treatments-xml/data/03/8E/AC/038EAC691072ED5CC2CFFDCFFF4CFD57.xml

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<mods:title id="6067F6ED43F6427F19F08C5218666697">The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Turkey with description of a new species, Rhyacophila anatolica Ekingen &amp; Valladolid, sp. nov., based on morphological, genetic, and ecological evidence</mods:title>
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<mods:namePart id="CA85E390B12EE7CEB369A428FB880B6F">Valladolid, Maria</mods:namePart>
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<mods:affiliation id="5FDF89DBA132E65F4F285D0F8BD62A62">Department of Biodiversity and Evolutionary Biology, Museo Nacional de Ciencias Naturales (MNCN-CSIC), c / José Gutiérrez Abascal, 2, 28006 Madrid, Spain</mods:affiliation>
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<mods:namePart id="13A5444B60000A9630E0690207A58466">Ekingen, Pinar</mods:namePart>
<mods:nameIdentifier id="40F355A3E8A463DA84E08036590E2FB4" type="ORCID">0000-0002-0528-9945</mods:nameIdentifier>
<mods:affiliation id="6A22FB3988F7C0B01C42D249E1332C67">Laboratory of Biomonitoring, Section of Hydrobiology, Department of Biology, Hacettepe University, 06800 Beytepe, Ankara, Turkey</mods:affiliation>
<mods:nameIdentifier id="A19FEB5EAF23F2A5EBF53CBCFF4E4749" type="email">pinarekin@hacettepe.edu.tr</mods:nameIdentifier>
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<mods:namePart id="EEB6102DDFF1148278170E2639F30335">Arauzo, Mercedes</mods:namePart>
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<mods:affiliation id="67EAC6680D763D00485113F8155660E2">Institute of Agricultural Sciences (ICA-CSIC), c / Serrano 115, dpdo., 28006 Madrid, Spain</mods:affiliation>
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<mods:namePart id="ADE2ECDF9DBE3A78EF2304C3F61E9465">Dorda, Beatriz A.</mods:namePart>
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<mods:affiliation id="4FBCEEA44E4DC947A5F3383538D09084">Department of Collections, Museo Nacional de Ciencias Naturales (MNCN-CSIC), c / José Gutiérrez Abascal, 2, 28006 Madrid, Spain</mods:affiliation>
<mods:nameIdentifier id="060E5FD801E603B623CA8DE7CA507DAF" type="email">balvarez@mncn.csic.es</mods:nameIdentifier>
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<mods:namePart id="D7A5FD3F941B263D17FC40B60EF38FF4">Rey, Isabel</mods:namePart>
<mods:nameIdentifier id="46ACCD608FB3886F09218E3135EBBF40" type="ORCID">0000-0002-2122-5124</mods:nameIdentifier>
<mods:affiliation id="6C7C4208903D427583FCBBFA5F3BE260">Department of Collections, Museo Nacional de Ciencias Naturales (MNCN-CSIC), c / José Gutiérrez Abascal, 2, 28006 Madrid, Spain</mods:affiliation>
<mods:nameIdentifier id="09D99BDFD843F5800F00629C6FC4292D" type="email">isabel.rey@csic.es</mods:nameIdentifier>
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<emphasis id="B953C16D1072ED4AC2CFFDCFFE04FDC4" bold="true" box="[151,424,543,570]" italics="true" pageId="5" pageNumber="306">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
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, 
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Introduction: This species has been found in northern 
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, mainly in the western and middle part of the Black Sea Region of 
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.
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<emphasis id="B953C16D1072ED4AC29FFD61FE38FD34" box="[199,404,689,714]" italics="true" pageId="5" pageNumber="306">Derivatio nominis</emphasis>
: Derived from the name of the area (
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) of 
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where the new species has been found.
</paragraph>
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<paragraph id="8B981D7F1072ED4AC29FFD28FE0EFCEC" blockId="5.[151,1437,616,1110]" box="[199,418,760,786]" pageId="5" pageNumber="306">Material examined:</paragraph>
<paragraph id="8B981D7F1072ED4AC29FFCCCFD8CFCA4" blockId="5.[151,1437,616,1110]" pageId="5" pageNumber="306">
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: 
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(ANA 6), 
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, 
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, 
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, 
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, 
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, 
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<quantity id="4CDFB09A1072ED4AC60EFCCCFB07FCC9" box="[1110,1195,796,823]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="306" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
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, 
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</date>
, leg. 
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[
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(
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)]
</materialsCitation>
</paragraph>
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<materialsCitation id="3B4F17221072ED4AC29FFCB5FE0CFC5C" ID-GBIF-Occurrence="4986858304" collectingDate="2020-10-06" collectionCode="HUTC" collectorName="P. Ekingen" country="Turkey" elevation="1000" isTailEnumeration="true" latitude="41.032" location="TURKEY" longLatPrecision="69" longitude="32.436" municipality="Citdere" pageId="5" pageNumber="306" specimenCount="71" specimenCount-female="7" specimenCount-male="60" stateProvince="Karabuk" typeStatus="paratype">
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: 
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[
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, 
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+ 
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+ 
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P + 
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+ 
<specimenCount id="9D21D6F61072ED4AC1CEFCB4FC6CFC81" box="[918,960,868,895]" pageId="5" pageNumber="306">3 L</specimenCount>
]; 
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+ 
<specimenCount id="9D21D6F61072ED4AC618FCB4FBC6FC81" box="[1088,1130,868,895]" count="1" pageId="5" pageNumber="306" type="female">1 ♀</specimenCount>
, same locality as holotype, 
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<collectingDate id="EFDDC2571072ED4AC2CFFC58FEA5FC5C" box="[151,265,904,930]" pageId="5" pageNumber="306" value="2020-10-06">06/x/2020</collectingDate>
</date>
(
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)
</materialsCitation>
; 
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<specimenCount id="9D21D6F61072ED4AC3F3FC58FE47FC5D" box="[427,491,904,931]" count="5" pageId="5" pageNumber="306" type="male">5 ♂♂</specimenCount>
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, 
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, 
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, 
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<collectingDate id="EFDDC2571072ED4AC146FC58FC23FC5C" box="[798,911,904,930]" pageId="5" pageNumber="306" value="2020-10-06">06/x/2020</collectingDate>
</date>
(
<collectorName id="26D278A91072ED4AC1C7FC58FBB3FC5C" box="[927,1055,904,930]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
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; 
<materialsCitation id="3B4F17221072ED4AC66AFC58FE14FC38" ID-GBIF-Occurrence="4986858312" collectingDate="2020-08-11" collectorName="P. Ekingen" country="Turkey" location="Yenice" municipality="Citdere" pageId="5" pageNumber="306" specimenCount="5" specimenCount-male="5" stateProvince="Karabuk" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC66AFC58FBDEFC5D" box="[1074,1138,904,931]" count="5" pageId="5" pageNumber="306" type="male">5 ♂♂</specimenCount>
<collectingMunicipality id="6BFC87051072ED4AC621FC58FB62FC5C" box="[1145,1230,904,930]" pageId="5" pageNumber="306">Çitdere</collectingMunicipality>
, 
<location id="8EF84BA41072ED4AC682FC58FA85FC5C" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC682FC58FA85FC5C" box="[1242,1321,904,930]" country="Turkey" municipality="Citdere" name="Yenice" pageId="5" pageNumber="306" stateProvince="Karabuk">Yenice</location>
, 
<collectingRegion id="49E3D39D1072ED4AC76DFC58FA34FC5C" box="[1333,1432,904,930]" country="Turkey" name="Karabuk" pageId="5" pageNumber="306">Karabük</collectingRegion>
, 
<date id="FF993BBF1072ED4AC2CFFC7CFEB3FC38" box="[151,287,940,966]" pageId="5" pageNumber="306" value="2020-08-11">
<collectingDate id="EFDDC2571072ED4AC2CFFC7CFEB3FC38" box="[151,287,940,966]" pageId="5" pageNumber="306" value="2020-08-11">11/viii/2020</collectingDate>
</date>
(
<collectorName id="26D278A91072ED4AC368FC7CFE1CFC38" box="[304,432,940,966]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="3B4F17221072ED4AC39CFC7CFB38FC38" ID-GBIF-Occurrence="4986858303" box="[452,1172,940,967]" collectingDate="2020-10-06" collectorName="P. Ekingen" country="Turkey" location="SimSir Stream" municipality="Yenice" pageId="5" pageNumber="306" specimenCount="2" specimenCount-male="2" stateProvince="Karabuk" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC39CFC7CFDA9FC39" box="[452,517,940,967]" count="2" pageId="5" pageNumber="306" type="male">2 ♂♂</specimenCount>
<location id="8EF84BA41072ED4AC056FC7CFD1DFC38" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC056FC7CFD1DFC38" box="[526,689,940,966]" country="Turkey" municipality="Yenice" name="SimSir Stream" pageId="5" pageNumber="306" stateProvince="Karabuk">ŞimŞir Stream</location>
, 
<collectingMunicipality id="6BFC87051072ED4AC0E5FC7CFCA0FC38" box="[701,780,940,966]" pageId="5" pageNumber="306">Yenice</collectingMunicipality>
, 
<collectingRegion id="49E3D39D1072ED4AC141FC7CFCD0FC38" box="[793,892,940,966]" country="Turkey" name="Karabuk" pageId="5" pageNumber="306">Karabük</collectingRegion>
, 
<date id="FF993BBF1072ED4AC1D1FC7CFC57FC38" box="[905,1019,940,966]" pageId="5" pageNumber="306" value="2020-10-06">
<collectingDate id="EFDDC2571072ED4AC1D1FC7CFC57FC38" box="[905,1019,940,966]" pageId="5" pageNumber="306" value="2020-10-06">06/x/2020</collectingDate>
</date>
(
<collectorName id="26D278A91072ED4AC654FC7CFB20FC38" box="[1036,1164,940,966]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="3B4F17221072ED4AC6F8FC7CFD48FC14" ID-GBIF-Occurrence="4986858301" collectingDate="2023-10-28" collectorName="P. Ekingen" country="Turkey" location="Yedigoller National Park" municipality="Selale" pageId="5" pageNumber="306" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bolu" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC6F8FC7CFB60FC39" box="[1184,1228,940,967]" count="1" pageId="5" pageNumber="306" type="male">1 ♂</specimenCount>
+ 
<specimenCount id="9D21D6F61072ED4AC6B5FC7CFAB5FC39" box="[1261,1305,940,967]" count="1" pageId="5" pageNumber="306" type="female">1 ♀</specimenCount>
, 
<location id="8EF84BA41072ED4AC77FFC7CFE99FC14" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC77FFC7CFE99FC14" country="Turkey" municipality="Selale" name="Yedigoller National Park" pageId="5" pageNumber="306" stateProvince="Bolu">Yedigöller National Park</location>
, 
<collectingMunicipality id="6BFC87051072ED4AC319FC00FE28FC14" box="[321,388,976,1002]" pageId="5" pageNumber="306">Şelale</collectingMunicipality>
, 
<collectingRegion id="49E3D39D1072ED4AC3C8FC00FE65FC14" box="[400,457,976,1002]" country="Turkey" name="Bolu" pageId="5" pageNumber="306">Bolu</collectingRegion>
, 
<date id="FF993BBF1072ED4AC38DFC00FDE5FC14" box="[469,585,976,1002]" pageId="5" pageNumber="306" value="2023-10-28">
<collectingDate id="EFDDC2571072ED4AC38DFC00FDE5FC14" box="[469,585,976,1002]" pageId="5" pageNumber="306" value="2023-10-28">28/x/2023</collectingDate>
</date>
, (
<collectorName id="26D278A91072ED4AC005FC00FD71FC14" box="[605,733,976,1002]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="3B4F17221072ED4AC0A8FC00FDB6FBF0" ID-GBIF-Occurrence="4986858309" collectingDate="2023-10-28" collectorName="P. Ekingen" country="Turkey" location="Yedigoller National Park" municipality="Selalenin" pageId="5" pageNumber="306" specimenCount="1" stateProvince="Bolu" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC0A8FC00FC83FC15" box="[752,815,976,1003]" count="3" pageId="5" pageNumber="306" type="male">3 ♂♂</specimenCount>
+ 
<specimenCount id="9D21D6F61072ED4AC117FC00FC23FC15" box="[847,911,976,1003]" count="4" pageId="5" pageNumber="306" type="female">4 ♀♀</specimenCount>
, 
<location id="8EF84BA41072ED4AC1C4FC00FB1BFC14" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC1C4FC00FB1BFC14" box="[924,1207,976,1002]" country="Turkey" municipality="Selalenin" name="Yedigoller National Park" pageId="5" pageNumber="306" stateProvince="Bolu">Yedigöller National Park</location>
, 
<collectingMunicipality id="6BFC87051072ED4AC69BFC00FA8BFC14" box="[1219,1319,976,1002]" pageId="5" pageNumber="306">Şelalenin</collectingMunicipality>
yanındaki kol, 
<collectingRegion id="49E3D39D1072ED4AC291FC24FEAEFBF0" box="[201,258,1012,1038]" country="Turkey" name="Bolu" pageId="5" pageNumber="306">Bolu</collectingRegion>
, 
<date id="FF993BBF1072ED4AC355FC24FE2DFBF0" box="[269,385,1012,1038]" pageId="5" pageNumber="306" value="2023-10-28">
<collectingDate id="EFDDC2571072ED4AC355FC24FE2DFBF0" box="[269,385,1012,1038]" pageId="5" pageNumber="306" value="2023-10-28">28/x/2023</collectingDate>
</date>
, (
<collectorName id="26D278A91072ED4AC3CCFC24FDBEFBF0" box="[404,530,1012,1038]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="3B4F17221072ED4AC07DFC24FEC5FBCC" ID-GBIF-Occurrence="4986858305" collectingDate="2023-10-28" collectionCode="HUTC" collectorName="P. Ekingen" country="Turkey" location="Yedigoller National Park" municipality="Derin Gol'e" pageId="5" pageNumber="306" specimenCount="1" stateProvince="Bolu" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC07DFC24FDFCFBF1" box="[549,592,1012,1039]" pageId="5" pageNumber="306">1 ♂</specimenCount>
, 
<location id="8EF84BA41072ED4AC003FC24FCD8FBF0" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC003FC24FCD8FBF0" box="[603,884,1012,1038]" country="Turkey" municipality="Derin Gol'e" name="Yedigoller National Park" pageId="5" pageNumber="306" stateProvince="Bolu">Yedigöller National Park</location>
, 
<collectingMunicipality id="6BFC87051072ED4AC127FC24FBA9FBF0" box="[895,1029,1012,1038]" pageId="5" pageNumber="306">Derin Göl’e</collectingMunicipality>
dökülen akarsu, 
<collectingRegion id="49E3D39D1072ED4AC69BFC24FB50FBF0" box="[1219,1276,1012,1038]" country="Turkey" name="Bolu" pageId="5" pageNumber="306">Bolu</collectingRegion>
, 
<date id="FF993BBF1072ED4AC75FFC24FAD5FBF0" box="[1287,1401,1012,1038]" pageId="5" pageNumber="306" value="2023-10-28">
<collectingDate id="EFDDC2571072ED4AC75FFC24FAD5FBF0" box="[1287,1401,1012,1038]" pageId="5" pageNumber="306" value="2023-10-28">28/x/2023</collectingDate>
</date>
(
<collectorName id="26D278A91072ED4AC7D0FC24FF5AFBCC" pageId="5" pageNumber="306">P. Ekingen</collectorName>
) [
<collectionCode id="ED3685BA1072ED4AC355FBC8FECEFBCC" box="[269,354,1048,1074]" pageId="5" pageNumber="306">HUTC</collectionCode>
]
</materialsCitation>
; 
<materialsCitation id="3B4F17221072ED4AC32DFBC8FB1DFBCC" ID-GBIF-Occurrence="4986858302" box="[373,1201,1048,1075]" collectingDate="2019-10-26" collectorName="P. Ekingen" country="Turkey" location="SimSir Stream" municipality="Yenice" pageId="5" pageNumber="306" specimenCount="2" stateProvince="Karabuk" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC32DFBC8FE03FBCD" box="[373,431,1048,1075]" pageId="5" pageNumber="306">1 ♂P</specimenCount>
+ 
<specimenCount id="9D21D6F61072ED4AC388FBC8FDA5FBCD" box="[464,521,1048,1075]" pageId="5" pageNumber="306">1 ♀P</specimenCount>
, 
<location id="8EF84BA41072ED4AC04FFBC8FD17FBCC" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC04FFBC8FD17FBCC" box="[535,699,1048,1074]" country="Turkey" municipality="Yenice" name="SimSir Stream" pageId="5" pageNumber="306" stateProvince="Karabuk">ŞimŞir Stream</location>
, 
<collectingMunicipality id="6BFC87051072ED4AC09FFBC8FCBAFBCC" box="[711,790,1048,1074]" pageId="5" pageNumber="306">Yenice</collectingMunicipality>
, 
<collectingRegion id="49E3D39D1072ED4AC17AFBC8FC29FBCC" box="[802,901,1048,1074]" country="Turkey" name="Karabuk" pageId="5" pageNumber="306">Karabük</collectingRegion>
, 
<date id="FF993BBF1072ED4AC1CAFBC8FBB8FBCC" box="[914,1044,1048,1074]" pageId="5" pageNumber="306" value="2019-10-26">
<collectingDate id="EFDDC2571072ED4AC1CAFBC8FBB8FBCC" box="[914,1044,1048,1074]" pageId="5" pageNumber="306" value="2019-10-26">26/10/2019</collectingDate>
</date>
, (
<collectorName id="26D278A91072ED4AC672FBC8FB06FBCC" box="[1066,1194,1048,1074]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="3B4F17221072ED4AC6E5FBC8FD28FBA8" ID-GBIF-Occurrence="4986858306" collectingDate="2020-08-11" collectorName="P. Ekingen" country="Turkey" location="SimSir Stream" municipality="Yenice" pageId="5" pageNumber="306" specimenCount="3" stateProvince="Karabuk" typeStatus="paratype">
<specimenCount id="9D21D6F61072ED4AC6E5FBC8FB4BFBCD" box="[1213,1255,1048,1075]" pageId="5" pageNumber="306">3 L</specimenCount>
, 
<location id="8EF84BA41072ED4AC6ACFBC8FA34FBCC" LSID="urn:lsid:plazi:treatment:038EAC691072ED5CC2CFFDCFFF4CFD57:8EF84BA41072ED4AC6ACFBC8FA34FBCC" box="[1268,1432,1048,1074]" country="Turkey" municipality="Yenice" name="SimSir Stream" pageId="5" pageNumber="306" stateProvince="Karabuk">ŞimŞir Stream</location>
, 
<collectingMunicipality id="6BFC87051072ED4AC2CFFBECFF4AFBA8" box="[151,230,1084,1110]" pageId="5" pageNumber="306">Yenice</collectingMunicipality>
, 
<collectingRegion id="49E3D39D1072ED4AC2AAFBECFEF9FBA8" box="[242,341,1084,1110]" country="Turkey" name="Karabuk" pageId="5" pageNumber="306">Karabük</collectingRegion>
, 
<date id="FF993BBF1072ED4AC338FBECFE47FBA8" box="[352,491,1084,1110]" pageId="5" pageNumber="306" value="2020-08-11">
<collectingDate id="EFDDC2571072ED4AC338FBECFE47FBA8" box="[352,491,1084,1110]" pageId="5" pageNumber="306" value="2020-08-11">11/viii/2020</collectingDate>
</date>
, (
<collectorName id="26D278A91072ED4AC3A6FBECFDD1FBA8" box="[510,637,1084,1110]" pageId="5" pageNumber="306">P. Ekingen</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41072ED48C2CFFB78FAC1FB3B" lastPageId="7" lastPageNumber="308" pageId="5" pageNumber="306" type="description">
<paragraph id="8B981D7F1072ED4AC2CFFB78FF56FB3C" blockId="5.[151,250,1192,1254]" box="[151,250,1192,1218]" pageId="5" pageNumber="306">
<heading id="D0D0AA131072ED4AC2CFFB78FF56FB3C" bold="true" box="[151,250,1192,1218]" fontSize="11" level="1" pageId="5" pageNumber="306" reason="1">
<emphasis id="B953C16D1072ED4AC2CFFB78FF56FB3C" bold="true" box="[151,250,1192,1218]" pageId="5" pageNumber="306">LARVA</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F1072ED4AC2CFFB1CFF54FB18" blockId="5.[151,250,1192,1254]" box="[151,248,1228,1254]" pageId="5" pageNumber="306">
<figureCitation id="131C01FA1072ED4AC2CFFB1CFF54FB18" box="[151,248,1228,1254]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Figs 2–7</figureCitation>
</paragraph>
<paragraph id="8B981D7F1072ED4AC2CFFADEFA27FAB5" blockId="5.[151,1437,1293,2039]" pageId="5" pageNumber="306">
<emphasis id="B953C16D1072ED4AC2CFFADEFDD9FAD9" box="[151,629,1293,1319]" italics="true" pageId="5" pageNumber="306">Biometrics of last instar larva and prepupa</emphasis>
: Length: 
<quantity id="4CDFB09A1072ED4AC0B8FADDFC2EFAD6" box="[736,898,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.65" metricValueMin="1.45" pageId="5" pageNumber="306" unit="mm" value="15.5" valueMax="16.5" valueMin="14.5">14.5–16.5 mm</quantity>
(x̄ = 15.8, 
<collectingRegion id="49E3D39D1072ED4AC1A1FADDFBB1FAD9" box="[1017,1053,1293,1319]" country="Serbia" name="Sid" pageId="5" pageNumber="306">SD</collectingRegion>
= 1.09, n = 3). Maximum width of larva at metathorax: 
<quantity id="4CDFB09A1072ED4AC326FAE1FDA9FAB2" box="[382,517,1329,1356]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="5" pageNumber="306" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3–2.8 mm</quantity>
(x̄ = 2.6, n = 3). Specimens preserved in ethanol yellowish, with dorsum darker.
</paragraph>
<paragraph id="8B981D7F1072ED4AC29FFA85FD8DF809" blockId="5.[151,1437,1293,2039]" pageId="5" pageNumber="306">
<emphasis id="B953C16D1072ED4AC29FFA85FEA8FA91" box="[199,260,1365,1391]" italics="true" pageId="5" pageNumber="306">Head</emphasis>
(
<figureCitation id="131C01FA1072ED4AC34AFA85FEF5FA91" box="[274,345,1365,1391]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Figs 2</figureCitation>
, 
<figureCitation id="131C01FA1072ED4AC33CFA85FE23FA91" box="[356,399,1365,1391]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="5" pageNumber="306">13a</figureCitation>
). Length: 
<quantity id="4CDFB09A1072ED4AC3A7FA85FD29FA8E" box="[511,645,1365,1392]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="5" pageNumber="306" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
(x̄ = 1.7, 
<collectingRegion id="49E3D39D1072ED4AC0B7FA85FCBFFA91" box="[751,787,1365,1391]" country="Serbia" name="Sid" pageId="5" pageNumber="306">SD</collectingRegion>
= 0.06, n = 3). Width: 
<quantity id="4CDFB09A1072ED4AC654FA85FB01FA8E" box="[1036,1197,1365,1392]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.24" metricValueMin="1.21" pageId="5" pageNumber="306" unit="mm" value="1.225" valueMax="1.24" valueMin="1.21">1.21–1.24 mm</quantity>
(x̄ = 1.22, 
<collectingRegion id="49E3D39D1072ED4AC77DFA85FAE5FA91" box="[1317,1353,1365,1391]" country="Serbia" name="Sid" pageId="5" pageNumber="306">SD</collectingRegion>
= 0.02, n = 3). Maximum width at posterior third of head, lateral margins slightly converging anteriorly, round posteriorly. Sclerotized areas of head generally pale with dark areas and spots, dorsal head pits with dark borders. Labrum with small transparent area in middle of anterior edge. Frontoclypeal apotome (
<figureCitation id="131C01FA1072ED4AC189FA11FBF9FA25" box="[977,1109,1473,1499]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Figs. 2a, 2d</figureCitation>
) pale, anterior edge concave; pair of small, transversely oval, pale brown, submesal muscle attachment spots in anterior third between setae S4; dark pigmentation in posterior third U-shaped, with truncate or rounded anterolateral edges on pits P4 but not around setae S6, anteromesal portion of dark pigmentation paler brown, somewhat square, main portion of dark pigmentation with four (two central circular and two lateral transversely oval) pale ring-shape muscle attachment spots (
<figureCitation id="131C01FA1072ED4AC287F9A5FEC8F971" box="[223,356,1653,1679]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Figs. 2a, 2d</figureCitation>
). Parietals of cephalic capsule (
<figureCitation id="131C01FA1072ED4AC0E7F9A5FC93F971" box="[703,831,1653,1679]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Figs 2a–2b</figureCitation>
), with pair of longitudinal subdorsal bands in posterior two-thirds, these longitudinal subdorsal bands each darker and somewhat triangular, acute anteriorly and without muscle attachment spots, gradually broader posteriorly, and with two lines of 4–6 transversely oval, dark brown muscle attachments spots perpendicular to edges, forming light circular area surrounding juncture of frontal and coronal sutures, including pairs of pits P9 and setae S17, these longitudinal subdorsal bands narrow, light brown posteriorly, converging mesally toward pale yellowish longitudinal stripe along middle portion of coronal suture, with three or four spots aligned along each side of coronal suture, and 6–8 spots next to posterior edge; two pairs of pale spots around bases of setae S15 and S16 (
<figureCitation id="131C01FA1072ED4AC09FF8A1FC8FF875" box="[711,803,1905,1931]" captionStart="FIGURE 1" captionStartId="4.[152,255,896,920]" captionTargetBox="[213,1377,184,863]" captionTargetId="figure-26@4.[204,1383,181,872]" captionTargetPageId="4" captionText="FIGURE 1. Setae (S) and sensory pits (P) of the head and thorax of Rhyacophila larvae. 1a, head, dorsal; 1b, prothorax, right sclerite, dorsal (from Williams &amp; Wiggins 1981)." figureDoi="http://doi.org/10.5281/zenodo.14239577" httpUri="https://zenodo.org/record/14239577/files/figure.png" pageId="5" pageNumber="306">Figs. 1a</figureCitation>
, 
<figureCitation id="131C01FA1072ED4AC169F8A1FCE2F875" box="[817,846,1905,1931]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">2a</figureCitation>
); laterally with longitudinally rectangular dark band in posterodorsal third, with oval dark spots (
<figureCitation id="131C01FA1072ED4AC0C4F845FD5EF851" box="[668,754,1941,1967]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Fig. 2b</figureCitation>
); ventrally pale except anteroventral edge, and submentum reddish brown (
<figureCitation id="131C01FA1072ED4AC31DF869FE3AF82D" box="[325,406,1977,2003]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="5" pageNumber="306">Fig. 2c</figureCitation>
). Posterior occipital foramen brown with black corners at anteroventral area, dark brown line surrounding remainder of foramen.
