treatments-xml/data/03/8E/8D/038E8D48617BA84CFF73F971F2D9F98B.xml
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<mods:title id="9D221DBDE446B7989A70031582CCA805">Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key</mods:title>
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<mods:namePart id="F5827205044BD153401D2B122522EC53">Telnov, Dmitry</mods:namePart>
<mods:affiliation id="615FCB61733521BC990C2680296AED63">Department of Life Sciences, Natural History Museum, SW 7 5 BD, London, United Kingdom. &amp; Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401, Daugavpils, Latvia &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004, Rīga, Latvia</mods:affiliation>
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<treatment id="038E8D48617BA84CFF73F971F2D9F98B" ID-DOI="http://doi.org/10.5281/zenodo.7415034" ID-GBIF-Taxon="205926629" ID-Zenodo-Dep="7415034" LSID="urn:lsid:zoobank.org:act:68B79D0D-502D-4536-9B65-CD90D7C41031" httpUri="http://treatment.plazi.org/id/038E8D48617BA84CFF73F971F2D9F98B" lastPageId="15" lastPageNumber="136" pageId="7" pageNumber="128">
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<taxonomicName id="4C2747DD617BA844FF73F971F787F968" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[151,400,1734,1761]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C617BA844FF73F971F787F968" bold="true" box="[151,400,1734,1761]" italics="true" pageId="7" pageNumber="128">Physciolagria smithi</emphasis>
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(
<figureCitation id="131C20DB617BA844FF7BF95CF737F88C" box="[159,288,1770,1797]" captionStart-0="FIGURES 1517" captionStart-1="FIGURES 1819" captionStartId-0="9.[152,270,1963,1987]" captionStartId-1="10.[152,270,1273,1297]" captionTargetBox-0="[188,1399,199,1938]" captionTargetBox-1="[152,1436,197,1236]" captionTargetId-0="figure-15@9.[173,1414,181,1939]" captionTargetId-1="figure-33@10.[151,1436,181,1249]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 1517. Physciolagria smithi sp. nov., holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale]." captionText-1="FIGURES 1819. Physciolagria smithi sp. nov., paratype ♀. 18—Habitus, dorsal view; 19—Forebody, anterodorsal view [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.7413360" figureDoi-1="http://doi.org/10.5281/zenodo.7413362" httpUri-0="https://zenodo.org/record/7413360/files/figure.png" httpUri-1="https://zenodo.org/record/7413362/files/figure.png" pageId="7" pageNumber="128">Figs 1519</figureCitation>
,
<figureCitation id="131C20DB617BA844FEC8F95DF789F88C" box="[300,414,1770,1797]" captionStart="FIGURES 2027" captionStartId="11.[152,270,1776,1800]" captionTargetBox="[159,1428,199,1738]" captionTargetId="figure-15@11.[151,1436,181,1752]" captionTargetPageId="11" captionText="FIGURES 2027. Male terminalia of Physciolagria Pic, 1930. 20—P. liturata Pic, 1930, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—P. smithi sp. nov., paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, spiculum gastrale; 27—P. smithi sp. nov., paratype, spiculum gastrale [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="7" pageNumber="128">2425, 27</figureCitation>
,
<figureCitation id="131C20DB617BA844FE4EF95DF7E5F88C" box="[426,498,1770,1797]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="7" pageNumber="128">3335</figureCitation>
,
<figureCitation id="131C20DB617BA844FE1AF95CF452F88C" box="[510,581,1771,1797]" captionStart="FIGURES 3644" captionStartId="13.[152,270,1906,1930]" captionTargetBox="[187,1403,196,1860]" captionTargetId="figure-15@13.[166,1422,181,1883]" captionTargetPageId="13" captionText="FIGURES 3644. Median lobe of Physciolagria Pic, 1930. 36—P. liturata Pic, 1930; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—P. smithi sp. nov., paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="7" pageNumber="128">4244</figureCitation>
)
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https://zoobank.org/
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[printed] //
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,
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, leg.
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—same labels as
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—same labels but
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<specimenCount id="9D21F7D76174A84BFE5AFF20F7CDFF38" box="[446,474,151,177]" pageId="8" pageNumber="129" type="male">1♂</specimenCount>
—same labels but
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—same labels but
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—same labels but
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[printed] //
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:2019.14, BMNH{
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E} 2020-19 [printed] //
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[printed] //
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:2019.14, BMNH{
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E} 2020-19 [printed] //
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LepiLED [printed] //
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:2019.14, BMNH{
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E} 2020-19 [printed] //
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[printed, provided with QR code];
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—same labels but
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD56174A84BFF23FDF1F2E5FD0C" pageId="8" pageNumber="129" type="etymology">
<paragraph id="8B983C5E6174A84BFF23FDF1F2E5FD0C" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="129">
<emphasis id="B953E04C6174A84BFF23FDF1F7B5FDE9" bold="true" box="[199,418,582,608]" pageId="8" pageNumber="129">Derivatio nominis.</emphasis>
Patronymic.This species is named for Richard Smith (The African Natural History Research Trust, Kingsland,
<collectingCountry id="F3307CCE6174A84BFE84FDDCF437FD0C" box="[352,544,619,645]" name="United Kingdom" pageId="8" pageNumber="129">United Kingdom</collectingCountry>
) to commemorate his investment in the study of African insects.
