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<document id="C2603ECE7D54F202F2568FC8F9FDE714" ID-CLB-Dataset="20418" ID-DOI="10.11646/megataxa.7.1.1" ID-GBIF-Dataset="bd1f1932-a0fa-4daf-af9f-cc52c932ed40" ID-ISSN="2703-3090" ID-Zenodo-Dep="5964441" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1643698274876" checkinUser="plazi" docAuthor="Iverson, John B." docDate="2022" docId="038F8788FFA0FFC8DAD1FD7CFB98230C" docLanguage="en" docName="44551-Article.Text-47635-51294-10-20220125.pdf.imf" docOrigin="Megataxa 7 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Manouria emys" docType="treatment" docVersion="11" lastPageNumber="55" masterDocId="FFB6FFF0FF96FFFEDA59FB57FF95276A" masterDocTitle="A review of Chelonian type specimens (order Testudines)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="55" updateTime="1732571001862" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="462DA06F8D1E5D639DBA50294562A842" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F96F3CC34F98205277FED3D9D7B3F852">A review of Chelonian type specimens (order Testudines)</mods:title>
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<mods:namePart id="5431773352CE3A2ECF95E31273706E66">Iverson, John B.</mods:namePart>
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<mods:title id="19CF454CBA58D830CD295F3A5D24227B">Megataxa</mods:title>
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<mods:date id="1D26239221CB3F2EA257F5AFE41A2F8A">2022</mods:date>
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<mods:identifier id="8C9A88CAA1519F4BC13FCE4FA5DBCF24" type="DOI">10.11646/megataxa.7.1.1</mods:identifier>
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<treatment id="038F8788FFA0FFC8DAD1FD7CFB98230C" ID-DOI="http://doi.org/10.5281/zenodo.6205126" ID-GBIF-Taxon="193856392" ID-Zenodo-Dep="6205126" LSID="urn:lsid:plazi:treatment:038F8788FFA0FFC8DAD1FD7CFB98230C" httpUri="http://treatment.plazi.org/id/038F8788FFA0FFC8DAD1FD7CFB98230C" lastPageNumber="55" pageId="54" pageNumber="55" scope_class="Testudines" scope_family="Testudinidae" scope_subOrder="Cryptodira">
<subSubSection id="C33C6515FFA0FFC8DAD1FD7CFD05202C" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="8B99369EFFA0FFC8DAD1FD7CFD00212E" blockId="54.[136,778,1578,1862]" box="[136,661,1578,1605]" pageId="54" pageNumber="55">
<heading id="D0D181F2FFA0FFC8DAD1FD7CFD00212E" box="[136,661,1578,1605]" fontSize="11" level="2" pageId="54" pageNumber="55" reason="2">
<taxonomicName id="4C264D1DFFA0FFC8DAD1FD7CFD00212E" ID-CoL="3XW2D" authority="(Schlegel &amp; Muller 1845: 34)" baseAuthorityName="Schlegel &amp; Muller" baseAuthorityPageNumber="34" baseAuthorityYear="1845" box="[136,661,1578,1605]" class="Reptilia" family="Testudinidae" genus="Manouria" kingdom="Animalia" order="Testudines" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="emys">
<emphasis id="B952EA8CFFA0FFC8DAD1FD7CFD00212E" bold="true" box="[136,661,1578,1605]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8DAD1FD7CFEAB212F" bold="true" box="[136,318,1579,1605]" italics="true" pageId="54" pageNumber="55">Manouria emys</emphasis>
(Schlegel &amp; Müller 1845:34)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B99369EFFA0FFC8DAD1FD1DFD05202C" blockId="54.[136,778,1578,1862]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8DAD1FD1DFF66210E" bold="true" box="[136,243,1610,1636]" pageId="54" pageNumber="55">
<typeStatus id="549D883CFFA0FFC8DAD1FD1DFF66210E" box="[136,243,1610,1636]" pageId="54" pageNumber="55" type="syntype">Syntypes</typeStatus>
</emphasis>