</paragraph>
<caption id="DF584DF71071ED49C2C0F895FACEF813" ID-DOI="http://doi.org/10.5281/zenodo.14239579" ID-Zenodo-Dep="14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="6" pageNumber="307" startId="6.[152,270,1861,1885]" targetBox="[214,1374,185,1830]" targetPageId="6" targetType="figure">
<paragraph id="8B981D7F1071ED49C2C0F895FACEF813" blockId="6.[151,1437,1861,2032]" pageId="6" pageNumber="307">
<emphasis id="B953C16D1071ED49C2C0F895FEEEF8A3" bold="true" box="[152,322,1861,1885]" pageId="6" pageNumber="307">FIGURES 2–7.</emphasis>
Larva of 
<taxonomicName id="4C2766FC1071ED49C3F0F895FCD9F8A3" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[424,885,1861,1886]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D1071ED49C3F0F895FD23F8A3" box="[424,655,1861,1885]" italics="true" pageId="6" pageNumber="307">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D1071ED49C125F896FC62F8A0" bold="true" box="[893,974,1862,1886]" pageId="6" pageNumber="307">
<taxonomicNameLabel id="A2607C161071ED49C125F896FC62F8A0" box="[893,974,1862,1886]" pageId="6" pageNumber="307" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (
<emphasis id="B953C16D1071ED49C64FF8B9FBB2F87F" box="[1047,1054,1897,1921]" italics="true" pageId="6" pageNumber="307">l</emphasis>
) and right (
<emphasis id="B953C16D1071ED49C6CCF8BAFB32F87F" box="[1172,1182,1898,1921]" italics="true" pageId="6" pageNumber="307">r</emphasis>
): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B981D7F1070ED48C29FFF47FB01FF07" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="308">
Mandibles (
<figureCitation id="131C01FA1070ED48C315FF47FE75FF4F" box="[333,473,151,177]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 3D, 3V</figureCitation>
) asymmetrical, as in other rhyacophilids; left mandible slightly longer than right one. Left mandible with mesal border of inner blade nearly straight, right mandible with inconspicuous, small tooth in middle, and concave subapically. Maxillolabium and labrum as in other 
<taxonomicName id="4C2766FC1070ED48C198FF0FFB01FF07" box="[960,1197,223,249]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B953C16D1070ED48C198FF0FFBE2FF07" box="[960,1102,223,249]" italics="true" pageId="7" pageNumber="308">Rhyacophila</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8B981D7F1070ED48C29FFED3FBC5FCEB" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="308">
<emphasis id="B953C16D1070ED48C29FFED3FEB4FEE3" box="[199,280,259,285]" italics="true" pageId="7" pageNumber="308">Thorax</emphasis>
(
<figureCitation id="131C01FA1070ED48C37EFED3FEC3FEE3" box="[294,367,259,285]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 4</figureCitation>
, 
<figureCitation id="131C01FA1070ED48C322FED3FE0BFEE3" box="[378,423,259,285]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="7" pageNumber="308">13b</figureCitation>
). Anterior half of pronotum pale or very light coloured. Anterior margin straight or slightly convex (
<figureCitation id="131C01FA1070ED48C2A1FEF7FEE3FEBF" box="[249,335,295,321]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 4a</figureCitation>
, 
<figureCitation id="131C01FA1070ED48C305FEF7FE26FEBF" box="[349,394,295,321]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="7" pageNumber="308">13b</figureCitation>
), light brown, anterolateral borders strongly angulate (
<figureCitation id="131C01FA1070ED48C1A1FEF7FBE3FEBF" box="[1017,1103,295,321]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Fig. 4b</figureCitation>
); each side with two or three dark brown spots and black band from setae S22 and S 
<quantity id="4CDFB09A1070ED48C151FE9BFCE8FE9B" box="[777,836,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="7" pageNumber="308" unit="in" value="23.0">23 in</quantity>
anterior edge, and narrow black band in lateral edge touching broad posterolateral band (
<figureCitation id="131C01FA1070ED48C077FEBFFD28FE77" box="[559,644,367,393]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Fig. 4b</figureCitation>
). Posterior half of pronotum (
<figureCitation id="131C01FA1070ED48C18CFEBFFB89FE77" box="[980,1061,367,393]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Fig. 4a</figureCitation>
) with three vaguely defined dark regions separated by light zones and by setae S2 and S3 and pit P2 near mid-length of pronotum; (1) central dark region trapezoidal, tapered posteriorly and reaching posterior edge of sclerite, anteriorly truncate near midlength of pronotum, slightly wider than posteriorly, its lateral edges slightly concave or straight, pair of submesal lines of six or seven slightly dark brown, longitudinally oval spots inside this central dark region, reaching posterior edges of region; (2 and 3) pair of lateral dark regions irregular, each with its mesal edge convex, posterior edge convex and parallel to posterior mid-height pale brown border, and anterior edge diagonal between its anterior subdorsal edge and its posterior sublateral edge. Each lateral dark region with four or five brown spots around setae S5, transversely oval spot near its diagonal edge, and two transverse rows of circular or slightly transversely oval brown spots parallel to its posterior border. Posterior margin of pronotum strongly sinuous, with posterior border of each half black, this border at mid-height pale brown anteriorly (
<figureCitation id="131C01FA1070ED48C177FD07FC02FD0F" box="[815,942,727,753]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 4a, 4b</figureCitation>
, 
<figureCitation id="131C01FA1070ED48C1E2FD07FC4BFD0F" box="[954,999,727,753]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="7" pageNumber="308">13b</figureCitation>
); each posterolateral margin projecting posterad and bordered by black from posterolateral corner, touching anterolateral band.
</paragraph>
<paragraph id="8B981D7F1070ED48C29FFCCFFE28FC37" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="308">
<emphasis id="B953C16D1070ED48C29FFCCFFF50FCC6" box="[199,252,799,824]" italics="true" pageId="7" pageNumber="308">Legs</emphasis>
. Similar to those of other 
<taxonomicName id="4C2766FC1070ED48C06FFCCFFC8BFCC7" box="[567,807,799,825]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B953C16D1070ED48C06FFCCFFD69FCC7" box="[567,709,799,825]" italics="true" pageId="7" pageNumber="308">Rhyacophila</emphasis>
species.
</taxonomicName>
Colour yellowish, tarsi light reddish brown, proximal regions of coxae, trochanters, femora, and tibiae each with narrow black band, small circular black spots in both faces of each femur distally. Forelegs each with anterior and ventral rows of spicules, extending along distal half of trochanter, proximal half of femur, distal half of tibia, and distal half of tarsus. Setae black or dark brown, some transparent ventrally.
</paragraph>
<paragraph id="8B981D7F1070ED48C29FFC03FAC1FB3B" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="308">
<emphasis id="B953C16D1070ED48C29FFC03FE9DFC13" box="[199,305,979,1005]" italics="true" pageId="7" pageNumber="308">Abdomen</emphasis>
(
<figureCitation id="131C01FA1070ED48C318FC03FE09FC13" box="[320,421,978,1005]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 5–7</figureCitation>
, 
<figureCitation id="131C01FA1070ED48C3EAFC03FDBAFC13" box="[434,534,979,1005]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="7" pageNumber="308">13c, 13d</figureCitation>
). Similar to that of other species of 
<taxonomicName id="4C2766FC1070ED48C1F2FC03FB94FC13" box="[938,1080,979,1005]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B953C16D1070ED48C1F2FC03FB94FC13" box="[938,1080,979,1005]" italics="true" pageId="7" pageNumber="308">Rhyacophila</emphasis>
</taxonomicName>
. Preserved larva reddish brown dorsally, two stripes of anterolateral triangular pale spots, connected anteriorly and posteriorly with pale bar in central area, and two central stripes of oval dark spots (
<figureCitation id="131C01FA1070ED48C146FBCBFCC8FBCB" box="[798,868,1050,1077]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Fig. 5</figureCitation>
), ventral side pale. Widths of anterior abdominal segments similar, narrower in posterior segments. Lateral abdominal gills in tufts of 20–35 fine filaments. Tergite IX (
<figureCitation id="131C01FA1070ED48C2C7FBB3FF4BFB83" box="[159,231,1123,1149]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 6</figureCitation>
, 
<figureCitation id="131C01FA1070ED48C2A9FBB3FEB6FB83" box="[241,282,1123,1149]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="7" pageNumber="308">13c</figureCitation>
) yellowish, with black band on anterior edge and two black areas around central setae on posterior edge. Anal prolegs typical of group, each with long sword process (
<figureCitation id="131C01FA1070ED48C11BFB57FC6EFB5F" box="[835,962,1159,1185]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 7a; 7b</figureCitation>
, sp; 13d). Anal claws each with three teeth on ventral edge (
<figureCitation id="131C01FA1070ED48C30BFB7BFE7DFB3B" box="[339,465,1195,1221]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 7a, 7c</figureCitation>
), its basoventral hook black basally and reddish brown apically (
<figureCitation id="131C01FA1070ED48C6FFFB7BFB52FB3B" box="[1191,1278,1195,1221]" captionStart="FIGURES 2–7" captionStartId="6.[152,270,1861,1885]" captionTargetBox="[214,1374,185,1830]" captionTargetId="figure-26@6.[204,1384,181,1837]" captionTargetPageId="6" captionText="FIGURES 2–7. Larva of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 2a–2d, head: 2a, dorsal; 2b, right lateral; 2c, ventral; 2d, frontoclypeus, dorsal. 3D–3V, mandibles from last instar larval exuviae, left (l) and right (r): 3D, dorsal; 3V, ventral. 4a–4b, prothorax: 4a, dorsal; 4b, left hemisclerite, left lateral. 5, abdominal segment III, dorsal, A = anterior edge. 6, abdominal tergite IX. 7a–7c, details of anal prolegs: 7a, right anal claw, right lateral; 7b, detail of basolateral plate with sword process (sp) and basoventral hook (bh), right lateral; 7c, detail of right claw, right lateral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14239579" httpUri="https://zenodo.org/record/14239579/files/figure.png" pageId="7" pageNumber="308">Figs 7b</figureCitation>
, bh; 13c).
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41070ED48C2CFFAC6FADBF967" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="8B981D7F1070ED48C2CFFAC6FE29FACF" blockId="7.[151,389,1302,1329]" box="[151,389,1302,1329]" pageId="7" pageNumber="308">
<emphasis id="B953C16D1070ED48C2CFFAC6FE29FACF" bold="true" box="[151,389,1302,1329]" pageId="7" pageNumber="308">Diagnosis of larvae</emphasis>
</paragraph>
<paragraph id="8B981D7F1070ED48C2CFFA8FFC75F9D3" blockId="7.[151,1437,1375,2013]" pageId="7" pageNumber="308">
<emphasis id="B953C16D1070ED48C2CFFA8FFF78FA87" box="[151,212,1375,1401]" italics="true" pageId="7" pageNumber="308">Head</emphasis>
(
<figureCitation id="131C01FA1070ED48C2BAFA8FFED0FA87" box="[226,380,1375,1401]" captionStart="FIGURES 13–14" captionStartId="13.[152,270,1908,1932]" captionTargetBox="[216,1350,196,1865]" captionTargetId="figure-19@13.[216,1372,181,1884]" captionTargetPageId="13" captionText="FIGURES 13–14. Larvae of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 13, R. anatolica: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, R. fasciata: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="7" pageNumber="308">Figs 13a, 14a</figureCitation>
). In 
<taxonomicName id="4C2766FC1070ED48C3F5FA8FFD99FA87" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[429,565,1375,1401]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1070ED48C3F5FA8FFD99FA87" box="[429,565,1375,1401]" italics="true" pageId="7" pageNumber="308">R. anatolica</emphasis>
</taxonomicName>
, the pair of dark brown patches outside the posterior edge of the frontal suture are more or less triangular in shape, inside each patch two rows of dark spots aligned almost perpendicular to the frontal suture and a row of three or four spots aligned along sides of coronal suture. In 
<taxonomicName id="4C2766FC1070ED48C636FA77FB4BFA3F" box="[1134,1255,1447,1473]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1070ED48C636FA77FBD3FA3E" box="[1134,1151,1447,1472]" italics="true" pageId="7" pageNumber="308">R</emphasis>
. 
<emphasis id="B953C16D1070ED48C6D6FA77FB4BFA3F" box="[1166,1255,1447,1473]" italics="true" pageId="7" pageNumber="308">fasciata</emphasis>
</taxonomicName>
the pair of dark brown patches outside the posterior edge of the frontal suture are in the shape of open wings, inside the darker anterior extension of each dark brown patch there is only a row of dark spots aligned parallel to the frontal suture, two groups of spots outside coronal suture (
<bibRefCitation id="EFB6608E1070ED48C0D8F9C3FCDAF9D3" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[640,886,1555,1581]" pageId="7" pageNumber="308" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1070ED48C0A4F9C3FC98F9D3" box="[764,820,1555,1581]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2021
</bibRefCitation>
, fig. 4a).
</paragraph>
<paragraph id="8B981D7F1070ED48C29FF9E7FADBF967" blockId="7.[151,1437,1375,2013]" pageId="7" pageNumber="308">
In 
<taxonomicName id="4C2766FC1070ED48C2B1F9E7FEDAF9AF" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[233,374,1591,1617]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1070ED48C2B1F9E7FEDAF9AF" box="[233,374,1591,1617]" italics="true" pageId="7" pageNumber="308">R. anatolica</emphasis>
</taxonomicName>
each side of the head has laterally a rectangular dark band with two lines of darker spots, extending one-third the distance from the posterior edge. In 
<taxonomicName id="4C2766FC1070ED48C166F98BFC1BF98B" box="[830,951,1627,1653]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1070ED48C166F98BFC1BF98B" box="[830,951,1627,1653]" italics="true" pageId="7" pageNumber="308">R. fasciata</emphasis>
</taxonomicName>
, each side of the head has laterally an oval dark area with darker spots, extending half the distance from the posterior edge (
<bibRefCitation id="EFB6608E1070ED48C645F9AFFABFF967" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[1053,1299,1663,1689]" pageId="7" pageNumber="308" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1070ED48C6C1F9AFFB66F967" box="[1177,1226,1663,1689]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. 2021
</bibRefCitation>
, fig. 4b).
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41070ED47C29FF973FB2DFB3B" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308" type="description">
<paragraph id="8B981D7F1070ED48C29FF973FE2DF8FB" blockId="7.[151,1437,1375,2013]" pageId="7" pageNumber="308">
In 
<taxonomicName id="4C2766FC1070ED48C2BDF973FEC2F943" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[229,366,1699,1725]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1070ED48C2BDF973FEC2F943" box="[229,366,1699,1725]" italics="true" pageId="7" pageNumber="308">R. anatolica</emphasis>
</taxonomicName>
the left mandible has its inner blade nearly straight, right mandible with inconspicuous tooth. In 
<taxonomicName id="4C2766FC1070ED48C2CFF917FEBCF91F" box="[151,272,1735,1761]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1070ED48C2CFF917FEBCF91F" box="[151,272,1735,1761]" italics="true" pageId="7" pageNumber="308">R. fasciata</emphasis>
</taxonomicName>
the left mandible has its inner blade slightly convex, right mandible with small tooth (
<bibRefCitation id="EFB6608E1070ED48C6BEF917FF7EF8FB" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." pageId="7" pageNumber="308" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1070ED48C73BF917FA31F91F" box="[1379,1437,1735,1761]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
2021
</bibRefCitation>
, figs 5Dl, 5Vl).
</paragraph>
<paragraph id="8B981D7F1070ED47C29FF8DFFB18FEBF" blockId="7.[151,1437,1375,2013]" lastBlockId="8.[151,1437,151,537]" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<emphasis id="B953C16D1070ED48C29FF8DFFEB4F8D7" box="[199,280,1807,1833]" italics="true" pageId="7" pageNumber="308">Thorax</emphasis>
: In 
<taxonomicName id="4C2766FC1070ED48C31DF8DFFE62F8D7" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[325,462,1807,1833]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1070ED48C31DF8DFFE62F8D7" box="[325,462,1807,1833]" italics="true" pageId="7" pageNumber="308">R. anatolica</emphasis>
</taxonomicName>
the anterior half of the pronotum is pale; each side with two or three dark brown spots, and black band from setae S22 and S 
<quantity id="4CDFB09A1070ED48C01EF8E3FD2FF8B3" box="[582,643,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="7" pageNumber="308" unit="in" value="23.0">23 in</quantity>
anterior edge, and narrow black band in lateral edge touching broad posterolateral band. In 
<taxonomicName id="4C2766FC1070ED48C3CFF887FDA2F88F" box="[407,526,1879,1905]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1070ED48C3CFF887FDA2F88F" box="[407,526,1879,1905]" italics="true" pageId="7" pageNumber="308">R. fasciata</emphasis>
</taxonomicName>
the black band in each anterolateral angle extends posterad to join with the broad, black band of the posterolateral margin (
<bibRefCitation id="EFB6608E1070ED48C035F8ABFCC7F86B" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[621,875,1915,1941]" pageId="7" pageNumber="308" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1070ED48C0B4F8ABFC8CF86B" box="[748,800,1915,1941]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. 2021
</bibRefCitation>
, figs 6a, 6b); there is a conspicuous black circle sublaterally one-third of the distance from the anterior margin. In 
<taxonomicName id="4C2766FC1070ED48C137F84FFC5BF847" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[879,1015,1951,1977]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1070ED48C137F84FFC5BF847" box="[879,1015,1951,1977]" italics="true" pageId="7" pageNumber="308">R. anatolica</emphasis>
</taxonomicName>
on the posterior half of the pronotum, the median dark stripe is truncate anteriorly and slightly wider than posteriorly, and with its lateral edges slightly concave or straight, pair of submesal lines of six or seven slightly dark brown, longitudinally oval spots inside this central dark region, reaching posterior edge. The posterior marginal band at mid-height is pale brown and black anteriorly and posteriorly, respectively. In 
<taxonomicName id="4C2766FC107FED47C02EFF0FFD43FF07" box="[630,751,223,249]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107FED47C02EFF0FFD43FF07" box="[630,751,223,249]" italics="true" pageId="8" pageNumber="309">R. fasciata</emphasis>
</taxonomicName>
on the posterior half of the pronotum the median dark stripe is convex anteriorly and with its lateral edges slightly convex. The posterior marginal band at mid-height is dark brown anteriorly and posteriorly, and black in the middle area (
<bibRefCitation id="EFB6608E107FED47C101FEF7FBE3FEBF" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[857,1103,295,321]" pageId="8" pageNumber="309" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107FED47C18DFEF7FBA2FEBF" box="[981,1038,295,321]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
2021
</bibRefCitation>
, fig. 6b).
</paragraph>
<paragraph id="8B981D7F107FED47C297FE9BFF7BFDE7" blockId="8.[151,1437,151,537]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C297FE9BFEEDFE9B" box="[207,321,331,357]" italics="true" pageId="8" pageNumber="309">Abdomen.</emphasis>
In 
<taxonomicName id="4C2766FC107FED47C33FFE9BFE5DFE9B" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[359,497,331,357]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107FED47C33FFE9BFE5DFE9B" box="[359,497,331,357]" italics="true" pageId="8" pageNumber="309">R. anatolica</emphasis>
</taxonomicName>
tergite IX yellowish, the band of the anterior edge is continuous, black; the central area of the tergite is pale; on the posterior edge a pair of black areas is on the posterior margin behind the bases of the central setae. In 
<taxonomicName id="4C2766FC107FED47C32CFE43FE47FE53" box="[372,491,403,429]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107FED47C32CFE43FE47FE53" box="[372,491,403,429]" italics="true" pageId="8" pageNumber="309">R. fasciata</emphasis>
</taxonomicName>
tergite IX (
<bibRefCitation id="EFB6608E107FED47C031FE43FCF0FE53" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[617,860,403,429]" pageId="8" pageNumber="309" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107FED47C0BCFE43FCB9FE53" box="[740,789,403,429]" italics="true" pageId="8" pageNumber="309">et al</emphasis>
. 2021
</bibRefCitation>
, fig. 7) the band of the anterior edge is continuously black in darker larvae, and has a pair of dark brown extensions surrounding the pair of pits P1; the central area of the tergite is uniformly light brown; on the posterior edge the pair of black areas surround the bases of the central setae.
</paragraph>
<paragraph id="8B981D7F107FED47C2CFFDBAFF4FFD7A" blockId="8.[151,266,618,681]" box="[151,227,618,644]" pageId="8" pageNumber="309">
<heading id="D0D0AA13107FED47C2CFFDBAFF4FFD7A" bold="true" box="[151,227,618,644]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<emphasis id="B953C16D107FED47C2CFFDBAFF4FFD7A" bold="true" box="[151,227,618,644]" pageId="8" pageNumber="309">PUPA</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F107FED47C2CFFD5FFEA6FD57" blockId="8.[151,266,618,681]" box="[151,266,655,681]" pageId="8" pageNumber="309">
<figureCitation id="131C01FA107FED47C2CFFD5FFEA6FD57" box="[151,266,655,681]" captionStart="FIGURE 8" captionStartId="9.[152,255,1903,1927]" captionTargetBox="[300,1286,181,1878]" captionTargetId="figure-19@9.[300,1287,181,1878]" captionTargetPageId="9" captionText="FIGURE 8. Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., paired abdominal hook plates of male pupa. 8l and 8r, left and right hook plates, respectively. A = anterior hook plates, dorsal, P = posterior hook plates, dorsal; III–VII = abdominal terga III through VII, dorsal. Scale bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14239581" httpUri="https://zenodo.org/record/14239581/files/figure.png" pageId="8" pageNumber="309">Figs 8l, 8r</figureCitation>
</paragraph>
<paragraph id="8B981D7F107FED47C2CFFD07FC1CFCA3" blockId="8.[151,1437,727,1221]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C2CFFD07FDA6FD0E" box="[151,522,727,753]" italics="true" pageId="8" pageNumber="309">Biometrics of pupa and cocoon.</emphasis>
Pupal length (total): 
<quantity id="4CDFB09A107FED47C15DFD07FC31FD0F" box="[773,925,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.23" metricValueMin="0.8699999999999999" pageId="8" pageNumber="309" unit="mm" value="10.5" valueMax="12.3" valueMin="8.7">8.7–12.3 mm</quantity>
(x̄ = 10.2, 
<collectingRegion id="49E3D39D107FED47C67FFD07FBE7FD0F" box="[1063,1099,727,753]" country="Serbia" name="Sid" pageId="8" pageNumber="309">SD</collectingRegion>
= 1.04, n = 13) [male pupa: 
<quantity id="4CDFB09A107FED47C2CFFD2AFE96FCEB" box="[151,314,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="9.725" metricValueMax="10.75" metricValueMin="8.7" pageId="8" pageNumber="309" unit="mm" value="9.725" valueMax="10.75" valueMin="8.7">8.7–10.75 mm</quantity>
(x̄ = 9.8, 
<collectingRegion id="49E3D39D107FED47C3FFFD2BFE67FCEB" box="[423,459,763,789]" country="Serbia" name="Sid" pageId="8" pageNumber="309">SD</collectingRegion>
= 0.77, n = 8); female pupa: 
<quantity id="4CDFB09A107FED47C157FD2BFC08FCEB" box="[783,932,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" metricValueMax="1.23" metricValueMin="0.93" pageId="8" pageNumber="309" unit="mm" value="10.8" valueMax="12.3" valueMin="9.3">9.3–12.3 mm</quantity>
(x̄ = 10.9, 
<collectingRegion id="49E3D39D107FED47C647FD2BFBEFFCEB" box="[1055,1091,763,789]" country="Serbia" name="Sid" pageId="8" pageNumber="309">SD</collectingRegion>
= 1.17, n = 5)]. Cocoon length (total): 11.7–16.0 mm (x̄ = 13.6, 
<collectingRegion id="49E3D39D107FED47C051FCCFFD81FCC7" box="[521,557,799,825]" country="Serbia" name="Sid" pageId="8" pageNumber="309">SD</collectingRegion>
= 1.14, n = 13) [male cocoon length: 11.7–14.0 mm (x̄ = 13.2, 
<collectingRegion id="49E3D39D107FED47C6AEFCCFFAB6FCC7" box="[1270,1306,799,825]" country="Serbia" name="Sid" pageId="8" pageNumber="309">SD</collectingRegion>
= 0.76, n = 8); female cocoon length: 12.0–16.0 mm (x̄ = 14.2, 
<collectingRegion id="49E3D39D107FED47C084FC93FCACFCA3" box="[732,768,835,861]" country="Serbia" name="Sid" pageId="8" pageNumber="309">SD</collectingRegion>
= 1.46, n = 5)].
</paragraph>
<paragraph id="8B981D7F107FED47C29FFCB7FDBBFBEF" blockId="8.[151,1437,727,1221]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C29FFCB7FEA8FC7F" box="[199,260,871,897]" italics="true" pageId="8" pageNumber="309">Head</emphasis>
. Mandibles reddish brown, with basis pale, left mandible with two preapical teeth, distal tooth larger, and right mandible with three preapical teeth, distal tooth larger, both mandibles with numerous fine teeth on inner blade. Antennae of variable length, reaching posterior edge of abdominal segment IV to posterior edge of segment VIII (from middle of segment VII to posterior edge of segment VIII in males, posterior edge of segment IV to middle of segment VI in females).
</paragraph>
<paragraph id="8B981D7F107FED47C29FFBCBFD2AFBA7" blockId="8.[151,1437,727,1221]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C29FFBCBFEB4FBCB" box="[199,280,1051,1077]" italics="true" pageId="8" pageNumber="309">Thorax</emphasis>
: Tubercles of prothorax each with 5 setae. Hind wing pads variously reaching posterior edge of abdominal segment III to posterior edge of segment IV.
</paragraph>
<paragraph id="8B981D7F107FED47C29FFBB3FB2DFB3B" blockId="8.[151,1437,727,1221]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C29FFBB3FE9DFB83" box="[199,305,1123,1149]" italics="true" pageId="8" pageNumber="309">Abdomen</emphasis>
: Paired anterior hook plates pedunculate, and present on abdominal terga IV to VII (
<figureCitation id="131C01FA107FED47C681FBB3FAB0FB83" box="[1241,1308,1123,1149]" captionStart="FIGURE 8" captionStartId="9.[152,255,1903,1927]" captionTargetBox="[300,1286,181,1878]" captionTargetId="figure-19@9.[300,1287,181,1878]" captionTargetPageId="9" captionText="FIGURE 8. Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., paired abdominal hook plates of male pupa. 8l and 8r, left and right hook plates, respectively. A = anterior hook plates, dorsal, P = posterior hook plates, dorsal; III–VII = abdominal terga III through VII, dorsal. Scale bars: 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14239581" httpUri="https://zenodo.org/record/14239581/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
), each with 5–10 spines (AIV, the smallest plates) and 10–20 spines (AV and AVII). Paired posterior hook plates PIII–PV sessile and circular, plates PIV slightly bigger, slightly transversely oval (length: width = 1: 1.2).
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF4107FED47C2CFFAC6FEE9F9F7" pageId="8" pageNumber="309" type="diagnosis">
<paragraph id="8B981D7F107FED47C2CFFAC6FE28FACF" blockId="8.[151,388,1302,1329]" box="[151,388,1302,1329]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C2CFFAC6FE28FACF" bold="true" box="[151,388,1302,1329]" pageId="8" pageNumber="309">Diagnosis of pupae</emphasis>
</paragraph>
<paragraph id="8B981D7F107FED47C2CFFA8FFEE9F9F7" blockId="8.[151,1437,1375,1545]" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C2CFFA8FFEADFA87" box="[151,257,1375,1401]" italics="true" pageId="8" pageNumber="309">Abdomen</emphasis>
. In 
<taxonomicName id="4C2766FC107FED47C376FA8FFE14FA87" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[302,440,1375,1401]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107FED47C376FA8FFE14FA87" box="[302,440,1375,1401]" italics="true" pageId="8" pageNumber="309">R. anatolica</emphasis>
</taxonomicName>
the anterior hook plates AIV are much smaller than the other plates; the paired posterior hook plates are almost circular except for oval PIV (length: width = 1: 1.2). In 
<taxonomicName id="4C2766FC107FED47C64FFA53FB3CFA63" box="[1047,1168,1411,1437]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107FED47C64FFA53FB3CFA63" box="[1047,1168,1411,1437]" italics="true" pageId="8" pageNumber="309">R. fasciata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB6608E107FED47C6C7FA53FA34FA63" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[1183,1432,1411,1437]" pageId="8" pageNumber="309" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107FED47C744FA53FAE3FA63" box="[1308,1359,1411,1437]" italics="true" pageId="8" pageNumber="309">et al</emphasis>
. 2021
</bibRefCitation>
, fig. 10), a pair of small flat anterior plates AIII without spines, and the paired anterior hook plates AIV are slightly smaller than the other plates; the paired posterior hook plates PIII–PV are all slightly transversely oval (length: width = 1: 1.3).
