex of the dentiform anterior angiocoxal processes (
<emphasis id="B952EAD1FF94FF86FE092582FD9D9DD9" bold="true" box="[499,529,1941,1967]" pageId="27" pageNumber="28">ap</emphasis>
,
<figureCitation id="131D2A46FF94FF86FDE42582FD119DC6" box="[542,669,1941,1968]" captionStart="FIGURES 45–48" captionStartId="29.[151,264,1004,1028]" captionTargetBox="[151,1435,181,980]" captionTargetId="figure-16@29.[151,1435,181,980]" captionTargetPageId="29" captionText="FIGURES 45–48. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 45–46) and Bolshekhekhtsirsky Nature Reserve (Figs 47–48). 45, 47, gonopods, front view; 46, 48, distal parts of colpocoxites; ap, anterior angiocoxal process; cl, colpocoxites. Scales: 20 μm (Figs 46, 48), 100 μm (Figs 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5606709" httpUri="https://zenodo.org/record/5606709/files/figure.png" pageId="27" pageNumber="28">Figs 45, 46</figureCitation>
) are covered with tiny tubercles (
<figureCitation id="131D2A46FF94FF86FBEE2582FBE59DC6" box="[1044,1129,1941,1968]" captionStart="FIGURES 45–48" captionStartId="29.[151,264,1004,1028]" captionTargetBox="[151,1435,181,980]" captionTargetId="figure-16@29.[151,1435,181,980]" captionTargetPageId="29" captionText="FIGURES 45–48. Pacifiosoma cristofer (Mikhaljova, 1993), males from Gassi Lake basin (Figs 45–46) and Bolshekhekhtsirsky Nature Reserve (Figs 47–48). 45, 47, gonopods, front view; 46, 48, distal parts of colpocoxites; ap, anterior angiocoxal process; cl, colpocoxites. Scales: 20 μm (Figs 46, 48), 100 μm (Figs 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5606709" httpUri="https://zenodo.org/record/5606709/files/figure.png" pageId="27" pageNumber="28">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="8B9936C3FF94FF86FF3D25ADFBB39D8E" blockId="27.[151,1437,1185,2040]" pageId="27" pageNumber="28">
It should be mentioned that
<taxonomicName id="4C264D40FF94FF86FDF825AEFD6B9DA5" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[514,743,1977,2003]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="acutum" status="sp. nov.">
<emphasis id="B952EAD1FF94FF86FDF825AEFD6B9DA5" box="[514,743,1977,2003]" italics="true" pageId="27" pageNumber="28">Pacifiosoma acutum</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF94FF86FD1525AEFCC59DA5" bold="true" box="[751,841,1977,2003]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A26157AAFF94FF86FD1525AEFCC59DA5" box="[751,841,1977,2003]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was also found near Sofiysk, Ulchsky District of the Khabarovskiy Krai, i.e. within the locality of the
<typeStatus id="549D8861FF94FF86FD4525CAFCAD9D81" box="[703,801,2013,2039]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C264D40FF94FF86FCBC25CAFBB49D81" baseAuthorityName="Mikhaljova" baseAuthorityYear="1993" box="[838,1080,2013,2039]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cristofer">
<emphasis id="B952EAD1FF94FF86FCBC25CAFBB49D81" box="[838,1080,2013,2039]" italics="true" pageId="27" pageNumber="28">Pacifiosoma cristofer</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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