treatments-xml/data/03/8E/C9/038EC92D79607E4AFC1B6DE524E8F932.xml

<document id="CA865A6A47BAA108BC5FCFAD2FEC55B5" ID-DOI="10.26107/RBZ-2023-0047" ID-ISSN="2345-7600" ID-Zenodo-Dep="10271307" ID-ZooBank="821BC4EC-5AF9-4727-84A3-C44839DFBE28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701819361795" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, Shane T. &amp; Castro, Peter" docDate="2023" docId="038EC92D79607E4AFC1B6DE524E8F932" docLanguage="en" docName="RafflesBZool.71.606-631.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Domeciidae Ortmann 1893" docType="treatment" docVersion="2" lastPageNumber="626" masterDocId="FFB7B15579727E5EFFBA6E582531FFA7" masterDocTitle="Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="631" masterPageNumber="606" pageNumber="624" updateTime="1701819819889" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6AF770AC5E6E8571914F796F7D39F038">Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)</mods:title>
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<heading id="D0D0CF5779607E4CFC1B6DE52006FC72" bold="true" box="[929,1335,957,981]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="624" reason="2">
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Family 
<taxonomicName id="4C2703B879607E4CFC406DE52006FC72" ID-CoL="7NJ99" authority="Ortmann, 1893" authorityName="Ortmann" authorityYear="1893" box="[1018,1335,957,981]" class="Malacostraca" family="Domeciidae" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="family">
Domeciidae 
<bibRefCitation id="EFB605CA79607E4CFB326DE52006FC72" author="Ortmann AE" box="[1160,1335,957,981]" pageId="18" pageNumber="624" pagination="411 - 495" refId="ref21184" refString="Ortmann AE (1893) Abtheilung: Brachyura (Brachyura genuina Boas) II. Unterabtheilung: Cancroidea, 2. Section: Cancrinea, 1. Gruppe: Cyclometopa. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen, Theil VII. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (3): 411 - 495, pl. 17." type="journal article" year="1893">Ortmann, 1893</bibRefCitation>
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<paragraph id="8B98783B79607E4AFC976DA52466FEB2" blockId="18.[811,1454,1021,2101]" lastBlockId="20.[113,752,189,277]" lastPageId="20" lastPageNumber="626" pageId="18" pageNumber="624">
<emphasis id="B953A42979607E4CFC976DA52690FBB2" bold="true" box="[813,929,1021,1045]" pageId="18" pageNumber="624">Diagnosis.</emphasis>
Carapace transversely hexagonal to almost ovate; antero- and posterolateral margins demarcated, anterolateral margin with 2 or 3 teeth, sometimes low, and additional acute granules or spinules (
<figureCitation id="131C64BE79607E4CFBEA6A0521FBFBD2" box="[1104,1226,1117,1141]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11I–K</figureCitation>
); front bilobed, lined with sharp granules or short spines; posterolateral margin gently convex to sinuous (
<figureCitation id="131C64BE79607E4CFBF56AC521FAFB12" box="[1103,1227,1181,1205]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11I–K</figureCitation>
); carapace relatively low, dorsal surface gently sinuous; posterior margin of epistome with small median lobe, lateral margin relatively short, gently concave, with triangular lobe just before pterygostomial lobe, separated by V-shaped cleft (
<figureCitation id="131C64BE79607E4CFAE96B452670FAF2" captionStart="Fig" captionStartId="8.[113,148,1170,1192]" captionTargetBox="[113,1451,189,1144]" captionTargetId="figure-13@8.[113,1451,189,1144]" captionTargetPageId="8" captionText="Fig. 5. Frontal view of cephalothorax showing epistome, orbits, antennules, and antenna. A, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; D, Calocarcinus africanus, male (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; E, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; F, Cherusius triunguiculatus, female (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271321" httpUri="https://zenodo.org/record/10271321/files/figure.png" pageId="18" pageNumber="624">Fig. 5E, F</figureCitation>
). Basal antennal article quadrate, mobile, peduncle entering hiatus of orbit. Maxilliped 3 merus very short, appearing compressed, anteroexternal part not auriculiform, narrower than large trapezoidal ischium; distal edge of exopod reaching to distal edge of merus (