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF4107FED43C2CFF98AFBAFFD9F" lastPageId="12" lastPageNumber="313" pageId="8" pageNumber="309" type="description">
<paragraph id="8B981D7F107FED47C2CFF98AFF57F98A" blockId="8.[151,276,1626,1689]" box="[151,251,1626,1652]" pageId="8" pageNumber="309">
<heading id="D0D0AA13107FED47C2CFF98AFF57F98A" bold="true" box="[151,251,1626,1652]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<emphasis id="B953C16D107FED47C2CFF98AFF57F98A" bold="true" box="[151,251,1626,1652]" pageId="8" pageNumber="309">ADULT</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F107FED47C2CFF9AFFEB8F967" blockId="8.[151,276,1626,1689]" box="[151,276,1663,1689]" pageId="8" pageNumber="309">
<figureCitation id="131C01FA107FED47C2CFF9AFFEB8F967" box="[151,276,1663,1689]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="10.[152,270,1062,1086]" captionStartId-1="11.[152,270,1787,1811]" captionTargetBox-0="[201,1382,198,1025]" captionTargetBox-1="[260,1320,192,1752]" captionTargetId-0="figure-26@10.[192,1395,198,1038]" captionTargetId-1="figure-19@11.[257,1331,181,1762]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 9–10. Wings of Rhyacophila fasciata Hagen 1859 and Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 9a, forewing of R. fasciata, lectotype [Museum of Comparative Zoology of Harvard University, Cambridge, MA, USA (MCZH), with permission]; 9b, forewing of R. fasciata male from Slovakia; 9c, hind wing of R. fasciata male from Slovakia. 10a, forewing of R. anatolica male. 10b, hind wing of R. anatolica male. St = stigma. Major veins: C = costa, Sc = subcosta, R 1 –R = 5 radius, M 1 –M 4 = media, Cu1–Cu2 = cubitus, A1–A3 = anal; m-cu = crossvein between medial and cubital veins, cu-a = crossvein between second cubital and first anal veins (from Holzenthal et al. 2007). Scale bars: 2 mm. 9a: specimen preserved dry, 9b–c, 10a–b: specimens preserved in ethanol." captionText-1="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.14239583" figureDoi-1="http://doi.org/10.5281/zenodo.14239585" httpUri-0="https://zenodo.org/record/14239583/files/figure.png" httpUri-1="https://zenodo.org/record/14239585/files/figure.png" pageId="8" pageNumber="309">Figs 10–12</figureCitation>
</paragraph>
<paragraph id="8B981D7F107FED45C2CFF910FB25FA34" blockId="8.[151,1436,1728,2048]" lastBlockId="10.[151,1436,1384,1482]" lastPageId="10" lastPageNumber="311" pageId="8" pageNumber="309">
<emphasis id="B953C16D107FED47C2CFF910FF4AF924" box="[151,230,1728,1754]" italics="true" pageId="8" pageNumber="309">Colour</emphasis>
. Specimens preserved in ethanol reddish brown (some specimens yellowish) dorsally, yellowish ventrally, spurs reddish brown, setae brown, females generally darker than males; small black or dark brown spots in dorsal area of abdomen, in lateral view, some specimens with large elongate black spots marking border between dorsal and ventral areas. Forewings (
<figureCitation id="131C01FA107FED47C3B4F8FCFDE8F8B8" box="[492,580,1836,1862]" captionStart="FIGURES 9–10" captionStartId="10.[152,270,1062,1086]" captionTargetBox="[201,1382,198,1025]" captionTargetId="figure-26@10.[192,1395,198,1038]" captionTargetPageId="10" captionText="FIGURES 9–10. Wings of Rhyacophila fasciata Hagen 1859 and Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 9a, forewing of R. fasciata, lectotype [Museum of Comparative Zoology of Harvard University, Cambridge, MA, USA (MCZH), with permission]; 9b, forewing of R. fasciata male from Slovakia; 9c, hind wing of R. fasciata male from Slovakia. 10a, forewing of R. anatolica male. 10b, hind wing of R. anatolica male. St = stigma. Major veins: C = costa, Sc = subcosta, R 1 –R = 5 radius, M 1 –M 4 = media, Cu1–Cu2 = cubitus, A1–A3 = anal; m-cu = crossvein between medial and cubital veins, cu-a = crossvein between second cubital and first anal veins (from Holzenthal et al. 2007). Scale bars: 2 mm. 9a: specimen preserved dry, 9b–c, 10a–b: specimens preserved in ethanol." figureDoi="http://doi.org/10.5281/zenodo.14239583" httpUri="https://zenodo.org/record/14239583/files/figure.png" pageId="8" pageNumber="309">Fig 10a</figureCitation>
) dark, each with small pale spots or uniformly coloured, one or two central transversal dark bands (fascia) in distal area more or less conspicuous, and one central dark spot in proximal area, with pale thick stigma (St) on leading edge between subcostal (Sc) and first radial (R 
<subScript id="17A31F3A107FED47C608F856FBF4F86A" attach="left" box="[1104,1112,1926,1940]" fontSize="6" pageId="8" pageNumber="309">1</subScript>
) veins, hyaline circular zone in connection area of medial vein (M) and 
<emphasis id="B953C16D107FED47C02AF849FD06F84C" box="[626,682,1945,1970]" italics="true" pageId="8" pageNumber="309">m-cu</emphasis>
crossvein; additional crossvein connecting costal (C) and subcostal (Sc) veins in some specimens, crossveins connecting R 
<subScript id="17A31F3A107FED47C0AFF81EFD53F822" attach="left" box="[759,767,1998,2012]" fontSize="6" pageId="8" pageNumber="309">1</subScript>
+R 
<subScript id="17A31F3A107FED47C17AF81EFC86F822" attach="left" box="[802,810,1998,2012]" fontSize="6" pageId="8" pageNumber="309">2</subScript>
(
<emphasis id="B953C16D107FED47C16FF86DFCEEF828" box="[823,834,1981,2006]" italics="true" pageId="8" pageNumber="309">r</emphasis>
) and fifth radial (R 
<subScript id="17A31F3A107FED47C64EF81EFBB2F822" attach="left" box="[1046,1054,1998,2012]" fontSize="6" pageId="8" pageNumber="309">5</subScript>
), and first medial (M 
<subScript id="17A31F3A107FED47C752F81EFABEF822" attach="right" box="[1290,1298,1998,2012]" fontSize="6" pageId="8" pageNumber="309">1</subScript>
) veins (
<emphasis id="B953C16D107FED47C733F86DFA3FF828" box="[1387,1427,1981,2006]" italics="true" pageId="8" pageNumber="309">r-m</emphasis>
) present, distal segment of second anal (A 
<subScript id="17A31F3A107FED47C038F822FDC4F7FE" attach="right" box="[608,616,2034,2048]" fontSize="6" pageId="8" pageNumber="309">2</subScript>
) connecting first and third anal (A 
<subScript id="17A31F3A107FED47C1B2F822FC5EF7FE" attach="left" box="[1002,1010,2034,2048]" fontSize="6" pageId="8" pageNumber="309">1</subScript>
and A 
<subScript id="17A31F3A107FED47C664F822FBE8F7FE" attach="right" box="[1084,1092,2034,2048]" fontSize="6" pageId="8" pageNumber="309">3</subScript>
) not conspicuous. Hind wings (
<figureCitation id="131C01FA107DED45C2C7FAB8FEACFA7C" box="[159,256,1384,1410]" captionStart="FIGURES 9–10" captionStartId="10.[152,270,1062,1086]" captionTargetBox="[201,1382,198,1025]" captionTargetId="figure-26@10.[192,1395,198,1038]" captionTargetPageId="10" captionText="FIGURES 9–10. Wings of Rhyacophila fasciata Hagen 1859 and Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. 9a, forewing of R. fasciata, lectotype [Museum of Comparative Zoology of Harvard University, Cambridge, MA, USA (MCZH), with permission]; 9b, forewing of R. fasciata male from Slovakia; 9c, hind wing of R. fasciata male from Slovakia. 10a, forewing of R. anatolica male. 10b, hind wing of R. anatolica male. St = stigma. Major veins: C = costa, Sc = subcosta, R 1 –R = 5 radius, M 1 –M 4 = media, Cu1–Cu2 = cubitus, A1–A3 = anal; m-cu = crossvein between medial and cubital veins, cu-a = crossvein between second cubital and first anal veins (from Holzenthal et al. 2007). Scale bars: 2 mm. 9a: specimen preserved dry, 9b–c, 10a–b: specimens preserved in ethanol." figureDoi="http://doi.org/10.5281/zenodo.14239583" httpUri="https://zenodo.org/record/14239583/files/figure.png" pageId="10" pageNumber="311">Fig. 10b</figureCitation>
) pale, each with pale thick stigma on leading edge between subcostal (Sc) and second radial (R2) veins; proximal portion of medial vein (M) not conspicuous, median crossvein 
<emphasis id="B953C16D107DED45C183FA5DFC43FA58" box="[987,1007,1421,1446]" italics="true" pageId="10" pageNumber="311">m</emphasis>
missing; first cubital (Cu 
<subScript id="17A31F3A107DED45C74FFA4EFA98FA52" attach="left" box="[1303,1332,1438,1452]" fontSize="6" pageId="10" pageNumber="311">1a-b</subScript>
) thicker; proximal crossvein 
<emphasis id="B953C16D107DED45C32CFA61FE0AFA34" box="[372,422,1457,1482]" italics="true" pageId="10" pageNumber="311">cu-a</emphasis>
connecting second cubital (Cu 2) and first anal (A 1) veins missing.
</paragraph>
<caption id="DF584DF7107EED46C2C0F8BFFDEFF831" ID-DOI="http://doi.org/10.5281/zenodo.14239581" ID-Zenodo-Dep="14239581" httpUri="https://zenodo.org/record/14239581/files/figure.png" pageId="9" pageNumber="310" startId="9.[152,255,1903,1927]" targetBox="[300,1286,181,1878]" targetPageId="9" targetType="figure">
<paragraph id="8B981D7F107EED46C2C0F8BFFDEFF831" blockId="9.[151,1436,1903,1999]" pageId="9" pageNumber="310">
<emphasis id="B953C16D107EED46C2C0F8BFFEB4F879" bold="true" box="[152,280,1903,1927]" pageId="9" pageNumber="310">FIGURE 8.</emphasis>
<taxonomicName id="4C2766FC107EED46C347F8BFFD45F879" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[287,745,1903,1927]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D107EED46C347F8BFFDAAF879" box="[287,518,1903,1927]" italics="true" pageId="9" pageNumber="310">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D107EED46C0A8F8BFFCEDF879" bold="true" box="[752,833,1903,1927]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="A2607C16107EED46C0A8F8BFFCEDF879" box="[752,833,1903,1927]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paired abdominal hook plates of male pupa. 8 
<emphasis id="B953C16D107EED46C746F8BFFA89F879" box="[1310,1317,1903,1927]" italics="true" pageId="9" pageNumber="310">l</emphasis>
and 8 
<emphasis id="B953C16D107EED46C73CF8A0FAC2F879" box="[1380,1390,1904,1927]" italics="true" pageId="9" pageNumber="310">r</emphasis>
, left and right hook plates, respectively. A = anterior hook plates, dorsal, P = posterior hook plates, dorsal; III–VII = abdominal terga III through VII, dorsal. Scale bars: 50 μm.
</paragraph>
</caption>
<caption id="DF584DF7107DED45C2C0FBF6FD8BFAE8" ID-DOI="http://doi.org/10.5281/zenodo.14239583" ID-Zenodo-Dep="14239583" httpUri="https://zenodo.org/record/14239583/files/figure.png" pageId="10" pageNumber="311" startId="10.[152,270,1062,1086]" targetBox="[201,1382,198,1025]" targetPageId="10" targetType="figure">
<paragraph id="8B981D7F107DED45C2C0FBF6FD8BFAE8" blockId="10.[151,1437,1062,1302]" pageId="10" pageNumber="311">
<emphasis id="B953C16D107DED45C2C0FBF6FEE3FBC0" bold="true" box="[152,335,1062,1086]" pageId="10" pageNumber="311">FIGURES 9–10.</emphasis>
Wings of 
<taxonomicName id="4C2766FC107DED45C3E4FBF6FCB5FBC0" authority="Hagen 1859" authorityName="Hagen" authorityYear="1859" box="[444,793,1062,1086]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107DED45C3E4FBF6FD38FBC0" box="[444,660,1062,1086]" italics="true" pageId="10" pageNumber="311">Rhyacophila fasciata</emphasis>
<bibRefCitation id="EFB6608E107DED45C0C4FBF6FCB5FBC0" author="Hagen, H." box="[668,793,1062,1086]" pageId="10" pageNumber="311" pagination="4 - 6" refId="ref13323" refString="Hagen, H. (1859) Die Phryganiden Pictet's. Entomologische Zeitung, 20 (4 - 6), 131 - 170. Available from: https: // www. biodiversitylibrary. org / item / 35933 # page / 141 / mode / 1 up (accessed 5 March 2024)" type="journal article" year="1859">Hagen 1859</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C2766FC107DED45C115FBF6FAB0FBC0" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[845,1308,1062,1086]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D107DED45C115FBF6FB99FBC0" box="[845,1077,1062,1086]" italics="true" pageId="10" pageNumber="311">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D107DED45C77CFBF6FADAFBC0" bold="true" box="[1316,1398,1062,1086]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A2607C16107DED45C77CFBF6FADAFBC0" box="[1316,1398,1062,1086]" pageId="10" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9a, forewing of 
<taxonomicName id="4C2766FC107DED45C34EFB9BFE2FFB9C" authorityName="Hagen" authorityYear="1859" box="[278,387,1098,1122]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107DED45C34EFB9BFE2FFB9C" box="[278,387,1098,1122]" italics="true" pageId="10" pageNumber="311">R. fasciata</emphasis>
</taxonomicName>
, lectotype [Museum of Comparative Zoology of Harvard University, Cambridge, MA, USA (MCZH), with permission]; 9b, forewing of 
<taxonomicName id="4C2766FC107DED45C05CFBBFFDD8FB78" authorityName="Hagen" authorityYear="1859" box="[516,628,1134,1158]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107DED45C05CFBBFFDD8FB78" box="[516,628,1134,1158]" italics="true" pageId="10" pageNumber="311">R. fasciata</emphasis>
</taxonomicName>
male from Slovakia; 9c, hind wing of 
<taxonomicName id="4C2766FC107DED45C640FBBFFB24FB78" authorityName="Hagen" authorityYear="1859" box="[1048,1160,1134,1158]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107DED45C640FBBFFB24FB78" box="[1048,1160,1134,1158]" italics="true" pageId="10" pageNumber="311">R. fasciata</emphasis>
</taxonomicName>
male from Slovakia. 10a, forewing of 
<taxonomicName id="4C2766FC107DED45C34BFB43FE23FB54" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[275,399,1170,1194]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107DED45C34BFB43FE23FB54" box="[275,399,1170,1194]" italics="true" pageId="10" pageNumber="311">R. anatolica</emphasis>
</taxonomicName>
male. 10b, hind wing of 
<taxonomicName id="4C2766FC107DED45C0D2FB43FCA9FB54" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[650,773,1170,1194]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107DED45C0D2FB43FCA9FB54" box="[650,773,1170,1194]" italics="true" pageId="10" pageNumber="311">R. anatolica</emphasis>
</taxonomicName>
male. St = stigma. Major veins: C = costa, Sc = subcosta, R 
<subScript id="17A31F3A107DED45C706FB72FAC9FB4E" attach="left" box="[1374,1381,1186,1200]" fontSize="6" pageId="10" pageNumber="311">1</subScript>
–R = 
<subScript id="17A31F3A107DED45C7DBFB73FA26FB4F" attach="left" box="[1411,1418,1187,1201]" fontSize="6" pageId="10" pageNumber="311">5</subScript>
radius, M 
<subScript id="17A31F3A107DED45C2A0FB16FF53FB2A" attach="left" box="[248,255,1222,1236]" fontSize="6" pageId="10" pageNumber="311">1</subScript>
–M 
<subScript id="17A31F3A107DED45C37BFB16FE86FB2A" attach="left" box="[291,298,1222,1236]" fontSize="6" pageId="10" pageNumber="311">4</subScript>
= media, Cu 
<subScript id="17A31F3A107DED45C3F4FB16FE1FFB2A" attach="left" box="[428,435,1222,1236]" fontSize="6" pageId="10" pageNumber="311">1</subScript>
–Cu 
<subScript id="17A31F3A107DED45C386FB16FE49FB2A" attach="left" box="[478,485,1222,1236]" fontSize="6" pageId="10" pageNumber="311">2</subScript>
= cubitus, A 
<subScript id="17A31F3A107DED45C03EFB16FDC1FB2A" attach="left" box="[614,621,1222,1236]" fontSize="6" pageId="10" pageNumber="311">1</subScript>
–A 
<subScript id="17A31F3A107DED45C0D4FB16FD3FFB2A" attach="left" box="[652,659,1222,1236]" fontSize="6" pageId="10" pageNumber="311">3</subScript>
= anal; 
<emphasis id="B953C16D107DED45C0BDFB67FCB4FB30" box="[741,792,1207,1230]" italics="true" pageId="10" pageNumber="311">m-cu</emphasis>
= crossvein between medial and cubital veins, 
<emphasis id="B953C16D107DED45C6ADFB67FA8EFB30" box="[1269,1314,1207,1230]" italics="true" pageId="10" pageNumber="311">cu-a</emphasis>
= crossvein between second cubital and first anal veins (from 
<bibRefCitation id="EFB6608E107DED45C0C9FB0AFCDBFB0C" author="Holzenthal, R. W. &amp; Blahnik, R. J. &amp; Prather, A. L. &amp; Kjer, K. M." box="[657,887,1242,1266]" pageId="10" pageNumber="311" pagination="639 - 698" refId="ref13691" refString="Holzenthal, R. W., Blahnik, R. J., Prather, A. L. &amp; Kjer, K. M. (2007) Order Trichoptera Kirby, 1813 (Insecta), Caddisflies. Zootaxa, 1668 (1), 639 - 698. https: // doi. org / 10.11646 / zootaxa. 1668.1.29" type="journal article" year="2007">
Holzenthal 
<emphasis id="B953C16D107DED45C15EFB0BFC96FB0C" box="[774,826,1242,1266]" italics="true" pageId="10" pageNumber="311">et al.</emphasis>
2007
</bibRefCitation>
). Scale bars: 2 mm. 9a: specimen preserved dry, 9b–c, 10a–b: specimens preserved in ethanol.
</paragraph>
</caption>
<paragraph id="8B981D7F107DED45C2CFF9CCFF43F9C8" blockId="10.[151,239,1564,1590]" box="[151,239,1564,1590]" pageId="10" pageNumber="311">
<heading id="D0D0AA13107DED45C2CFF9CCFF43F9C8" bold="true" box="[151,239,1564,1590]" fontSize="11" level="1" pageId="10" pageNumber="311" reason="1">
<emphasis id="B953C16D107DED45C2CFF9CCFF43F9C8" bold="true" box="[151,239,1564,1590]" pageId="10" pageNumber="311">MALE</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F107DED45C2CFF9B5FA22F939" blockId="10.[151,1437,1636,1879]" pageId="10" pageNumber="311">
<emphasis id="B953C16D107DED45C2CFF9B5FEBCF980" box="[151,272,1637,1662]" italics="true" pageId="10" pageNumber="311">Biometrics</emphasis>
. 
<emphasis id="B953C16D107DED45C346F9B5FE36F980" box="[286,410,1637,1662]" italics="true" pageId="10" pageNumber="311">Type series</emphasis>
: Lengths: Body (distance from front of head to distal edge of segment IX): 
<quantity id="4CDFB09A107DED45C6B0F9B4FAD1F981" box="[1256,1405,1636,1663]" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="10.2" metricValueMin="7.5" pageId="10" pageNumber="311" unit="mm" value="8.85" valueMax="10.2" valueMin="7.5">7.5–10.2 mm</quantity>
(x̄ = 9.0, 
<collectingRegion id="49E3D39D107DED45C287F958FEAFF95C" box="[223,259,1672,1698]" country="Serbia" name="Sid" pageId="10" pageNumber="311">SD</collectingRegion>
= 0.67, n = 43), each forewing 9.6–12.5 (x̄ = 11.2, 
<collectingRegion id="49E3D39D107DED45C119F958FCC9F95C" box="[833,869,1672,1698]" country="Serbia" name="Sid" pageId="10" pageNumber="311">SD</collectingRegion>
= 0.69, n = 43), each hind wing 
<quantity id="4CDFB09A107DED45C68AF958FACAF95D" box="[1234,1382,1672,1699]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.9" metricValueMin="8.3" pageId="10" pageNumber="311" unit="mm" value="9.6" valueMax="10.9" valueMin="8.3">8.3–10.9 mm</quantity>
(x̄ = 9.7, 
<collectingRegion id="49E3D39D107DED45C290F97CFF40F938" box="[200,236,1708,1734]" country="Serbia" name="Sid" pageId="10" pageNumber="311">SD</collectingRegion>
= 0.61, n = 43). 
<emphasis id="B953C16D107DED45C3F0F97CFDE0F938" box="[424,588,1708,1734]" italics="true" pageId="10" pageNumber="311">
<typeStatus id="549CA3DD107DED45C3F0F97CFDA3F938" box="[424,527,1708,1734]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
male
</emphasis>
Lengths: Body, 
<quantity id="4CDFB09A107DED45C15DF97CFCF1F939" box="[773,861,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="311" unit="mm" value="9.7">9.7 mm</quantity>
, each forewing 
<quantity id="4CDFB09A107DED45C657F97CFBD8F939" box="[1039,1140,1708,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="10" pageNumber="311" unit="mm" value="11.1">11.1 mm</quantity>
, each hind wing 
<quantity id="4CDFB09A107DED45C769F97CFA26F939" box="[1329,1418,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="311" unit="mm" value="9.7">9.7 mm</quantity>
.
</paragraph>
<paragraph id="8B981D7F107DED45C29FF900FBE4F8A9" blockId="10.[151,1437,1636,1879]" pageId="10" pageNumber="311">
<emphasis id="B953C16D107DED45C29FF900FE9DF914" box="[199,305,1744,1770]" italics="true" pageId="10" pageNumber="311">Genitalia</emphasis>
(
<figureCitation id="131C01FA107DED45C367F900FE39F914" box="[319,405,1744,1770]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="10" pageNumber="311">Figs 11</figureCitation>
, 
<figureCitation id="131C01FA107DED45C3F8F900FE11F915" box="[416,445,1744,1771]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="10" pageNumber="311">15</figureCitation>
) as for 
<taxonomicName id="4C2766FC107DED45C04AF901FD26F914" authorityName="Hagen" authorityYear="1859" box="[530,650,1744,1770]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107DED45C04AF901FD26F914" box="[530,650,1744,1770]" italics="true" pageId="10" pageNumber="311">R. fasciata</emphasis>
</taxonomicName>
except as follows: 2nd segment of each inferior appendage (
<figureCitation id="131C01FA107DED45C776F900FA3BF914" box="[1326,1431,1744,1770]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="10" pageNumber="311">Figs 11A</figureCitation>
, 
<figureCitation id="131C01FA107DED45C2CFF924FF6DF8F1" box="[151,193,1780,1807]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="10" pageNumber="311">15a</figureCitation>
) with basal and distal edges diverging, dorsal and ventral edges diverging, posterior edge straight or slightly convex, ventral edge slightly concave, 2 times longer that dorsal edge; apicodorsal vertex of 2nd segment ~ 120° and round, apicoventral angle ~ 50°, projecting as thick lobe tapering to round apex.
</paragraph>
<caption id="DF584DF7107CED44C2C0F92BFDC9F815" ID-DOI="http://doi.org/10.5281/zenodo.14239585" ID-Zenodo-Dep="14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="11" pageNumber="312" startId="11.[152,270,1787,1811]" targetBox="[260,1320,192,1752]" targetPageId="11" targetType="figure">
<paragraph id="8B981D7F107CED44C2C0F92BFDC9F815" blockId="11.[151,1437,1787,2029]" pageId="11" pageNumber="312">
<emphasis id="B953C16D107CED44C2C0F92BFEF5F8ED" bold="true" box="[152,345,1787,1811]" pageId="11" pageNumber="312">FIGURES 11–12.</emphasis>
Male and female genitalia of 
<taxonomicName id="4C2766FC107CED44C0D1F92BFBFFF8ED" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[649,1107,1787,1811]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D107CED44C0D1F92BFCDCF8ED" box="[649,880,1787,1811]" italics="true" pageId="11" pageNumber="312">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D107CED44C602F92BFB06F8ED" bold="true" box="[1114,1194,1787,1811]" pageId="11" pageNumber="312">
<taxonomicNameLabel id="A2607C16107CED44C602F92BFB06F8ED" box="[1114,1194,1787,1811]" pageId="11" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm.
</paragraph>
</caption>
<paragraph id="8B981D7F107BED43C29FFF47FBF3FE53" blockId="12.[151,1437,150,609]" pageId="12" pageNumber="313">
Parameres in ventral view curved posteromesad in distal half (
<figureCitation id="131C01FA107BED43C1DFFF47FC74FF4F" box="[903,984,151,177]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Figs 11</figureCitation>
BV, p; 15c). In lateral view (
<figureCitation id="131C01FA107BED43C745FF47FADCFF4F" box="[1309,1392,151,177]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Figs 11</figureCitation>
BL, 15d) each constricted 1/3 distance beyond base, dilated at middle, with round dorsal and ventral margins, almost parallel; along midventral margin abundant spines; midlateral surface with long spicules or setae of similar size, from midventral to dorsal edges. Aedeagus (phallicata) in lateral view (
<figureCitation id="131C01FA107BED43C19FFED3FBBAFEE3" box="[967,1046,259,285]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
CL) with anterodorsal margin and posterior corner of dorsal concavity hooked anterad, apical edge of aedeagus concave; ventral lobe of aedeagus in ventral view triangular (
<figureCitation id="131C01FA107BED43C3FFFE9BFE58FE9B" box="[423,500,331,357]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
BV, vl), in lateral view vertical edge concave, posteroventral angle broadly convex and obtuse (
<figureCitation id="131C01FA107BED43C345FEBFFEC6FE77" box="[285,362,367,393]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
CL). In dorsal view posterior edges of lateroventral lobes of phallus apically convex, apicolateral margins straight or slightly convex, lateral margins diverging anterad (
<figureCitation id="131C01FA107BED43C1F4FE43FC56FE53" box="[940,1018,403,429]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
CD, lvl).