(6): MNHN 9422; RMNH 3808, 6005, 6030 (illustrated in
<bibRefCitation id="EFB74B6FFFA0FFC8DBDEFD3CFDE121EF" author="Lidth de Jeude, T. W. van" box="[391,628,1643,1669]" pageId="54" pageNumber="55" pagination="197 - 204" refId="ref76389" refString="Lidth de Jeude, T. W. van (1896) On Testudo emys Schleg. and Mull. and its affinities. Notes Leyden Museum, 17, 197 - 204." type="journal article" year="1896">Lidth de Jeude 1896</bibRefCitation>
:Pl. 5), 17967 (illustrated in
<bibRefCitation id="EFB74B6FFFA0FFC8DBCBFDDCFD1A21CF" author="Lidth de Jeude, T. W. van" box="[402,655,1675,1701]" pageId="54" pageNumber="55" pagination="197 - 204" refId="ref76389" refString="Lidth de Jeude, T. W. van (1896) On Testudo emys Schleg. and Mull. and its affinities. Notes Leyden Museum, 17, 197 - 204." type="journal article" year="1896">Lidth de Jeude 1896</bibRefCitation>
:Pl. 5; and photographed in
<bibRefCitation id="EFB74B6FFFA0FFC8DBF1FDFCFD5921AF" author="Hoogmoed, M. S. &amp; Gasso Miracle, M. E. &amp; van den Hoek Ostende, L. W." box="[424,716,1707,1733]" pageId="54" pageNumber="55" pagination="1 - 41" refId="ref74490" refString="Hoogmoed, M. S., Gasso Miracle, M. E. &amp; van den Hoek Ostende, L. W. (2010) Type specimens of recent and fossil Testudines and Crocodylia in the collections of the Netherlands Centre for Biodiversity Naturalis, Leiden, the Netherlands. Zoologische Mededelingen Leiden, 84, 1 - 41." type="journal article" year="2010">
Hoogmoed
<emphasis id="B952EA8CFFA0FFC8D875FDFBFDF021AF" box="[556,613,1707,1733]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2010:18
</bibRefCitation>
), and one unlocated
<typeStatus id="549D883CFFA0FFC8DBDAFD9CFE4F218F" box="[387,474,1739,1765]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
(
<bibRefCitation id="EFB74B6FFFA0FFC8DBBCFD9CFD68218F" author="Hoogmoed, M. S. &amp; Gasso Miracle, M. E. &amp; van den Hoek Ostende, L. W." box="[485,765,1739,1765]" pageId="54" pageNumber="55" pagination="1 - 41" refId="ref74490" refString="Hoogmoed, M. S., Gasso Miracle, M. E. &amp; van den Hoek Ostende, L. W. (2010) Type specimens of recent and fossil Testudines and Crocodylia in the collections of the Netherlands Centre for Biodiversity Naturalis, Leiden, the Netherlands. Zoologische Mededelingen Leiden, 84, 1 - 41." type="journal article" year="2010">
Hoogmoed
<emphasis id="B952EA8CFFA0FFC8D83FFD9BFD0F218F" box="[614,666,1739,1765]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2010:19
</bibRefCitation>
); RMNH 3808 designated
<typeStatus id="549D883CFFA0FFC8DBA7FDBCFDF0206F" box="[510,613,1771,1797]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="EFB74B6FFFA0FFC8D8D4FDBCFE4D204C" author="Hoogmoed, M. S. &amp; Crumly, C. R." pageId="54" pageNumber="55" pagination="241 - 259" refId="ref74391" refString="Hoogmoed, M. S. &amp; Crumly, C. R. (1984) Land tortoise types in the Rijksmuseum van Natuurlijke Historie with comments on nomenclature and systematics (Reptilia: Testudines: Testudinidae). Zoologische Mededelingen, Leiden, 58, 241 - 259." type="journal article" year="1984">Hoogmoed &amp; Crumly (1984:251</bibRefCitation>
, and photographed therein, p. 253, Fig. 6, and in
<bibRefCitation id="EFB74B6FFFA0FFC8DB81FC7CFD16202C" author="Bour, R." box="[472,643,1835,1862]" pageId="54" pageNumber="55" pagination="25 - 40" refId="ref66449" refString="Bour, R. (1998 a) Histoire du genre Manouria Gray, 1854, et des especes actuelles incluses. Manouria, 1, 25 - 40." type="journal article" year="1998">Bour 1998a:26</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33C6515FFA0FFC8DAF2FC3BFA8B254F" pageId="54" pageNumber="55" type="description">