<figureCitation id="131C64BE79607E4CFB976BE5214FFA72" box="[1069,1150,1469,1493]" captionStart="Fig" captionStartId="9.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1545]" captionTargetId="figure-7@9.[136,1474,189,1545]" captionTargetPageId="9" captionText="Fig. 6. Right maxilliped 3 (A–F); right major chela (G, H). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus, male (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271323" httpUri="https://zenodo.org/record/10271323/files/figure.png" pageId="18" pageNumber="624">Fig. 6F</figureCitation>
). Chelipeds relatively short, heterochelous, larger cheliped shorter than twice carapace width; merus quadrate in dorsal view, flexor margin smooth or finely granular, extensor margin with sharp spines or tubercles; carpus inner angle with spine or tooth; propodus outer surface with prominent rounded or pointed tubercles (
<figureCitation id="131C64BE79607E4CFC8F68252685F932" box="[821,948,1661,1685]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11I–K</figureCitation>
). P2–5 short to very short, longest pereopod slightly longer than carapace width; merus relatively stout (
<figureCitation id="131C64BE79607E4CFC8F68E52685F972" box="[821,948,1725,1749]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11I–K</figureCitation>
). Sternopleonal cavity reaching anteriorly to level of posterior margin of P2 coxae (
<figureCitation id="131C64BE79607E4CFB686885200EF952" box="[1234,1343,1757,1781]" captionStart="Fig" captionStartId="11.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1546]" captionTargetId="figure-8@11.[136,1474,189,1546]" captionTargetPageId="11" captionText="Fig.7. Male anterior thoracic sternum and pleon.A,Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; D, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271327" httpUri="https://zenodo.org/record/10271327/files/figure.png" pageId="18" pageNumber="624">Fig. 7H, I</figureCitation>
); tubercle of male pleonal-locking mechanism distinct, on posterior third of thoracic sternite 5, may be near suture with sternite 6 (
<figureCitation id="131C64BE79607E4CFCF56965269DF8F2" box="[847,940,1853,1877]" captionStart="Fig" captionStartId="13.[136,171,1937,1959]" captionTargetBox="[144,1464,189,1923]" captionTargetId="figure-8@13.[144,1464,189,1923]" captionTargetPageId="13" captionText="Fig. 8. Posterior part of male thoracic sternum and pleon (A–F); sternopleonal cavity showing tubercle of pleonal-locking mechanism on sternite 5 (G–L). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, I, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, L. Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; K, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia." figureDoi="http://doi.org/10.5281/zenodo.10271329" httpUri="https://zenodo.org/record/10271329/files/figure.png" pageId="18" pageNumber="624">Fig. 8L</figureCitation>
). 
<collectingRegion id="49E3B6D979607E4CFC7F696526CEF8F2" box="[965,1023,1853,1877]" country="Maldives" name="Maale" pageId="18" pageNumber="624">Male</collectingRegion>
pleon triangular, lateral margins of somites 3–6 converging, all somites and telson free (
<figureCitation id="131C64BE79607E4CFACC69052659F832" captionStart="Fig" captionStartId="11.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1546]" captionTargetId="figure-8@11.[136,1474,189,1546]" captionTargetPageId="11" captionText="Fig.7. Male anterior thoracic sternum and pleon.A,Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; D, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271327" httpUri="https://zenodo.org/record/10271327/files/figure.png" pageId="18" pageNumber="624">Figs. 7H, I</figureCitation>
, 
<figureCitation id="131C64BE79607E4CFCCC692526A4F832" box="[886,917,1917,1941]" captionStart="Fig" captionStartId="13.[136,171,1937,1959]" captionTargetBox="[144,1464,189,1923]" captionTargetId="figure-8@13.[144,1464,189,1923]" captionTargetPageId="13" captionText="Fig. 8. Posterior part of male thoracic sternum and pleon (A–F); sternopleonal cavity showing tubercle of pleonal-locking mechanism on sternite 5 (G–L). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, I, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, L. Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; K, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia." figureDoi="http://doi.org/10.5281/zenodo.10271329" httpUri="https://zenodo.org/record/10271329/files/figure.png" pageId="18" pageNumber="624">8F</figureCitation>