</paragraph>
<paragraph id="8B981D7F107BED43C29FFE67FBAFFD9F" blockId="12.[151,1437,150,609]" pageId="12" pageNumber="313">
Apicodorsal lobe of segment IX in dorsal view dilated subapicolaterally, wider than long (
<figureCitation id="131C01FA107BED43C6EDFE67FAABFE2F" box="[1205,1287,439,465]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Figs 11</figureCitation>
DD, al; 15b), with broad apicomesal excision; preanal appendages (
<figureCitation id="131C01FA107BED43C0A1FE0BFCEBFE0B" box="[761,839,475,501]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DD, pa) rectangular, 2/3 as long as apicodorsal lobe, posterior edges convex. In ventral view, apical band oval (
<figureCitation id="131C01FA107BED43C145FE2FFCC5FDE7" box="[797,873,511,537]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DV, ab), much longer than wide, its posterior arms parallel and directed posterad, and round apically, its non-sclerotized mesal area triangular (
<figureCitation id="131C01FA107BED43C6EBFDF3FAADFDC3" box="[1203,1281,547,573]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DV, va); anal sclerites triangular, posterior edges straight or slightly convex (
<figureCitation id="131C01FA107BED43C101FD97FC0AFD9F" box="[857,934,583,609]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DV, as).
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF4107BED43C2CFFD62FED0F91F" pageId="12" pageNumber="313" type="diagnosis">
<paragraph id="8B981D7F107BED43C2CFFD62FED3FD33" blockId="12.[151,383,690,717]" box="[151,383,690,717]" pageId="12" pageNumber="313">
<emphasis id="B953C16D107BED43C2CFFD62FED3FD33" bold="true" box="[151,383,690,717]" pageId="12" pageNumber="313">Diagnosis of males</emphasis>
</paragraph>
<paragraph id="8B981D7F107BED43C2CFFD2BFDF2FCA3" blockId="12.[151,1437,763,1761]" pageId="12" pageNumber="313">
<emphasis id="B953C16D107BED43C2CFFD2BFEADFCEB" box="[151,257,763,789]" italics="true" pageId="12" pageNumber="313">Genitalia</emphasis>
. In 
<taxonomicName id="4C2766FC107BED43C374FD2BFE19FCEB" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[300,437,763,789]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107BED43C374FD2BFE19FCEB" box="[300,437,763,789]" italics="true" pageId="12" pageNumber="313">R. anatolica</emphasis>
</taxonomicName>
the posterior edge of the second segment of each inferior appendage is straight or slightly convex. In 
<emphasis id="B953C16D107BED43C342FCCFFE37FCC7" box="[282,411,799,825]" italics="true" pageId="12" pageNumber="313">
<taxonomicName id="4C2766FC107BED43C342FCCFFE34FCC7" authorityName="Hagen" authorityYear="1859" box="[282,408,799,825]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="fasciata">R. fasciata</taxonomicName>
,
</emphasis>
the posterior edge of the second segment of each inferior appendage is slightly concave dorsally, and convex ventrally (
<figureCitation id="131C01FA107BED43C3AFFC93FDFDFCA3" box="[503,593,835,861]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="12" pageNumber="313">Fig 16a</figureCitation>
).
</paragraph>
<paragraph id="8B981D7F107BED43C29FFCB7FEF7FBA7" blockId="12.[151,1437,763,1761]" pageId="12" pageNumber="313">
In 
<taxonomicName id="4C2766FC107BED43C2BFFCB7FEDDFC7F" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[231,369,871,897]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107BED43C2BFFCB7FEDDFC7F" box="[231,369,871,897]" italics="true" pageId="12" pageNumber="313">R. anatolica</emphasis>
</taxonomicName>
the posterior edge of the aedeagus is concave in lateral view; the ventral lobe of the aedeagus is subtriangular, in lateral view round, obtuse; posterior edges of the lateroventral lobes of phallus are convex, the apicolateral margins are straight or slightly convex, and the lateral margins are diverging anteriorly. In 
<taxonomicName id="4C2766FC107BED43C745FC7FFA3AFC37" authorityName="Hagen" authorityYear="1859" box="[1309,1430,943,969]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107BED43C745FC7FFA3AFC37" box="[1309,1430,943,969]" italics="true" pageId="12" pageNumber="313">R. fasciata</emphasis>
</taxonomicName>
, the posterior edge of the aedeagus is straight, projected posteroventrad, and rounded at its posteroventral apex; the ventral lobe of the aedeagus is subtriangular and round posteroventrally (
<figureCitation id="131C01FA107BED43C1A6FC27FBF2FBEF" box="[1022,1118,1015,1041]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="12" pageNumber="313">Fig. 16c</figureCitation>
), its lateral edges are nearly straight, almost parallel; the lateroventral lobes of the phallus are straight, its posterior edges convex, apicolateral margins rounded.
</paragraph>
<paragraph id="8B981D7F107BED43C29FFBB3FC96FA1B" blockId="12.[151,1437,763,1761]" pageId="12" pageNumber="313">
In 
<taxonomicName id="4C2766FC107BED43C2B2FBB3FEDBFB83" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[234,375,1123,1149]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107BED43C2B2FBB3FEDBFB83" box="[234,375,1123,1149]" italics="true" pageId="12" pageNumber="313">R. anatolica</emphasis>
</taxonomicName>
the apicodorsal lobe of segment IX in dorsal view is dilated subapicolaterally, wider than long (
<figureCitation id="131C01FA107BED43C281FB57FE93FB5F" box="[217,319,1158,1185]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="12" pageNumber="313">Figs 15b</figureCitation>
; 
<figureCitation id="131C01FA107BED43C314FB57FEC9FB5F" box="[332,357,1159,1185]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">11</figureCitation>
DD, al), with broad apicomesal excision; the preanal appendages are elongate rectangular, with posterior edges convex; the apical band in ventral view (
<figureCitation id="131C01FA107BED43C177FB7BFCD3FB3B" box="[815,895,1195,1221]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DV, ab) is oval, 1.4 times as long as wide, its posterior arms are parallel and directed posterad; the non-sclerotized area between these arms (
<figureCitation id="131C01FA107BED43C68FFB1FFA89FB17" box="[1239,1317,1231,1257]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DV, va) is triangular; the anal sclerites (
<figureCitation id="131C01FA107BED43C3B9FB23FD83FAF3" box="[481,559,1267,1293]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
DV, as) are straight or slightly convex apically. In 
<taxonomicName id="4C2766FC107BED43C628FB23FB45FAF3" authorityName="Hagen" authorityYear="1859" box="[1136,1257,1267,1293]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107BED43C628FB23FB45FAF3" box="[1136,1257,1267,1293]" italics="true" pageId="12" pageNumber="313">R. fasciata</emphasis>
</taxonomicName>
the apicodorsal lobe of segment IX in dorsal view (
<figureCitation id="131C01FA107BED43C042FAC7FDD6FACF" box="[538,634,1303,1329]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="12" pageNumber="313">Fig. 16b</figureCitation>
; 
<bibRefCitation id="EFB6608E107BED43C0DCFAC7FCD8FACF" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[644,884,1303,1329]" pageId="12" pageNumber="313" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107BED43C0A5FAC7FC9FFACF" box="[765,819,1303,1329]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
2021
</bibRefCitation>
, fig. 12DD, al) is dilated subapicolaterally, almost round, with a shallow apicomesal excision in some specimens; the preanal appendages (
<bibRefCitation id="EFB6608E107BED43C62CFAEBFAC7FAAB" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[1140,1387,1339,1365]" pageId="12" pageNumber="313" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107BED43C6A8FAEBFA85FAAB" box="[1264,1321,1339,1365]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
2021
</bibRefCitation>
, fig. 12DD, pa) are round, convex laterally; the apical band in ventral view (
<bibRefCitation id="EFB6608E107BED43C1EFFA8FFB01FA87" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[951,1197,1375,1401]" pageId="12" pageNumber="313" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107BED43C66BFA8FFBC0FA87" box="[1075,1132,1375,1401]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
2021
</bibRefCitation>
, fig. 12DV, ab) is 1.2 times as long as wide, its posterior arms widely separate and straight; the non-sclerotized area between these arms (
<bibRefCitation id="EFB6608E107BED43C2C6FA77FE35FA3F" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[158,409,1447,1473]" pageId="12" pageNumber="313" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107BED43C344FA77FEFAFA3F" box="[284,342,1447,1473]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
2021
</bibRefCitation>
, fig. 12DV, va) is heart-shaped, and with a wider posteromesal excision; the anal sclerites (
<bibRefCitation id="EFB6608E107BED43C2C6FA1BFE39FA1B" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[158,405,1483,1509]" pageId="12" pageNumber="313" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107BED43C343FA1BFEFFFA1B" box="[283,339,1483,1509]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
2021
</bibRefCitation>
, fig. 12DV, as) are truncated apically.
</paragraph>
<paragraph id="8B981D7F107BED43C29FFA3FFED0F91F" blockId="12.[151,1437,763,1761]" pageId="12" pageNumber="313">
In 
<taxonomicName id="4C2766FC107BED43C2BFFA3FFEDDF9F7" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[231,369,1519,1545]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107BED43C2BFFA3FFEDDF9F7" box="[231,369,1519,1545]" italics="true" pageId="12" pageNumber="313">R. anatolica</emphasis>
</taxonomicName>
the parameres in lateral view (
<figureCitation id="131C01FA107BED43C08AFA3FFC8CF9F7" box="[722,800,1519,1545]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
BL) are convex on dorsal and ventral margins, each has two rows of long spines along all its midventral margin; the midlateral surface has spicules or setae, similar in size from middle ventral to dorsal edges. In 
<taxonomicName id="4C2766FC107BED43C00FF9E7FD7CF9AF" authorityName="Hagen" authorityYear="1859" box="[599,720,1591,1617]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107BED43C00FF9E7FD7CF9AF" box="[599,720,1591,1617]" italics="true" pageId="12" pageNumber="313">R. fasciata</emphasis>
</taxonomicName>
the parameres in lateral view (
<figureCitation id="131C01FA107BED43C675F9E7FB23F9AF" box="[1069,1167,1591,1617]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="12" pageNumber="313">Fig. 16d</figureCitation>
; 
<bibRefCitation id="EFB6608E107BED43C6C3F9E7FA34F9AF" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[1179,1432,1591,1617]" pageId="12" pageNumber="313" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D107BED43C74EF9E7FAE3F9AF" box="[1302,1359,1591,1617]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
, 2021
</bibRefCitation>
, fig 12BL) are convex basally, each has two rows of long, thick spines on the posterior midventral margin, each row with more than three spines pointing laterad; the midlateral surface has sparse fine setae from the middle of the anteroventral edge to the posterodorsal edge, covering most of the apex, but absent in the middle anteroventral and posterodorsal edges.
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF4107BED40C2CFF8E2FAC6FE77" lastPageId="15" lastPageNumber="316" pageId="12" pageNumber="313" type="description">
<paragraph id="8B981D7F107BED43C2CFF8E2FEB8F8B2" blockId="12.[151,276,1842,1868]" box="[151,276,1842,1868]" pageId="12" pageNumber="313">
<heading id="D0D0AA13107BED43C2CFF8E2FEB8F8B2" bold="true" box="[151,276,1842,1868]" fontSize="11" level="1" pageId="12" pageNumber="313" reason="1">
<emphasis id="B953C16D107BED43C2CFF8E2FEB8F8B2" bold="true" box="[151,276,1842,1868]" pageId="12" pageNumber="313">FEMALE</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F107BED43C2CFF8ABFE9BF823" blockId="12.[151,1436,1914,2013]" pageId="12" pageNumber="313">
<emphasis id="B953C16D107BED43C2CFF8ABFEBCF86A" box="[151,272,1915,1940]" italics="true" pageId="12" pageNumber="313">Biometrics</emphasis>
. Body length (distance from front of head to distal edge of segment VIII): 
<quantity id="4CDFB09A107BED43C604F8AAFB5EF86B" box="[1116,1266,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="9.55" metricValueMax="10.6" metricValueMin="8.5" pageId="12" pageNumber="313" unit="mm" value="9.55" valueMax="10.6" valueMin="8.5">8.5–10.6 mm</quantity>
(x̄ = 9.6, 
<collectingRegion id="49E3D39D107BED43C739F8ABFA29F86B" box="[1377,1413,1915,1941]" country="Serbia" name="Sid" pageId="12" pageNumber="313">SD</collectingRegion>
= 0.83, n = 6), each forewing 
<quantity id="4CDFB09A107BED43C397F84EFDC9F847" box="[463,613,1950,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.115" metricValueMax="1.28" metricValueMin="0.95" pageId="12" pageNumber="313" unit="mm" value="11.15" valueMax="12.8" valueMin="9.5">9.5–12.8 mm</quantity>
(x̄ = 11.6, 
<collectingRegion id="49E3D39D107BED43C0B9F84FFCA9F847" box="[737,773,1951,1977]" country="Serbia" name="Sid" pageId="12" pageNumber="313">SD</collectingRegion>
= 1.13, n = 6), each hind wing 8.7–11.0 mm (x̄ = 10.2, 
<collectingRegion id="49E3D39D107BED43C720F84FFA30F847" box="[1400,1436,1951,1977]" country="Serbia" name="Sid" pageId="12" pageNumber="313">SD</collectingRegion>
= 0.81, n = 6).
</paragraph>
<caption id="DF584DF7107AED42C2C0F8A4FEBCF806" ID-DOI="http://doi.org/10.5281/zenodo.14239587" ID-Zenodo-Dep="14239587" httpUri="https://zenodo.org/record/14239587/files/figure.png" pageId="13" pageNumber="314" startId="13.[152,270,1908,1932]" targetBox="[216,1350,196,1865]" targetPageId="13" targetType="figure">
<paragraph id="8B981D7F107AED42C2C0F8A4FEBCF806" blockId="13.[151,1437,1908,2041]" pageId="13" pageNumber="314">
<emphasis id="B953C16D107AED42C2C0F8A4FEF6F872" bold="true" box="[152,346,1908,1932]" pageId="13" pageNumber="314">FIGURES 13–14.</emphasis>
Larvae of 
<taxonomicName id="4C2766FC107AED42C390F8A4FC3CF872" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[456,912,1908,1933]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D107AED42C390F8A4FD02F872" box="[456,686,1908,1932]" italics="true" pageId="13" pageNumber="314">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D107AED42C1CFF8A5FC4BF873" bold="true" box="[919,999,1909,1933]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="A2607C16107AED42C1CFF8A5FC4BF873" box="[919,999,1909,1933]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C2766FC107AED42C640F8A4FADFF872" authority="Hagen 1859" authorityName="Hagen" authorityYear="1859" box="[1048,1395,1908,1933]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107AED42C640F8A4FB43F873" box="[1048,1263,1908,1933]" italics="true" pageId="13" pageNumber="314">Rhyacophila fasciata</emphasis>
<bibRefCitation id="EFB6608E107AED42C6ADF8A5FADFF872" author="Hagen, H." box="[1269,1395,1908,1933]" pageId="13" pageNumber="314" pagination="4 - 6" refId="ref13323" refString="Hagen, H. (1859) Die Phryganiden Pictet's. Entomologische Zeitung, 20 (4 - 6), 131 - 170. Available from: https: // www. biodiversitylibrary. org / item / 35933 # page / 141 / mode / 1 up (accessed 5 March 2024)" type="journal article" year="1859">Hagen 1859</bibRefCitation>
</taxonomicName>
. 13, 
<taxonomicName id="4C2766FC107AED42C2CFF849FEB8F84E" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[151,276,1944,1968]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D107AED42C2CFF849FEB8F84E" box="[151,276,1944,1968]" italics="true" pageId="13" pageNumber="314">R. anatolica</emphasis>
</taxonomicName>
: 13a, head, dorsal; 13b, prothorax, dorsal; 13c, abdominal tergite IX, dorsal; 13d, left anal proleg, left lateral. 14, 
<taxonomicName id="4C2766FC107AED42C2CFF86DFEA8F82B" authorityName="Hagen" authorityYear="1859" box="[151,260,1981,2005]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D107AED42C2CFF86DFEA8F82B" box="[151,260,1981,2005]" italics="true" pageId="13" pageNumber="314">R. fasciata</emphasis>
</taxonomicName>
: 14a, head, dorsal. 14b, prothorax, dorsal; 14c, abdominal tergite IX, dorsal; 14d, left anal proleg, left lateral. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF584DF71079ED41C2C0F908FE29F812" ID-DOI="http://doi.org/10.5281/zenodo.14239589" ID-Zenodo-Dep="14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="14" pageNumber="315" startId="14.[152,270,1752,1776]" targetBox="[284,1303,192,1717]" targetPageId="14" targetType="figure">
<paragraph id="8B981D7F1079ED41C2C0F908FE29F812" blockId="14.[151,1437,1752,2028]" pageId="14" pageNumber="315">
<emphasis id="B953C16D1079ED41C2C0F908FEF2F90E" bold="true" box="[152,350,1752,1776]" pageId="14" pageNumber="315">FIGURES 15–18.</emphasis>
Males and females of 
<taxonomicName id="4C2766FC1079ED41C00DF908FB80F90E" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[597,1068,1752,1776]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D1079ED41C00DF908FC93F90E" box="[597,831,1752,1776]" italics="true" pageId="14" pageNumber="315">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D1079ED41C66EF908FB26F90E" bold="true" box="[1078,1162,1752,1776]" pageId="14" pageNumber="315">
<taxonomicNameLabel id="A2607C161079ED41C66EF908FB26F90E" box="[1078,1162,1752,1776]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C2766FC1079ED41C69AF908FEB9F8EA" authority="Hagen 1859" authorityName="Hagen" authorityYear="1859" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1079ED41C69AF908FA30F90E" box="[1218,1436,1752,1776]" italics="true" pageId="14" pageNumber="315">Rhyacophila fasciata</emphasis>
<bibRefCitation id="EFB6608E1079ED41C2CFF92DFEB9F8EA" author="Hagen, H." box="[151,277,1788,1813]" pageId="14" pageNumber="315" pagination="4 - 6" refId="ref13323" refString="Hagen, H. (1859) Die Phryganiden Pictet's. Entomologische Zeitung, 20 (4 - 6), 131 - 170. Available from: https: // www. biodiversitylibrary. org / item / 35933 # page / 141 / mode / 1 up (accessed 5 March 2024)" type="journal article" year="1859">Hagen 1859</bibRefCitation>
</taxonomicName>
. 15, male of 
<taxonomicName id="4C2766FC1079ED41C3CCF92DFDA3F8EA" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[404,527,1788,1812]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1079ED41C3CCF92DFDA3F8EA" box="[404,527,1788,1812]" italics="true" pageId="14" pageNumber="315">R. anatolica</emphasis>
</taxonomicName>
: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of 
<taxonomicName id="4C2766FC1079ED41C301F895FE68F8A2" authorityName="Hagen" authorityYear="1859" box="[345,452,1860,1884]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1079ED41C301F895FE68F8A2" box="[345,452,1860,1884]" italics="true" pageId="14" pageNumber="315">R. fasciata</emphasis>
</taxonomicName>
: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of 
<taxonomicName id="4C2766FC1079ED41C3B1F85DFDCAF85A" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[489,614,1932,1956]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1079ED41C3B1F85DFDCAF85A" box="[489,614,1932,1956]" italics="true" pageId="14" pageNumber="315">R. anatolica</emphasis>
</taxonomicName>
: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of 
<taxonomicName id="4C2766FC1079ED41C7DEF85DFF4BF836" authorityName="Hagen" authorityYear="1859" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1079ED41C7DEF85DFF4BF836" italics="true" pageId="14" pageNumber="315">R. fasciata</emphasis>
</taxonomicName>
: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm.
</paragraph>
</caption>
<paragraph id="8B981D7F1078ED40C29FFF47FAC6FE77" blockId="15.[151,1437,151,393]" pageId="15" pageNumber="316">
<emphasis id="B953C16D1078ED40C29FFF47FE9DFF4F" box="[199,305,151,177]" italics="true" pageId="15" pageNumber="316">Genitalia</emphasis>
(
<figureCitation id="131C01FA1078ED40C31BFF47FE32FF4F" box="[323,414,151,177]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C3F6FF47FE61FF4F" box="[430,461,151,177]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17</figureCitation>
). In lateral view (
<figureCitation id="131C01FA1078ED40C0F8FF47FCA0FF4F" box="[672,780,151,177]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12L</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C144FF47FCE0FF4F" box="[796,844,151,177]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17L</figureCitation>
), posterior margin of segment VIII valves on each side triangular, with dorsolateral projection directed dorsomesad, posterior dorsolateral and posteroventral margins concave, posterolateral projection round. In dorsal view (
<figureCitation id="131C01FA1078ED40C147FF0FFC20FF07" box="[799,908,223,249]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12D</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C1C1FF0FFC60FF07" box="[921,972,223,249]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17D</figureCitation>
), with indentation between segment VIII valves nearly enclosed by pair of dorsolateral suboval projections directed dorsomesad, slightly dilated at apices, with rounded apical and subapical corners, delimiting bean-shape space. In ventral view (
<figureCitation id="131C01FA1078ED40C6D1FEF7FB5EFEBF" box="[1161,1266,295,321]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12V</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C6A6FEF7FA83FEBF" box="[1278,1327,295,321]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17V</figureCitation>
) segment VIII valves forming two elongate sclerites, concave mesally, and diverging in apical halves, separated anteriorly; intersegmental membrane distally with pair of dark, ovoid or slightly elongate sclerites, converging anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41078ED40C2CFFE0AFC2FFC37" pageId="15" pageNumber="316" type="diagnosis">
<paragraph id="8B981D7F1078ED40C2CFFE0AFE39FE0A" blockId="15.[151,405,474,500]" box="[151,405,474,500]" pageId="15" pageNumber="316">
<emphasis id="B953C16D1078ED40C2CFFE0AFE39FE0A" bold="true" box="[151,405,474,500]" pageId="15" pageNumber="316">Diagnosis of females</emphasis>
</paragraph>
<paragraph id="8B981D7F1078ED40C2CFFDF3FBE8FCEB" blockId="15.[151,1437,547,1221]" pageId="15" pageNumber="316">
<emphasis id="B953C16D1078ED40C2CFFDF3FEADFDC3" box="[151,257,547,573]" italics="true" pageId="15" pageNumber="316">Genitalia</emphasis>
. In 
<taxonomicName id="4C2766FC1078ED40C369FDF3FE17FDC3" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[305,443,547,573]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1078ED40C369FDF3FE17FDC3" box="[305,443,547,573]" italics="true" pageId="15" pageNumber="316">R. anatolica</emphasis>
</taxonomicName>
lateral view (
<figureCitation id="131C01FA1078ED40C001FDF3FD6FFDC3" box="[601,707,547,573]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12L</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C089FDF3FD53FDC3" box="[721,767,547,573]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17L</figureCitation>
) each of the segment VIII lateral valves is triangular, with an apicodorsal projection; the oblique posterodorsal and posteroventral margins are concave, meeting at a round apex. In 
<taxonomicName id="4C2766FC1078ED40C2A1FDBBFEDEFD7B" authorityName="Hagen" authorityYear="1859" box="[249,370,619,645]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C2A1FDBBFEDEFD7B" box="[249,370,619,645]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
lateral view (
<figureCitation id="131C01FA1078ED40C054FDBBFDDDFD7B" box="[524,625,619,645]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">Fig. 18L</figureCitation>
; 
<bibRefCitation id="EFB6608E1078ED40C025FDBBFCDEFD7B" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[637,882,619,645]" pageId="15" pageNumber="316" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1078ED40C0AFFDBBFC9CFD7B" box="[759,816,619,645]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2021
</bibRefCitation>
, fig. 13L) each of the segment VIII lateral valves is quadrate with an apicodorsal projection; the upper posterior margin is more nearly vertical, and with an irregular edge having one or two posterior sinuosities (
<bibRefCitation id="EFB6608E1078ED40C0CDFD63FC20FD33" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[661,908,691,717]" pageId="15" pageNumber="316" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1078ED40C14AFD63FCE6FD33" box="[786,842,691,717]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2021
</bibRefCitation>
, figs 13La, 13Lb), the posteroventral margin is slightly concave sub-basally, and straight or convex distally, in some specimens slightly indented posteriorly, the upper posterior margin and the posteroventral margin meet at a round, obtuse angle.
</paragraph>
<paragraph id="8B981D7F1078ED40C29FFCCFFC2FFC37" blockId="15.[151,1437,547,1221]" pageId="15" pageNumber="316">
In 
<taxonomicName id="4C2766FC1078ED40C2BCFCCFFEC0FCC7" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[228,364,799,825]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1078ED40C2BCFCCFFEC0FCC7" box="[228,364,799,825]" italics="true" pageId="15" pageNumber="316">R. anatolica</emphasis>
</taxonomicName>
dorsal view (
<figureCitation id="131C01FA1078ED40C058FCCFFDC6FCC7" box="[512,618,799,825]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12D</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C02CFCCFFD0BFCC7" box="[628,679,799,825]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17D</figureCitation>
), the indentation between the segment VIII valves is nearly enclosed by pair of suboval apicodorsal projections, slightly dilated at their apices, delimiting a bean-shape membranous space. In 
<taxonomicName id="4C2766FC1078ED40C35AFCB7FED6FC7F" authorityName="Hagen" authorityYear="1859" box="[258,378,871,897]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C35AFCB7FED6FC7F" box="[258,378,871,897]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
dorsal view (
<figureCitation id="131C01FA1078ED40C049FCB7FDD5FC7F" box="[529,633,871,897]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">Fig. 18D</figureCitation>
; 
<bibRefCitation id="EFB6608E1078ED40C0DCFCB7FCD4FC7F" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[644,888,871,897]" pageId="15" pageNumber="316" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1078ED40C0A6FCB7FC9AFC7F" box="[766,822,871,897]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2021
</bibRefCitation>
, fig. 13Da), the indentation between the segment VIII valves is nearly enclosed by a pair of thick and tapered apicodorsal projections delimiting an oval membranous space; a small projection occurs anteromesally in some specimens.