</paragraph>
</subSubSection>
<caption id="DF586CD66174A84BFF7CF801F4E8F87B" ID-DOI="http://doi.org/10.5281/zenodo.7413358" ID-Zenodo-Dep="7413358" httpUri="https://zenodo.org/record/7413358/files/figure.png" pageId="8" pageNumber="129" startId="8.[152,270,1974,1998]" targetBox="[151,1435,669,1949]" targetPageId="8">
<paragraph id="8B983C5E6174A84BFF7CF801F4E8F87B" blockId="8.[151,1436,1974,2034]" pageId="8" pageNumber="129">
<emphasis id="B953E04C6174A84BFF7CF801F74DF847" bold="true" box="[152,346,1974,1998]" pageId="8" pageNumber="129">FIGURES 1214.</emphasis>
<taxonomicName id="4C2747DD6174A84BFE84F801F51EF847" authority="(Pic, 1935)" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1935" box="[352,777,1974,1998]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C6174A84BFE84F801F487F847" box="[352,656,1974,1998]" italics="true" pageId="8" pageNumber="129">Physciolagria singularicornis</emphasis>
(
<bibRefCitation id="EFB641AF6174A84BFD79F801F516F847" author="Pic, M." box="[669,769,1974,1998]" pageId="8" pageNumber="129" pagination="1 - 36" refId="ref8308" refString="Pic, M. (1935) Nouveautes diverses. Melanges exotico-entomologiques 65, 1 - 36." type="journal article" year="1935">Pic, 1935</bibRefCitation>
)
</taxonomicName>
, ♂ from Sebele, Botswana. 12—Habitus, dorsal view; 13—ditto, anterodorsal view; 14—Forebody, dorsal view [not to scale].
</paragraph>
</caption>
<caption id="DF586CD66175A84AFF7CF81CF7C9F86E" ID-DOI="http://doi.org/10.5281/zenodo.7413360" ID-Zenodo-Dep="7413360" httpUri="https://zenodo.org/record/7413360/files/figure.png" pageId="9" pageNumber="130" startId="9.[152,270,1963,1987]" targetBox="[188,1399,199,1938]" targetPageId="9">
<paragraph id="8B983C5E6175A84AFF7CF81CF7C9F86E" blockId="9.[151,1437,1963,2023]" pageId="9" pageNumber="130">
<emphasis id="B953E04C6175A84AFF7CF81CF74EF84A" bold="true" box="[152,345,1963,1987]" pageId="9" pageNumber="130">FIGURES 1517.</emphasis>
<taxonomicName id="4C2747DD6175A84AFEB9F81CF439F84A" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[349,558,1963,1987]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="130" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C6175A84AFEB9F81CF439F84A" box="[349,558,1963,1987]" italics="true" pageId="9" pageNumber="130">Physciolagria smithi</emphasis>
</taxonomicName>
<emphasis id="B953E04C6175A84AFDD7F81CF495F84A" bold="true" box="[563,642,1963,1987]" pageId="9" pageNumber="130">
<taxonomicNameLabel id="A2605D376175A84AFDD7F81CF495F84A" box="[563,642,1963,1987]" pageId="9" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale].
</paragraph>
</caption>
<caption id="DF586CD66176A849FF7CFB4EF6E4FABC" ID-DOI="http://doi.org/10.5281/zenodo.7413362" ID-Zenodo-Dep="7413362" httpUri="https://zenodo.org/record/7413362/files/figure.png" pageId="10" pageNumber="131" startId="10.[152,270,1273,1297]" targetBox="[152,1436,197,1236]" targetPageId="10">
<paragraph id="8B983C5E6176A849FF7CFB4EF6E4FABC" blockId="10.[151,1436,1273,1333]" pageId="10" pageNumber="131">
<emphasis id="B953E04C6176A849FF7CFB4EF74EFA98" bold="true" box="[152,345,1273,1297]" pageId="10" pageNumber="131">FIGURES 1819.</emphasis>
<taxonomicName id="4C2747DD6176A849FEBBFB4EF427FA98" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[351,560,1273,1297]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="131" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C6176A849FEBBFB4EF427FA98" box="[351,560,1273,1297]" italics="true" pageId="10" pageNumber="131">Physciolagria smithi</emphasis>
</taxonomicName>
<emphasis id="B953E04C6176A849FDD2FB4EF491FA98" bold="true" box="[566,646,1273,1297]" pageId="10" pageNumber="131">
<taxonomicNameLabel id="A2605D376176A849FDD2FB4EF491FA98" box="[566,646,1273,1297]" pageId="10" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype ♀. 18—Habitus, dorsal view; 19—Forebody, anterodorsal view [not to scale].
