289 lines
41 KiB
XML
289 lines
41 KiB
XML
<document id="D0232AC19BC65FC42B8B0BA689469D81" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard" docDate="2024" docId="038E87DAFFFDE41AFC6EFA8EFE7FF85F" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Timbellus radulfiensis Merle, Pacaud, Ledon & Goret, 2024, n. sp." docType="treatment" docUuid="ADEDF3CF-00EE-4FBA-8D11-3435CD75F8A5" docUuidSource="ZooBank" docVersion="9" lastPageNumber="521" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="520" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="1AE5EA4A9CDD7502113C605E236232B8">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:name id="CEF68E15E27FF0DE9B4C48323588D06F" type="personal">
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<mods:namePart id="60B88E5638367593E5792E08FA056B61">Merle, Didier</mods:namePart>
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<mods:namePart id="99589E8E973EC0D5203C0D8FEBE42501">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="B5F824B2632971E7EC2F806482127EA2">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="DF3150C4412CD21046CCE91C91CA22B2">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:nameIdentifier id="06CAB63007A3768B8A2B7055396B6565" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="8F0CAB17F1B3C3AFE9A5A08201DDE5A5">Goret, Bernard</mods:namePart>
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<mods:affiliation id="EBEFFE2C6E49A5DC0AB20ABF80FF33C2">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:nameIdentifier id="038752E8936E7FC1BAA66EF09626CE71" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="DFF66C833DAA8BA313CFAB7665A19109">Geodiversitas</mods:title>
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<mods:part id="94F7C3FF6777723DD1AB94150E58CB41">
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<mods:date id="582E7DBD11053581509CD502D0AE3583">2024</mods:date>
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<mods:number id="92C7E0B2848EC93D191503BDDCD4A169">2024-10-24</mods:number>
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<mods:number id="CCE898897AF428F41F69DC89880C8550">46</mods:number>
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<mods:url id="F682F54369553AF585683133B0980E7E">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf</mods:url>
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<treatment id="038E87DAFFFDE41AFC6EFA8EFE7FF85F" ID-DOI="http://doi.org/10.5281/zenodo.13989278" ID-GBIF-Taxon="244735062" ID-Zenodo-Dep="13989278" LSID="urn:lsid:zoobank.org:act:ADEDF3CF-00EE-4FBA-8D11-3435CD75F8A5" httpUri="http://treatment.plazi.org/id/038E87DAFFFDE41AFC6EFA8EFE7FF85F" lastPageId="28" lastPageNumber="521" pageId="27" pageNumber="520" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae">
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<subSubSection id="C33D6547FFFDE41DFC6EFA8EFA87FAC2" box="[982,1286,1367,1394]" pageId="27" pageNumber="520" type="nomenclature">
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<taxonomicName id="4C274D4FFFFDE41DFC6EFA8EFB42FAC1" ID-CoL="CN5LZ" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[982,1219,1368,1394]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="radulfiensis" status="sp. nov.">
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<emphasis id="B953EADEFFFDE41DFC6EFA8EFB42FAC1" bold="true" box="[982,1219,1368,1394]" italics="true" pageId="27" pageNumber="520">Timbellus radulfiensis</emphasis>
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<taxonomicNameLabel id="A26057A5FFFDE41DFB73FA81FA87FAC2" box="[1227,1286,1367,1393]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C33D6547FFFDE41DFBAAFAA1FA2EFA7E" pageId="27" pageNumber="520" type="description">
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<paragraph id="8B9836CCFFFDE41DFBAAFAA1FB4BFA22" blockId="27.[982,1286,1367,1425]" box="[1042,1226,1398,1425]" pageId="27" pageNumber="520">
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(
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<figureCitation id="131C2A49FFFDE41DFBA3FAA1FBE8FA22" box="[1051,1129,1398,1425]" captionStart="FIG" captionStartId="28.[132,143,796,813]" captionTargetBox="[132,1455,215,753]" captionTargetId="figure-713@28.[176,842,231,736]" captionTargetPageId="28" captionText="FIG. 15. — Spiral cords and internal denticles of the outer lip in Timbellus radulfiensis n. sp.: A, B, holotype, MNHN.F.A77754 (MNHN. coll.), Rauville-la-Place, Manche, Normandy, France, late Eocene (Priabonian); C, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; D, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: 5 mm. Credits: A-C, P. Loubry (MNHN/CNRS); D, L. Cazes (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985796" httpUri="https://zenodo.org/record/13985796/files/figure.png" pageId="27" pageNumber="520">Figs 15</figureCitation>
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;
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<figureCitation id="131C2A49FFFDE41DFBCFFAA0FB12FA23" box="[1143,1171,1398,1424]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="27" pageNumber="520">26</figureCitation>
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N-P)
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</paragraph>
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urn:lsid:zoobank.org:act:
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. —
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.