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41078ED40C29FFC03FE33FB3B" pageId="15" pageNumber="316" type="description">
<paragraph id="8B981D7F1078ED40C29FFC03FE33FB3B" blockId="15.[151,1437,547,1221]" pageId="15" pageNumber="316">
In 
<taxonomicName id="4C2766FC1078ED40C2BFFC03FEDDFC13" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[231,369,979,1005]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1078ED40C2BFFC03FEDDFC13" box="[231,369,979,1005]" italics="true" pageId="15" pageNumber="316">R. anatolica</emphasis>
</taxonomicName>
ventral view (
<figureCitation id="131C01FA1078ED40C04EFC03FDD3FC13" box="[534,639,979,1005]" captionStart="FIGURES 11–12" captionStartId="11.[152,270,1787,1811]" captionTargetBox="[260,1320,192,1752]" captionTargetId="figure-19@11.[257,1331,181,1762]" captionTargetPageId="11" captionText="FIGURES 11–12. Male and female genitalia of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov., respectively. 11, male: 11A, 2nd segment of left inferior appendage, left lateral; 11BV, 11BL, parameres: 11BV, parameres (p) and ventral lobe of aedeagus (phallicata) (vl), ventral; 11BL, left paramere, left lateral. 11CL, 11CD, aedeagus (phallicata) and lateroventral lobes: 11CL, aedeagus and its left lateroventral lobe, left lateral; 11CD, aedeagus and its lateroventral lobes (lvl), dorsal. 10DD, apicodorsal lobe of segment IX (al) and preanal appendages (pa), dorsal; 11DV, segment X, ventral, ab = apical band, as = anal sclerites, va = non-sclerotized ventral area. 12, female segments VIII–XI: 12L, left lateral; 12D, dorsal; 12V, ventral. Scale bars: unlabelled = 1 mm; ● = 0.5 mm; * = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14239585" httpUri="https://zenodo.org/record/14239585/files/figure.png" pageId="15" pageNumber="316">Figs 12V</figureCitation>
, 
<figureCitation id="131C01FA1078ED40C0D3FC03FD13FC13" box="[651,703,979,1005]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">17V</figureCitation>
), the segment VIII valves form two elongate sclerites, each with concave and convex edges in posterior and anterior areas respectively, widely separated basally; the intersegmental membrane has a pair of ovoid elongate sclerites, converging anteriorly. In 
<taxonomicName id="4C2766FC1078ED40C65CFBCBFB2CFBCB" authorityName="Hagen" authorityYear="1859" box="[1028,1152,1051,1077]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C65CFBCBFB2CFBCB" box="[1028,1152,1051,1077]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
ventral view (
<figureCitation id="131C01FA1078ED40C776FBCBFA3BFBCB" box="[1326,1431,1051,1077]" captionStart="FIGURES 15–18" captionStartId="14.[152,270,1752,1776]" captionTargetBox="[284,1303,192,1717]" captionTargetId="figure-26@14.[275,1312,181,1727]" captionTargetPageId="14" captionText="FIGURES 15–18. Males and females of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. and Rhyacophila fasciata Hagen 1859. 15, male of R. anatolica: 15a, 2nd segment of left inferior appendage, left lateral; 15b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 15c, parameres and ventral lobe of aedeagus (phallicata), ventral; 15d, left paramere, left lateral. 16, male of R. fasciata: 16a, 2nd segment of left inferior appendage, left lateral; 16b, apicodorsal lobe of segment IX and preanal appendages, dorsal; 16c, parameres and ventral lobe of aedeagus (phallicata), ventral; 16d, left paramere, left lateral. 17, female segments VIII–XI of R. anatolica: 17L, left lateral; 17D, dorsal; 17V, ventral. 18, female segments VIII–XI of R. fasciata: 18L, left lateral; 18D, dorsal; 18V, ventral. Scale bars: Figures 15a–c, 16a–c = 0.5 mm, Figures 15d, 16d = 200 μm, Figures 17, 18 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14239589" httpUri="https://zenodo.org/record/14239589/files/figure.png" pageId="15" pageNumber="316">Fig. 18V</figureCitation>
; 
<bibRefCitation id="EFB6608E1078ED40C2CFFBEFFE23FBA7" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[151,399,1087,1113]" pageId="15" pageNumber="316" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">
Valladolid 
<emphasis id="B953C16D1078ED40C34BFBEFFEE0FBA7" box="[275,332,1087,1113]" italics="true" pageId="15" pageNumber="316">et al.</emphasis>
2021
</bibRefCitation>
, fig. 13V), the segment VIII valves form two elongate sclerites that are narrowly separated basally, the mesal margins are parallel in their basomesal 1/4, and straight and divergent ~ 47° in their apicomesal 3/4, their lateral margins convex; their intersegmental membrane has a pair of ovoid elongate sclerites separated, that are almost parallel.
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41078ED5CC2CFFAC6FCC1FF07" lastPageId="19" lastPageNumber="320" pageId="15" pageNumber="316" type="discussion">
<paragraph id="8B981D7F1078ED40C2CFFAC6FE5FFACE" blockId="15.[151,499,1302,1328]" box="[151,499,1302,1328]" pageId="15" pageNumber="316">
<heading id="D0D0AA131078ED40C2CFFAC6FE5FFACE" bold="true" box="[151,499,1302,1328]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
<emphasis id="B953C16D1078ED40C2CFFAC6FE5FFACE" bold="true" box="[151,499,1302,1328]" pageId="15" pageNumber="316">Phylogenetic considerations</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F1078ED40C2CFFA8FFE9EF91F" blockId="15.[151,1437,1375,1905]" pageId="15" pageNumber="316">
The combined BI + ML tree (
<figureCitation id="131C01FA1078ED40C380FA8FFD84FA87" box="[472,552,1375,1401]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19</figureCitation>
) shows clear differences between 
<taxonomicName id="4C2766FC1078ED40C1F8FA8FFBBAFA87" authorityName="Hagen" authorityYear="1859" box="[928,1046,1375,1401]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C1F8FA8FFBBAFA87" box="[928,1046,1375,1401]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
and the other species included, that previously were considered as 
<taxonomicName id="4C2766FC1078ED40C3B5FA53FDC8FA63" authorityName="Hagen" authorityYear="1859" box="[493,612,1411,1437]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C3B5FA53FDC8FA63" box="[493,612,1411,1437]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
or subspecies of 
<taxonomicName id="4C2766FC1078ED40C17AFA53FC35FA63" authorityName="Hagen" authorityYear="1859" box="[802,921,1411,1437]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C17AFA53FC35FA63" box="[802,921,1411,1437]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
. ML (bootstrap support, BS) and BI (posterior probabilities, PP) phylogenetic analyses recovered similar topologies, supporting two major clades: monophyletic 
<taxonomicName id="4C2766FC1078ED40C2CFFA1BFE8DFA1B" authorityName="Malicky &amp; Sipahiler" authorityYear="1993" box="[151,289,1483,1509]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="kykladica">
<emphasis id="B953C16D1078ED40C2CFFA1BFE8DFA1B" box="[151,289,1483,1509]" italics="true" pageId="15" pageNumber="316">R. kykladica</emphasis>
</taxonomicName>
(BS = 86, PP = 1) (
<figureCitation id="131C01FA1078ED40C3A1FA1BFDF6FA1B" box="[505,602,1483,1509]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19k</figureCitation>
), and the rest of species of the “ 
<taxonomicName id="4C2766FC1078ED40C1E2FA1BFB9EFA1B" authorityName="Hagen" authorityYear="1859" box="[954,1074,1483,1509]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C1E2FA1BFB9EFA1B" box="[954,1074,1483,1509]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
Species Complex” (BS = 69, PP = 0.97). Inside this last clade, we found six well supported groups, and three paraphyletic groups: (1) paraphyletic 
<taxonomicName id="4C2766FC1078ED40C2CFF9C3FEE5F9D2" authorityName="McLachlan" authorityYear="1865" box="[151,329,1555,1580]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="septentrionis">
<emphasis id="B953C16D1078ED40C2CFF9C3FEE5F9D2" box="[151,329,1555,1580]" italics="true" pageId="15" pageNumber="316">R. septentrionis</emphasis>
</taxonomicName>
(BS = 69, PP = 0.97) (
<figureCitation id="131C01FA1078ED40C00AF9C3FD18F9D3" box="[594,692,1555,1581]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19b</figureCitation>
), (2) paraphyletic 
<taxonomicName id="4C2766FC1078ED40C1DEF9C3FC50F9D3" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[902,1020,1555,1581]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1078ED40C1DEF9C3FC50F9D3" box="[902,1020,1555,1581]" italics="true" pageId="15" pageNumber="316">R. viteceki</emphasis>
</taxonomicName>
(BS = 51, PP = 0.82) (
<figureCitation id="131C01FA1078ED40C75DF9C3FAC9F9D3" box="[1285,1381,1555,1581]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19c</figureCitation>
), (3) monophyletic 
<taxonomicName id="4C2766FC1078ED40C36FF9E7FE02F9AF" box="[311,430,1591,1617]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B953C16D1078ED40C36FF9E7FE02F9AF" box="[311,430,1591,1617]" italics="true" pageId="15" pageNumber="316">R. loeffleri</emphasis>
</taxonomicName>
(BS = 75, PP = 0.98) (
<figureCitation id="131C01FA1078ED40C0FEF9E7FCABF9AF" box="[678,775,1591,1617]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19d</figureCitation>
), (4) paraphyletic 
<taxonomicName id="4C2766FC1078ED40C189F9E7FBDAF9AF" authorityName="Karaouzas, Valladolid &amp; Ibrahimi" authorityYear="2022" box="[977,1142,1591,1617]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="macedonica">
<emphasis id="B953C16D1078ED40C189F9E7FBDAF9AF" box="[977,1142,1591,1617]" italics="true" pageId="15" pageNumber="316">R. macedonica</emphasis>
</taxonomicName>
(BS = 76, PP = 0.98) (
<figureCitation id="131C01FA1078ED40C737F9E7FF6FF98B" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19e</figureCitation>
), and (5) monophyletic 
<taxonomicName id="4C2766FC1078ED40C388F98BFDE4F98B" authorityName="Hagen" authorityYear="1859" box="[464,584,1627,1653]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1078ED40C388F98BFDE4F98B" box="[464,584,1627,1653]" italics="true" pageId="15" pageNumber="316">R. fasciata</emphasis>
</taxonomicName>
(BS = 73, PP = 0.97) (
<figureCitation id="131C01FA1078ED40C111F98BFC08F98B" box="[841,932,1627,1653]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19f</figureCitation>
) as sister species. (6) Monophyletic 
<taxonomicName id="4C2766FC1078ED40C767F98BFA30F98B" authorityName="Kucinic &amp; Valladolid" authorityYear="2020" box="[1343,1436,1627,1653]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="delici">
<emphasis id="B953C16D1078ED40C767F98BFA30F98B" box="[1343,1436,1627,1653]" italics="true" pageId="15" pageNumber="316">R. delici</emphasis>
</taxonomicName>
(BS = 85, PP = 0.99) (
<figureCitation id="131C01FA1078ED40C3D6F9AFFE43F967" box="[398,495,1663,1689]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">
Fig. 
<quantity id="4CDFB09A1078ED40C39AF9AFFE43F967" box="[450,495,1663,1689]" metricMagnitude="-2" metricUnit="kg" metricValue="1.9" pageId="15" pageNumber="316" unit="g" value="19.0">19g</quantity>
</figureCitation>
), (7) monophyletic 
<taxonomicName id="4C2766FC1078ED40C093F9AFFCC4F967" authorityName="McLachlan" authorityYear="1879" box="[715,872,1663,1689]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B953C16D1078ED40C093F9AFFCC4F967" box="[715,872,1663,1689]" italics="true" pageId="15" pageNumber="316">R. denticulata</emphasis>
</taxonomicName>
(BS = 85, PP = 0.99) (
<figureCitation id="131C01FA1078ED40C63EF9AFFB6BF967" box="[1126,1223,1663,1689]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19h</figureCitation>
), (8) monophyletic 
<taxonomicName id="4C2766FC1078ED40C2CFF973FEA5F942" authorityName="Navas" authorityYear="1916" box="[151,265,1699,1724]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="sociata">
<emphasis id="B953C16D1078ED40C2CFF973FEA5F942" box="[151,265,1699,1724]" italics="true" pageId="15" pageNumber="316">R. sociata</emphasis>
</taxonomicName>
(BS = 90, PP = 0.97) (
<figureCitation id="131C01FA1078ED40C04DF973FDDCF943" box="[533,624,1699,1725]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19i</figureCitation>
), and (9) monophyletic 
<taxonomicName id="4C2766FC1078ED40C1DCF973FBA3F943" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[900,1039,1699,1725]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1078ED40C1DCF973FBA3F943" box="[900,1039,1699,1725]" italics="true" pageId="15" pageNumber="316">R. anatolica</emphasis>
</taxonomicName>
(BS = 99, PP =0.99) (
<figureCitation id="131C01FA1078ED40C74AF973FAC2F943" box="[1298,1390,1699,1725]" captionStart="FIGURE 19" captionStartId="16.[152,255,1685,1709]" captionTargetBox="[276,1302,190,1661]" captionTargetId="figure-26@16.[276,1312,181,1661]" captionTargetPageId="16" captionText="FIGURE 19. Phylogenetic relationships among the species of the “Rhyacophila fasciata Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, Rhyacophila cf. obliterata, outgroup; 19b–19k, R. fasciata Complex: 19b, Rhyacophila septentrionis McLachlan 1865; 19c, R. viteceki Valladolid &amp; Kučinić 2020 (by Valladolid et al. 2020); 19d, R. loeffleri Valladolid &amp; Waringer (by Valladolid et al. 2023); 19e, R. macedonica Karaouzas, Valladolid &amp; Ibrahimi 2022 (by Valladolid et al. 2022); 19f, R. fasciata Hagen 1859; 19g, R. delici Kučinić &amp; Valladolid 2020 (by Valladolid et al. 2020); 19h, R. denticulata McLachlan 1879; 19i, R. sociata Navás 1916; 19j, R. anatolica sp. nov.; 19k, R. kykladica Malicky &amp; Sipahiler 1993. Data for specimens are summarized in Table 1 (R. anatolica) and by Valladolid et al 2024. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch." figureDoi="http://doi.org/10.5281/zenodo.14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="15" pageNumber="316">Fig. 19j</figureCitation>
), as sister species.
</paragraph>
<paragraph id="8B981D7F1078ED40C29FF93BFC61F88F" blockId="15.[151,1437,1375,1905]" pageId="15" pageNumber="316">
Intraspecific distances (maximum composite likelihood model) are lower than 1% (
<tableCitation id="C6A528C41078ED40C601F93BFB17F8FB" box="[1113,1211,1771,1797]" captionStart="TABLE 2" captionStartId="17.[152,244,151,175]" captionTargetPageId="17" captionText="TABLE 2. Genetic distances inand among “Rhyacophila fasciata Species Complex” taxa (maximum composite likelihood model). 2a: intraspecific distances, n/c = not computed (1 specimen only). 2b: interspecific distances. Table 2a" httpUri="http://table.plazi.org/id/DF584DF71066ED5EC2C0FF47FEACFF05" pageId="15" pageNumber="316" tableUuid="DF584DF71066ED5EC2C0FF47FEACFF05">Table 2a</tableCitation>
), being the lowest in 
<emphasis id="B953C16D1078ED40C2CFF8DFFE64F8D7" box="[151,456,1807,1833]" italics="true" pageId="15" pageNumber="316">
<taxonomicName id="4C2766FC1078ED40C2CFF8DFFEE5F8D6" authorityName="McLachlan" authorityYear="1865" box="[151,329,1807,1832]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="septentrionis">R. septentrionis</taxonomicName>
, 
<taxonomicName id="4C2766FC1078ED40C30CF8DFFE64F8D7" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[340,456,1807,1833]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="viteceki">R. viteceki</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C2766FC1078ED40C05CF8DFFD22F8D7" authorityName="Malicky &amp; Sipahiler" authorityYear="1993" box="[516,654,1807,1833]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="kykladica">
<emphasis id="B953C16D1078ED40C05CF8DFFD22F8D7" box="[516,654,1807,1833]" italics="true" pageId="15" pageNumber="316">R. kykladica</emphasis>
</taxonomicName>
(
<geoCoordinate id="EE137BB81078ED40C0C4F8DFFD41F8D7" box="[668,749,1807,1833]" degrees="0.0020" direction="north" orientation="latitude" pageId="15" pageNumber="316" precision="5" value="0.002">0.0020</geoCoordinate>
, 
<geoCoordinate id="EE137BB81078ED40C0A1F8DFFCE5F8D7" box="[761,841,1807,1833]" degrees="0.0026" direction="east" orientation="longitude" pageId="15" pageNumber="316" precision="5" value="0.0026">0.0026</geoCoordinate>
, and 0.0030, respectively). Interspecific distances are higher than 1% (
<tableCitation id="C6A528C41078ED40C309F8E3FE19F8B3" box="[337,437,1843,1869]" captionStart="TABLE 2" captionStartId="17.[152,244,151,175]" captionTargetPageId="17" captionText="TABLE 2. Genetic distances inand among “Rhyacophila fasciata Species Complex” taxa (maximum composite likelihood model). 2a: intraspecific distances, n/c = not computed (1 specimen only). 2b: interspecific distances. Table 2a" httpUri="http://table.plazi.org/id/DF584DF71066ED5EC2C0FF47FEACFF05" pageId="15" pageNumber="316" tableUuid="DF584DF71066ED5EC2C0FF47FEACFF05">Table 2b</tableCitation>
) among all the species of the Complex. The lower distances are between 
<taxonomicName id="4C2766FC1078ED40C6B4F8E3FA31F8B2" authorityName="McLachlan" authorityYear="1865" box="[1260,1437,1843,1868]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="septentrionis">
<emphasis id="B953C16D1078ED40C6B4F8E3FA31F8B2" box="[1260,1437,1843,1868]" italics="true" pageId="15" pageNumber="316">R. septentrionis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2766FC1078ED40C29FF887FE90F88F" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[199,316,1879,1905]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1078ED40C29FF887FE90F88F" box="[199,316,1879,1905]" italics="true" pageId="15" pageNumber="316">R. viteceki</emphasis>
</taxonomicName>
(0.0100) and between 
<taxonomicName id="4C2766FC1078ED40C065F887FD19F88F" box="[573,693,1879,1905]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B953C16D1078ED40C065F887FD19F88F" box="[573,693,1879,1905]" italics="true" pageId="15" pageNumber="316">R. loeffleri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2766FC1078ED40C0B3F887FCCCF88F" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[747,864,1879,1905]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1078ED40C0B3F887FCCCF88F" box="[747,864,1879,1905]" italics="true" pageId="15" pageNumber="316">R. viteceki</emphasis>
</taxonomicName>
(0.0163).
</paragraph>
<caption id="DF584DF71067ED5FC2C0F945FB7BF80F" ID-DOI="http://doi.org/10.5281/zenodo.14239591" ID-Zenodo-Dep="14239591" httpUri="https://zenodo.org/record/14239591/files/figure.png" pageId="16" pageNumber="317" startId="16.[152,255,1685,1709]" targetBox="[276,1302,190,1661]" targetPageId="16" targetType="figure">
<paragraph id="8B981D7F1067ED5FC2C0F945FB7BF80F" blockId="16.[151,1437,1685,2033]" pageId="16" pageNumber="317">
<emphasis id="B953C16D1067ED5FC2C0F945FE8BF953" bold="true" box="[152,295,1685,1709]" pageId="16" pageNumber="317">FIGURE 19.</emphasis>
Phylogenetic relationships among the species of the “ 
<taxonomicName id="4C2766FC1067ED5FC100F945FB9CF953" authorityName="Hagen" authorityYear="1859" box="[856,1072,1685,1709]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1067ED5FC100F945FB9CF953" box="[856,1072,1685,1709]" italics="true" pageId="16" pageNumber="317">Rhyacophila fasciata</emphasis>
</taxonomicName>
Species Complex” included in this study. The ML tree represents a combination of Bayesian inference and maximum likelihood trees based on COI. Support for each node is represented by the posterior probabilities (PP) resulting from the Bayesian inference analysis and the bootstrap support values (BS) obtained for the maximum likelihood tree (PP/BS, respectively). 19a, 
<taxonomicName id="4C2766FC1067ED5FC67DF8D1FA82F8E7" authorityName="McLachlan" authorityYear="1863" box="[1061,1326,1793,1817]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" isUncertain="true" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="obliterata">
<emphasis id="B953C16D1067ED5FC67DF8D1FA82F8E7" box="[1061,1326,1793,1817]" italics="true" pageId="16" pageNumber="317">Rhyacophila cf. obliterata</emphasis>
</taxonomicName>
, outgroup; 19b–19k, 
<taxonomicName id="4C2766FC1067ED5FC2A6F8F6FEC7F8C3" authorityName="Hagen" authorityYear="1859" box="[254,363,1829,1853]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1067ED5FC2A6F8F6FEC7F8C3" box="[254,363,1829,1853]" italics="true" pageId="16" pageNumber="317">R. fasciata</emphasis>
</taxonomicName>
Complex: 19b, 
<taxonomicName id="4C2766FC1067ED5FC04BF8F5FC7AF8C3" authority="McLachlan 1865" authorityName="McLachlan" authorityYear="1865" box="[531,982,1829,1853]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="septentrionis">
<emphasis id="B953C16D1067ED5FC04BF8F5FCB2F8C3" box="[531,798,1829,1853]" italics="true" pageId="16" pageNumber="317">Rhyacophila septentrionis</emphasis>
<bibRefCitation id="EFB6608E1067ED5FC17DF8F5FC7AF8C3" author="McLachlan, R." box="[805,982,1829,1853]" pageId="16" pageNumber="317" pagination="1 - 183" refId="ref14141" refString="McLachlan, R. (1865) Trichoptera Britannica: A monograph of the British species of caddis-flies. Transactions of the Entomological Society of London, Series 3, 5, 1 - 183. https: // doi. org / 10.1111 / j. 1365 - 2311.1967. tb 01433. x" type="journal article" year="1865">McLachlan 1865</bibRefCitation>
</taxonomicName>
; 19c, 
<taxonomicName id="4C2766FC1067ED5FC64CF8F6FA31F8C3" authority="Valladolid &amp; Kucinic 2020" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[1044,1437,1829,1853]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1067ED5FC64CF8F6FBD3F8C3" box="[1044,1151,1829,1853]" italics="true" pageId="16" pageNumber="317">R. viteceki</emphasis>
<bibRefCitation id="EFB6608E1067ED5FC6DEF8F5FA31F8C3" author="Valladolid, M. &amp; Kucinic, M. &amp; Arauzo, M. &amp; Cerjanec, D. &amp; Cuk, R. &amp; Dorda, B. A. &amp; Lodovici, O. &amp; Stanic Kostroman, S. &amp; Vuckovic, I. &amp; Rey, I." box="[1158,1437,1829,1853]" pageId="16" pageNumber="317" refId="ref16645" refString="Valladolid, M., Kucinic, M., Arauzo, M., Cerjanec, D., Cuk, R., Dorda, B. A., Lodovici, O., Stanic Kostroman, S., Vuckovic, I. &amp; Rey, I. (2020) The Rhyacophila fasciata Group in Croatia and Bosnia and Herzegovina: Rhyacophila f. fasciata Hagen, 1859 and the description of two new subspecies, Rhyacophila fasciata delici Kucinic &amp; Valladolid (ssp. nov.) from Croatia and Bosnia and Herzegovina and Rhyacophila fasciata viteceki Valladolid &amp; Kucinic (ssp. nov.) from Bosnia and" type="journal volume" year="2020">Valladolid &amp; Kučinić 2020</bibRefCitation>
</taxonomicName>
(by 
<bibRefCitation id="EFB6608E1067ED5FC2E7F899FE32F89F" author="Valladolid, M. &amp; Kucinic, M. &amp; Arauzo, M. &amp; Cerjanec, D. &amp; Cuk, R. &amp; Dorda, B. A. &amp; Lodovici, O. &amp; Stanic Kostroman, S. &amp; Vuckovic, I. &amp; Rey, I." box="[191,414,1865,1889]" pageId="16" pageNumber="317" refId="ref16645" refString="Valladolid, M., Kucinic, M., Arauzo, M., Cerjanec, D., Cuk, R., Dorda, B. A., Lodovici, O., Stanic Kostroman, S., Vuckovic, I. &amp; Rey, I. (2020) The Rhyacophila fasciata Group in Croatia and Bosnia and Herzegovina: Rhyacophila f. fasciata Hagen, 1859 and the description of two new subspecies, Rhyacophila fasciata delici Kucinic &amp; Valladolid (ssp. nov.) from Croatia and Bosnia and Herzegovina and Rhyacophila fasciata viteceki Valladolid &amp; Kucinic (ssp. nov.) from Bosnia and" type="journal volume" year="2020">
Valladolid 
<emphasis id="B953C16D1067ED5FC376F89AFECDF89F" box="[302,353,1865,1889]" italics="true" pageId="16" pageNumber="317">et al.</emphasis>
2020
</bibRefCitation>
); 19d, 
<taxonomicName id="4C2766FC1067ED5FC3BBF89AFC91F89F" authority="Valladolid &amp; Waringer" authorityName="Valladolid &amp; Waringer" authorityYear="2023" box="[483,829,1865,1889]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B953C16D1067ED5FC3BBF89AFDFCF89F" box="[483,592,1865,1889]" italics="true" pageId="16" pageNumber="317">R. loeffleri</emphasis>
Valladolid &amp; Waringer
</taxonomicName>
(by 
<bibRefCitation id="EFB6608E1067ED5FC133F899FBE6F89F" author="Valladolid, M. &amp; Waringer, J. &amp; Arauzo, M. &amp; Chvojka, P. &amp; Dorda, B. A. &amp; Komzak, P. &amp; Lodovici, O. &amp; Rey, I." box="[875,1098,1865,1889]" pageId="16" pageNumber="317" pagination="451 - 482" refId="ref17056" refString="Valladolid, M., Waringer, J., Arauzo, M., Chvojka, P., Dorda, B. A., Komzak, P., Lodovici, O., Rey, I. (2023) The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Central Europe with description of a new species, Rhyacophila loeffleri Valladolid &amp; Waringer, n. sp., based on morphological, genetic and ecological evidence. Zootaxa, 5325 (4), 451 - 482. https: // doi. org / 10.11646 / zootaxa. 5325.4.1" type="journal article" year="2023">