<paragraph id="8B99369EFFA0FFC8DAF2FC3BFD9820EC" blockId="54.[171,754,1899,1990]" box="[171,525,1899,1926]" pageId="54" pageNumber="55">
<taxonomicName id="4C264D1DFFA0FFC8DAF2FC3BFD9820EC" authority="Gray 1854: 134" authorityName="Gray" authorityYear="1854" box="[171,525,1899,1926]" class="Reptilia" family="Testudinidae" genus="Manouria" kingdom="Animalia" order="Testudines" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="fusca">
<emphasis id="B952EA8CFFA0FFC8DAF2FC3BFEC820EF" box="[171,349,1899,1925]" italics="true" pageId="54" pageNumber="55">Manouria fusca</emphasis>
Gray 1854:134
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFFA0FFC8DAF2FCDCFB7227AC" blockId="54.[171,754,1899,1990]" lastBlockId="54.[845,1452,171,293]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8DAF2FCDCFE8D20CF" bold="true" box="[171,280,1931,1957]" pageId="54" pageNumber="55">Holoype:</emphasis>
BMNH 1947.3.4.57 (illustrated in
<bibRefCitation id="EFB74B6FFFA0FFC8D8FAFCDCFEB020AC" author="Gray, J. E. &amp; Catalogue of Shield Reptiles &amp; Collection of the British Museum &amp; Part I." pageId="54" pageNumber="55" refId="ref73300" refString="Gray, J. E. (1856) [&quot; 1855 &quot;]. Catalogue of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). British Museum, London, 79 pp. [Published March 1856]." type="book" year="1856">Gray 1856</bibRefCitation>
:Pl. 4, and reproduced in
<bibRefCitation id="EFB74B6FFFA0FFC8D863FCFCFD7020AF" author="Bour, R." box="[570,741,1963,1989]" pageId="54" pageNumber="55" pagination="25 - 40" refId="ref66449" refString="Bour, R. (1998 a) Histoire du genre Manouria Gray, 1854, et des especes actuelles incluses. Manouria, 1, 25 - 40." type="journal article" year="1998">Bour 1998a:27</bibRefCitation>
).
<taxonomicName id="4C264D1DFFA0FFC8D914FBFCFB7227AC" authority="Le Conte 1854: 187" authorityName="Le Conte" authorityPageNumber="187" authorityYear="1854" box="[845,1255,171,198]" class="Reptilia" family="Testudinidae" genus="Teleopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="luxatus">
<emphasis id="B952EA8CFFA0FFC8D914FBFCFB9227AF" box="[845,1031,171,197]" italics="true" pageId="54" pageNumber="55">Teleopus luxatus</emphasis>
Le Conte 1854:187
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFFA0FFC8D914FB9DFC47264F" blockId="54.[845,1452,171,293]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8D914FB9DFC2D278E" bold="true" box="[845,952,202,228]" pageId="54" pageNumber="55">
<typeStatus id="549D883CFFA0FFC8D914FB9DFC2D278E" box="[845,952,202,228]" pageId="54" pageNumber="55" type="syntype">Syntypes</typeStatus>
</emphasis>
(number uncertain): Originally in the ANSP (
<bibRefCitation id="EFB74B6FFFA0FFC8D9C8FBBCFB80266F" author="Baur, G." box="[913,1045,235,261]" pageId="54" pageNumber="55" pagination="155 - 159" refId="ref65207" refString="Baur, G. (1892) Bemerkungen uber verschiedene Arten von Schildkroten. Zoologischer Anzeiger, 15, 155 - 159." type="journal article" year="1892">Baur 1892</bibRefCitation>
), but no longer present (
<bibRefCitation id="EFB74B6FFFA0FFC8DF17FBBCFC53264F" author="Malnate, E. V." pageId="54" pageNumber="55" pagination="345 - 375" refId="ref77071" refString="Malnate, E. V. (1971) A catalog of primary types in the herpetological collections of the Academy of Natural Sciences, Philadelphia (ANSP). Proceedings of the Academy of Natural Sciences of Philadelphia, 123 (9), 345 - 375." type="journal article" year="1971">Malnate 1971</bibRefCitation>
).