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<collectingCountry id="F330765CFFFDE41DFBFBF9D7FB03F9AA" box="[1091,1154,1537,1561]" name="France" pageId="27" pageNumber="520">France</collectingCountry>
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</emphasis>
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•
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,
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<collectingMunicipality id="6BFCACB6FFFDE41DFABBF9D6FAD3F9AB" box="[1283,1362,1536,1560]" pageId="27" pageNumber="520">Manche</collectingMunicipality>
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,
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<location id="8EF86017FFFDE41DFAE2F9D6FCFAF980" LSID="urn:lsid:plazi:treatment:038E87DAFFFDE41AFC6EFA8EFE7FF85F:8EF86017FFFDE41DFAE2F9D6FCFAF980" country="France" municipality="Manche" name="Rauville-la-Place" pageId="27" pageNumber="520" stateProvince="Haute-Normandie">Rauville-la-Place</location>
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;
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<collectorName id="26D2531AFFFDE41DFC3DF9CDFC46F980" box="[901,967,1563,1587]" pageId="27" pageNumber="520">Argiles</collectorName>
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à
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<location id="8EF86017FFFDE41DFC59F9CDFB2AF980" LSID="urn:lsid:plazi:treatment:038E87DAFFFDE41AFC6EFA8EFE7FF85F:8EF86017FFFDE41DFC59F9CDFB2AF980" box="[993,1195,1563,1587]" country="France" name="Corbules Formation" pageId="27" pageNumber="520" stateProvince="Haute-Normandie">Corbules Formation</location>
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; late Eocene (Priabonian);
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<collectionCode id="ED36AE09FFFDE41DFC96F9E0FC01F9FE" box="[814,896,1590,1613]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
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.F.A77754 (
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<collectionCode id="ED36AE09FFFDE41DFC55F9E3FBC1F9FE" box="[1005,1088,1589,1613]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
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coll.), H:
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(
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<figureCitation id="131C2A49FFFDE41DFB47F9E3FAEDF9FE" box="[1279,1388,1589,1614]" captionStart="FIG" captionStartId="28.[132,143,796,813]" captionTargetBox="[132,1455,215,753]" captionTargetId="figure-713@28.[176,842,231,736]" captionTargetPageId="28" captionText="FIG. 15. — Spiral cords and internal denticles of the outer lip in Timbellus radulfiensis n. sp.: A, B, holotype, MNHN.F.A77754 (MNHN. coll.), Rauville-la-Place, Manche, Normandy, France, late Eocene (Priabonian); C, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; D, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: 5 mm. Credits: A-C, P. Loubry (MNHN/CNRS); D, L. Cazes (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985796" httpUri="https://zenodo.org/record/13985796/files/figure.png" pageId="27" pageNumber="520">Figs 15A, B</figureCitation>
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;
|
||
<figureCitation id="131C2A49FFFDE41DFACBF9E3FA25F9FD" box="[1395,1444,1589,1614]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="27" pageNumber="520">26N</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4F3C91FFFDE41DFC95F987FBE3F937" ID-GBIF-Occurrence="4958671375" collectionCode="MNHN" country="France" location="France" pageId="27" pageNumber="520" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B953EADEFFFDE41DFC95F987FC4FF9DA" bold="true" box="[813,974,1617,1641]" pageId="27" pageNumber="520">