). G1 relatively stout, with distal portion slender, straight to curved, distal margins lined with short spinules or setae (
<figureCitation id="131C64BE79607E4CFC2869E5213CF872" box="[914,1037,1981,2005]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Fig. 9O, Q</figureCitation>
). G2 relatively stout, elongated, more than half to subequal length of G1, distal portion flattened, spatuliform (
<figureCitation id="131C64BE79607E4CFC0C69A52116F7B2" box="[950,1063,2045,2069]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Fig. 9P, R</figureCitation>
). Vulvae relatively large, positioned on anterior half of sternite 6, positioned close to median line, opening directed anteriorly with lateral sternal vulvar cover directed anteriorly (
<figureCitation id="131C64BE79667E4AFE296E8524C9FF52" box="[403,504,221,245]" captionStart="Fig" captionStartId="17.[136,171,1911,1933]" captionTargetBox="[136,1474,227,1896]" captionTargetId="figure-115@17.[136,1474,227,1896]" captionTargetPageId="17" captionText="Fig. 10. Vulvae. A, Tetralia nigrolineata (9.8 × 8.6 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (8.3 × 6.7 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (12.2 × 10.1 mm) (ZRC 2007.0079), Singapore; D, Quadrella maculosa (5.9 × 5.0 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.5 × 5.7 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (12.0 × 10.1 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (14.5 × 10.0 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; J, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion." figureDoi="http://doi.org/10.5281/zenodo.10271335" httpUri="https://zenodo.org/record/10271335/files/figure.png" pageId="20" pageNumber="626">Fig. 10H</figureCitation>
) or opening obliquely inwards (
<figureCitation id="131C64BE79667E4AFF6C6EA5247BFEB2" box="[214,330,253,277]" captionStart="Fig" captionStartId="17.[136,171,1911,1933]" captionTargetBox="[136,1474,227,1896]" captionTargetId="figure-115@17.[136,1474,227,1896]" captionTargetPageId="17" captionText="Fig. 10. Vulvae. A, Tetralia nigrolineata (9.8 × 8.6 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (8.3 × 6.7 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (12.2 × 10.1 mm) (ZRC 2007.0079), Singapore; D, Quadrella maculosa (5.9 × 5.0 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.5 × 5.7 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (12.0 × 10.1 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (14.5 × 10.0 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; J, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion." figureDoi="http://doi.org/10.5281/zenodo.10271335" httpUri="https://zenodo.org/record/10271335/files/figure.png" pageId="20" pageNumber="626">Fig. 10I, J</figureCitation>
).
</paragraph>
<caption id="DF5828B379617E4DFF3269D82728F7E4" ID-DOI="http://doi.org/10.5281/zenodo.10271337" ID-Zenodo-Dep="10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="19" pageNumber="625" startId="19.[136,171,1920,1942]" targetBox="[174,1436,189,1904]" targetPageId="19" targetType="figure">
<paragraph id="8B98783B79617E4DFF3269D82728F7E4" blockId="19.[136,1475,1920,2115]" pageId="19" pageNumber="625">
Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, 
<taxonomicName id="4C2703B879617E4DFB7269D820B4F832" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[1224,1413,1920,1941]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="cymodoce">
<emphasis id="B953A42979617E4DFB7269D820B4F832" box="[1224,1413,1920,1941]" italics="true" pageId="19" pageNumber="625">Trapezia cymodoce</emphasis>
</taxonomicName>
, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, 
<taxonomicName id="4C2703B879617E4DFD3C69C5267FF815" authorityName="Alcock" authorityYear="1898" box="[646,846,1949,1970]" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="B953A42979617E4DFD3C69C5267FF815" box="[646,846,1949,1970]" italics="true" pageId="19" pageNumber="625">Quadrella maculosa</emphasis>
</taxonomicName>
, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, 
<taxonomicName id="4C2703B879617E4DFF7D69E124F4F869" authorityName="Komai, Higashiji &amp; Castro" authorityYear="2010" box="[199,453,1977,1998]" class="Malacostraca" family="Trapeziidae" genus="Hexagonaloides" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="bathyalis">
<emphasis id="B953A42979617E4DFF7D69E124F4F869" box="[199,453,1977,1998]" italics="true" pageId="19" pageNumber="625">Hexagonaloides bathyalis</emphasis>