Valladolid 
<emphasis id="B953C16D1067ED5FC182F89AFBABF89F" box="[986,1031,1865,1889]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
. 2023
</bibRefCitation>
); 19e, 
<taxonomicName id="4C2766FC1067ED5FC6D6F89AFE1DF87B" authority="Karaouzas, Valladolid &amp; Ibrahimi 2022" authorityName="Karaouzas, Valladolid &amp; Ibrahimi" authorityYear="2022" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="macedonica">
<emphasis id="B953C16D1067ED5FC6D6F89AFA88F89F" box="[1166,1316,1865,1889]" italics="true" pageId="16" pageNumber="317">R. macedonica</emphasis>
Karaouzas, 
<bibRefCitation id="EFB6608E1067ED5FC2CFF8BDFE1DF87B" author="Valladolid, M. &amp; Karaouzas, I. &amp; Ibrahimi, H. &amp; Arauzo, M. &amp; Slavevska Stamenkovic, V. &amp; Dorda, B. A. &amp; Hinic, J. &amp; Bilalli, A. &amp; Musliu, M. &amp; Rey, I." box="[151,433,1901,1925]" pageId="16" pageNumber="317" pagination="101 - 130" refId="ref16941" refString="Valladolid, M., Karaouzas, I., Ibrahimi, H., Arauzo, M., Slavevska Stamenkovic, V., Dorda, B. A., Hinic, J., Bilalli, A., Musliu, M. &amp; Rey, I. (2022) The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas, Valladolid &amp; Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae). Zootaxa, 5125 (2), 101 - 130. https: // doi. org / 10.11646 / zootaxa. 5125.2.1" type="journal article" year="2022">Valladolid &amp; Ibrahimi 2022</bibRefCitation>
</taxonomicName>
(by 
<bibRefCitation id="EFB6608E1067ED5FC387F8BDFD13F87B" author="Valladolid, M. &amp; Karaouzas, I. &amp; Ibrahimi, H. &amp; Arauzo, M. &amp; Slavevska Stamenkovic, V. &amp; Dorda, B. A. &amp; Hinic, J. &amp; Bilalli, A. &amp; Musliu, M. &amp; Rey, I." box="[479,703,1901,1925]" pageId="16" pageNumber="317" pagination="101 - 130" refId="ref16941" refString="Valladolid, M., Karaouzas, I., Ibrahimi, H., Arauzo, M., Slavevska Stamenkovic, V., Dorda, B. A., Hinic, J., Bilalli, A., Musliu, M. &amp; Rey, I. (2022) The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas, Valladolid &amp; Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae). Zootaxa, 5125 (2), 101 - 130. https: // doi. org / 10.11646 / zootaxa. 5125.2.1" type="journal article" year="2022">
Valladolid 
<emphasis id="B953C16D1067ED5FC016F8BEFD2EF87B" box="[590,642,1901,1925]" italics="true" pageId="16" pageNumber="317">et al.</emphasis>
2022
</bibRefCitation>
); 19f, 
<taxonomicName id="4C2766FC1067ED5FC0A7F8BEFC5DF87B" authority="Hagen 1859" authorityName="Hagen" authorityYear="1859" box="[767,1009,1901,1925]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1067ED5FC0A7F8BEFCC0F87B" box="[767,876,1901,1925]" italics="true" pageId="16" pageNumber="317">R. fasciata</emphasis>
<bibRefCitation id="EFB6608E1067ED5FC12BF8BDFC5DF87B" author="Hagen, H." box="[883,1009,1901,1925]" pageId="16" pageNumber="317" pagination="4 - 6" refId="ref13323" refString="Hagen, H. (1859) Die Phryganiden Pictet's. Entomologische Zeitung, 20 (4 - 6), 131 - 170. Available from: https: // www. biodiversitylibrary. org / item / 35933 # page / 141 / mode / 1 up (accessed 5 March 2024)" type="journal article" year="1859">Hagen 1859</bibRefCitation>
</taxonomicName>
; 19g, 
<taxonomicName id="4C2766FC1067ED5FC677F8BEFA31F87B" authority="Kucinic &amp; Valladolid 2020" authorityName="Kucinic &amp; Valladolid" authorityYear="2020" box="[1071,1437,1901,1925]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="delici">
<emphasis id="B953C16D1067ED5FC677F8BEFB2FF87B" box="[1071,1155,1901,1925]" italics="true" pageId="16" pageNumber="317">R. delici</emphasis>
Kučinić &amp; 
<bibRefCitation id="EFB6608E1067ED5FC6A3F8BDFA31F87B" author="Valladolid, M. &amp; Kucinic, M. &amp; Arauzo, M. &amp; Cerjanec, D. &amp; Cuk, R. &amp; Dorda, B. A. &amp; Lodovici, O. &amp; Stanic Kostroman, S. &amp; Vuckovic, I. &amp; Rey, I." box="[1275,1437,1901,1925]" pageId="16" pageNumber="317" refId="ref16645" refString="Valladolid, M., Kucinic, M., Arauzo, M., Cerjanec, D., Cuk, R., Dorda, B. A., Lodovici, O., Stanic Kostroman, S., Vuckovic, I. &amp; Rey, I. (2020) The Rhyacophila fasciata Group in Croatia and Bosnia and Herzegovina: Rhyacophila f. fasciata Hagen, 1859 and the description of two new subspecies, Rhyacophila fasciata delici Kucinic &amp; Valladolid (ssp. nov.) from Croatia and Bosnia and Herzegovina and Rhyacophila fasciata viteceki Valladolid &amp; Kucinic (ssp. nov.) from Bosnia and" type="journal volume" year="2020">Valladolid 2020</bibRefCitation>
</taxonomicName>
(by 
<bibRefCitation id="EFB6608E1067ED5FC2E6F841FE30F857" author="Valladolid, M. &amp; Kucinic, M. &amp; Arauzo, M. &amp; Cerjanec, D. &amp; Cuk, R. &amp; Dorda, B. A. &amp; Lodovici, O. &amp; Stanic Kostroman, S. &amp; Vuckovic, I. &amp; Rey, I." box="[190,412,1937,1961]" pageId="16" pageNumber="317" refId="ref16645" refString="Valladolid, M., Kucinic, M., Arauzo, M., Cerjanec, D., Cuk, R., Dorda, B. A., Lodovici, O., Stanic Kostroman, S., Vuckovic, I. &amp; Rey, I. (2020) The Rhyacophila fasciata Group in Croatia and Bosnia and Herzegovina: Rhyacophila f. fasciata Hagen, 1859 and the description of two new subspecies, Rhyacophila fasciata delici Kucinic &amp; Valladolid (ssp. nov.) from Croatia and Bosnia and Herzegovina and Rhyacophila fasciata viteceki Valladolid &amp; Kucinic (ssp. nov.) from Bosnia and" type="journal volume" year="2020">
Valladolid 
<emphasis id="B953C16D1067ED5FC375F842FEF3F857" box="[301,351,1937,1961]" italics="true" pageId="16" pageNumber="317">et al.</emphasis>
2020
</bibRefCitation>
); 19h, 
<taxonomicName id="4C2766FC1067ED5FC387F842FC8EF857" authority="McLachlan 1879" authorityName="McLachlan" authorityYear="1879" box="[479,802,1937,1961]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B953C16D1067ED5FC387F842FDC1F857" box="[479,621,1937,1961]" italics="true" pageId="16" pageNumber="317">R. denticulata</emphasis>
<bibRefCitation id="EFB6608E1067ED5FC02BF841FC8EF857" author="McLachlan, R." box="[627,802,1937,1961]" pageId="16" pageNumber="317" pagination="429 - 500" refId="ref14247" refString="McLachlan, R. (1879) A Monographic Revision and Synopsis of the Trichoptera of the European Fauna. Part 8. John Van Voorst, London, pp. 429 - 500, pls. 45 - 51. [1874 - 1880] https: // doi. org / 10.5962 / bhl. title. 28556" type="book chapter" year="1879">McLachlan 1879</bibRefCitation>
</taxonomicName>
; 19i, 
<taxonomicName id="4C2766FC1067ED5FC101F842FB93F857" authority="Navas 1916" authorityName="Navas" authorityYear="1916" box="[857,1087,1937,1961]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="sociata">
<emphasis id="B953C16D1067ED5FC101F842FC11F857" box="[857,957,1938,1961]" italics="true" pageId="16" pageNumber="317">R. sociata</emphasis>
<bibRefCitation id="EFB6608E1067ED5FC19BF841FB93F857" author="Navas, L." box="[963,1087,1937,1961]" pageId="16" pageNumber="317" pagination="5 - 11" refId="ref14509" refString="Navas, L. (1916) Tricopteros nuevos de Espana. 1 ª Serie. Broteria, Serie Zoologica, 14, 5 - 11." type="journal article" year="1916">Navás 1916</bibRefCitation>
</taxonomicName>
; 19j, 
<taxonomicName id="4C2766FC1067ED5FC62DF842FB5DF857" authorityName="Valladolid &amp; Ekingen &amp; Arauzo &amp; Dorda &amp; Rey" authorityYear="2024" box="[1141,1265,1937,1961]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D1067ED5FC62DF842FB5DF857" box="[1141,1265,1937,1961]" italics="true" pageId="16" pageNumber="317">R. anatolica</emphasis>
</taxonomicName>
<emphasis id="B953C16D1067ED5FC6AFF841FAEBF857" bold="true" box="[1271,1351,1937,1961]" pageId="16" pageNumber="317">
<taxonomicNameLabel id="A2607C161067ED5FC6AFF841FAEBF857" box="[1271,1351,1937,1961]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 19k, 
<taxonomicName id="4C2766FC1067ED5FC7DEF842FDA5F833" authority="Malicky &amp; Sipahiler 1993" authorityName="Malicky &amp; Sipahiler" authorityYear="1993" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="kykladica">
<emphasis id="B953C16D1067ED5FC7DEF842FF54F833" italics="true" pageId="16" pageNumber="317">R. kykladica</emphasis>
<bibRefCitation id="EFB6608E1067ED5FC2A6F865FDA5F833" author="Malicky, H. &amp; Sipahiler, F." box="[254,521,1973,1997]" pageId="16" pageNumber="317" pagination="457 - 468" refId="ref14009" refString="Malicky, H. &amp; Sipahiler, F. (1993) Kocherfliegen (Trichoptera) aus der Turkei, mit Bemerkungen zu weiteren mediterranean Kocherfliegen. Bulletin de la societe entomologique Suisse, 66, 457 - 468." type="journal article" year="1993">Malicky &amp; Sipahiler 1993</bibRefCitation>
</taxonomicName>
. Data for specimens are summarized in Table 1 (
<taxonomicName id="4C2766FC1067ED5FC1A9F866FBC1F833" authorityName="Valladolid &amp; Ekingen &amp; Arauzo &amp; Dorda &amp; Rey" authorityYear="2024" box="[1009,1133,1973,1997]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1067ED5FC1A9F866FBC1F833" box="[1009,1133,1973,1997]" italics="true" pageId="16" pageNumber="317">R. anatolica</emphasis>
</taxonomicName>
) and by 
<bibRefCitation id="EFB6608E1067ED5FC69CF865FA35F833" author="Valladolid, M. &amp; Arauzo, M. &amp; Dorda, B. A. &amp; Pais, M. &amp; Rey, I." box="[1220,1433,1973,1997]" pageId="16" pageNumber="317" refId="ref17163" refString="Valladolid, M., Arauzo, M., Dorda, B. A., Pais, M. &amp; Rey, I. (2024) Complete data of the DNA sequences used for the study of the Rhyacophila fasciata Complex (Insecta, Trichoptera, Rhyacophilidae) in Europe. Dataset, Digital CSIC. Available from: https: // digital. csic. es / handle / 10261 / 260864 (accessed 5 March 2024)" type="url" year="2024">
Valladolid 
<emphasis id="B953C16D1067ED5FC76AF866FAF2F833" box="[1330,1374,1973,1997]" italics="true" pageId="16" pageNumber="317">et al</emphasis>
2024
</bibRefCitation>
. Scale bar: mean number of nucleotide substitutions per site or nucleotide position on the respective branch.
</paragraph>
</caption>
<caption id="DF584DF71066ED5EC2C0FF47FEACFF05" ID-Table-UUID="DF584DF71066ED5EC2C0FF47FEACFF05" httpUri="http://table.plazi.org/id/DF584DF71066ED5EC2C0FF47FEACFF05" pageId="17" pageNumber="318" startId="17.[152,244,151,175]" targetBox="[163,875,266,734]" targetIsTable="true" targetPageId="17" targetType="table">
<paragraph id="8B981D7F1066ED5EC2C0FF47FB7DFF2D" blockId="17.[151,1436,151,251]" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC2C0FF47FEA0FF51" bold="true" box="[152,268,151,175]" pageId="17" pageNumber="318">TABLE 2.</emphasis>
Genetic distances in and among “ 
<taxonomicName id="4C2766FC1066ED5EC032FF47FCE5FF4E" authorityName="Hagen" authorityYear="1859" box="[618,841,151,176]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1066ED5EC032FF47FCE5FF4E" box="[618,841,151,176]" italics="true" pageId="17" pageNumber="318">Rhyacophila fasciata</emphasis>
</taxonomicName>
Species Complex” taxa (maximum composite likelihood model). 2a: intraspecific distances, n/c = not computed (1 specimen only). 2b: interspecific distances.
</paragraph>
<paragraph id="8B981D7F1066ED5EC2FBFF33FEACFF05" blockId="17.[151,1436,151,251]" box="[163,256,227,251]" pageId="17" pageNumber="318">Table 2a</paragraph>
</caption>
<paragraph id="8B981D7F1066ED5EC2FBFEDAFCE8FD20" pageId="17" pageNumber="318">
<table id="F927EFDF106612B0C2FBFEDAFCC7FD20" box="[163,875,266,734]" gridcols="2" gridrows="12" pageId="17" pageNumber="318">
<tr id="35171F3D106612B0C2FBFEDAFCC7FEDC" box="[163,875,266,290]" gridrow="0" pageId="17" pageNumber="318">
<th id="76C67641106612B0C2FBFEDAFCC7FEDC" box="[163,875,266,290]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC2FBFEDAFE87FEDC" bold="true" box="[163,299,266,290]" pageId="17" pageNumber="318">Intraspecific</emphasis>
</th>
</tr>
<tr id="35171F3D106612B0C2FBFEE3FCC7FEB5" box="[163,875,307,331]" gridrow="1" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFEE3FEEEFEB5" box="[163,322,307,331]" gridcol="0" gridrow="1" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFEE4FEB3FEB5" authorityName="Valladolid &amp; Ekingen &amp; Arauzo &amp; Dorda &amp; Rey" authorityYear="2024" box="[163,287,307,331]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1066ED5EC2FBFEE4FEB3FEB5" box="[163,287,307,331]" italics="true" pageId="17" pageNumber="318">R. anatolica</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFEE3FCC7FEB5" box="[805,875,307,331]" gridcol="1" gridrow="1" pageId="17" pageNumber="318">0.0039</td>
</tr>
<tr id="35171F3D106612B0C2FBFE8BFCC7FE8D" box="[163,875,347,371]" gridrow="2" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFE8BFEEEFE8D" box="[163,322,347,371]" gridcol="0" gridrow="2" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFE8CFF5BFE8D" authorityName="Kucinic &amp; Valladolid" authorityYear="2020" box="[163,247,347,371]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="delici">
<emphasis id="B953C16D1066ED5EC2FBFE8CFF5BFE8D" box="[163,247,347,371]" italics="true" pageId="17" pageNumber="318">R. delici</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFE8BFCC7FE8D" box="[805,875,347,371]" gridcol="1" gridrow="2" pageId="17" pageNumber="318">0.0086</td>
</tr>
<tr id="35171F3D106612B0C2FBFE53FCC7FE65" box="[163,875,387,411]" gridrow="3" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFE53FEEEFE65" box="[163,322,387,411]" gridcol="0" gridrow="3" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFE54FE9DFE65" authorityName="McLachlan" authorityYear="1879" box="[163,305,387,411]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B953C16D1066ED5EC2FBFE54FE9DFE65" box="[163,305,387,411]" italics="true" pageId="17" pageNumber="318">R. denticulata</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFE53FCC7FE65" box="[805,875,387,411]" gridcol="1" gridrow="3" pageId="17" pageNumber="318">0.0086</td>
</tr>
<tr id="35171F3D106612B0C2FBFE7CFCC7FE3A" box="[163,875,428,452]" gridrow="4" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFE7CFEEEFE3A" box="[163,322,428,452]" gridcol="0" gridrow="4" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFE7CFEA3FE3A" authorityName="Hagen" authorityYear="1859" box="[163,271,428,452]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1066ED5EC2FBFE7CFEA3FE3A" box="[163,271,428,452]" italics="true" pageId="17" pageNumber="318">R. fasciata</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFE7CFCC7FE3A" box="[805,875,428,452]" gridcol="1" gridrow="4" pageId="17" pageNumber="318">0.0071</td>
</tr>
<tr id="35171F3D106612B0C2FBFE04FCC7FE12" box="[163,875,468,492]" gridrow="5" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFE04FEEEFE12" box="[163,322,468,492]" gridcol="0" gridrow="5" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFE05FE8CFE12" authorityName="Malicky &amp; Sipahiler" authorityYear="1993" box="[163,288,468,492]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="kykladica">
<emphasis id="B953C16D1066ED5EC2FBFE05FE8CFE12" box="[163,288,468,492]" italics="true" pageId="17" pageNumber="318">R. kykladica</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFE04FCC7FE12" box="[805,875,468,492]" gridcol="1" gridrow="5" pageId="17" pageNumber="318">0.0030</td>
</tr>
<tr id="35171F3D106612B0C2FBFE2CFCC7FDEA" box="[163,875,508,532]" gridrow="6" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFE2CFEEEFDEA" box="[163,322,508,532]" gridcol="0" gridrow="6" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFE2DFEA3FDEA" authorityName="Valladolid &amp; Waringer" authorityYear="2023" box="[163,271,508,532]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B953C16D1066ED5EC2FBFE2DFEA3FDEA" box="[163,271,508,532]" italics="true" pageId="17" pageNumber="318">R. loeffleri</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFE2CFCC7FDEA" box="[805,875,508,532]" gridcol="1" gridrow="6" pageId="17" pageNumber="318">0.0081</td>
</tr>
<tr id="35171F3D106612B0C2FBFDF4FCC7FDC2" box="[163,875,548,572]" gridrow="7" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFDF4FEEEFDC2" box="[163,322,548,572]" gridcol="0" gridrow="7" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFDF5FE95FDC2" authorityName="Karaouzas, Valladolid &amp; Ibrahimi" authorityYear="2022" box="[163,313,548,572]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="macedonica">
<emphasis id="B953C16D1066ED5EC2FBFDF5FE95FDC2" box="[163,313,548,572]" italics="true" pageId="17" pageNumber="318">R. macedonica</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFDF4FCC7FDC2" box="[805,875,548,572]" gridcol="1" gridrow="7" pageId="17" pageNumber="318">0.0062</td>
</tr>
<tr id="35171F3D106612B0C2FBFD9DFCC7FD9B" box="[163,875,589,613]" gridrow="8" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFD9DFEEEFD9B" box="[163,322,589,613]" gridcol="0" gridrow="8" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFD9DFEEEFD9A" authorityName="McLachlan" authorityYear="1865" box="[163,322,589,612]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="septentrionis">
<emphasis id="B953C16D1066ED5EC2FBFD9DFEEEFD9A" box="[163,322,589,612]" italics="true" pageId="17" pageNumber="318">R. septentrionis</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFD9DFCC7FD9B" box="[805,875,589,613]" gridcol="1" gridrow="8" pageId="17" pageNumber="318">0.0020</td>
</tr>
<tr id="35171F3D106612B0C2FBFDA5FCC7FD73" box="[163,875,629,653]" gridrow="9" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFDA5FEEEFD73" box="[163,322,629,653]" gridcol="0" gridrow="9" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFDA6FEA4FD73" authorityName="Navas" authorityYear="1916" box="[163,264,630,653]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sociata">
<emphasis id="B953C16D1066ED5EC2FBFDA6FEA4FD73" box="[163,264,630,653]" italics="true" pageId="17" pageNumber="318">R. sociata</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFDA5FCC7FD73" box="[805,875,629,653]" gridcol="1" gridrow="9" pageId="17" pageNumber="318">0.0062</td>
</tr>
<tr id="35171F3D106612B0C2FBFD4DFCC7FD4B" box="[163,875,669,693]" gridrow="10" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFD4DFEEEFD4B" box="[163,322,669,693]" gridcol="0" gridrow="10" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFD4EFEA0FD4B" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[163,268,669,693]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1066ED5EC2FBFD4EFEA0FD4B" box="[163,268,669,693]" italics="true" pageId="17" pageNumber="318">R. viteceki</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFD4DFCC7FD4B" box="[805,875,669,693]" gridcol="1" gridrow="10" pageId="17" pageNumber="318">0.0026</td>
</tr>
<tr id="35171F3D106612B0C2FBFD15FCC7FD20" box="[163,875,709,734]" gridrow="11" pageId="17" pageNumber="318">
<td id="76C67641106612B0C2FBFD15FEEEFD20" box="[163,322,709,734]" gridcol="0" gridrow="11" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFD16FEEEFD23" authorityName="McLachlan" authorityYear="1863" box="[163,322,709,734]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" isUncertain="true" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="obliterata">
<emphasis id="B953C16D1066ED5EC2FBFD16FEEEFD23" box="[163,322,709,734]" italics="true" pageId="17" pageNumber="318">R. cf. obliterata</emphasis>
</taxonomicName>
</td>
<td id="76C67641106612B0C17DFD15FCC7FD20" box="[805,875,709,734]" gridcol="1" gridrow="11" pageId="17" pageNumber="318">n/c</td>
</tr>
</table>
</paragraph>
<caption id="DF584DF71066ED5EC2FBFCC2FEAEFCD4" ID-Table-UUID="DF584DF71066ED5EC2FBFCC2FEAEFCD4" box="[163,258,786,810]" httpUri="http://table.plazi.org/id/DF584DF71066ED5EC2FBFCC2FEAEFCD4" pageId="17" pageNumber="318" startId="17.[163,225,786,810]" targetBox="[163,1406,826,1253]" targetIsTable="true" targetPageId="17">
<paragraph id="8B981D7F1066ED5EC2FBFCC2FEAEFCD4" blockId="17.[163,258,786,810]" box="[163,258,786,810]" pageId="17" pageNumber="318">Table 2b</paragraph>
</caption>
<paragraph id="8B981D7F1066ED5EC2FBFCEAFAD2FB1B" pageId="17" pageNumber="318">
<table id="F927EFDF106612B0C2FBFCEAFAD2FB1B" box="[163,1406,826,1253]" gridcols="11" gridrows="11" pageId="17" pageNumber="318">
<tr id="35171F3D106612B0C2FBFCEAFAD2FCAC" box="[163,1406,826,850]" gridrow="0" isHeader="true" pageId="17" pageNumber="318">
<th id="76C67641106612B0C2FBFCEAFEEEFCAC" box="[163,322,826,850]" gridcol="0" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC2FBFCEAFE86FCAC" bold="true" box="[163,298,826,850]" pageId="17" pageNumber="318">Interspecific</emphasis>
</th>
<th id="76C67641106612B0C330FCEAFE1FFCAC" box="[360,435,826,850]" gridcol="1" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FCEAFE1FFCAF" bold="true" box="[360,435,826,849]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC330FCEAFE1CFCAF" box="[360,432,826,849]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="ana">R. ana</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C382FCEAFD8CFCAC" box="[474,544,826,850]" gridcol="2" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC382FCEAFDB9FCAC" box="[474,533,826,850]" italics="true" pageId="17" pageNumber="318">R.del.</emphasis>
</th>
<th id="76C67641106612B0C01DFCEAFD38FCAC" box="[581,660,826,850]" gridcol="3" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC01DFCEBFD38FCAC" box="[581,660,826,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC01DFCEBFD3CFCAC" box="[581,656,826,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="dent">R. dent</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C0EEFCEAFD50FCAC" box="[694,764,826,850]" gridcol="4" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC0EEFCEBFD5BFCAC" box="[694,759,826,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC0EEFCEBFD5FFCAC" authorityName="Milne" authorityYear="1936" box="[694,755,826,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fas">R. fas</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C179FCEAFCCBFCAC" box="[801,871,826,850]" gridcol="5" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC179FCEBFCCAFCAC" box="[801,870,826,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC179FCEBFCCEFCAC" box="[801,866,826,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="kyk">R. kyk</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C1D4FCEAFC7EFCAC" box="[908,978,826,850]" gridcol="6" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC1D4FCEBFC62FCAC" box="[908,974,826,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC1D4FCEBFC66FCAC" box="[908,970,826,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="loe">R. loe</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C1AFFCEAFBE8FCAC" box="[1015,1092,826,850]" gridcol="7" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC1AFFCEBFBE8FCAC" box="[1015,1092,827,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC1AFFCEBFBECFCAC" box="[1015,1088,827,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="mac">R. mac</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C63AFCEAFB04FCAC" box="[1122,1192,826,850]" gridcol="8" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC63AFCEBFB0CFCAC" box="[1122,1184,827,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC63AFCEBFB31FCAC" box="[1122,1181,827,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sep">R sep</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C695FCEAFABFFCAC" box="[1229,1299,826,850]" gridcol="9" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC695FCEBFABEFCAC" box="[1229,1298,827,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC695FCEBFAA2FCAC" box="[1229,1294,827,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="soc">R. soc</taxonomicName>
.
</emphasis>
</th>
<th id="76C67641106612B0C760FCEAFAD2FCAC" box="[1336,1406,826,850]" gridcol="10" gridrow="0" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC760FCEBFAD9FCAC" box="[1336,1397,827,850]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC760FCEBFADCFCAC" authorityName="Milne" authorityYear="1936" box="[1336,1392,827,850]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="vit">R. vit</taxonomicName>
.