|
||
<typeStatus id="549C886EFFFDE41DFC95F987FC08F9DA" box="[813,905,1617,1641]" pageId="27" pageNumber="520" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<collectingCountry id="F330765CFFFDE41DFC28F987FC4FF9DA" box="[912,974,1617,1641]" name="France" pageId="27" pageNumber="520">France</collectingCountry>
|
||
</emphasis>
|
||
• 1 spm; same as for the holotype;
|
||
<collectionCode id="ED36AE09FFFDE41DFB45F987FACFF9DB" box="[1277,1358,1617,1640]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A77755 (
|
||
<collectionCode id="ED36AE09FFFDE41DFC8CF9BAFC09F937" box="[820,904,1644,1668]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
coll.) (
|
||
<figureCitation id="131C2A49FFFDE41DFC73F9BDFBA3F937" box="[971,1058,1643,1668]" captionStart="FIG" captionStartId="28.[132,143,796,813]" captionTargetBox="[132,1455,215,753]" captionTargetId="figure-713@28.[176,842,231,736]" captionTargetPageId="28" captionText="FIG. 15. — Spiral cords and internal denticles of the outer lip in Timbellus radulfiensis n. sp.: A, B, holotype, MNHN.F.A77754 (MNHN. coll.), Rauville-la-Place, Manche, Normandy, France, late Eocene (Priabonian); C, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; D, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: 5 mm. Credits: A-C, P. Loubry (MNHN/CNRS); D, L. Cazes (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985796" httpUri="https://zenodo.org/record/13985796/files/figure.png" pageId="27" pageNumber="520">Figs 15C</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFFDE41DFB94F9BDFBD8F937" box="[1068,1113,1643,1668]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="27" pageNumber="520">26O</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4F3C91FFFDE41DFBCDF9BAFC3EF92D" ID-GBIF-Occurrence="4958671353" collectionCode="MNHN" country="France" location="France" pageId="27" pageNumber="520" specimenCount="1" typeStatus="paratype">
|
||
3 spm; idem;
|
||
<collectionCode id="ED36AE09FFFDE41DFB4DF9BAFACBF930" box="[1269,1354,1644,1667]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A77756 (
|
||
<collectionCode id="ED36AE09FFFDE41DFC8CF950FC09F92D" box="[820,904,1670,1694]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
coll.)
|
||
</materialsCitation>
|
||
• 3 spm;
|
||
<materialsCitation id="3B4F3C91FFFDE41DFBA3F950FA26F90A" ID-GBIF-Occurrence="4958671323" collectionCode="MNHN" country="France" location="France" pageId="27" pageNumber="520" specimenCount="1" typeStatus="paratype">
|
||
idem;
|
||
<collectionCode id="ED36AE09FFFDE41DFBE0F950FB2CF92E" box="[1112,1197,1670,1693]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A24118 (
|
||
<collectorName id="26D2531AFFFDE41DFAA5F950FAE9F92D" box="[1309,1384,1670,1694]" pageId="27" pageNumber="520">Vasseur</collectorName>
|
||
coll.) • 1 spm; idem;
|
||
<collectionCode id="ED36AE09FFFDE41DFC0BF977FB89F90B" box="[947,1032,1697,1720]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A90559 (
|
||
<collectorName id="26D2531AFFFDE41DFBC1F976FB45F90B" box="[1145,1220,1696,1720]" pageId="27" pageNumber="520">Vasseur</collectorName>
|
||
coll.) (
|
||