</paragraph>
</caption>
<subSubSection id="C33D6FD56176A84CFF23FAD4F30AFAA2" lastPageId="15" lastPageNumber="136" pageId="10" pageNumber="131" type="description">
<paragraph id="8B983C5E6176A849FF23FAD4F474FA60" blockId="10.[151,1437,1379,2018]" pageId="10" pageNumber="131">
<emphasis id="B953E04C6176A849FF23FAD4F769FAF4" bold="true" box="[199,382,1379,1405]" pageId="10" pageNumber="131">Measurements.</emphasis>
<typeStatus id="549C82FC6176A849FE61FAD4F7FAFAF4" box="[389,493,1379,1405]" pageId="10" pageNumber="131" type="holotype">Holotype</typeStatus>
<specimenCount id="9D21F7D76176A849FE12FAD3F41CFAF7" box="[502,523,1380,1406]" pageId="10" pageNumber="131" type="male"></specimenCount>
(
<figureCitation id="131C20DB6176A849FDFFFAD4F48AFAF7" box="[539,669,1379,1406]" captionStart="FIGURES 1517" captionStartId="9.[152,270,1963,1987]" captionTargetBox="[188,1399,199,1938]" captionTargetId="figure-15@9.[173,1414,181,1939]" captionTargetPageId="9" captionText="FIGURES 1517. Physciolagria smithi sp. nov., holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413360" httpUri="https://zenodo.org/record/7413360/files/figure.png" pageId="10" pageNumber="131">Figs 1517</figureCitation>
), total body length
<quantity id="4CDF91BB6176A849FC9FFAD4F5C2FAF7" box="[891,981,1379,1406]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="131" unit="mm" value="3.7">3.7 mm</quantity>
; head length
<quantity id="4CDF91BB6176A849FB88FAD4F2D2FAF7" box="[1132,1221,1379,1406]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="131" unit="mm" value="0.6">0.6 mm</quantity>
, head width across eyes
<quantity id="4CDF91BB6176A849FF34FA30F73FFA2B" box="[208,296,1415,1442]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="131" unit="mm" value="0.9">0.9 mm</quantity>
, pronotal length
<quantity id="4CDF91BB6176A849FE02FA30F429FA2B" box="[486,574,1415,1442]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="131" unit="mm" value="0.6">0.6 mm</quantity>
, maximum pronotal width
<quantity id="4CDF91BB6176A849FC89FA30F5D1FA2B" box="[877,966,1415,1442]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="131" unit="mm" value="0.8">0.8 mm</quantity>
, minimum pronotal width
<quantity id="4CDF91BB6176A849FB15FA30F35EFA2B" box="[1265,1353,1415,1442]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="131" unit="mm" value="0.7">0.7 mm</quantity>
, elytral length
<quantity id="4CDF91BB6176A849FF03FA1CF754FA4F" box="[231,323,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="131" unit="mm" value="2.5">2.5 mm</quantity>
, combined maximum elytral width in apical third
<quantity id="4CDF91BB6176A849FC6AFA1CF5FCFA4F" box="[910,1003,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="131" unit="mm" value="1.3">1.3 mm</quantity>
. Selected
<specimenCount id="9D21F7D76176A849FB85FA1BF29CFA4F" box="[1121,1163,1452,1478]" pageId="10" pageNumber="131" type="male">♂♂</specimenCount>
<typeStatus id="549C82FC6176A849FB72FA1BF316FA4F" box="[1174,1281,1452,1478]" pageId="10" pageNumber="131" type="paratype">paratypes</typeStatus>
<quantity id="4CDF91BB6176A849FAEFFA1CF38FFA4F" box="[1291,1432,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="10" pageNumber="131" unit="mm" value="3.9" valueMax="4.3" valueMin="3.5">3.54.3 mm</quantity>
, selected
<specimenCount id="9D21F7D76176A849FF1CFA67F735FA63" box="[248,290,1488,1514]" pageId="10" pageNumber="131" type="female">♀♀</specimenCount>
<typeStatus id="549C82FC6176A849FECDFA67F783FA63" box="[297,404,1488,1514]" pageId="10" pageNumber="131" type="paratype">paratypes</typeStatus>
<quantity id="4CDF91BB6176A849FE78FA78F434FA63" box="[412,547,1487,1514]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="10" pageNumber="131" unit="mm" value="5.05" valueMax="5.2" valueMin="4.9">4.95.2 mm</quantity>
long.
</paragraph>
<paragraph id="8B983C5E6176A84CFF23FA44F7A1FC61" blockId="10.[151,1437,1379,2018]" lastBlockId="15.[151,1437,151,1539]" lastPageId="15" lastPageNumber="136" pageId="10" pageNumber="131">
<emphasis id="B953E04C6176A849FF23FA44F746F984" bold="true" box="[199,337,1523,1549]" pageId="10" pageNumber="131">Description</emphasis>
,
<specimenCount id="9D21F7D76176A849FEBBFA43F760F987" box="[351,375,1524,1550]" pageId="10" pageNumber="131" type="male"></specimenCount>
: Dorsal forebody black, pronotal basal area narrowly paler, clypeus and labrum dirty yellow to yellowish brown. Elytra ochre with black, strongly elongate sutural spot in apical third leaving suture and apex of elytra narrowly yellow. Antennae black. Maxillary palps pale brown. Legs black-brown to brown, tarsi yellowish brown, trochanters yellowish brown. Venter generally brown, ventral and posterolateral parts of prothoracic hypomeron yellow to yellowish brown. Head (
<figureCitation id="131C20DB6176A849FD4BF934F526F917" box="[687,817,1667,1694]" captionStart="FIGURES 1517" captionStartId="9.[152,270,1963,1987]" captionTargetBox="[188,1399,199,1938]" captionTargetId="figure-15@9.[173,1414,181,1939]" captionTargetPageId="9" captionText="FIGURES 1517. Physciolagria smithi sp. nov., holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413360" httpUri="https://zenodo.org/record/7413360/files/figure.png" pageId="10" pageNumber="131">Figs 1517</figureCitation>
) transverse, glossy dorsally, flattened, with extremely large and globose hemispherical compound eyes which occupy most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, rather long. Frontoclypeal suture or impression not indicated. Labrum broadly emarginate at anterior margin.Frons extremely narrow between compound eyes (at its narrowest point frons is narrower than length of basal antennomere), widens posteriad.Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures large, dense and rather coarse, irregular. Intervening spaces glossy on frons. Occiput more regularly punctate, punctures circular, intervening spaces glossy. Head dorsal setation yellowish, inconspicuous, suberect.Antenna (
<figureCitation id="131C20DB6176A849FC9FF814F5EFF834" box="[891,1016,1955,1982]" captionStart="FIGURES 1517" captionStartId="9.[152,270,1963,1987]" captionTargetBox="[188,1399,199,1938]" captionTargetId="figure-15@9.[173,1414,181,1939]" captionTargetPageId="9" captionText="FIGURES 1517. Physciolagria smithi sp. nov., holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413360" httpUri="https://zenodo.org/record/7413360/files/figure.png" pageId="10" pageNumber="131">Figs 15, 17</figureCitation>
) extending slightly beyond mesocoxa when directed backwards. Basal antennomere short, about 1.21.3× as long as wide. Antennomere three about 1.51.7× as long as antennomere two. Antennomeres 58 distinctly asymmetrical, 410 strongly shortened, of them 510 strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, rounded apically, at anterior margin slightly narrowed apicad from about midlength where there is an obtuse angulation, about 1.41.5× as long as combined length of antennomeres 110. Terminal maxillary palpomere slightly fusiform, narrow. Pronotum (