</taxonomicName>
, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, 
<taxonomicName id="4C2703B879617E4DFBF769E12002F869" authorityName=", Calman" authorityYear="1909" box="[1101,1331,1977,1998]" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B953A42979617E4DFBF769E12002F869" box="[1101,1331,1977,1998]" italics="true" pageId="19" pageNumber="625">Calocarcinus africanus</emphasis>
</taxonomicName>
, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, 
<taxonomicName id="4C2703B879617E4DFDC5698E26B3F84C" authorityName="Garth &amp; Kim" authorityYear="1983" box="[639,898,2006,2027]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="oviformis">
<emphasis id="B953A42979617E4DFDC5698E26B3F84C" box="[639,898,2006,2027]" italics="true" pageId="19" pageNumber="625">Philippicarcinus oviformis</emphasis>
</taxonomicName>
, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, 
<taxonomicName id="4C2703B879617E4DFF3269AC245FF7AF" baseAuthorityName="Desbonne in Desbonne &amp; Schramm" baseAuthorityYear="1867" box="[136,366,2035,2057]" class="Malacostraca" family="Domeciidae" genus="Domecia" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="acanthophora">
<emphasis id="B953A42979617E4DFF3269AC245FF7AF" box="[136,366,2035,2057]" italics="true" pageId="19" pageNumber="625">Domecia acanthophora</emphasis>
</taxonomicName>
, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, 
<taxonomicName id="4C2703B879617E4DFB9969AB2011F7AF" baseAuthorityName="Borradaile" baseAuthorityYear="1902" box="[1059,1312,2035,2056]" class="Malacostraca" family="Domeciidae" genus="Cherusius" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="triunguiculatus">
<emphasis id="B953A42979617E4DFB9969AB2011F7AF" box="[1059,1312,2035,2056]" italics="true" pageId="19" pageNumber="625">Cherusius triunguiculatus</emphasis>
</taxonomicName>
(6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, 
<taxonomicName id="4C2703B879617E4DFE57664827F3F782" baseAuthorityName="Rathbun in Edmondson" baseAuthorityYear="1923" box="[493,706,2064,2085]" class="Malacostraca" family="Domeciidae" genus="Palmyria" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="625" phylum="Arthropoda" rank="species" species="palmyrensis">
<emphasis id="B953A42979617E4DFE57664827F3F782" box="[493,706,2064,2085]" italics="true" pageId="19" pageNumber="625">Palmyria palmyrensis</emphasis>
</taxonomicName>
(4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after 
<bibRefCitation id="EFB605CA79617E4DFAF96648258DF7E5" author="Komai T &amp; Higashiji T &amp; Castro P" pageId="19" pageNumber="625" pagination="62 - 68" refId="ref20225" refString="Komai T, Higashiji T &amp; Castro P (2010) A new genus and new species of deep-water trapeziid crab (Crustacea: Decapoda: Brachyura: Trapezioidea) from the Ryukyu Islands, Japan. Zootaxa, 2555: 62 - 68." type="journal article" year="2010">Komai et al., 2010</bibRefCitation>
: fig. 2B; D, courtesy of T. Komai).
</paragraph>
</caption>
<paragraph id="8B98783B79667E4AFFCB6F6524E2FE32" blockId="20.[113,753,317,405]" pageId="20" pageNumber="626">
<emphasis id="B953A42979667E4AFFCB6F65240AFEF2" bold="true" box="[113,315,317,341]" pageId="20" pageNumber="626">Genera included.</emphasis>
<taxonomicName id="4C2703B879667E4AFEFF6F6527B1FEF2" authority="Low &amp; Ng, 2012" authorityName="Low &amp; Ng" authorityYear="2012" box="[325,640,317,342]" class="Malacostraca" family="Domeciidae" genus="Cherusius" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B953A42979667E4AFEFF6F652483FEF2" box="[325,434,317,341]" italics="true" pageId="20" pageNumber="626">Cherusius</emphasis>
<bibRefCitation id="EFB605CA79667E4AFE066F6627B1FEF2" author="Low MEY &amp; Ng PKL" box="[444,640,317,342]" pageId="20" pageNumber="626" pagination="59 - 62" refId="ref20496" refString="Low MEY &amp; Ng PKL (2012) The decapod crustaceans described by C. Sankarankutty, and a replacement name for Jonesius Sankarankutty, 1962 (Crustacea: Brachyura: Domeciidae), preoccupied by Jonesius Yamaguti, 1959 (Cestoda: Cyclophyllidea: Hymenolepididae). Zootaxa, 3363: 59 - 62." type="journal article" year="2012">Low &amp; Ng, 2012</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2703B879667E4AFD356F6624BEFED2" authority="Eydoux &amp; Souleyet, 1842" authorityName="Eydoux &amp; Souleyet" authorityYear="1842" class="Malacostraca" family="Domeciidae" genus="Domecia" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B953A42979667E4AFD356F6627C1FEF2" box="[655,752,318,341]" italics="true" pageId="20" pageNumber="626">Domecia</emphasis>