<figureCitation id="131C2A49FFFDE41DFAB2F976FAE6F90B" box="[1290,1383,1696,1720]" captionStart="FIG" captionStartId="28.[132,143,796,813]" captionTargetBox="[132,1455,215,753]" captionTargetId="figure-713@28.[176,842,231,736]" captionTargetPageId="28" captionText="FIG. 15. — Spiral cords and internal denticles of the outer lip in Timbellus radulfiensis n. sp.: A, B, holotype, MNHN.F.A77754 (MNHN. coll.), Rauville-la-Place, Manche, Normandy, France, late Eocene (Priabonian); C, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; D, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: 5 mm. Credits: A-C, P. Loubry (MNHN/CNRS); D, L. Cazes (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985796" httpUri="https://zenodo.org/record/13985796/files/figure.png" pageId="27" pageNumber="520">Figs 15D</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFFDE41DFAC9F976FA1AF90A" box="[1393,1435,1696,1721]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="27" pageNumber="520">26P</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFC95F905FC62F8B6" pageId="27" pageNumber="520" type="etymology">
|
||
<paragraph id="8B9836CCFFFDE41DFC95F905FC62F8B6" blockId="27.[813,1451,1747,1797]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC95F905FC27F95A" baselines="1764,1765" box="[813,934,1747,1770]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="27" pageNumber="520">ETYMOLOGY</smallCapsWord>
|
||
. — From the ancient name of the
|
||
<typeStatus id="549C886EFFFDE41DFB48F905FA9BF958" box="[1264,1306,1747,1771]" pageId="27" pageNumber="520">type</typeStatus>
|
||
locality
|
||
<emphasis id="B953EADEFFFDE41DFAD0F905FCD6F8B6" italics="true" pageId="27" pageNumber="520">Radulfi villa</emphasis>
|
||
around 1083.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFC95F8F6FBB1F80A" pageId="27" pageNumber="520" type="materials_examined">
|
||
<paragraph id="8B9836CCFFFDE41DFC95F8F6FC51F8DF" blockId="27.[813,1455,1823,1900]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC95F8F6FCE1F886" baselines="1841,1841" box="[813,864,1824,1847]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="520">
|
||
<typeStatus id="549C886EFFFDE41DFC95F8F6FCE1F886" box="[813,864,1824,1847]" pageId="27" pageNumber="520">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFFDE41DFCD0F8F3FC46F885" baselines="1842" box="[872,967,1829,1846]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="27" pageNumber="520">HORIZON</smallCapsWord>
|
||
. — Argiles à Corbules Formation, late Eocene (Priabonian), stratigraphic data given byCavelier (1979) and
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFAC5F8ECFC4BF8DF" author="CHATEAUNEUF J. - J." pageId="27" pageNumber="520" pagination="1 - 350" refId="ref43345" refString="CHATEAUNEUF J. - J. 1980. - Palynostratigraphie et paleoclimatologie de l'Eocene superieur et de l'Oligocene du bassin de Paris. Memoires du BRGM 116: 1 - 350." type="journal article" year="1980">Chateauneuf (1980)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFDE41DFC95F851FBB1F80A" blockId="27.[813,1455,1927,1977]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC95F851FCDCF82F" baselines="1944,1944" box="[813,861,1927,1950]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="520">
|
||
<typeStatus id="549C886EFFFDE41DFC95F851FCDCF82F" box="[813,861,1927,1950]" pageId="27" pageNumber="520">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFFDE41DFCDCF85AFC3EF82E" baselines="1945" box="[868,959,1932,1949]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="520">LOCALITY</smallCapsWord>
|
||
. —
|
||
<collectingCountry id="F330765CFFFDE41DFC54F851FBB1F82D" box="[1004,1072,1927,1950]" name="France" pageId="27" pageNumber="520">France</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E3F82EFFFDE41DFB82F851FB27F82C" box="[1082,1190,1927,1951]" country="France" name="Haute-Normandie" pageId="27" pageNumber="520">Normandy</collectingRegion>
|
||
, Manche, Rauville-la-Place (Cauvin),