<figureCitation id="131C20DB6172A84DFAF4F87AF384F861" box="[1296,1427,1997,2024]" captionStart="FIGURES 1517" captionStartId="9.[152,270,1963,1987]" captionTargetBox="[188,1399,199,1938]" captionTargetId="figure-15@9.[173,1414,181,1939]" captionTargetPageId="9" captionText="FIGURES 1517. Physciolagria smithi sp. nov., holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413360" httpUri="https://zenodo.org/record/7413360/files/figure.png" pageId="14" pageNumber="135">Figs 1517</figureCitation>
) glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate to broadly rounded at anterior margin. Lateral margins subparallel. Anterolateral angles obsolete, posterolateral angles broadly obtuse to rounded. Pronotal base broadly rounded, with narrow bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, as wide as to 3× as wide as punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, erect to suberect. Scutellar shield triangular, rounded apically. Elytra (
<figureCitation id="131C20DB6173A84CFAF7FE90F383FEC8" box="[1299,1428,294,321]" captionStart="FIGURES 1517" captionStartId="9.[152,270,1963,1987]" captionTargetBox="[188,1399,199,1938]" captionTargetId="figure-15@9.[173,1414,181,1939]" captionTargetPageId="9" captionText="FIGURES 1517. Physciolagria smithi sp. nov., holotype ♂. 15—Habitus, dorsal view; 16—ditto, lateral view; 17—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413360" httpUri="https://zenodo.org/record/7413360/files/figure.png" pageId="15" pageNumber="136">Figs 1516</figureCitation>
) elongate, nearly parallel-sided, moderately glossy, flattened dorsally, somewhat malacoderm-like thin and flimsy. Humerus broadly rounded. Punctures on elytral disc coarser and denser than those on pronotum, with irregular, glossy intervening spaces which are generally narrower than punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, suberect, moderately long, directed posteriad. Sutural striae not indicated. Epipleuron short, moderately broad, present from humerus towards about posterior margin of mesocoxal cavity. Metathoracic wings fully developed.All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Pro- and mesothoracic tibia about as long as corresponding femur, metathoracic tibia distinctly longer than metathoracic femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII truncate at posterior margin (
<figureCitation id="131C20DB6173A84CFF1DFD07F758FD43" box="[249,335,688,714]" captionStart="FIGURES 2027" captionStartId="11.[152,270,1776,1800]" captionTargetBox="[159,1428,199,1738]" captionTargetId="figure-15@11.[151,1436,181,1752]" captionTargetPageId="11" captionText="FIGURES 2027. Male terminalia of Physciolagria Pic, 1930. 20—P. liturata Pic, 1930, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—P. smithi sp. nov., paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, spiculum gastrale; 27—P. smithi sp. nov., paratype, spiculum gastrale [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="15" pageNumber="136">Fig. 24</figureCitation>
), morphological sternite VII short and broad, subtruncate at posterior margin (
<figureCitation id="131C20DB6173A84CFB04FD07F32FFD42" box="[1248,1336,688,715]" captionStart="FIGURES 2027" captionStartId="11.[152,270,1776,1800]" captionTargetBox="[159,1428,199,1738]" captionTargetId="figure-15@11.[151,1436,181,1752]" captionTargetPageId="11" captionText="FIGURES 2027. Male terminalia of Physciolagria Pic, 1930. 20—P. liturata Pic, 1930, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—P. smithi sp. nov., paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, spiculum gastrale; 27—P. smithi sp. nov., paratype, spiculum gastrale [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="15" pageNumber="136">Fig. 25</figureCitation>
). Tergite VIII short and broad, subtruncate at posterior margin, morphological sternite VIII of two sclerites interconnected by narrow, transparent membrane, subtruncate at posterior margin, short and broad. Spiculum gastrale as in
<figureCitation id="131C20DB6173A84CFA94FD4FF6A1FCBF" captionStart="FIGURES 2027" captionStartId="11.[152,270,1776,1800]" captionTargetBox="[159,1428,199,1738]" captionTargetId="figure-15@11.[151,1436,181,1752]" captionTargetPageId="11" captionText="FIGURES 2027. Male terminalia of Physciolagria Pic, 1930. 20—P. liturata Pic, 1930, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—P. smithi sp. nov., paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, spiculum gastrale; 27—P. smithi sp. nov., paratype, spiculum gastrale [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="15" pageNumber="136">Fig. 27</figureCitation>
. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally, gradually narrowed in apical half, broadly rounded apically, provided with irregular spines on its narrow part (
<figureCitation id="131C20DB6173A84CFAE8FC88F386FCD0" box="[1292,1425,831,857]" captionStart="FIGURES 3644" captionStartId="13.[152,270,1906,1930]" captionTargetBox="[187,1403,196,1860]" captionTargetId="figure-15@13.[166,1422,181,1883]" captionTargetPageId="13" captionText="FIGURES 3644. Median lobe of Physciolagria Pic, 1930. 36—P. liturata Pic, 1930; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—P. smithi sp. nov., paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="15" pageNumber="136">Figs 4244</figureCitation>
). Gonopore armature of numerous long, parallel baculi and one thicker, median rod (
<figureCitation id="131C20DB6173A84CFBB1FCD4F2BBFCF4" box="[1109,1196,867,893]" captionStart="FIGURES 3644" captionStartId="13.[152,270,1906,1930]" captionTargetBox="[187,1403,196,1860]" captionTargetId="figure-15@13.[166,1422,181,1883]" captionTargetPageId="13" captionText="FIGURES 3644. Median lobe of Physciolagria Pic, 1930. 36—P. liturata Pic, 1930; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—P. smithi sp. nov., paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="15" pageNumber="136">Fig. 44</figureCitation>
). Tegmen stout, with paramere externally curved in apical third only; paramere apex hook-like, rounded (
<figureCitation id="131C20DB6173A84CFBD0FC30F2A1FC28" box="[1076,1206,902,929]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="15" pageNumber="136">Figs 3335</figureCitation>
). Ventral surface and external margin of paramere with several long setae most dense on apical area (
<figureCitation id="131C20DB6173A84CFBF5FC1DF273FC4C" box="[1041,1124,938,965]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="15" pageNumber="136">Fig. 35</figureCitation>
). Outer and inner margin of paramere broad (
<figureCitation id="131C20DB6173A84CFEB1FC79F7BEFC61" box="[341,425,974,1000]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="15" pageNumber="136">Fig. 33</figureCitation>
).