</emphasis>
</th>
</tr>
<tr id="35171F3D106612B0C2FBFCB2FAD2FC84" box="[163,1406,866,890]" gridrow="1" pageId="17" pageNumber="318" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFCB2FEEEFC84" box="[163,322,866,890]" gridcol="0" gridrow="1" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFCB3FF5BFC84" authorityName="Kucinic &amp; Valladolid" authorityYear="2020" box="[163,247,866,890]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="delici">
<emphasis id="B953C16D1066ED5EC2FBFCB3FF5BFC84" box="[163,247,866,890]" italics="true" pageId="17" pageNumber="318">R. delici</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FCB2FE1FFC84" box="[360,435,866,890]" gridcol="1" gridrow="1" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FCB2FE02FC84" bold="true" box="[360,430,866,890]" pageId="17" pageNumber="318">0.0352</emphasis>
</td>
</tr>
<tr id="35171F3D106612B0C2FBFC5AFAD2FC5D" box="[163,1406,906,931]" gridrow="2" pageId="17" pageNumber="318" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFC5AFEEEFC5D" box="[163,322,906,931]" gridcol="0" gridrow="2" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFC5CFE9DFC5D" authorityName="McLachlan" authorityYear="1879" box="[163,305,907,931]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B953C16D1066ED5EC2FBFC5CFE9DFC5D" box="[163,305,907,931]" italics="true" pageId="17" pageNumber="318">R. denticulata</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FC5AFE1FFC5D" box="[360,435,906,931]" gridcol="1" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FC5AFE02FC5C" bold="true" box="[360,430,906,930]" pageId="17" pageNumber="318">0.0605</emphasis>
</td>
<td id="76C67641106612B0C382FC5AFD8CFC5D" box="[474,544,906,931]" gridcol="2" gridrow="2" pageId="17" pageNumber="318">0.0592</td>
</tr>
<tr id="35171F3D106612B0C2FBFC63FAD2FC35" box="[163,1406,947,971]" gridrow="3" pageId="17" pageNumber="318" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFC63FEEEFC35" box="[163,322,947,971]" gridcol="0" gridrow="3" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFC64FEA3FC35" authorityName="Hagen" authorityYear="1859" box="[163,271,947,971]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1066ED5EC2FBFC64FEA3FC35" box="[163,271,947,971]" italics="true" pageId="17" pageNumber="318">R. fasciata</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FC63FE1FFC35" box="[360,435,947,971]" gridcol="1" gridrow="3" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FC63FE02FC35" bold="true" box="[360,430,947,971]" pageId="17" pageNumber="318">0.0419</emphasis>
</td>
<td id="76C67641106612B0C382FC63FD8CFC35" box="[474,544,947,971]" gridcol="2" gridrow="3" pageId="17" pageNumber="318">0.0432</td>
<td id="76C67641106612B0C01DFC63FD38FC35" box="[581,660,947,971]" gridcol="3" gridrow="3" pageId="17" pageNumber="318">0.0635</td>
</tr>
<tr id="35171F3D106612B0C2FBFC0BFAD2FC0D" box="[163,1406,987,1011]" gridrow="4" pageId="17" pageNumber="318" rowspan-10="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFC0BFEEEFC0D" box="[163,322,987,1011]" gridcol="0" gridrow="4" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFC0CFE8CFC0D" authorityName="Malicky &amp; Sipahiler" authorityYear="1993" box="[163,288,987,1011]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="kykladica">
<emphasis id="B953C16D1066ED5EC2FBFC0CFE8CFC0D" box="[163,288,987,1011]" italics="true" pageId="17" pageNumber="318">R. kykladica</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FC0BFE1FFC0D" box="[360,435,987,1011]" gridcol="1" gridrow="4" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FC0BFE02FC0D" bold="true" box="[360,430,987,1011]" pageId="17" pageNumber="318">0.0572</emphasis>
</td>
<td id="76C67641106612B0C382FC0BFD8CFC0D" box="[474,544,987,1011]" gridcol="2" gridrow="4" pageId="17" pageNumber="318">0.0547</td>
<td id="76C67641106612B0C01DFC0BFD38FC0D" box="[581,660,987,1011]" gridcol="3" gridrow="4" pageId="17" pageNumber="318">0.0671</td>
<td id="76C67641106612B0C0EEFC0BFD50FC0D" box="[694,764,987,1011]" gridcol="4" gridrow="4" pageId="17" pageNumber="318">0.0664</td>
</tr>
<tr id="35171F3D106612B0C2FBFBD3FAD2FBE2" box="[163,1406,1027,1052]" gridrow="5" pageId="17" pageNumber="318" rowspan-10="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFBD3FEEEFBE2" box="[163,322,1027,1052]" gridcol="0" gridrow="5" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFBD4FEA3FBE2" authorityName="Valladolid &amp; Waringer" authorityYear="2023" box="[163,271,1028,1052]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B953C16D1066ED5EC2FBFBD4FEA3FBE2" box="[163,271,1028,1052]" italics="true" pageId="17" pageNumber="318">R. loeffleri</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FBD3FE1FFBE2" box="[360,435,1027,1052]" gridcol="1" gridrow="5" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FBD3FE02FBE5" bold="true" box="[360,430,1027,1051]" pageId="17" pageNumber="318">0.0427</emphasis>
</td>
<td id="76C67641106612B0C382FBD3FD8CFBE2" box="[474,544,1027,1052]" gridcol="2" gridrow="5" pageId="17" pageNumber="318">0.0332</td>
<td id="76C67641106612B0C01DFBD3FD38FBE2" box="[581,660,1027,1052]" gridcol="3" gridrow="5" pageId="17" pageNumber="318">0.0595</td>
<td id="76C67641106612B0C0EEFBD3FD50FBE2" box="[694,764,1027,1052]" gridcol="4" gridrow="5" pageId="17" pageNumber="318">0.0317</td>
<td id="76C67641106612B0C179FBD3FCCBFBE2" box="[801,871,1027,1052]" gridcol="5" gridrow="5" pageId="17" pageNumber="318">0.0495</td>
</tr>
<tr id="35171F3D106612B0C2FBFBFBFAD2FBBA" box="[163,1406,1067,1092]" gridrow="6" pageId="17" pageNumber="318" rowspan-10="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFBFBFEEEFBBA" box="[163,322,1067,1092]" gridcol="0" gridrow="6" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFBFDFE95FBBA" authorityName="Karaouzas, Valladolid &amp; Ibrahimi" authorityYear="2022" box="[163,313,1068,1092]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="macedonica">
<emphasis id="B953C16D1066ED5EC2FBFBFDFE95FBBA" box="[163,313,1068,1092]" italics="true" pageId="17" pageNumber="318">R. macedonica</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FBFBFE1FFBBA" box="[360,435,1067,1092]" gridcol="1" gridrow="6" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FBFBFE02FBBD" bold="true" box="[360,430,1067,1091]" pageId="17" pageNumber="318">0.0336</emphasis>
</td>
<td id="76C67641106612B0C382FBFBFD8CFBBA" box="[474,544,1067,1092]" gridcol="2" gridrow="6" pageId="17" pageNumber="318">0.0318</td>
<td id="76C67641106612B0C01DFBFBFD38FBBA" box="[581,660,1067,1092]" gridcol="3" gridrow="6" pageId="17" pageNumber="318">0.0553</td>
<td id="76C67641106612B0C0EEFBFBFD50FBBA" box="[694,764,1067,1092]" gridcol="4" gridrow="6" pageId="17" pageNumber="318">0.0173</td>
<td id="76C67641106612B0C179FBFBFCCBFBBA" box="[801,871,1067,1092]" gridcol="5" gridrow="6" pageId="17" pageNumber="318">0.0545</td>
<td id="76C67641106612B0C1D4FBFBFC7EFBBA" box="[908,978,1067,1092]" gridcol="6" gridrow="6" pageId="17" pageNumber="318">0.0239</td>
</tr>
<tr id="35171F3D106612B0C2FBFB84FAD2FB92" box="[163,1406,1108,1132]" gridrow="7" pageId="17" pageNumber="318" rowspan-10="1" rowspan-8="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFB84FEEEFB92" box="[163,322,1108,1132]" gridcol="0" gridrow="7" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFB85FEEEFB92" authorityName="McLachlan" authorityYear="1865" box="[163,322,1109,1132]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="septentrionis">
<emphasis id="B953C16D1066ED5EC2FBFB85FEEEFB92" box="[163,322,1109,1132]" italics="true" pageId="17" pageNumber="318">R. septentrionis</emphasis>
</taxonomicName>
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<td id="76C67641106612B0C330FB84FE1FFB92" box="[360,435,1108,1132]" gridcol="1" gridrow="7" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FB84FE02FB92" bold="true" box="[360,430,1108,1132]" pageId="17" pageNumber="318">0.0314</emphasis>
</td>
<td id="76C67641106612B0C382FB84FD8CFB92" box="[474,544,1108,1132]" gridcol="2" gridrow="7" pageId="17" pageNumber="318">0.0261</td>
<td id="76C67641106612B0C01DFB84FD38FB92" box="[581,660,1108,1132]" gridcol="3" gridrow="7" pageId="17" pageNumber="318">0.0522</td>
<td id="76C67641106612B0C0EEFB84FD50FB92" box="[694,764,1108,1132]" gridcol="4" gridrow="7" pageId="17" pageNumber="318">0.0297</td>
<td id="76C67641106612B0C179FB84FCCBFB92" box="[801,871,1108,1132]" gridcol="5" gridrow="7" pageId="17" pageNumber="318">0.0433</td>
<td id="76C67641106612B0C1D4FB84FC7EFB92" box="[908,978,1108,1132]" gridcol="6" gridrow="7" pageId="17" pageNumber="318">0.0224</td>
<td id="76C67641106612B0C1AFFB84FBE8FB92" box="[1015,1092,1108,1132]" gridcol="7" gridrow="7" pageId="17" pageNumber="318">0.0205</td>
</tr>
<tr id="35171F3D106612B0C2FBFBACFAD2FB6A" box="[163,1406,1148,1172]" gridrow="8" pageId="17" pageNumber="318" rowspan-10="1" rowspan-9="1">
<th id="76C67641106612B0C2FBFBACFEEEFB6A" box="[163,322,1148,1172]" gridcol="0" gridrow="8" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFBADFEA4FB6A" authorityName="Navas" authorityYear="1916" box="[163,264,1149,1172]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sociata">
<emphasis id="B953C16D1066ED5EC2FBFBADFEA4FB6A" box="[163,264,1149,1172]" italics="true" pageId="17" pageNumber="318">R. sociata</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FBACFE1FFB6A" box="[360,435,1148,1172]" gridcol="1" gridrow="8" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FBACFE02FB6A" bold="true" box="[360,430,1148,1172]" pageId="17" pageNumber="318">0.0585</emphasis>
</td>
<td id="76C67641106612B0C382FBACFD8CFB6A" box="[474,544,1148,1172]" gridcol="2" gridrow="8" pageId="17" pageNumber="318">0.0615</td>
<td id="76C67641106612B0C01DFBACFD38FB6A" box="[581,660,1148,1172]" gridcol="3" gridrow="8" pageId="17" pageNumber="318">0.0432</td>
<td id="76C67641106612B0C0EEFBACFD50FB6A" box="[694,764,1148,1172]" gridcol="4" gridrow="8" pageId="17" pageNumber="318">0.0679</td>
<td id="76C67641106612B0C179FBACFCCBFB6A" box="[801,871,1148,1172]" gridcol="5" gridrow="8" pageId="17" pageNumber="318">0.0783</td>
<td id="76C67641106612B0C1D4FBACFC7EFB6A" box="[908,978,1148,1172]" gridcol="6" gridrow="8" pageId="17" pageNumber="318">0.0566</td>
<td id="76C67641106612B0C1AFFBACFBE8FB6A" box="[1015,1092,1148,1172]" gridcol="7" gridrow="8" pageId="17" pageNumber="318">0.0587</td>
<td id="76C67641106612B0C63AFBACFB04FB6A" box="[1122,1192,1148,1172]" gridcol="8" gridrow="8" pageId="17" pageNumber="318">0.0553</td>
</tr>
<tr id="35171F3D106612B0C2FBFB74FAD2FB43" box="[163,1406,1188,1213]" gridrow="9" pageId="17" pageNumber="318" rowspan-10="1">
<th id="76C67641106612B0C2FBFB74FEEEFB43" box="[163,322,1188,1213]" gridcol="0" gridrow="9" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFB75FEA0FB43" authorityName="Valladolid &amp; Kucinic" authorityYear="2020" box="[163,268,1189,1213]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1066ED5EC2FBFB75FEA0FB43" box="[163,268,1189,1213]" italics="true" pageId="17" pageNumber="318">R. viteceki</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FB74FE1FFB43" box="[360,435,1188,1213]" gridcol="1" gridrow="9" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FB74FE02FB42" bold="true" box="[360,430,1188,1212]" pageId="17" pageNumber="318">0.0293</emphasis>
</td>
<td id="76C67641106612B0C382FB74FD8CFB43" box="[474,544,1188,1213]" gridcol="2" gridrow="9" pageId="17" pageNumber="318">0.0196</td>
<td id="76C67641106612B0C01DFB74FD38FB43" box="[581,660,1188,1213]" gridcol="3" gridrow="9" pageId="17" pageNumber="318">0.0500</td>
<td id="76C67641106612B0C0EEFB74FD50FB43" box="[694,764,1188,1213]" gridcol="4" gridrow="9" pageId="17" pageNumber="318">0.0258</td>
<td id="76C67641106612B0C179FB74FCCBFB43" box="[801,871,1188,1213]" gridcol="5" gridrow="9" pageId="17" pageNumber="318">0.0508</td>
<td id="76C67641106612B0C1D4FB74FC7EFB43" box="[908,978,1188,1213]" gridcol="6" gridrow="9" pageId="17" pageNumber="318">0.0163</td>
<td id="76C67641106612B0C1AFFB74FBE8FB43" box="[1015,1092,1188,1213]" gridcol="7" gridrow="9" pageId="17" pageNumber="318">0.0178</td>
<td id="76C67641106612B0C63AFB74FB04FB43" box="[1122,1192,1188,1213]" gridcol="8" gridrow="9" pageId="17" pageNumber="318">0.0100</td>
<td id="76C67641106612B0C695FB74FABFFB43" box="[1229,1299,1188,1213]" gridcol="9" gridrow="9" pageId="17" pageNumber="318">0.0495</td>
</tr>
<tr id="35171F3D106612B0C2FBFB1DFAD2FB1B" box="[163,1406,1229,1253]" gridrow="10" pageId="17" pageNumber="318">
<th id="76C67641106612B0C2FBFB1DFEEEFB1B" box="[163,322,1229,1253]" gridcol="0" gridrow="10" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC2FBFB1EFEEEFB1B" authorityName="McLachlan" authorityYear="1863" box="[163,322,1229,1253]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" isUncertain="true" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="obliterata">
<emphasis id="B953C16D1066ED5EC2FBFB1EFEEEFB1B" box="[163,322,1229,1253]" italics="true" pageId="17" pageNumber="318">R. cf. obliterata</emphasis>
</taxonomicName>
</th>
<td id="76C67641106612B0C330FB1DFE1FFB1B" box="[360,435,1229,1253]" gridcol="1" gridrow="10" pageId="17" pageNumber="318">
<emphasis id="B953C16D1066ED5EC330FB1DFE02FB1B" bold="true" box="[360,430,1229,1253]" pageId="17" pageNumber="318">0.1043</emphasis>
</td>
<td id="76C67641106612B0C382FB1DFD8CFB1B" box="[474,544,1229,1253]" gridcol="2" gridrow="10" pageId="17" pageNumber="318">0.1025</td>
<td id="76C67641106612B0C01DFB1DFD38FB1B" box="[581,660,1229,1253]" gridcol="3" gridrow="10" pageId="17" pageNumber="318">0.0999</td>
<td id="76C67641106612B0C0EEFB1DFD50FB1B" box="[694,764,1229,1253]" gridcol="4" gridrow="10" pageId="17" pageNumber="318">0.0982</td>
<td id="76C67641106612B0C179FB1DFCCBFB1B" box="[801,871,1229,1253]" gridcol="5" gridrow="10" pageId="17" pageNumber="318">0.0937</td>
<td id="76C67641106612B0C1D4FB1DFC7EFB1B" box="[908,978,1229,1253]" gridcol="6" gridrow="10" pageId="17" pageNumber="318">0.0913</td>
<td id="76C67641106612B0C1AFFB1DFBE8FB1B" box="[1015,1092,1229,1253]" gridcol="7" gridrow="10" pageId="17" pageNumber="318">0.0943</td>
<td id="76C67641106612B0C63AFB1DFB04FB1B" box="[1122,1192,1229,1253]" gridcol="8" gridrow="10" pageId="17" pageNumber="318">0.0825</td>
<td id="76C67641106612B0C695FB1DFABFFB1B" box="[1229,1299,1229,1253]" gridcol="9" gridrow="10" pageId="17" pageNumber="318">0.1118</td>
<td id="76C67641106612B0C760FB1DFAD2FB1B" box="[1336,1406,1229,1253]" gridcol="10" gridrow="10" pageId="17" pageNumber="318">0.0848</td>
</tr>
</table>
</paragraph>
<paragraph id="8B981D7F1066ED5EC2CFFAEBFEB7FAAB" blockId="17.[151,283,1339,1365]" box="[151,283,1339,1365]" pageId="17" pageNumber="318">
<heading id="D0D0AA131066ED5EC2CFFAEBFEB7FAAB" bold="true" box="[151,283,1339,1365]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
<emphasis id="B953C16D1066ED5EC2CFFAEBFEB7FAAB" bold="true" box="[151,283,1339,1365]" pageId="17" pageNumber="318">Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8B981D7F1066ED5EC2CFFA53FC20F8FB" blockId="17.[151,1437,1411,2013]" pageId="17" pageNumber="318">
The use of the combination of selected characters in males (e.g., inferior appendages, dorsal lobes, parameres, and aedeagus), and females (shape of the sclerotized lateral valves of segment VIII in dorsal, ventral, and lateral views) has allowed morphological descriptions of new species of the “ 
<taxonomicName id="4C2766FC1066ED5EC120FA1CFC58FA1B" authorityName="Hagen" authorityYear="1859" box="[888,1012,1483,1509]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1066ED5EC120FA1CFC58FA1B" box="[888,1012,1483,1509]" italics="true" pageId="17" pageNumber="318">R. fasciata</emphasis>
</taxonomicName>
Species Complex” (
<bibRefCitation id="EFB6608E1066ED5EC6BAFA1BFF7FF9F7" author="Valladolid, M. &amp; Karaouzas, I. &amp; Arauzo, M. &amp; Dorda, B. A. &amp; Rey, I." pageId="17" pageNumber="318" pagination="503 - 522" refId="ref16560" refString="Valladolid, M., Karaouzas, I., Arauzo, M., Dorda, B. A. &amp; Rey, I. (2019) The Rhyacophila fasciata Group in Greece: Rhyacophila kykladica Malicky &amp; Sipahiler 1993 (stat. prom.) (Trichoptera: Rhyacophilidae). Morphological description, genetic and ecological features, Zootaxa, 4657 (3), 503 - 522. https: // doi. org / 10.11646 / zootaxa. 4657.3.5" type="journal article" year="2019">
Valladolid 
<emphasis id="B953C16D1066ED5EC739FA1CFA31FA1B" box="[1377,1437,1483,1509]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2019
</bibRefCitation>
), the resurrection of species considered synonyms of 
<taxonomicName id="4C2766FC1066ED5EC17CFA20FC30F9F7" authorityName="Hagen" authorityYear="1859" box="[804,924,1519,1545]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1066ED5EC17CFA20FC30F9F7" box="[804,924,1519,1545]" italics="true" pageId="17" pageNumber="318">R. fasciata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB6608E1066ED5EC1F1FA3FFB31F9F7" author="Valladolid, M. &amp; Arauzo, M. &amp; Basaguren, A. &amp; Dorda, B. A. &amp; Rey, I." box="[937,1181,1519,1545]" pageId="17" pageNumber="318" pagination="526 - 544" refId="ref16463" refString="Valladolid, M., Arauzo, M., Basaguren, A., Dorda, B. A. &amp; Rey, I. (2018) The Rhyacophila fasciata Group in Western Europe: Confirmation of Rhyacophila denticulata McLachlan 1879 (stat. prom.) and Rhyacophila sociata Navas 1916 (stat. res.), based on morphological and molecular genetic evidence (Trichoptera: Rhyacophilidae), Zootaxa, 4418 (6), 526 - 544. https: // doi. org / 10.11646 / zootaxa. 4418.6.2" type="journal article" year="2018">
Valladolid 
<emphasis id="B953C16D1066ED5EC67CFA20FBF0F9F7" box="[1060,1116,1519,1545]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2018
</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1066ED5EC6FFFA3FFB4FF9F7" author="Valladolid, M. &amp; Arauzo, M. &amp; Chertoprud, M. &amp; Chvojka, P. &amp; Czachorowski, S. &amp; Dorda, B. &amp; Hinic, J. &amp; Ibrahimi, H. &amp; Karaouzas, I. &amp; Krpac, V. &amp; Kucinic, M. &amp; Lodovici, O. &amp; Salokannel, J. &amp; Slavevska &amp; Stamenkovic, V. &amp; Stojanovic, K. &amp; Wallace, I. &amp; Rey, I." box="[1191,1251,1519,1545]" pageId="17" pageNumber="318" pagination="1 - 57" refId="ref16790" refString="Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. &amp; Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1" type="journal article" year="2021">2021</bibRefCitation>
), and changes of status from subspecies to species (
<bibRefCitation id="EFB6608E1066ED5EC04BF9C3FCABF9D3" author="Valladolid, M. &amp; Arauzo, M. &amp; Basaguren, A. &amp; Dorda, B. A. &amp; Rey, I." box="[531,775,1555,1581]" pageId="17" pageNumber="318" pagination="526 - 544" refId="ref16463" refString="Valladolid, M., Arauzo, M., Basaguren, A., Dorda, B. A. &amp; Rey, I. (2018) The Rhyacophila fasciata Group in Western Europe: Confirmation of Rhyacophila denticulata McLachlan 1879 (stat. prom.) and Rhyacophila sociata Navas 1916 (stat. res.), based on morphological and molecular genetic evidence (Trichoptera: Rhyacophilidae), Zootaxa, 4418 (6), 526 - 544. https: // doi. org / 10.11646 / zootaxa. 4418.6.2" type="journal article" year="2018">
Valladolid 
<emphasis id="B953C16D1066ED5EC0D6F9C4FD6AF9D3" box="[654,710,1555,1581]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2018
</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1066ED5EC149F9C3FCE0F9D3" author="Valladolid, M. &amp; Karaouzas, I. &amp; Arauzo, M. &amp; Dorda, B. A. &amp; Rey, I." box="[785,844,1555,1581]" pageId="17" pageNumber="318" pagination="503 - 522" refId="ref16560" refString="Valladolid, M., Karaouzas, I., Arauzo, M., Dorda, B. A. &amp; Rey, I. (2019) The Rhyacophila fasciata Group in Greece: Rhyacophila kykladica Malicky &amp; Sipahiler 1993 (stat. prom.) (Trichoptera: Rhyacophilidae). Morphological description, genetic and ecological features, Zootaxa, 4657 (3), 503 - 522. https: // doi. org / 10.11646 / zootaxa. 4657.3.5" type="journal article" year="2019">2019</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1066ED5EC10FF9C3FC38F9D3" author="Valladolid, M. &amp; Karaouzas, I. &amp; Ibrahimi, H. &amp; Arauzo, M. &amp; Slavevska Stamenkovic, V. &amp; Dorda, B. A. &amp; Hinic, J. &amp; Bilalli, A. &amp; Musliu, M. &amp; Rey, I." box="[855,916,1555,1581]" pageId="17" pageNumber="318" pagination="101 - 130" refId="ref16941" refString="Valladolid, M., Karaouzas, I., Ibrahimi, H., Arauzo, M., Slavevska Stamenkovic, V., Dorda, B. A., Hinic, J., Bilalli, A., Musliu, M. &amp; Rey, I. (2022) The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas, Valladolid &amp; Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae). Zootaxa, 5125 (2), 101 - 130. https: // doi. org / 10.11646 / zootaxa. 5125.2.1" type="journal article" year="2022">2022</bibRefCitation>
). In the successive studies that we have carried out, our hypothesis has been supported that the morphological differences, which until now have been considered simple intraspecific variations, are actually interspecific differences, since, in those species with a wide distribution, and that coincide in an area with another species of the 
<emphasis id="B953C16D1066ED5EC143F9AFFC08F967" box="[795,932,1663,1689]" italics="true" pageId="17" pageNumber="318">
“ 
<taxonomicName id="4C2766FC1066ED5EC17FF9AFFC08F967" authorityName="Hagen" authorityYear="1859" box="[807,932,1663,1689]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="fasciata">R. fasciata</taxonomicName>
</emphasis>
Species Complex”, the differences between them and the specimens with which they share territory are greater than the differences with other specimens of their species that are further away, sometimes hundreds or thousands of kilometers. In this study, these characters show the presence of a new species of this group, 
<emphasis id="B953C16D1066ED5EC0DEF93BFC20F8FB" box="[646,908,1771,1797]" italics="true" pageId="17" pageNumber="318">
<taxonomicName id="4C2766FC1066ED5EC0DEF93BFC2BF8FB" authorityName="Valladolid &amp; Ekingen &amp; Arauzo &amp; Dorda &amp; Rey" authorityYear="2024" box="[646,903,1771,1797]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="anatolica">Rhyacophila anatolica</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B981D7F1066ED5DC29FF8DFFB41FEE3" blockId="17.[151,1437,1411,2013]" lastBlockId="18.[151,1436,151,285]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
Mitochondrial DNA (mtDNA) sequences can be useful as a first indicator in species delineation, and, combined with careful morphological, behavioural, distributional, and ecological analysis, can help to establish accurate species boundaries (
<bibRefCitation id="EFB6608E1066ED5EC323F887FDCBF88F" author="Bickford, D. &amp; Lohman, D. J. &amp; Sodhi, N. S. &amp; Ng, P. K. &amp; Meier, R. &amp; Winker, K. &amp; Ingram, K. K. &amp; Das, I." box="[379,615,1879,1905]" pageId="17" pageNumber="318" pagination="148 - 155" refId="ref12381" refString="Bickford, D., Lohman, D. J., Sodhi, N. S., Ng, P. K., Meier, R., Winker, K., Ingram, K. K. &amp; Das, I. (2007) Cryptic species as a window on diversity and conservation. Trends in Ecology and Evolution, 22, 148 - 155. https: // doi. org / 10.1016 / j. tree. 2006.11.004" type="journal article" year="2007">
Bickford 
<emphasis id="B953C16D1066ED5EC3B2F888FD88F88F" box="[490,548,1879,1905]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2007
</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1066ED5EC02CF887FC65F88F" author="Dasmahapatra, K. &amp; Mallet, J." box="[628,969,1879,1905]" pageId="17" pageNumber="318" pagination="254 - 255" refId="ref12754" refString="Dasmahapatra, K. &amp; Mallet, J. (2006) DNA barcodes: Recent successes and future prospects. Heredity, 97, 254 - 255. https: // doi. org / 10.1038 / sj. hdy. 6800858" type="journal article" year="2006">Dasmahapatra &amp; Mallet 2006</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1066ED5EC18EF887FB07F88F" author="Galtier, N. &amp; Nabholz, B. &amp; Glemin, S. &amp; Hurst, G. D. D." box="[982,1195,1879,1905]" pageId="17" pageNumber="318" pagination="4541 - 4550" refId="ref13144" refString="Galtier, N., Nabholz, B., Glemin, S. &amp; Hurst, G. D. D. (2009) Mitochondrial DNA as a marker of molecular diversity: A reappraisal. Molecular Ecology, 18, 4541 - 4550. https: // doi. org / 10.1111 / j. 1365 - 294 x. 2009.04380. x" type="journal article" year="2009">
Galtier 
<emphasis id="B953C16D1066ED5EC674F888FBCAF88F" box="[1068,1126,1879,1905]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2009
</bibRefCitation>
). Most recent studies show the importance of mtDNA in the emergence of new species through mitochondrial-nuclear (mitonuclear) coevolution. Among the mtDNA, the nucleotide sequence of the COI gene has been established as a highly effective DNA barcode for diagnosing the species boundaries of animals (
<bibRefCitation id="EFB6608E1066ED5EC13DF813FB92F823" author="Bucklin, A. &amp; Steinke, D. &amp; Blanco-Bercial, L." box="[869,1086,1987,2013]" pageId="17" pageNumber="318" pagination="471 - 508" refId="ref12506" refString="Bucklin, A., Steinke, D. &amp; Blanco-Bercial, L. (2011) DNA barcoding of marine Metazoa. Annual Review of Marine Science, 3, 471 - 508. https: // doi. org / 10.1146 / annurev-marine- 120308 - 080950" type="journal article" year="2011">
Bucklin 
<emphasis id="B953C16D1066ED5EC19EF814FC52F823" box="[966,1022,1987,2013]" italics="true" pageId="17" pageNumber="318">et al.</emphasis>
2011
</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1066ED5EC612F813FA34F823" author="Dasmahapatra, K. &amp; Mallet, J." box="[1098,1432,1987,2013]" pageId="17" pageNumber="318" pagination="254 - 255" refId="ref12754" refString="Dasmahapatra, K. &amp; Mallet, J. (2006) DNA barcodes: Recent successes and future prospects. Heredity, 97, 254 - 255. https: // doi. org / 10.1038 / sj. hdy. 6800858" type="journal article" year="2006">Dasmahapatra &amp; Mallet 2006</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1065ED5DC2CFFF47FED2FF4F" author="Hebert, P. D. N. &amp; Ratnasingham, S. &amp; de Waard, J. R." box="[151,382,151,177]" pageId="18" pageNumber="319" refId="ref13448" refString="Hebert, P. D. N., Ratnasingham, S. &amp; de Waard, J. R. (2003 b) Barcoding animal life: Cytochrome c oxidase subunit 1 divergences among closely related species. Proceedings of the Royal Society of London, Series B, 270 (Supplement 1), S 96 - S 99. https: // doi. org / 10.1098 / rsbl. 2003.0025" type="journal volume" year="2003">
Hebert 
<emphasis id="B953C16D1065ED5DC2B6FF47FE8FFF4F" box="[238,291,151,177]" italics="true" pageId="18" pageNumber="319">et al</emphasis>
. 2003b
</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1065ED5DC3D5FF47FDADFF4F" author="Hill, G. E." box="[397,513,151,177]" pageId="18" pageNumber="319" pagination="5831 - 5842" refId="ref13644" refString="Hill, G. E. (2016) Mitonuclear coevolution as the genesis of speciation and the mitochondrial DNA barcode gap. Ecology and Evolution, 6 (16), 5831 - 5842. https: // doi. org / 10.1002 / ece 3.2338" type="journal article" year="2016">Hill 2016</bibRefCitation>
). In most metazoan populations, there is a correspondence between the species designated by COI and species previously described morphologically (
<bibRefCitation id="EFB6608E1065ED5DC1F5FF6BFB01FF2B" author="Tavares, E. S. &amp; Baker, A. J." box="[941,1197,187,213]" pageId="18" pageNumber="319" pagination="1 - 14" refId="ref16153" refString="Tavares, E. S. &amp; Baker, A. J. (2008) Single mitochondrial gene barcodes reliably identify sister-species in diverse clades of birds. BMC Evolutionary Biology, 8, Art. No. 81, 1 - 14. https: // doi. org / 10.1186 / 1471 - 2148 - 8 - 81" type="book chapter" year="2008">Tavares &amp; Baker 2008</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1065ED5DC6E0FF6BFA23FF2B" author="Tavares, E. S. &amp; Goncalves, P. &amp; Miyaki, C. Y. &amp; Baker, A. J." box="[1208,1423,187,213]" pageId="18" pageNumber="319" refId="ref16214" refString="Tavares, E. S., Goncalves, P., Miyaki, C. Y. &amp; Baker, A. J. (2011) DNA barcode detects high genetic structure within Neotropical bird species. PLoS ONE, 6, e 28543. https: // doi. org / 10.1371 / journal. pone. 0028543" type="journal volume" year="2011">
Tavares 
<emphasis id="B953C16D1065ED5DC74EFF6BFAE2FF2B" box="[1302,1358,187,213]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2011
</bibRefCitation>
); furthermore, discovery of barcode gaps often leads to the morphological/behavioural/distributional/ecological recognition of new species upon further study (
<bibRefCitation id="EFB6608E1065ED5DC0FFFED3FC21FEE3" author="Bickford, D. &amp; Lohman, D. J. &amp; Sodhi, N. S. &amp; Ng, P. K. &amp; Meier, R. &amp; Winker, K. &amp; Ingram, K. K. &amp; Das, I." box="[679,909,259,285]" pageId="18" pageNumber="319" pagination="148 - 155" refId="ref12381" refString="Bickford, D., Lohman, D. J., Sodhi, N. S., Ng, P. K., Meier, R., Winker, K., Ingram, K. K. &amp; Das, I. (2007) Cryptic species as a window on diversity and conservation. Trends in Ecology and Evolution, 22, 148 - 155. https: // doi. org / 10.1016 / j. tree. 2006.11.004" type="journal article" year="2007">
Bickford 
<emphasis id="B953C16D1065ED5DC14BFED3FCE0FEE3" box="[787,844,259,285]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2007
</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1065ED5DC1C1FED3FBCBFEE3" author="Hebert, P. D. &amp; Penton, E. H. &amp; Burns, J. M. &amp; Janzen, D. H. &amp; Hallwachs, W." box="[921,1127,259,285]" pageId="18" pageNumber="319" pagination="14812 - 14817" refId="ref13523" refString="Hebert, P. D., Penton, E. H., Burns, J. M., Janzen, D. H. &amp; Hallwachs, W. (2004) Ten species in one: DNA barcoding reveals cryptic species in the Neotropical skipper butterfly Astraptes fulgerator. Proceedings of the National Academy of Sciences of the United States of America, 101, 14812 - 14817. https: // doi. org / 10.1073 / pnas. 0406166101" type="journal article" year="2004">
Hebert 
<emphasis id="B953C16D1065ED5DC1B4FED3FB88FEE3" box="[1004,1060,259,285]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2004
</bibRefCitation>
; 
<bibRefCitation id="EFB6608E1065ED5DC62AFED3FB4EFEE3" author="Hill, G. E." box="[1138,1250,259,285]" pageId="18" pageNumber="319" pagination="5831 - 5842" refId="ref13644" refString="Hill, G. E. (2016) Mitonuclear coevolution as the genesis of speciation and the mitochondrial DNA barcode gap. Ecology and Evolution, 6 (16), 5831 - 5842. https: // doi. org / 10.1002 / ece 3.2338" type="journal article" year="2016">Hill 2016</bibRefCitation>
).