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFC29F877FBABF80A" author="CAVELIER C." box="[913,1066,1953,1977]" pageId="27" pageNumber="520" pagination="1 - 280" refId="ref43197" refString="CAVELIER C. 1979. - La limite eocene-oligocene en Europe occidentale. Sciences Geologiques (Institut de Geologie de Strasbourg), Memoire 54: 1 - 280. https: // www. persee. fr / doc / sgeol _ 0302 - 2684 _ 1979 _ mon _ 54 _ 1" type="journal article" year="1979">Cavelier (1979)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFC95F802FAB2F858" box="[813,1331,2003,2028]" pageId="27" pageNumber="520" type="distribution">
|
||
<paragraph id="8B9836CCFFFDE41DFC95F802FAB2F858" blockId="27.[813,1331,2003,2028]" box="[813,1331,2003,2028]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC95F802FC3BF859" baselines="2021,2022" box="[813,954,2004,2027]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="27" pageNumber="520">DISTRIBUTION</smallCapsWord>
|
||
. — Only know from the
|
||
<typeStatus id="549C886EFFFDE41DFB0FF802FB60F85F" box="[1207,1249,2004,2028]" pageId="27" pageNumber="520">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF586644FFFAE41AFF3CFCCAFB9EFCE8" ID-DOI="http://doi.org/10.5281/zenodo.13985796" ID-Zenodo-Dep="13985796" httpUri="https://zenodo.org/record/13985796/files/figure.png" pageId="28" pageNumber="521" startId="28.[132,143,796,813]" targetBox="[132,1455,215,753]" targetPageId="28" targetType="figure">
|
||
<paragraph id="8B9836CCFFFAE41AFF3CFCCAFB9EFCE8" blockId="28.[132,1457,796,859]" pageId="28" pageNumber="521">
|
||
<smallCapsWord id="8D7EA010FFFAE41AFF3CFCCAFF1CFC9E" baselines="809,810" box="[132,157,796,813]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="28" pageNumber="521">FIG</smallCapsWord>
|
||
. 15. — Spiral cords and internal denticles of the outer lip in
|
||
<taxonomicName id="4C274D4FFFFAE41AFD21FCCAFCCDFC9E" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[665,844,796,813]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="radulfiensis" status="sp. nov.">
|
||
<emphasis id="B953EADEFFFAE41AFD21FCCAFCCDFC9E" box="[665,844,796,813]" italics="true" pageId="28" pageNumber="521">Timbellus radulfiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFAE41AFCEAFCCAFC04FC9E" box="[850,901,796,813]" pageId="28" pageNumber="521" rank="species">n. sp.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B953EADEFFFAE41AFC34FCCAFC18FC9E" bold="true" box="[908,921,796,813]" pageId="28" pageNumber="521">A</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFFAE41AFC1CFCCAFC30FC9E" bold="true" box="[932,945,796,813]" pageId="28" pageNumber="521">B</emphasis>
|
||
, holotype, MNHN.F.A77754 (MNHN. coll.), Rauville-la-Place, Manche, Normandy, France, late Eocene (Priabonian);
|
||
<emphasis id="B953EADEFFFAE41AFDEBFCE5FDE0FCF7" bold="true" box="[595,609,819,836]" pageId="28" pageNumber="521">C</emphasis>
|
||
, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place;
|
||
<emphasis id="B953EADEFFFAE41AFBDCFCE5FBF3FCF7" bold="true" box="[1124,1138,819,836]" pageId="28" pageNumber="521">D</emphasis>
|
||
, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: 5 mm. Credits: A-C, P. Loubry (MNHN/CNRS); D, L. Cazes (MNHN/CNRS).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D6547FFFAE41AFF3CFC43FE7FF85F" pageId="28" pageNumber="521" type="description">
|
||
<paragraph id="8B9836CCFFFAE41AFF3CFC43FE94FC1E" blockId="28.[130,777,917,1645]" box="[132,277,917,943]" pageId="28" pageNumber="521">
|
||
<smallCapsWord id="8D7EA010FFFAE41AFF3CFC43FE94FC1E" baselines="936,936" box="[132,277,917,943]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="28" pageNumber="521">DESCRIPTION</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFAE41AFF3CFC63FEE7F9DE" blockId="28.[130,777,917,1645]" pageId="28" pageNumber="521">