</paragraph>
<caption id="DF586CD66177A848FF7CF947F53BF811" ID-DOI="http://doi.org/10.5281/zenodo.7413364" ID-Zenodo-Dep="7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="11" pageNumber="132" startId="11.[152,270,1776,1800]" targetBox="[159,1428,199,1738]" targetPageId="11">
<paragraph id="8B983C5E6177A848FF7CF947F53BF811" blockId="11.[151,1436,1776,1945]" pageId="11" pageNumber="132">
<emphasis id="B953E04C6177A848FF7CF947F74CF881" bold="true" box="[152,347,1776,1800]" pageId="11" pageNumber="132">FIGURES 2027.</emphasis>
Male terminalia of
<taxonomicName id="4C2747DD6177A848FDC3F947F536F880" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[551,801,1776,1801]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C6177A848FDC3F947F4A2F881" box="[551,693,1776,1800]" italics="true" pageId="11" pageNumber="132">Physciolagria</emphasis>
<bibRefCitation id="EFB641AF6177A848FD59F947F536F880" author="Pic, M." box="[701,801,1776,1801]" pageId="11" pageNumber="132" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
. 20—
<taxonomicName id="4C2747DD6177A848FCBAF946F23AF880" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[862,1069,1776,1801]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="132" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C6177A848FCBAF946F5D4F881" box="[862,963,1776,1800]" italics="true" pageId="11" pageNumber="132">P. liturata</emphasis>
<bibRefCitation id="EFB641AF6177A848FC2EF947F23AF880" author="Pic, M." box="[970,1069,1776,1801]" pageId="11" pageNumber="132" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—
<taxonomicName id="4C2747DD6177A848FD9BF8A2F5A1F8A5" authority="(Pic, 1935)" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1935" box="[639,950,1812,1836]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="132" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C6177A848FD9BF8A2F520F8A5" box="[639,823,1812,1836]" italics="true" pageId="11" pageNumber="132">P. singularicornis</emphasis>
(
<bibRefCitation id="EFB641AF6177A848FCA3F8A3F5B8F8A5" author="Pic, M." box="[839,943,1812,1836]" pageId="11" pageNumber="132" pagination="1 - 36" refId="ref8308" refString="Pic, M. (1935) Nouveautes diverses. Melanges exotico-entomologiques 65, 1 - 36." type="journal article" year="1935">Pic, 1935</bibRefCitation>
)
</taxonomicName>
, ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—
<taxonomicName id="4C2747DD6177A848FCF9F88EF564F8D9" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[797,883,1848,1872]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="132" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C6177A848FCF9F88EF564F8D9" box="[797,883,1848,1872]" italics="true" pageId="11" pageNumber="132">P. smithi</emphasis>
</taxonomicName>
<emphasis id="B953E04C6177A848FC9EF88EF5DDF8D8" bold="true" box="[890,970,1849,1873]" pageId="11" pageNumber="132">
<taxonomicNameLabel id="A2605D376177A848FC9EF88EF5DDF8D8" box="[890,970,1849,1873]" pageId="11" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—
<taxonomicName id="4C2747DD6177A848FD9BF8EAF5A2F8FD" authority="(Pic, 1935)" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1935" box="[639,949,1884,1908]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="132" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C6177A848FD9BF8EAF521F8FD" box="[639,822,1884,1908]" italics="true" pageId="11" pageNumber="132">P. singularicornis</emphasis>
(
<bibRefCitation id="EFB641AF6177A848FCA2F8EBF5BAF8FD" author="Pic, M." box="[838,941,1884,1908]" pageId="11" pageNumber="132" pagination="1 - 36" refId="ref8308" refString="Pic, M. (1935) Nouveautes diverses. Melanges exotico-entomologiques 65, 1 - 36." type="journal article" year="1935">Pic, 1935</bibRefCitation>
)
</taxonomicName>
, ♂ from Sebele, Botswana, spiculum gastrale; 27—
<taxonomicName id="4C2747DD6177A848FF2EF836F736F811" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[202,289,1920,1944]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="132" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C6177A848FF2EF836F736F811" box="[202,289,1920,1944]" italics="true" pageId="11" pageNumber="132">P. smithi</emphasis>
</taxonomicName>
<emphasis id="B953E04C6177A848FEC3F836F760F810" bold="true" box="[295,375,1921,1945]" pageId="11" pageNumber="132">
<taxonomicNameLabel id="A2605D376177A848FEC3F836F760F810" box="[295,375,1921,1945]" pageId="11" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, spiculum gastrale [not to scale].