<bibRefCitation id="EFB605CA79667E4AFFCB6F0524BEFED2" author="Eydoux F &amp; Souleyet LFA" box="[113,399,349,373]" pageId="20" pageNumber="626" pagination="107 - 328" refId="ref19437" refString="Eydoux F &amp; Souleyet LFA (1842) Zoologie. In: Vaillant AN (ed.) Voyage autour du monde execute pendant les annees 1836 et 1837 sur la corvette la Bonite, commandee par M. Vaillant, Tome Premier, Seconde Partie. Arthus Bertrand, Paris, pp. 107 - 328." type="book chapter" year="1842">Eydoux &amp; Souleyet, 1842</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2703B879667E4AFE246F05278CFED2" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[414,701,349,373]" class="Malacostraca" family="Domeciidae" genus="Maldivia" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B953A42979667E4AFE246F0524CFFED2" box="[414,510,349,373]" italics="true" pageId="20" pageNumber="626">Maldivia</emphasis>
<bibRefCitation id="EFB605CA79667E4AFDBF6F05278CFED2" author="Borradaile LA" box="[517,701,349,373]" pageId="20" pageNumber="626" pagination="237 - 271" refId="ref18066" refString="Borradaile LA (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner JS (ed.) The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, Volume 1, Part 3. Cambridge University Press, London, pp. 237 - 271, figs. 41 - 60." type="book chapter" year="1902">Borradaile, 1902</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C2703B879667E4AFFC96F2524FEFE32" authority="Galil &amp; Takeda, 1986" authorityName="Galil &amp; Takeda" authorityYear="1986" box="[115,463,381,405]" class="Malacostraca" family="Domeciidae" genus="Palmyria" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B953A42979667E4AFFC96F2525E7FE32" box="[115,214,381,405]" italics="true" pageId="20" pageNumber="626">Palmyria</emphasis>
<bibRefCitation id="EFB605CA79667E4AFF676F2524FEFE32" author="Galil B &amp; Takeda M" box="[221,463,381,405]" pageId="20" pageNumber="626" pagination="163 - 171" refId="ref19759" refString="Galil B &amp; Takeda M (1986) Resurrection of the genus Jonesius and the establishment of a new genus: commensal crabs associated with corals from the Indo-Pacific Ocean. Bulletin of National Science Museum, Series A (Zoology), Tokyo, 12: 163 - 171." type="journal article" year="1986">Galil &amp; Takeda, 1986</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D2BB079667E4AFFCB6FE524E8F932" pageId="20" pageNumber="626" type="discussion">
<paragraph id="8B98783B79667E4AFFCB6FE52459FA12" blockId="20.[112,754,445,1461]" pageId="20" pageNumber="626">
<emphasis id="B953A42979667E4AFFCB6FE525D0FE72" bold="true" box="[113,225,445,469]" pageId="20" pageNumber="626">Remarks.</emphasis>
Rathbun (in 
<bibRefCitation id="EFB605CA79667E4AFEC06FE52772FE72" author="Edmondson CH" box="[378,579,445,469]" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson, 1923</bibRefCitation>
) described two new species from 
<taxonomicName id="4C2703B879667E4AFE8D6F8524A1FE52" authorityName="Savigny" authorityYear="1818" box="[311,400,477,501]" class="Polychaeta" family="Aphroditidae" genus="Palmyra" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="614" phylum="Annelida" rank="genus">Palmyra</taxonomicName>
Island in the central Pacific, the domeciid 
<taxonomicName id="4C2703B879667E4AFF606FA524F0FDB2" authorityName="Rathbun in Edmondson" authorityYear="1923" box="[218,449,509,533]" class="Malacostraca" family="Domeciidae" genus="Maldivia" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="species" species="palmyrensis">
<emphasis id="B953A42979667E4AFF606FA524F0FDB2" box="[218,449,509,533]" italics="true" pageId="20" pageNumber="626">Maldivia palmyrensis</emphasis>
</taxonomicName>
(presently in 
<taxonomicName id="4C2703B879667E4AFDEB6FA52414FD92" authority="Galil &amp; Takeda, 1986" authorityName="Galil &amp; Takeda" authorityYear="1986" class="Malacostraca" family="Domeciidae" genus="Palmyria" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B953A42979667E4AFDEB6FA52785FDB2" box="[593,692,509,533]" italics="true" pageId="20" pageNumber="626">Palmyria</emphasis>