</paragraph>
<caption id="DF584DF71065ED5DC2C0FBBAFB84FB58" ID-DOI="http://doi.org/10.5281/zenodo.14239593" ID-Zenodo-Dep="14239593" httpUri="https://zenodo.org/record/14239593/files/figure.png" pageId="18" pageNumber="319" startId="18.[152,255,1130,1154]" targetBox="[186,1402,309,1107]" targetPageId="18" targetType="figure">
<paragraph id="8B981D7F1065ED5DC2C0FBBAFB84FB58" blockId="18.[151,1436,1130,1191]" pageId="18" pageNumber="319">
<emphasis id="B953C16D1065ED5DC2C0FBBAFE88FB7C" bold="true" box="[152,292,1130,1154]" pageId="18" pageNumber="319">FIGURE 20.</emphasis>
Spatial distribution of 
<taxonomicName id="4C2766FC1065ED5DC050FBBAFC61FB7C" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[520,973,1130,1155]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D1065ED5DC050FBBAFD42FB7C" box="[520,750,1130,1154]" italics="true" pageId="18" pageNumber="319">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
<emphasis id="B953C16D1065ED5DC18AFBBBFB8DFB7D" bold="true" box="[978,1057,1131,1155]" pageId="18" pageNumber="319">
<taxonomicNameLabel id="A2607C161065ED5DC18AFBBBFB8DFB7D" box="[978,1057,1131,1155]" pageId="18" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Turkey. Star: specimens with DNA information. B. Bulgaria, C: Cyprus, Ge: Georgia, Gr: Greece, I: Iraq, R: Russia, S: Syria.
</paragraph>
</caption>
<paragraph id="8B981D7F1065ED5DC29FFB3BFEBAF9B4" blockId="18.[151,1437,1259,2042]" pageId="18" pageNumber="319">
The corresponding differences observed at genetic and morphological levels corroborate the hypothesis that 
<taxonomicName id="4C2766FC1065ED5DC7DCFB3CFF5CFAD7" authorityName="Hagen" authorityYear="1859" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1065ED5DC7DCFB3CFF5CFAD7" italics="true" pageId="18" pageNumber="319">R. fasciata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2766FC1065ED5DC37CFAC0FE00FAD7" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" box="[292,428,1295,1321]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1065ED5DC37CFAC0FE00FAD7" box="[292,428,1295,1321]" italics="true" pageId="18" pageNumber="319">R. anatolica</emphasis>
</taxonomicName>
are two different species: The intraspecific distances were always less than 1% divergence, and interspecific distances were higher than 1% (
<tableCitation id="C6A528C41065ED5DC09AFAE3FCB6FAB0" box="[706,794,1331,1358]" captionStart="TABLE 2" captionStartId="17.[152,244,151,175]" captionTargetPageId="17" captionText="TABLE 2. Genetic distances inand among “Rhyacophila fasciata Species Complex” taxa (maximum composite likelihood model). 2a: intraspecific distances, n/c = not computed (1 specimen only). 2b: interspecific distances. Table 2a" httpUri="http://table.plazi.org/id/DF584DF71066ED5EC2C0FF47FEACFF05" pageId="18" pageNumber="319" tableUuid="DF584DF71066ED5EC2C0FF47FEACFF05">Table 2</tableCitation>
). This result is similar to what we found in specimens of 
<taxonomicName id="4C2766FC1065ED5DC2CFFA88FE9FFA8F" authorityName="McLachlan" authorityYear="1879" box="[151,307,1367,1393]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B953C16D1065ED5DC2CFFA88FE9FFA8F" box="[151,307,1367,1393]" italics="true" pageId="18" pageNumber="319">R. denticulata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2766FC1065ED5DC330FA88FE7BFA8F" authorityName="Navas" authorityYear="1916" box="[360,471,1368,1393]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="sociata">
<emphasis id="B953C16D1065ED5DC330FA88FE7BFA8F" box="[360,471,1368,1393]" italics="true" pageId="18" pageNumber="319">R. sociata</emphasis>
</taxonomicName>
(Valladolid 
<emphasis id="B953C16D1065ED5DC038FA88FD32FA8F" box="[608,670,1367,1393]" italics="true" pageId="18" pageNumber="319">et. al.</emphasis>
2018), 
<taxonomicName id="4C2766FC1065ED5DC0ABFA88FB2AFA8C" authority="(Valladolid et al. 2019)" baseAuthorityName="Valladolid" baseAuthorityYear="2019" box="[755,1158,1367,1394]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="kykladica">
<emphasis id="B953C16D1065ED5DC0ABFA88FCD0FA8F" box="[755,892,1367,1393]" italics="true" pageId="18" pageNumber="319">R. kykladica</emphasis>
(
<bibRefCitation id="EFB6608E1065ED5DC1D1FA87FBD2FA8C" author="Valladolid, M. &amp; Karaouzas, I. &amp; Arauzo, M. &amp; Dorda, B. A. &amp; Rey, I." box="[905,1150,1367,1394]" pageId="18" pageNumber="319" pagination="503 - 522" refId="ref16560" refString="Valladolid, M., Karaouzas, I., Arauzo, M., Dorda, B. A. &amp; Rey, I. (2019) The Rhyacophila fasciata Group in Greece: Rhyacophila kykladica Malicky &amp; Sipahiler 1993 (stat. prom.) (Trichoptera: Rhyacophilidae). Morphological description, genetic and ecological features, Zootaxa, 4657 (3), 503 - 522. https: // doi. org / 10.11646 / zootaxa. 4657.3.5" type="journal article" year="2019">
Valladolid 
<emphasis id="B953C16D1065ED5DC65DFA88FB90FA8F" box="[1029,1084,1367,1393]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C2766FC1065ED5DC6C8FA88FB41FA8F" authorityName="Kucinic &amp; Valladolid" authorityYear="2020" box="[1168,1261,1367,1393]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="delici">
<emphasis id="B953C16D1065ED5DC6C8FA88FB41FA8F" box="[1168,1261,1367,1393]" italics="true" pageId="18" pageNumber="319">R. delici</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C2766FC1065ED5DC771FA88FE0CFA68" authority="(Valladolid et al. 2020)" baseAuthorityName="Valladolid" baseAuthorityYear="2020" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="viteceki">
<emphasis id="B953C16D1065ED5DC771FA88FA31FA8F" box="[1321,1437,1367,1393]" italics="true" pageId="18" pageNumber="319">R. viteceki</emphasis>
(
<bibRefCitation id="EFB6608E1065ED5DC2C6FAABFE34FA68" author="Valladolid, M. &amp; Kucinic, M. &amp; Arauzo, M. &amp; Cerjanec, D. &amp; Cuk, R. &amp; Dorda, B. A. &amp; Lodovici, O. &amp; Stanic Kostroman, S. &amp; Vuckovic, I. &amp; Rey, I." box="[158,408,1403,1430]" pageId="18" pageNumber="319" refId="ref16645" refString="Valladolid, M., Kucinic, M., Arauzo, M., Cerjanec, D., Cuk, R., Dorda, B. A., Lodovici, O., Stanic Kostroman, S., Vuckovic, I. &amp; Rey, I. (2020) The Rhyacophila fasciata Group in Croatia and Bosnia and Herzegovina: Rhyacophila f. fasciata Hagen, 1859 and the description of two new subspecies, Rhyacophila fasciata delici Kucinic &amp; Valladolid (ssp. nov.) from Croatia and Bosnia and Herzegovina and Rhyacophila fasciata viteceki Valladolid &amp; Kucinic (ssp. nov.) from Bosnia and" type="journal volume" year="2020">
Valladolid 
<emphasis id="B953C16D1065ED5DC343FAACFEF8FA6B" box="[283,340,1403,1429]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2020
</bibRefCitation>
)
</taxonomicName>
, or 
<taxonomicName id="4C2766FC1065ED5DC393FAACFCF8FA68" authority="(Valladolid et al. 2023)" baseAuthorityName="Valladolid" baseAuthorityYear="2023" box="[459,852,1403,1430]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B953C16D1065ED5DC393FAACFDE8FA6B" box="[459,580,1403,1429]" italics="true" pageId="18" pageNumber="319">R. loeffleri</emphasis>
(
<bibRefCitation id="EFB6608E1065ED5DC00AFAABFCE0FA68" author="Valladolid, M. &amp; Waringer, J. &amp; Arauzo, M. &amp; Chvojka, P. &amp; Dorda, B. A. &amp; Komzak, P. &amp; Lodovici, O. &amp; Rey, I." box="[594,844,1403,1430]" pageId="18" pageNumber="319" pagination="451 - 482" refId="ref17056" refString="Valladolid, M., Waringer, J., Arauzo, M., Chvojka, P., Dorda, B. A., Komzak, P., Lodovici, O., Rey, I. (2023) The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in Central Europe with description of a new species, Rhyacophila loeffleri Valladolid &amp; Waringer, n. sp., based on morphological, genetic and ecological evidence. Zootaxa, 5325 (4), 451 - 482. https: // doi. org / 10.11646 / zootaxa. 5325.4.1" type="journal article" year="2023">
Valladolid 
<emphasis id="B953C16D1065ED5DC097FAACFCA4FA6B" box="[719,776,1403,1429]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
2023
</bibRefCitation>
)
</taxonomicName>
. Other authors that have analysed the divergence of COI sequences between different species of several taxonomic groups found that more than 98% of the species pairs show divergences greater than 2% (e.g., 
<bibRefCitation id="EFB6608E1065ED5DC0FEFA13FC24FA23" author="Hebert, P. D. N. &amp; Ratnasingham, S. &amp; de Waard, J. R." box="[678,904,1475,1502]" pageId="18" pageNumber="319" refId="ref13448" refString="Hebert, P. D. N., Ratnasingham, S. &amp; de Waard, J. R. (2003 b) Barcoding animal life: Cytochrome c oxidase subunit 1 divergences among closely related species. Proceedings of the Royal Society of London, Series B, 270 (Supplement 1), S 96 - S 99. https: // doi. org / 10.1098 / rsbl. 2003.0025" type="journal volume" year="2003">
Hebert 
<emphasis id="B953C16D1065ED5DC0A3FA14FC82FA23" box="[763,814,1475,1501]" italics="true" pageId="18" pageNumber="319">et al</emphasis>
. 2003b
</bibRefCitation>
), and that the intraspecific divergences usually are less than 1% (
<bibRefCitation id="EFB6608E1065ED5DC302FA37FE73F9FC" author="Avise, J. C." box="[346,479,1511,1538]" pageId="18" pageNumber="319" refId="ref12338" refString="Avise, J. C. (2000) Phylogeography. The History and Formation of Species. University Press, Cambridge, Massachusetts, 447 pp. https: // doi. org / 10.2307 / j. ctv 1 nzfgj 7" type="book" year="2000">Avise 2000</bibRefCitation>
). We also agree with the premise of 
<bibRefCitation id="EFB6608E1065ED5DC128FA37FBFBF9FC" author="Hebert, P. D. N. &amp; Cywinska, A. &amp; Ball, S. L. &amp; de Waard, J. R." box="[880,1111,1511,1538]" pageId="18" pageNumber="319" pagination="313 - 321" refId="ref13378" refString="Hebert, P. D. N., Cywinska, A., Ball, S. L. &amp; de Waard, J. R. (2003 a) Biological identifications through DNA barcodes. Proceedings of the Royal Society of London, Series B, 270 (1512), 313 - 321. https: // doi. org / 10.1098 / rspb. 2002.2218" type="journal article" year="2003">
Hebert 
<emphasis id="B953C16D1065ED5DC19AFA38FC56F9FF" box="[962,1018,1511,1537]" italics="true" pageId="18" pageNumber="319">et al.</emphasis>
(2003a)
</bibRefCitation>
that we can find new species by their genetic divergences from known species assemblages. In this case, the species was identified originally as 
<taxonomicName id="4C2766FC1065ED5DC2CFF9E0FEA3F9B7" authorityName="Hagen" authorityYear="1859" box="[151,271,1583,1609]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1065ED5DC2CFF9E0FEA3F9B7" box="[151,271,1583,1609]" italics="true" pageId="18" pageNumber="319">R. fasciata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B981D7F1065ED5DC29FF984FC2EF900" blockId="18.[151,1437,1259,2042]" pageId="18" pageNumber="319">
In future studies we intend to prepare Cytochrome B gene sequences for these specimens. This gene, used extensively and particularly for vertebrates, has enabled researchers to resolve relationships among closely related taxa (Tauzt 
<emphasis id="B953C16D1065ED5DC344F94CFEE2F94B" box="[284,334,1691,1717]" italics="true" pageId="18" pageNumber="319">et al</emphasis>
. 2003) and, combined with the COI findings, could lead us to a better knowledge of this group of 
<taxonomicName id="4C2766FC1065ED5DC2CFF96FFEB5F927" authorityName=", McLachlan" authorityYear="1865" box="[151,281,1727,1753]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
in Europe. Another option that we would like to explore is the inclusion of a nuclear gene in the study, which complements the information from the mitochondrial DNA.
</paragraph>
<paragraph id="8B981D7F1065ED5DC29FF8D8FE89F807" blockId="18.[151,1437,1259,2042]" pageId="18" pageNumber="319">
Our successive studies of the “ 
<taxonomicName id="4C2766FC1065ED5DC01CF8D8FD6FF8DF" authorityName="Hagen" authorityYear="1859" box="[580,707,1799,1825]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1065ED5DC01CF8D8FD6FF8DF" box="[580,707,1799,1825]" italics="true" pageId="18" pageNumber="319">R. fasciata</emphasis>
</taxonomicName>
Species Complex” have increased the number of sequences (haplotypes), and provided more information about the historical relationships among the species previously described and possible new species, mainly in those species apparently sharing a recent common ancestor. Based on our morphological and genetic evidence, we therefore propose a new species: 
<taxonomicName id="4C2766FC1065ED5DC66FF8A3FE8BF84F" authority="Ekingen &amp; Valladolid" authorityName="Ekingen &amp; Valladolid" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B953C16D1065ED5DC66FF8A3FA9BF873" box="[1079,1335,1907,1933]" italics="true" pageId="18" pageNumber="319">Rhyacophila anatolica</emphasis>
Ekingen &amp; Valladolid
</taxonomicName>
(
<emphasis id="B953C16D1065ED5DC361F848FE3FF84C" bold="true" box="[313,403,1944,1970]" pageId="18" pageNumber="319">sp. nov.</emphasis>
) from 
<collectingCountry id="F3305DEF1065ED5DC3BAF847FD98F84F" box="[482,564,1943,1969]" name="Turkey" pageId="18" pageNumber="319">Turkey</collectingCountry>
. All the specimens from this country cited as 
<taxonomicName id="4C2766FC1065ED5DC662F848FABFF84F" box="[1082,1299,1943,1969]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="subSpecies" species="fasciata" subSpecies="fasciata">
<emphasis id="B953C16D1065ED5DC662F848FABFF84F" box="[1082,1299,1943,1969]" italics="true" pageId="18" pageNumber="319">R. fasciata fasciata</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFB6608E1065ED5DC767F847FEC0F828" author="Malicky, H. &amp; Sipahiler, F." pageId="18" pageNumber="319" pagination="457 - 468" refId="ref14009" refString="Malicky, H. &amp; Sipahiler, F. (1993) Kocherfliegen (Trichoptera) aus der Turkei, mit Bemerkungen zu weiteren mediterranean Kocherfliegen. Bulletin de la societe entomologique Suisse, 66, 457 - 468." type="journal article" year="1993">Malicky &amp; Sipahiler (1993)</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1065ED5DC320F86BFD8BF828" author="Sipahiler, F." box="[376,551,1979,2006]" pageId="18" pageNumber="319" pagination="36 - 42" refId="ref15525" refString="Sipahiler, F. (2007) The Trichoptera fauna of North-western Turkey with descriptions of a new species and of some previously unknown females (Philopotamidae, Sericostomatidae). Braueria, 34, 36 - 42. Available from: https: // www. zobodat. at / pdf / BRA _ 34 _ 0036 - 0042. pdf (accessed 11 January 2024)" type="journal article" year="2007">Sipahiler (2007</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1065ED5DC06AF86CFDC1F828" author="Sipahiler, F." box="[562,621,1980,2006]" pageId="18" pageNumber="319" pagination="93 - 109" refId="ref15593" refString="Sipahiler, F. (2008) Zoological characteristics of the Trichoptera Fauna of Turkey. In: Meyer, M. &amp; Neu, P. (Eds) Proceedings of the First Conference on Faunistics and Zoogeography of European Trichoptera. Ferrantia, 55, 93 - 109. Available from: https: // ps. mnhn. lu / ferrantia / publications / Ferrantia 55. pdf (accessed 11 January 2024)" type="journal article" year="2008">2008</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1065ED5DC020F86CFD1FF828" author="Sipahiler, F." box="[632,691,1980,2006]" pageId="18" pageNumber="319" pagination="1 - 17" refId="ref15670" refString="Sipahiler, F. (2012) Five new species of Trichoptera with the faunistic list of Sinop and Samsun Provinces in Turkey (Glossosomatidae, Philopotamidae, Hydropsychidae, Sericostomatidae). Munis Entomology &amp; Zoology, 7 (1), 1 - 17. Available from: https: // www. munisentzool. org / Issue / 2012 - vol- 7 - number- 1 - 14 (accessed 11 January 2024)" type="journal article" year="2012">2012</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1065ED5DC0E6F86CFD55F828" author="Sipahiler, F." box="[702,761,1980,2006]" pageId="18" pageNumber="319" pagination="43 - 55" refId="ref15746" refString="Sipahiler, F. (2013) Revision of the Rhyacopghila stigmata Species Group in Turkey with descriptions of three new species (Trichoptera, Rhyacophilidae). Zootaxa, 3669 (1), 43 - 55. https: // doi. org / 10.11646 / zootaxa. 3669.1.5" type="journal article" year="2013">2013</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1065ED5DC15BF86CFC92F828" author="Sipahiler, F." box="[771,830,1980,2006]" pageId="18" pageNumber="319" pagination="542 - 553" refId="ref15797" refString="Sipahiler, F. (2014) Three new species of Trichoptera (Odontoceridae, Leptoceridae) and the faunistic list for Zonguldak and Karabuk provinces in northwestern Turkey. Munis Entomology &amp; Zoology, 9 (1), 542 - 553. Available from: https: // www. munisentzool. org / Issue / 2014 - vol- 9 - number- 1 - 18 (accessed 11 January 2024)" type="journal article" year="2014">2014</bibRefCitation>
, 
<bibRefCitation id="EFB6608E1065ED5DC111F86BFC29F82B" author="Sipahiler, F." box="[841,901,1979,2005]" pageId="18" pageNumber="319" pagination="11 - 16" refId="ref15870" refString="Sipahiler, F. (2016) Faunistic studies on the Trichoptera fauna of northwestern Turkey and Thrace. Braueria, 43, 11 - 16. Available from: https: // www. zobodat. at / pdf / BRA _ 43 _ 0011 - 0016. pdf (accessed 11 January 2024)" type="journal article" year="2016">2016</bibRefCitation>
), and 
<bibRefCitation id="EFB6608E1065ED5DC19FF86BFB52F828" author="Sipahiler, F. &amp; Malicky, H." box="[967,1278,1979,2006]" pageId="18" pageNumber="319" pagination="77 - 165" refId="ref15926" refString="Sipahiler, F. &amp; Malicky, H. (1987) Die Kocherfliegen der Turkei (Trichoptera). Entomofauna, 8, 77 - 165. Available from: https: // www. zobodat. at / pdf / ENT _ 0008 _ 0077 - 0165. pdf (accessed 11 January 2024)" type="journal article" year="1987">Sipahiler &amp; Malicky (1987)</bibRefCitation>
belong to this new species.
</paragraph>
<paragraph id="8B981D7F1064ED5CC29FFF47FCC1FF07" blockId="19.[151,1436,151,249]" pageId="19" pageNumber="320">
Finally, the descriptions of species included in the “ 
<taxonomicName id="4C2766FC1064ED5CC14DFF47FC23FF4F" authorityName="Hagen" authorityYear="1859" box="[789,911,151,177]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B953C16D1064ED5CC14DFF47FC23FF4F" box="[789,911,151,177]" italics="true" pageId="19" pageNumber="320">R. fasciata</emphasis>
</taxonomicName>
Species Complex”, together with the addition of a few more descriptions of new species in this group (in preparation) will allow us to prepare a synopsis of the diagnostic characteristics of the Complex in a future publication.
</paragraph>
</subSubSection>
<subSubSection id="C33D4EF41064ED5CC2CFFE9AFF4CFD57" pageId="19" pageNumber="320" type="distribution">
<paragraph id="8B981D7F1064ED5CC2CFFE9AFE67FE9B" blockId="19.[151,459,330,357]" box="[151,459,330,357]" pageId="19" pageNumber="320">
<emphasis id="B953C16D1064ED5CC2CFFE9AFE67FE9B" bold="true" box="[151,459,330,357]" pageId="19" pageNumber="320">Distribution and ecology</emphasis>
</paragraph>
<paragraph id="8B981D7F1064ED5CC2CFFE43FF4CFD57" blockId="19.[151,1437,403,681]" pageId="19" pageNumber="320">
<figureCitation id="131C01FA1064ED5CC2CFFE43FEAFFE53" box="[151,259,403,429]" captionStart="FIGURE 20" captionStartId="18.[152,255,1130,1154]" captionTargetBox="[186,1402,309,1107]" captionTargetId="figure-87@18.[186,1402,309,1107]" captionTargetPageId="18" captionText="FIGURE 20. Spatial distribution of Rhyacophila anatolica Ekingen &amp; Valladolid sp. nov. in Turkey. Star: specimens with DNA information. B. Bulgaria, C: Cyprus, Ge: Georgia, Gr: Greece, I: Iraq, R: Russia, S: Syria." figureDoi="http://doi.org/10.5281/zenodo.14239593" httpUri="https://zenodo.org/record/14239593/files/figure.png" pageId="19" pageNumber="320">Figure 20</figureCitation>
presents the current knowledge of the distribution of 
<taxonomicName id="4C2766FC1064ED5CC10DFE43FC72FE53" authorityName="Valladolid &amp; Ekingen &amp; Arauzo &amp; Dorda &amp; Rey" authorityYear="2024" box="[853,990,403,429]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B953C16D1064ED5CC10DFE43FC72FE53" box="[853,990,403,429]" italics="true" pageId="19" pageNumber="320">R. anatolica</emphasis>
</taxonomicName>
<emphasis id="B953C16D1064ED5CC1BCFE43FB90FE53" bold="true" box="[996,1084,403,429]" pageId="19" pageNumber="320">
<taxonomicNameLabel id="A2607C161064ED5CC1BCFE43FB90FE53" box="[996,1084,403,429]" pageId="19" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in 
<collectingCountry id="F3305DEF1064ED5CC606FE43FB1CFE53" box="[1118,1200,403,429]" name="Turkey" pageId="19" pageNumber="320">Turkey</collectingCountry>
. The species is found in the western and central Black Sea Region, and east of the Marmara Region, in small forest streams flowing into the Black Sea and the Marmara Sea. The substrate consists of boulder, cobble, pebble, and gravel, with sand in very small amounts, with rich stream bank vegetation, and consequently entrained fallen leaves and twigs. Sampling sites are relatively isolated from human influence, and range from 
<quantity id="4CDFB09A1064ED5CC11DFDF2FC75FDC3" box="[837,985,546,573]" metricMagnitude="2" metricUnit="m" metricValue="7.845" metricValueMax="15.34" metricValueMin="0.35" pageId="19" pageNumber="320" unit="m" value="784.5" valueMax="1534.0" valueMin="35.0">35 to 1534 m</quantity>
a.s.l. Flight activity occurs usually from late May to November. In 
<collectingRegion id="49E3D39D1064ED5CC39EFD97FD85FD9F" box="[454,553,583,609]" country="Turkey" name="Karabuk" pageId="19" pageNumber="320">Karabük</collectingRegion>
, larvae are found in alkaline waters (pH 7.58–8.65), in streams 
<quantity id="4CDFB09A1064ED5CC758FD97FAFBFD9F" box="[1280,1367,583,609]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="19" pageNumber="320" unit="m" value="7.0" valueMax="12.0" valueMin="2.0">2–12 m</quantity>
wide, and with water temperatures from 10.41 to 19.53ºC, during three seasons (spring, summer, and autumn) (
<bibRefCitation id="EFB6608E1064ED5CC765FDBBFF78FD57" author="Ekingen, P." pageId="19" pageNumber="320" pagination="395 - 406" refId="ref12797" refString="Ekingen, P. (2022) Preliminary study of the relationship between caddisfly larvae and environmental variables in SimSir Stream (Karabuk, Turkey). Journal of the Entomological Research Society, 24 (3), 395 - 406. https: // doi. org / 10.51963 / jers. v 24 i 3.2333" type="journal article" year="2022">Ekingen 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>