</paragraph>
<paragraph id="8B99369EFFA0FFC8D973FA1CFA9F260E" blockId="54.[810,1451,330,549]" box="[810,1290,330,357]" pageId="54" pageNumber="55">
<taxonomicName id="4C264D1DFFA0FFC8D973FA1CFA9F260E" authority="(Blyth 1853: 639)" baseAuthorityName="Blyth" baseAuthorityPageNumber="639" baseAuthorityYear="1853" box="[810,1290,330,357]" class="Reptilia" family="Testudinidae" genus="Manouria" kingdom="Animalia" order="Testudines" pageId="54" pageNumber="55" phylum="Chordata" rank="subSpecies" species="emys" subSpecies="phayrei">
<emphasis id="B952EA8CFFA0FFC8D973FA1CFA9F260E" bold="true" box="[810,1290,330,357]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8D973FA1CFBA8260F" bold="true" box="[810,1085,330,357]" italics="true" pageId="54" pageNumber="55">Manouria emys phayrei</emphasis>
(Blyth 1853:639)
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFFA0FFC8D973FA3DFA8B254F" blockId="54.[810,1451,330,549]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8D973FA3DFC0026EE" bold="true" box="[810,917,362,388]" pageId="54" pageNumber="55">
<typeStatus id="549D883CFFA0FFC8D973FA3DFC0026EE" box="[810,917,362,388]" pageId="54" pageNumber="55" type="syntype">Syntypes</typeStatus>
</emphasis>
(2;
<bibRefCitation id="EFB74B6FFFA0FFC8D99CFA3CFB3226EF" author="Das, I. &amp; Dattagupta, B. &amp; Gayen, N. C." box="[965,1191,363,389]" pageId="54" pageNumber="55" pagination="121 - 172" refId="ref69833" refString="Das, I., Dattagupta, B. &amp; Gayen, N. C. (1998) History and catalogue of reptile types in the collection of the Zoological Survey of India, Calcutta. Journal of South Asian Natural History, 3 (2), 121 - 172." type="journal article" year="1998">
Das
<emphasis id="B952EA8CFFA0FFC8D9A0FA3BFBA726EF" box="[1017,1074,363,389]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
1998:127
</bibRefCitation>
;
<bibRefCitation id="EFB74B6FFFA0FFC8DEEDFA3CFAAF26EF" author="Das, I." box="[1204,1338,363,389]" pageId="54" pageNumber="55" pagination="1 - 14" refId="ref69800" refString="Das, I. (2009) The discovery of Indian turtles, with notes on publications, type localities and type repositories. ENVIS Bulletin, 12, 1 - 14." type="journal article" year="2009">Das 2009:4</bibRefCitation>
): ZSI 813 (photographed in
<bibRefCitation id="EFB74B6FFFA0FFC8DE3DFADCFAF026CF" author="Kundu, S. &amp; Kumar, V. &amp; Murthy, B. H. C. &amp; Chandra, K." box="[1124,1381,395,421]" pageId="54" pageNumber="55" refId="ref75926" refString="Kundu, S., Kumar, V., Murthy, B. H. C. K &amp; Chandra, K. (2018) Chelonian types of National Zoological Collections. Zoological Survey of India, Kolkata, 70 pp. https: // doi. org / 10.26515 / rzsi / v 118 / i 1 / 2018 / 122300" type="book" year="2018">
Kundu
<emphasis id="B952EA8CFFA0FFC8DE98FADBFA9126CF" box="[1217,1284,395,421]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2018:8
</bibRefCitation>
), and 15492 (
<bibRefCitation id="EFB74B6FFFA0FFC8D986FAFCFB5226AC" author="Kundu, S. &amp; Kumar, V. &amp; Murthy, B. H. C. &amp; Chandra, K." box="[991,1223,427,454]" pageId="54" pageNumber="55" refId="ref75926" refString="Kundu, S., Kumar, V., Murthy, B. H. C. K &amp; Chandra, K. (2018) Chelonian types of National Zoological Collections. Zoological Survey of India, Kolkata, 70 pp. https: // doi. org / 10.26515 / rzsi / v 118 / i 1 / 2018 / 122300" type="book" year="2018">
Kundu
<emphasis id="B952EA8CFFA0FFC8DE6CFAFBFBFB26AF" box="[1077,1134,427,453]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2018:6
</bibRefCitation>
). A skull illustrated in
<bibRefCitation id="EFB74B6FFFA0FFC8D9FBFA9CFBCD268F" author="Anderson, J." box="[930,1112,459,485]" pageId="54" pageNumber="55" pagination="132 - 144" refId="ref64173" refString="Anderson, J. (1872) On Manouria and Scapia, two genera of landtortoises. Proceedings of the Zoological Society of London, 1872, 132 - 144." type="journal article" year="1872">Anderson (1872</bibRefCitation>
; reproduced in
<bibRefCitation id="EFB74B6FFFA0FFC8DEA4FA9CFA36268F" author="Bour, R." box="[1277,1443,459,485]" pageId="54" pageNumber="55" pagination="25 - 40" refId="ref66449" refString="Bour, R. (1998 a) Histoire du genre Manouria Gray, 1854, et des especes actuelles incluses. Manouria, 1, 25 - 40." type="journal article" year="1998">Bour 1998a:29</bibRefCitation>