</paragraph>
</caption>
<caption id="DF586CD66170A84FFF7CF888F797F86E" ID-DOI="http://doi.org/10.5281/zenodo.7413366" ID-Zenodo-Dep="7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="12" pageNumber="133" startId="12.[152,270,1855,1879]" targetBox="[170,1426,197,1817]" targetPageId="12">
<paragraph id="8B983C5E6170A84FFF7CF888F797F86E" blockId="12.[151,1436,1855,2023]" pageId="12" pageNumber="133">
<emphasis id="B953E04C6170A84FFF7CF888F74BF8DE" bold="true" box="[152,348,1855,1879]" pageId="12" pageNumber="133">FIGURES 2835.</emphasis>
Tegmen of
<taxonomicName id="4C2747DD6170A84FFE3CF888F4C4F8DE" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[472,723,1855,1879]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C6170A84FFE3CF888F471F8DE" box="[472,614,1855,1879]" italics="true" pageId="12" pageNumber="133">Physciolagria</emphasis>
<bibRefCitation id="EFB641AF6170A84FFD8AF888F4C4F8DE" author="Pic, M." box="[622,723,1855,1879]" pageId="12" pageNumber="133" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
. 28—
<taxonomicName id="4C2747DD6170A84FFCF4F8F7F5F5F8DE" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[784,994,1855,1879]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="133" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C6170A84FFCF4F8F7F561F8DE" box="[784,886,1855,1879]" italics="true" pageId="12" pageNumber="133">P. liturata</emphasis>
<bibRefCitation id="EFB641AF6170A84FFC9AF888F5F5F8DE" author="Pic, M." box="[894,994,1855,1879]" pageId="12" pageNumber="133" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—
<taxonomicName id="4C2747DD6170A84FFDC2F8D3F54DF8F2" authority="(Pic, 1935)" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1935" box="[550,858,1891,1915]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="133" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C6170A84FFDC2F8D3F4CBF8F2" box="[550,732,1891,1915]" italics="true" pageId="12" pageNumber="133">P. singularicornis</emphasis>
(
<bibRefCitation id="EFB641AF6170A84FFD08F8D4F544F8F2" author="Pic, M." box="[748,851,1891,1915]" pageId="12" pageNumber="133" pagination="1 - 36" refId="ref8308" refString="Pic, M. (1935) Nouveautes diverses. Melanges exotico-entomologiques 65, 1 - 36." type="journal article" year="1935">Pic, 1935</bibRefCitation>
)
</taxonomicName>
, ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—
<taxonomicName id="4C2747DD6170A84FFB3FF83FF321F816" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[1243,1334,1927,1951]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="133" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C6170A84FFB3FF83FF321F816" box="[1243,1334,1927,1951]" italics="true" pageId="12" pageNumber="133">P. smithi</emphasis>
</taxonomicName>
<emphasis id="B953E04C6170A84FFAA5F830F382F816" bold="true" box="[1345,1429,1927,1951]" pageId="12" pageNumber="133">
<taxonomicNameLabel id="A2605D376170A84FFAA5F830F382F816" box="[1345,1429,1927,1951]" pageId="12" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale].
</paragraph>
</caption>
<caption id="DF586CD66171A84EFF7CF8C5F466F87E" ID-DOI="http://doi.org/10.5281/zenodo.7413368" ID-Zenodo-Dep="7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="13" pageNumber="134" startId="13.[152,270,1906,1930]" targetBox="[187,1403,196,1860]" targetPageId="13">
<paragraph id="8B983C5E6171A84EFF7CF8C5F466F87E" blockId="13.[151,1436,1906,2039]" pageId="13" pageNumber="134">
<emphasis id="B953E04C6171A84EFF7CF8C5F74BF803" bold="true" box="[152,348,1906,1930]" pageId="13" pageNumber="134">FIGURES 3644.</emphasis>
Median lobe of
<taxonomicName id="4C2747DD6171A84EFDEDF8C4F514F802" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[521,771,1907,1931]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C6171A84EFDEDF8C4F480F802" box="[521,663,1907,1931]" italics="true" pageId="13" pageNumber="134">Physciolagria</emphasis>
<bibRefCitation id="EFB641AF6171A84EFD7BF8C4F514F802" author="Pic, M." box="[671,771,1907,1931]" pageId="13" pageNumber="134" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
. 36—
<taxonomicName id="4C2747DD6171A84EFCA5F8C4F203F802" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[833,1044,1907,1931]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="134" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C6171A84EFCA5F8C4F5B0F802" box="[833,935,1907,1931]" italics="true" pageId="13" pageNumber="134">P. liturata</emphasis>
<bibRefCitation id="EFB641AF6171A84EFC4BF8C4F203F802" author="Pic, M." box="[943,1044,1907,1931]" pageId="13" pageNumber="134" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—
<taxonomicName id="4C2747DD6171A84EFDB0F820F594F827" authority="(Pic, 1935)" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1935" box="[596,899,1942,1967]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="134" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C6171A84EFDB0F820F51FF826" box="[596,776,1943,1967]" italics="true" pageId="13" pageNumber="134">P. singularicornis</emphasis>
(
<bibRefCitation id="EFB641AF6171A84EFCF2F820F56CF827" author="Pic, M." box="[790,891,1942,1967]" pageId="13" pageNumber="134" pagination="1 - 36" refId="ref8308" refString="Pic, M. (1935) Nouveautes diverses. Melanges exotico-entomologiques 65, 1 - 36." type="journal article" year="1935">Pic, 1935</bibRefCitation>
)
</taxonomicName>
, ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—
<taxonomicName id="4C2747DD6171A84EFD0EF80CF553F85A" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[746,836,1979,2003]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="134" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="B953E04C6171A84EFD0EF80CF553F85A" box="[746,836,1979,2003]" italics="true" pageId="13" pageNumber="134">P. smithi</emphasis>
</taxonomicName>
<emphasis id="B953E04C6171A84EFCA9F80CF5B6F85A" bold="true" box="[845,929,1979,2003]" pageId="13" pageNumber="134">
<taxonomicNameLabel id="A2605D376171A84EFCA9F80CF5B6F85A" box="[845,929,1979,2003]" pageId="13" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale].