|
||
Bulbous protoconch of one whorl.Teleoconch up to
|
||
<quantity id="4CDF9B29FFFAE41AFD1EFC62FC86FC7C" box="[678,775,948,975]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="28" pageNumber="521" unit="mm" value="20.5">20.5 mm</quantity>
|
||
in height, up to
|
||
<quantity id="4CDF9B29FFFAE41AFE84FC02FE15FC5C" box="[316,404,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="28" pageNumber="521" unit="mm" value="7.5">7.5 mm</quantity>
|
||
in width, subfusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 74% of total length. Apical angle 61°. Spiral sculpture with moderately marked primary cords. First whorl: no cord. Second whorl: appearance of P2. Third whorl: appearance of P1 (weaker than P2) and s2. Fourth and fifth whorl: no change. Sixth whorl: P1 to P6, s1; P6 not atrophied; ADP, MP and ABP on siphonal canal. From first whorl to second whorl: ten protovarices. On third whorl: seven protovarices. Fourth whorl: appearance of varices; three varices and one intervarix intercalated. From fifth to sixth whorl: no change. Higher relief of intervarices on P2. Varices sublamellose. Appearance of short P1 spine on third whorl. Abapically spinelets from P1 to P6 and ADP. Ovate aperture up to 34% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, poorly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with developed denticles including ID, D1 to D6. Outer lip not crenulate. Siphonal canal open, straight, up to 47% of aperture length. Pseudoumbilicus narrow.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFAE41AFF3CF944FE9EF918" blockId="28.[130,775,1682,2028]" box="[132,287,1682,1708]" pageId="28" pageNumber="521">
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</paragraph>
|
||
<paragraph id="8B9836CCFFFAE41AFF3CF964FE7FF85F" blockId="28.[130,775,1682,2028]" pageId="28" pageNumber="521">
|
||
This species can be compared to other Priabonian and Rupelian species from Northern Europe (
|
||
<collectingCountry id="F330765CFFFAE41AFDB6F904FDF6F95F" box="[526,631,1746,1772]" name="Germany" pageId="28" pageNumber="521">Germany</collectingCountry>
|
||
) and Eastern Europe (
|
||
<collectingCountry id="F330765CFFFAE41AFF58F924FEBAF8BF" box="[224,315,1778,1804]" name="Ukraine" pageId="28" pageNumber="521">Ukraine</collectingCountry>
|
||
) with which it shares a narrow shape. It differs from
|
||
<taxonomicName id="4C274D4FFFFAE41AFF5FF8C4FDABF89F" authority="(von Koenen, 1890)" baseAuthorityName="von Koenen" baseAuthorityYear="1890" box="[231,554,1810,1836]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="detritus">
|
||
<emphasis id="B953EADEFFFAE41AFF5FF8C4FECAF89F" box="[231,331,1810,1836]" italics="true" pageId="28" pageNumber="521">T detritus</emphasis>
|
||
(von Koenen, 1890)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFFAE41AFD8CF8C4FF45F8FF" authority="(Steuer, 1912)" baseAuthorityName="Steuer" baseAuthorityYear="1912" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="arenarius">
|
||
<emphasis id="B953EADEFFFAE41AFD8CF8C4FD2FF898" box="[564,686,1810,1835]" italics="true" pageId="28" pageNumber="521">T. arenarius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFAE41AFD04F8C4FF3DF8FF" author="STEUER A." pageId="28" pageNumber="521" pagination="1 - 65" refId="ref48658" refString="STEUER A. 1912. - Marine Conchylien aus dem Mainzer Becken, I. Abhandlungen der Grossherzoglich Hessischen Geologischen Landeshanstalt zu Darmstadt 6 (1): 1 - 65." type="journal article" year="1912">Steuer, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFFAE41AFF47F8E4FD9DF8FF" authority="Pacaud (2018)" authorityName="Pacaud" authorityYear="2018" box="[255,540,1841,1868]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="danapris">