<bibRefCitation id="EFB605CA79667E4AFD016FA52414FD92" author="Galil B &amp; Takeda M" pageId="20" pageNumber="626" pagination="163 - 171" refId="ref19759" refString="Galil B &amp; Takeda M (1986) Resurrection of the genus Jonesius and the establishment of a new genus: commensal crabs associated with corals from the Indo-Pacific Ocean. Bulletin of National Science Museum, Series A (Zoology), Tokyo, 12: 163 - 171." type="journal article" year="1986">Galil &amp; Takeda, 1986</bibRefCitation>
</taxonomicName>
) and the portunid 
<taxonomicName id="4C2703B879667E4AFE4C6C45258BFDF2" authorityName="Rathbun in Edmondson" authorityYear="1923" class="Malacostraca" family="Portunidae" genus="Charybdis" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="species" species="cookei" subGenus="Charybdis">
<emphasis id="B953A42979667E4AFE4C6C452754FD92" box="[502,613,541,565]" italics="true" pageId="20" pageNumber="626">Charybdis</emphasis>
(
<emphasis id="B953A42979667E4AFDC26C4527D6FD92" box="[632,743,541,565]" italics="true" pageId="20" pageNumber="626">Charybdis</emphasis>
) 
<emphasis id="B953A42979667E4AFFCB6C65258BFDF2" box="[113,186,573,597]" italics="true" pageId="20" pageNumber="626">cookei</emphasis>
</taxonomicName>
. The description was part of a larger article by 
<bibRefCitation id="EFB605CA79667E4AFFCB6C052471FDD2" author="Edmondson CH" box="[113,320,605,629]" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson (1923)</bibRefCitation>
on the Crustacea from 
<taxonomicName id="4C2703B879667E4AFDFA6C0527A8FDD2" authorityName="Savigny" authorityYear="1818" box="[576,665,605,629]" class="Polychaeta" family="Aphroditidae" genus="Palmyra" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="614" phylum="Annelida" rank="genus">Palmyra</taxonomicName>
and the Fanning Islands. The format of this publication is such that the taxa could be cited as Rathbun (1923) or Rathbun (in 
<bibRefCitation id="EFB605CA79667E4AFFCB6CE52406FD72" author="Edmondson CH" box="[113,311,701,725]" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson, 1923</bibRefCitation>
). The article is the only one included in a volume of the P. Bishop Museum Bulletin, and the title on the cover page: “Crustacea from 
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and Fanning Islands by Charles Howard Edmondson with descriptions of new species of crabs from 
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Island by Mary J. Rathbun” leaves no doubt of Rathbun’s contribution. The main text and the treatment of all the previously described taxa were by 
<bibRefCitation id="EFB605CA79667E4AFF296DC524AEFC12" author="Edmondson CH" box="[147,415,925,949]" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson (1923: 1–37)</bibRefCitation>
. In his introduction, 
<bibRefCitation id="EFB605CA79667E4AFDCE6DC525E3FC72" author="Edmondson CH" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson (1923: 7)</bibRefCitation>
stated: “In addition to previous acknowledgments, credit is due Drs. Mary J. Rathbun and Waldo L. Schmitt of the 
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National Museum for services rendered in the identification of certain species included in this paper. Rathbun also described two new species of crabs from 
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Island. (See p. 38).” There is a new heading on page 38 that reads: “Descriptions of new species of crabs from 
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Island by Mary J. Rathbun”, and the two new species were described in the next two pages (
<bibRefCitation id="EFB605CA79667E4AFDD26AE525CEFB52" author="Edmondson CH" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson, 1923: 38–40</bibRefCitation>
). After Rathbun’s section, 
<bibRefCitation id="EFB605CA79667E4AFD9E6A8525F1FAB2" author="Edmondson CH" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson (1923: 41–43)</bibRefCitation>
provided an index followed by the plates. One of the authors (PC) checked both the original “Bulletin of the Bernice P. Bishop Museum” as well as the reprint, and the information is identical. Placed in the complete context, the best way to cite authorship of the two species is “Rathbun in 
<bibRefCitation id="EFB605CA79667E4AFF2B6BC5246BFA12" author="Edmondson CH" box="[145,346,1437,1461]" pageId="20" pageNumber="626" pagination="1 - 43" refId="ref19392" refString="Edmondson CH (1923) Crustacea from Palmyra and Fanning Islands, with descriptions of new species of crabs from Palmyra Island by Mary Rathbun. Bernice P. Bishop Museum Bulletin, 5: 1 - 43, pls. 1, 2." type="book chapter" year="1923">Edmondson, 1923</bibRefCitation>
”.