</paragraph>
</caption>
<caption id="DF586CD66172A84DFF7CF964F5CAF887" ID-DOI="http://doi.org/10.5281/zenodo.7413370" ID-Zenodo-Dep="7413370" httpUri="https://zenodo.org/record/7413370/files/figure.png" pageId="14" pageNumber="135" startId="14.[152,270,1747,1771]" targetBox="[160,1419,203,1715]" targetPageId="14">
<paragraph id="8B983C5E6172A84DFF7CF964F5CAF887" blockId="14.[151,1436,1747,1807]" pageId="14" pageNumber="135">
<emphasis id="B953E04C6172A84DFF7CF964F74BF962" bold="true" box="[152,348,1747,1771]" pageId="14" pageNumber="135">FIGURES 4548.</emphasis>
<taxonomicName id="4C2747DD6172A84DFE86F964F518F962" authority="(Pic, 1935)" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1935" box="[354,783,1747,1771]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="135" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C6172A84DFE86F964F484F962" box="[354,659,1747,1771]" italics="true" pageId="14" pageNumber="135">Physciolagria singularicornis</emphasis>
(
<bibRefCitation id="EFB641AF6172A84DFD46F964F510F962" author="Pic, M." box="[674,775,1747,1771]" pageId="14" pageNumber="135" pagination="1 - 36" refId="ref8308" refString="Pic, M. (1935) Nouveautes diverses. Melanges exotico-entomologiques 65, 1 - 36." type="journal article" year="1935">Pic, 1935</bibRefCitation>
)
</taxonomicName>
, holotype ♀. 45—Habitus, dorsal view; 46—ditto, lateral view; 47—ditto, ventral view; 48—Labels [images courtesy Christophe Rivier, MNHN].
</paragraph>
</caption>
<paragraph id="8B983C5E6173A84CFF23FC45F2EAFB6B" blockId="15.[151,1437,151,1539]" pageId="15" pageNumber="136">
<emphasis id="B953E04C6173A84CFF23FC45F746FB85" bold="true" box="[199,337,1010,1036]" pageId="15" pageNumber="136">Description</emphasis>
,
<specimenCount id="9D21F7D76173A84CFE82FC45F76BFB85" box="[358,380,1010,1036]" pageId="15" pageNumber="136" type="female"></specimenCount>
. Female (
<figureCitation id="131C20DB6173A84CFE12FC45F46BFB85" box="[502,636,1010,1036]" captionStart="FIGURES 1819" captionStartId="10.[152,270,1273,1297]" captionTargetBox="[152,1436,197,1236]" captionTargetId="figure-33@10.[151,1436,181,1249]" captionTargetPageId="10" captionText="FIGURES 1819. Physciolagria smithi sp. nov., paratype ♀. 18—Habitus, dorsal view; 19—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413362" httpUri="https://zenodo.org/record/7413362/files/figure.png" pageId="15" pageNumber="136">Figs 1819</figureCitation>
) generally longer and slenderer than male (consider Measurements above). Frons comparatively wider between compound eyes. Antenna (
<figureCitation id="131C20DB6173A84CFC55FBA2F226FBB9" box="[945,1073,1045,1072]" captionStart="FIGURES 1819" captionStartId="10.[152,270,1273,1297]" captionTargetBox="[152,1436,197,1236]" captionTargetId="figure-33@10.[151,1436,181,1249]" captionTargetPageId="10" captionText="FIGURES 1819. Physciolagria smithi sp. nov., paratype ♀. 18—Habitus, dorsal view; 19—Forebody, anterodorsal view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413362" httpUri="https://zenodo.org/record/7413362/files/figure.png" pageId="15" pageNumber="136">Figs 1819</figureCitation>
) comparatively shorter than that of male. Antennomere three about as long as antennomere two. Antennomeres 48 distinctly asymmetrical, 510 strongly transverse, 910 distinctly broadly emarginate at posterior margin. Terminal antennomere of similar shape as of male but subtruncate apically, about as long as 44.5 preceding antennomeres. Terminal maxillary palpomere subtriangular, slightly widened apically. Basal metathoracic tarsomere longer than combined length of remaining metathoracic tarsomeres. Tergite VII and morphological sternite VII subtruncate at posterior margin.
</paragraph>
<paragraph id="8B983C5E6173A84CFF23FB5BF30AFAA2" blockId="15.[151,1437,151,1539]" pageId="15" pageNumber="136">
<emphasis id="B953E04C6173A84CFF23FB5BF7F0FA8F" bold="true" box="[199,487,1260,1286]" pageId="15" pageNumber="136">Intraspecific variability.</emphasis>
Colour of pronotum varies from entirely black to yellowish-brown. Colour of venter vary from yellowish brown to dark brown. Pronotal disc shallowly concave dorsally in
<specimenCount id="9D21F7D76173A84CFB8EFAA6F2B9FAA2" box="[1130,1198,1297,1323]" pageId="15" pageNumber="136" type="male">one ♂</specimenCount>
<typeStatus id="549C82FC6173A84CFB51FAA6F30EFAA2" box="[1205,1305,1297,1323]" pageId="15" pageNumber="136" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD56173A84CFF23FA83F476FAFB" pageId="15" pageNumber="136" type="diagnosis">
<paragraph id="8B983C5E6173A84CFF23FA83F476FAFB" blockId="15.[151,1437,151,1539]" pageId="15" pageNumber="136">
<emphasis id="B953E04C6173A84CFF23FA83F7DAFAC7" bold="true" box="[199,461,1332,1358]" pageId="15" pageNumber="136">Differential diagnosis.</emphasis>
The new species is readily distinguished from other congeners primarily in the shape of the male genital organs (see Key below).