) was recorded as the ‘type’ (i.e.,
<typeStatus id="549D883CFFA0FFC8DF4BFABCFAEE256C" box="[1298,1403,491,518]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
) by Anderson (see also
<bibRefCitation id="EFB74B6FFFA0FFC8DE3FF95CFA84254F" author="Bour, R." box="[1126,1297,523,549]" pageId="54" pageNumber="55" pagination="25 - 40" refId="ref66449" refString="Bour, R. (1998 a) Histoire du genre Manouria Gray, 1854, et des especes actuelles incluses. Manouria, 1, 25 - 40." type="journal article" year="1998">Bour 1998a:28</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33C6515FFA0FFC8D914F91CFB54240F" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="8B99369EFFA0FFC8D914F91CFAB1250F" blockId="54.[845,1458,587,870]" box="[845,1316,587,613]" pageId="54" pageNumber="55">
<taxonomicName id="4C264D1DFFA0FFC8D914F91CFAB1250F" authority="Gray 1869: 169" authorityName="Gray" authorityPageNumber="169" authorityYear="1869" box="[845,1316,587,613]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="falconeri" subGenus="Scapia">
<emphasis id="B952EA8CFFA0FFC8D914F91CFBE0250F" box="[845,1141,587,613]" italics="true" pageId="54" pageNumber="55">Testudo (Scapia) falconeri</emphasis>
Gray 1869:169
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFFA0FFC8D914F93DFB54240F" blockId="54.[845,1458,587,870]" pageId="54" pageNumber="55">
<emphasis id="B952EA8CFFA0FFC8D914F93DFC2D25EE" bold="true" box="[845,952,618,644]" pageId="54" pageNumber="55">
<typeStatus id="549D883CFFA0FFC8D914F93DFC2D25EE" box="[845,952,618,644]" pageId="54" pageNumber="55" type="syntype">Syntypes</typeStatus>
</emphasis>
(2): One originally in the BMNH (skull illustrated in the original description, p. 170, Fig. 1), and reproduced in
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), but later removed (
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;
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); currently unlocated. That skull is also a
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of
<emphasis id="B952EA8CFFA0FFC8DE10F85BFB74244F" box="[1097,1249,779,805]" italics="true" pageId="54" pageNumber="55">M. e. phayrei</emphasis>
(
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). Second
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is in the ZSI according to
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, but not located.
</paragraph>
</subSubSection>
<subSubSection id="C33C6515FFA0FFC8D918F8DCFB98230C" pageId="54" pageNumber="55" type="description">
<paragraph id="8B99369EFFA0FFC8D918F8DCFC6E24AF" blockId="54.[833,1451,907,1126]" pageId="54" pageNumber="55">
<taxonomicName id="4C264D1DFFA0FFC8D918F8DCFA8424CF" authority="Reimann &amp; Nutaphand" authorityName="Reimann &amp; Nutaphand" box="[833,1297,907,933]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="nutapundi">
<emphasis id="B952EA8CFFA0FFC8D918F8DCFB9B24CF" box="[833,1038,907,933]" italics="true" pageId="54" pageNumber="55">Testudo nutapundi</emphasis>
Reimann &amp; Nutaphand
</taxonomicName>
in Nutaphand 1979:193.
</paragraph>
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<emphasis id="B952EA8CFFA0FFC8D914F89DFC38248E" bold="true" box="[845,941,970,996]" pageId="54" pageNumber="55">
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</emphasis>
(number uncertain): Not located (
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); the specimen illustrated in the original description (p. 142, Figs. 105–106) was not considered to be the
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by
<bibRefCitation id="EFB74B6FFFA0FFC8DF2FFF7CFB9C230C" author="Obst" pageId="54" pageNumber="55" pagination="247 - 256" refId="ref78306" refString="Obst, F. J. (1983) Beitrag zur Kenntnis der Landschildkroten- Gattung Manouria Gray, 1852 (Reptilia, Testudines, Testudinata). Zoologische Abhandlungen, Staatliches Museum fur Tierkunde Dresden, 38 (15), 247 - 256." type="journal article" year="1983">Obst (1983:250)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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