</paragraph>
<paragraph id="8B98783B79667E4AFFCB6B8524E8F932" blockId="20.[112,753,1501,1685]" pageId="20" pageNumber="626">
All domeciids are symbionts of zooxanthellate (hermatypic) scleractininan corals and, less commonly, hydrozoans and sponges (
<bibRefCitation id="EFB605CA79667E4AFEA86845249BF992" author="Castro P" box="[274,426,1565,1589]" pageId="20" pageNumber="626" pagination="543 - 581" refId="ref18613" refString="Castro P (2015) Symbiotic Brachyura. In: Castro P, Davie P, Guinot D, Schram FR &amp; von Vaupel Klein JC (eds.) Treatise on zoology - anatomy, taxonomy, biology. The Crustacea. Decapoda: Brachyura (Part 2), 9 C-II. Brill, Leiden, The Netherlands, pp. 543 - 581." type="book chapter" year="2015">Castro, 2015</bibRefCitation>
; 
<bibRefCitation id="EFB605CA79667E4AFE00684627D5F992" author="van der Meij SET &amp; Bravo H &amp; Scholten YJH &amp; Dromard CR" box="[442,740,1565,1589]" pageId="20" pageNumber="626" pagination="239 - 246" refId="ref22067" refString="van der Meij SET, Bravo H, Scholten YJH &amp; Dromard CR (2022) Host use of the elkhorn coral crab Domecia acanthophora (Brachyura: Domeciidae), with a phylogeny of the genus. Cahiers de Biologie Marine, 63: 239 - 246." type="journal article" year="2022">van der Meij et al., 2022</bibRefCitation>
). Phylogenetic relationships of 
<taxonomicName id="4C2703B879667E4AFE1368662738F9F2" authorityName="Eydoux &amp; Souleyet" authorityYear="1842" box="[425,521,1598,1621]" class="Malacostraca" family="Domeciidae" genus="Domecia" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="626" phylum="Arthropoda" rank="genus">
<emphasis id="B953A42979667E4AFE1368662738F9F2" box="[425,521,1598,1621]" italics="true" pageId="20" pageNumber="626">Domecia</emphasis>
</taxonomicName>
and its mitochondrial genome were analysed by 
<bibRefCitation id="EFB605CA79667E4AFE206806278CF9D2" author="van der Meij SET &amp; Bravo H &amp; Scholten YJH &amp; Dromard CR" box="[410,701,1629,1653]" pageId="20" pageNumber="626" pagination="239 - 246" refId="ref22067" refString="van der Meij SET, Bravo H, Scholten YJH &amp; Dromard CR (2022) Host use of the elkhorn coral crab Domecia acanthophora (Brachyura: Domeciidae), with a phylogeny of the genus. Cahiers de Biologie Marine, 63: 239 - 246." type="journal article" year="2022">van der Meij et al. (2022)</bibRefCitation>
and 
<bibRefCitation id="EFB605CA79667E4AFFCB68262472F932" author="Bravo H &amp; Baeza JA &amp; van der Meij S" box="[113,323,1661,1685]" pageId="20" pageNumber="626" pagination="3" refId="ref18148" refString="Bravo H, Baeza JA &amp; van der Meij S (2023) Genomic survey sequencing and complete mitochondrial genome of the elkhorn coral crab Domecia acanthophora (Desbonne in Desbonne &amp; Schramm, 1867) (Decapoda: Brachyura: Domeciidae). Journal of Crustacean Biology, 43 (3): ruad 046." type="journal article" year="2023">Bravo et al. (2023)</bibRefCitation>
, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>