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD56173A84CFF23FACAF48EFA56" pageId="15" pageNumber="136" type="biology_ecology">
<paragraph id="8B983C5E6173A84CFF23FACAF48EFA56" blockId="15.[151,1437,151,1539]" pageId="15" pageNumber="136">
<emphasis id="B953E04C6173A84CFF23FACAF73BFA1E" bold="true" box="[199,300,1405,1431]" pageId="15" pageNumber="136">Ecology.</emphasis>
<materialsCitation id="3B4F36036173A84CFED6FACBF482FA56" ID-GBIF-Occurrence="3988003330" collectionCode="LED, MV" collectorName="Grassland" elevation="165" location="Western African Mesic Woodland" pageId="15" pageNumber="136" specimenCount="2">
Savannah with patches or gully forest. All available specimens were attracted to various light sources (actinic,
<collectionCode id="ED36A49B6173A84CFF1FFA16F723FA32" box="[251,308,1441,1467]" pageId="15" pageNumber="136">LED</collectionCode>
,
<collectionCode id="ED36A49B6173A84CFEA5FA16F767FA32" box="[321,368,1441,1467]" country="0" name="University of Montana Museum" pageId="15" pageNumber="136">MV</collectionCode>
) at ~
<quantity id="4CDF91BB6173A84CFE4FFA16F408FA32" box="[427,543,1441,1467]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.0" metricValueMin="0.3" pageId="15" pageNumber="136" unit="m" value="165.0" valueMax="300.0" valueMin="30.0">
<elevation id="000ADB6D6173A84CFE4FFA16F408FA32" box="[427,543,1441,1467]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.0" metricValueMin="0.3" pageId="15" pageNumber="136" unit="m" value="165.0" valueMax="300.0" valueMin="30.0">30300 m</elevation>
</quantity>
elevation. The
<typeStatus id="549C82FC6173A84CFD36FA16F515FA32" box="[722,770,1441,1467]" pageId="15" pageNumber="136">type</typeStatus>
locality is within the
<location id="8EF86A856173A84CFC1AFA16F36AFA33" LSID="urn:lsid:plazi:treatment:038E8D48617BA84CFF73F971F2D9F98B:8EF86A856173A84CFC1AFA16F36AFA33" box="[1022,1405,1440,1467]" name="Western African Mesic Woodland" pageId="15" pageNumber="136">Western African Mesic Woodland</location>
&amp;
<collectorName id="26D259886173A84CFF73FA73F710FA57" box="[151,263,1476,1502]" pageId="15" pageNumber="136">Grassland</collectorName>
vegetation
<typeStatus id="549C82FC6173A84CFE6EFA72F7ADFA56" box="[394,442,1477,1503]" pageId="15" pageNumber="136">type</typeStatus>
(
<bibRefCitation id="EFB641AF6173A84CFE2DFA72F49AFA56" author="Sayre, R. &amp; Comer, P. &amp; Hak, J. &amp; Josse, C. &amp; Bow, J. &amp; Warner, H. &amp; Larwanou, M. &amp; Kelbessa, E. &amp; Bekele, T. &amp; Kehl, H. &amp; Amena, R. &amp; Andriamasimanana, R. &amp; Ba, T. &amp; Benson, L. &amp; Boucher, T. &amp; Brown, M. &amp; Cress, J. &amp; Dassering, O. &amp; Friesen, B. &amp; Gachathi, F. &amp; Houcine, S. &amp; Keita, M. &amp; Khamala, E. &amp; Marangu, D. &amp; Mokua, F. &amp; Morou, B. &amp; Mucina, L. &amp; Mugisha, S. &amp; Mwavu, E. &amp; Rutherford, M. &amp; Sanou, P. &amp; Syampungani, S. &amp; Tomor, B. &amp; Vall, A. &amp; Vande &amp; Weghe, J. &amp; Wangui, E. &amp; Waruingi, L." box="[457,653,1476,1503]" pageId="15" pageNumber="136" pagination="3 - 22" refId="ref8534" refString="Sayre, R., Comer, P., Hak, J., Josse, C., Bow, J., Warner, H., Larwanou, M., Kelbessa, E., Bekele, T., Kehl, H., Amena, R., Andriamasimanana, R., Ba, T., Benson, L., Boucher, T., Brown, M., Cress, J., Dassering, O., Friesen, B., Gachathi, F., Houcine, S., Keita, M., Khamala, E., Marangu, D., Mokua, F., Morou, B., Mucina, L., Mugisha, S., Mwavu, E., Rutherford, M., Sanou, P., Syampungani, S., Tomor, B., Vall, A., Vande, Weghe, J., Wangui, E. &amp; Waruingi, L. (2013) A New Map of Standardized Terrestrial Ecosystems of Africa. Association of American Geographers, Washington, D. C., pp. 3 - 22." type="book chapter" year="2013">
Sayre
<emphasis id="B953E04C6173A84CFDF5FA72F455FA57" box="[529,578,1476,1502]" italics="true" pageId="15" pageNumber="136">et al</emphasis>
. 2013
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD56173A84CFF23FA5FF2D9F98B" box="[199,1230,1512,1539]" pageId="15" pageNumber="136" type="distribution">
<paragraph id="8B983C5E6173A84CFF23FA5FF2D9F98B" blockId="15.[151,1437,151,1539]" box="[199,1230,1512,1539]" pageId="15" pageNumber="136">
<emphasis id="B953E04C6173A84CFF23FA5FF776F98B" bold="true" box="[199,353,1512,1538]" pageId="15" pageNumber="136">Distribution.</emphasis>
Only known from
<typeStatus id="549C82FC6173A84CFDD3FA5EF470F98A" box="[567,615,1513,1539]" pageId="15" pageNumber="136">type</typeStatus>
locality—Niokolo-Koba National Park in SE
<collectingCountry id="F3307CCE6173A84CFB89FA5FF2DEF98B" box="[1133,1225,1512,1538]" name="Senegal" pageId="15" pageNumber="136">Senegal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>