|
||
<emphasis id="B953EADEFFFAE41AFF47F8E4FEF7F8F8" box="[255,374,1841,1867]" italics="true" pageId="28" pageNumber="521">T. danapris</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFAE41AFEC5F8E7FD9DF8FF" author="PACAUD J. - M." box="[381,540,1841,1868]" pageId="28" pageNumber="521" pagination="107 - 116" refId="ref48075" refString="PACAUD J. - M. 2018. - Timbellus danapris n. sp., une espece nouvelle de Muricidae (Mollusca, Gastropoda) du Priabonien (Eocene superieur) de Dnipro (Oblast de Dnipropetrovsk, Ukraine). Novapex 19 (4): 107 - 116." type="journal article" year="2018">Pacaud (2018)</bibRefCitation>
|
||
</taxonomicName>
|
||
in having finer spiral cords. The spiral sculpture resembles more that of
|
||
<taxonomicName id="4C274D4FFFFAE41AFD32F884FEBFF838" authority="Pacaud, 2018" authorityName="Pacaud" authorityYear="2018" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="priabonicus">
|
||
<emphasis id="B953EADEFFFAE41AFD32F884FF22F838" italics="true" pageId="28" pageNumber="521">T. priabonicus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFAE41AFF13F8A7FEBFF838" author="PACAUD J. - M." box="[171,318,1905,1931]" pageId="28" pageNumber="521" pagination="107 - 116" refId="ref48075" refString="PACAUD J. - M. 2018. - Timbellus danapris n. sp., une espece nouvelle de Muricidae (Mollusca, Gastropoda) du Priabonien (Eocene superieur) de Dnipro (Oblast de Dnipropetrovsk, Ukraine). Novapex 19 (4): 107 - 116." type="journal article" year="2018">Pacaud, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
(nom. n. for
|
||
<taxonomicName id="4C274D4FFFFAE41AFE6CF8A4FF3EF818" authority="von Koenen, 1889" authorityName="von Koenen" authorityYear="1889" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="trialatus">
|
||
<emphasis id="B953EADEFFFAE41AFE6CF8A4FDF4F838" box="[468,629,1905,1931]" italics="true" pageId="28" pageNumber="521">Murex trialatus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFAE41AFDC4F8A4FF3EF818" author="VON KOENEN A." pageId="28" pageNumber="521" pagination="1 - 280" refId="ref49077" refString="VON KOENEN A. 1889. - Das Norddeutsche Unter-Oligocan und seine Mollusken-Fauna. Abhandlungen zur geologischen Specialkarte von Preussen und den Thuringischen Staaten 10 (1): 1 - 280." type="journal article" year="1889">von Koenen, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B953EADEFFFAE41AFF70F844FF6EF818" box="[200,239,1938,1963]" italics="true" pageId="28" pageNumber="521">non</emphasis>
|
||
G. B. Sowerby (II), 1834), but
|
||
<taxonomicName id="4C274D4FFFFAE41AFD80F844FD48F818" authorityName="Pacaud" authorityYear="2018" box="[568,713,1937,1963]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="priabonicus">
|
||
<emphasis id="B953EADEFFFAE41AFD80F844FD48F818" box="[568,713,1937,1963]" italics="true" pageId="28" pageNumber="521">T. priabonicus</emphasis>
|
||
</taxonomicName>
|
||
bears strongly developed P1 and P2 spines, whereas only P1 spine is developed in
|
||
<taxonomicName id="4C274D4FFFFAE41AFE94F804FE3AF858" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[300,443,2001,2027]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="521" phylum="Mollusca" rank="species" species="radulfiensis" status="n. sp.">
|
||
<emphasis id="B953EADEFFFAE41AFE94F804FE3AF858" box="[300,443,2001,2027]" italics="true" pageId="28" pageNumber="521">T. radulfiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFAE41AFE7BF804FE7FF85F" box="[451,510,2002,2028]" pageId="28" pageNumber="521" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |