499 lines
70 KiB
XML
499 lines
70 KiB
XML
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<mods:titleInfo id="34C8A8AA2AEFB41F0EB91CCA01408C0A">
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<mods:title id="B9C2DB74339110334438DCCCA722AFEC">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="D7C1BDF83EC86DA8C24F8D22EFC2FB03">Merle, Didier</mods:namePart>
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<mods:namePart id="0848A6A0AC5D77D3F1DC03F95EA2ED88">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="189BA46D1760F20DB63B04EC27186982">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="02E154B70AA5575ACBC082713B9AF4EF">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:nameIdentifier id="D683902671B30F22E17CB5745EC30FD5" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="7A9E8AFD3476DE2B4EC570DF3DF39112">Goret, Bernard</mods:namePart>
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<mods:affiliation id="473C3E1B1628F1C2B8F513A07F71EEBB">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:nameIdentifier id="65CA5C3D844628D6547D022158D9911A" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="74D36BE67F2B3DEE41B39AE1426EFBDD">Geodiversitas</mods:title>
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<mods:part id="F94E6768EDF7EE60074348AFE2B004C7">
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<mods:date id="121EFC60EA574FEBFC6820F0AC90A8B9">2024</mods:date>
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<mods:number id="1F690993E9848DDC9404898F8274EF51">2024-10-24</mods:number>
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<mods:number id="59DF6B5B85365DBD03D47BFF66E76F32">46</mods:number>
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<treatment id="038E87DAFFFCE41DFC63FA22FB82FAA3" ID-DOI="http://doi.org/10.5281/zenodo.13991571" ID-GBIF-Taxon="244735058" ID-Zenodo-Dep="13991571" LSID="urn:lsid:zoobank.org:act:00EFA23B-DFE2-4272-AC88-BF1C6253EBA7" httpUri="http://treatment.plazi.org/id/038E87DAFFFCE41DFC63FA22FB82FAA3" lastPageId="27" lastPageNumber="520" pageId="26" pageNumber="519" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae">
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<heading id="D0D081A0FFFCE41CFC63FA22FA80F9BE" box="[987,1281,1523,1550]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="519" reason="2">
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<taxonomicName id="4C274D4FFFFCE41CFC63FA22FB3EF9BE" ID-CoL="CN5MD" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[987,1215,1523,1550]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="519" phylum="Mollusca" rank="species" species="curvispina" status="sp. nov.">
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<emphasis id="B953EADEFFFCE41CFC63FA22FB3EF9BE" bold="true" box="[987,1215,1523,1550]" italics="true" pageId="26" pageNumber="519">Timbellus curvispina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26057A5FFFCE41CFB7EFA25FA80F9BE" box="[1222,1281,1523,1549]" pageId="26" pageNumber="519" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C33D6547FFFCE41CFC45F9C5FA31F9D9" pageId="26" pageNumber="519" type="description">
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<paragraph id="8B9836CCFFFCE41CFC45F9C5FB5FF99E" blockId="26.[987,1281,1523,1581]" box="[1021,1246,1554,1581]" pageId="26" pageNumber="519">
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(
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<figureCitation id="131C2A49FFFCE41CFBBEF9C4FBD5F99E" box="[1030,1108,1554,1581]" captionStart="FIG" captionStartId="26.[132,143,1378,1395]" captionTargetBox="[132,1455,215,1334]" captionTargetId="figure-411@26.[698,1267,213,775]" captionTargetPageId="26" captionText="FIG. 14. — Spiral cords and internal denticles of the outer lip in Timbellus crenulatus tricarinatus (Lamarck, 1803) (A-C) and T. curvispina n. sp. (D-H): A, original figure of Lamarck (1816: pl. 418, fig. 5a), Thiverval-Grignon, Yvelines, Paris Basin, France, middle Eocene (Lutetian); B, C, neotype designated herein, MNHN.F.A90547 (MNHN coll.), Thiverval-Grignon (Falunière); D, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin (Bois-du-But), Eure, Paris Basin, France,middle Eocene (Lutetian); E, F, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, Oise, middle Eocene (Lutetian);G, H, paratype,MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin. Scale bars: B-H, 10 mm. Credits: P. Loubry." figureDoi="http://doi.org/10.5281/zenodo.13985794" httpUri="https://zenodo.org/record/13985794/files/figure.png" pageId="26" pageNumber="519">Figs 14</figureCitation>
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D-H; 26J-L)
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</paragraph>
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<paragraph id="8B9836CCFFFCE41CFC95F98FFA31F9D9" blockId="26.[813,1456,1625,1642]" box="[813,1456,1625,1642]" pageId="26" pageNumber="519">
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<uri id="FFB63ACEFFFCE41CFC95F98FFA31F9D9" box="[813,1456,1625,1642]" pageId="26" pageNumber="519">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C33D6547FFFCE41CFC95F94AFB07F8AA" pageId="26" pageNumber="519" type="reference_group">
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<paragraph id="8B9836CCFFFCE41CFC95F94AFC4EF97D" blockId="26.[813,1455,1692,1742]" pageId="26" pageNumber="519">
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<treatmentCitationGroup id="AB3711E2FFFCE41CFC95F94AFC4EF97D" pageId="26" pageNumber="519">
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<taxonomicName id="4C274D4FFFFCE41CFC95F94AFC5EF900" ID-CoL="44MK4" authorityName="Lamarck" authorityYear="1803" box="[813,991,1692,1715]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="519" phylum="Mollusca" rank="species" species="tricarinatus">
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||
<emphasis id="B953EADEFFFCE41CFC95F94AFC5EF900" box="[813,991,1692,1715]" italics="true" pageId="26" pageNumber="519">Murex tricarinatus</emphasis>
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</taxonomicName>
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||
–
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||
<treatmentCitation id="0A8610DDFFFCE41CFC44F94AFB49F907" author="DESHAYES G. - P." box="[1020,1224,1692,1716]" page="597" pageId="26" pageNumber="519" year="1835">
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<bibRefCitation id="EFB64B3DFFFCE41CFC44F94AFB49F907" author="DESHAYES G. - P." box="[1020,1224,1692,1716]" pageId="26" pageNumber="519" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes 1835: 597</bibRefCitation>
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||
</treatmentCitation>
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||
, pl. 82, figs 9, 10 [
|
||
<emphasis id="B953EADEFFFCE41CFA33F94AFA2EF900" box="[1419,1455,1692,1715]" italics="true" pageId="26" pageNumber="519">non</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFCE41CFC95F960FC40F97D" author="LAMARCK J. - B." box="[813,961,1718,1742]" pageId="26" pageNumber="519" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
|
||
].
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9836CCFFFCE41CFC95F931FB07F8AA" blockId="26.[813,1455,1767,1818]" pageId="26" pageNumber="519">
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<treatmentCitationGroup id="AB3711E2FFFCE41CFC95F931FB07F8AA" pageId="26" pageNumber="519">
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<taxonomicName id="4C274D4FFFFCE41CFC95F931FBE4F94C" ID-CoL="5F882" baseAuthorityName="Lamarck" baseAuthorityYear="1803" box="[813,1125,1767,1791]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="519" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
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<emphasis id="B953EADEFFFCE41CFC95F931FBE4F94C" box="[813,1125,1767,1791]" italics="true" pageId="26" pageNumber="519">Timbellus crenulatus tricarinatus</emphasis>
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||
</taxonomicName>
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||
–
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||
<treatmentCitation id="0A8610DDFFFCE41CFB39F931FAE3F94C" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[1153,1378,1767,1791]" page="440" pageId="26" pageNumber="519" year="2011">
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<bibRefCitation id="EFB64B3DFFFCE41CFB39F931FAE3F94C" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[1153,1378,1767,1791]" pageId="26" pageNumber="519" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
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||
Merle
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||
<emphasis id="B953EADEFFFCE41CFB7AF93EFB70F94C" box="[1218,1265,1767,1791]" italics="true" pageId="26" pageNumber="519">et al.</emphasis>
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||
2011: 440
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 98, figs 6, 7 only [
|
||
<emphasis id="B953EADEFFFCE41CFC01F8D5FC5CF8A9" box="[953,989,1795,1818]" italics="true" pageId="26" pageNumber="519">non</emphasis>
|
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<bibRefCitation id="EFB64B3DFFFCE41CFC5BF8D4FBF9F8A9" author="LAMARCK J. - B." box="[995,1144,1794,1818]" pageId="26" pageNumber="519" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
|
||
].
|
||
</treatmentCitationGroup>
|
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</paragraph>
|
||
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<subSubSection id="C33D6547FFFCE41DFC95F8E2FEB8FE1B" lastPageId="27" lastPageNumber="520" pageId="26" pageNumber="519" type="materials_examined">
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<paragraph id="8B9836CCFFFCE41CFC95F8E2FC63F828" blockId="26.[813,1456,1843,2028]" pageId="26" pageNumber="519">
|
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<materialsCitation id="3B4F3C91FFFCE41CFC95F8E2FC63F828" ID-GBIF-Occurrence="4958671322" collectionCode="MNHN" collectorName="de Parnes" country="France" location="Falun de Parnes" pageId="26" pageNumber="519" specimenCount="1" typeStatus="holotype">
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<smallCapsWord id="8D7EA010FFFCE41CFC95F8E2FCDCF8FA" baselines="1861,1861" box="[813,861,1844,1867]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="26" pageNumber="519">TYPE</smallCapsWord>
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. —
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<emphasis id="B953EADEFFFCE41CFC4DF8E5FB23F8F8" bold="true" box="[1013,1186,1843,1867]" pageId="26" pageNumber="519">
|
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<typeStatus id="549C886EFFFCE41CFC4DF8E5FBD7F8F8" box="[1013,1110,1843,1867]" pageId="26" pageNumber="519" type="holotype">Holotype</typeStatus>
|
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.
|
||
<collectingCountry id="F330765CFFFCE41CFBD8F8E5FB23F8F8" box="[1120,1186,1843,1867]" name="France" pageId="26" pageNumber="519">France</collectingCountry>
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||
</emphasis>
|
||
• Paris Basin, Oise, Parnes (Les Bôves);
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||
<location id="8EF86017FFFCE41CFC0AF89BFBD6F8D6" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFCE41CFC0AF89BFBD6F8D6" box="[946,1111,1869,1893]" country="France" name="Falun de Parnes" pageId="26" pageNumber="519">
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Falun
|
||
<collectorName id="26D2531AFFFCE41CFC4CF89BFBD6F8D6" box="[1012,1111,1869,1893]" pageId="26" pageNumber="519">de Parnes</collectorName>
|
||
</location>
|
||
(biozone NP15); middle Eocene (middle Lutetian);
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||
<collectionCode id="ED36AE09FFFCE41CFC52F8BEFBBFF8CC" box="[1002,1086,1896,1919]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="519">MNHN</collectionCode>
|
||
.F.A90541 (ex B63594), H:
|
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<quantity id="4CDF9B29FFFCE41CFAECF8BEFA2EF8CC" box="[1364,1455,1896,1920]" metricMagnitude="-2" metricUnit="m" metricValue="5.18" pageId="26" pageNumber="519" unit="mm" value="51.8">51.8 mm</quantity>
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(
|
||
<figureCitation id="131C2A49FFFCE41CFC8DF854FC26F829" box="[821,935,1922,1947]" captionStart="FIG" captionStartId="26.[132,143,1378,1395]" captionTargetBox="[132,1455,215,1334]" captionTargetId="figure-411@26.[698,1267,213,775]" captionTargetPageId="26" captionText="FIG. 14. — Spiral cords and internal denticles of the outer lip in Timbellus crenulatus tricarinatus (Lamarck, 1803) (A-C) and T. curvispina n. sp. (D-H): A, original figure of Lamarck (1816: pl. 418, fig. 5a), Thiverval-Grignon, Yvelines, Paris Basin, France, middle Eocene (Lutetian); B, C, neotype designated herein, MNHN.F.A90547 (MNHN coll.), Thiverval-Grignon (Falunière); D, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin (Bois-du-But), Eure, Paris Basin, France,middle Eocene (Lutetian); E, F, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, Oise, middle Eocene (Lutetian);G, H, paratype,MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin. Scale bars: B-H, 10 mm. Credits: P. Loubry." figureDoi="http://doi.org/10.5281/zenodo.13985794" httpUri="https://zenodo.org/record/13985794/files/figure.png" pageId="26" pageNumber="519">Figs 14E, F</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFFCE41CFC09F854FC54F828" box="[945,981,1922,1947]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="26" pageNumber="519">26J</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFCE41DFC95F849FEB8FE1B" blockId="26.[813,1456,1843,2028]" lastBlockId="27.[131,776,215,424]" lastPageId="27" lastPageNumber="520" pageId="26" pageNumber="519">
|
||
<materialsCitation id="3B4F3C91FFFCE41DFC95F849FDF9FF5C" ID-GBIF-Occurrence="4958671317" collectorName="Paris Basin" country="France" lastPageId="27" lastPageNumber="520" location="France" pageId="26" pageNumber="519" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B953EADEFFFCE41CFC95F849FC5CF804" bold="true" box="[813,989,1951,1975]" pageId="26" pageNumber="519">
|
||
<typeStatus id="549C886EFFFCE41CFC95F849FC11F804" box="[813,912,1951,1975]" pageId="26" pageNumber="519" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<collectingCountry id="F330765CFFFCE41CFC23F849FC5CF804" box="[923,989,1951,1975]" name="France" pageId="26" pageNumber="519">France</collectingCountry>
|
||
</emphasis>
|
||
• 3 spm
|
||
<collectorName id="26D2531AFFFCE41CFB85F848FB2DF805" box="[1085,1196,1950,1974]" pageId="26" pageNumber="519">Paris Basin</collectorName>
|
||
;
|
||
<location id="8EF86017FFFCE41CFB00F849FB68F805" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFCE41CFB00F849FB68F805" box="[1208,1257,1951,1974]" country="France" name="Eure" pageId="26" pageNumber="519">Eure</location>
|
||
;
|
||
<location id="8EF86017FFFCE41CFB4CF848FA2EF805" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFCE41CFB4CF848FA2EF805" box="[1268,1455,1950,1974]" country="France" name="Fontenay-en-Vexin" pageId="26" pageNumber="519">Fontenay-en-Vexin</location>
|
||
(Bois du but);
|
||
<location id="8EF86017FFFCE41CFC7CF86FFBE2F863" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFCE41CFC7CF86FFBE2F863" box="[964,1123,1977,2001]" country="France" name="Falun de Parnes" pageId="26" pageNumber="519">Falun de Parnes</location>
|
||
(biozone NP15); middle
|
||
<location id="8EF86017FFFCE41CFAD1F86FFA31F863" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFCE41CFAD1F86FFA31F863" box="[1385,1456,1977,2000]" country="France" name="Eocene" pageId="26" pageNumber="519">Eocene</location>
|
||
(
|
||
<collectorName id="26D2531AFFFCE41CFC8DF805FC0AF858" box="[821,907,2003,2027]" pageId="26" pageNumber="519">Lutetian</collectorName>
|
||
);
|
||
<collectionCode id="ED36AE09FFFCE41CFC2FF802FC6BF858" box="[919,1002,2004,2027]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="519">MNHN</collectionCode>
|
||
.F.A73848 (
|
||
<figureCitation id="131C2A49FFFCE41CFBE1F805FB30F858" box="[1113,1201,2003,2027]" captionStart="FIG" captionStartId="26.[132,143,1378,1395]" captionTargetBox="[132,1455,215,1334]" captionTargetId="figure-411@26.[698,1267,213,775]" captionTargetPageId="26" captionText="FIG. 14. — Spiral cords and internal denticles of the outer lip in Timbellus crenulatus tricarinatus (Lamarck, 1803) (A-C) and T. curvispina n. sp. (D-H): A, original figure of Lamarck (1816: pl. 418, fig. 5a), Thiverval-Grignon, Yvelines, Paris Basin, France, middle Eocene (Lutetian); B, C, neotype designated herein, MNHN.F.A90547 (MNHN coll.), Thiverval-Grignon (Falunière); D, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin (Bois-du-But), Eure, Paris Basin, France,middle Eocene (Lutetian); E, F, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, Oise, middle Eocene (Lutetian);G, H, paratype,MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin. Scale bars: B-H, 10 mm. Credits: P. Loubry." figureDoi="http://doi.org/10.5281/zenodo.13985794" httpUri="https://zenodo.org/record/13985794/files/figure.png" pageId="26" pageNumber="519">Figs 14D</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFFCE41CFB02F805FB65F85F" box="[1210,1252,2003,2028]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="26" pageNumber="519">26L</figureCitation>
|
||
),
|
||
<collectionCode id="ED36AE09FFFCE41CFB4BF802FAC7F858" box="[1267,1350,2004,2027]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="519">MNHN</collectionCode>
|
||
.F.A90542,
|
||
<collectionCode id="ED36AE09FFFDE41DFF3CFF01FF58FF5D" box="[132,217,215,238]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A90543 (
|
||
<collectorName id="26D2531AFFFDE41DFEF2FF01FEFBFF5C" box="[330,378,215,239]" pageId="27" pageNumber="520">Pons</collectorName>
|
||
coll.) (
|
||
<figureCitation id="131C2A49FFFDE41DFE07FF01FDBBFF5D" box="[447,570,215,239]" captionStart="FIG" captionStartId="26.[132,143,1378,1395]" captionTargetBox="[132,1455,215,1334]" captionTargetId="figure-411@26.[698,1267,213,775]" captionTargetPageId="26" captionText="FIG. 14. — Spiral cords and internal denticles of the outer lip in Timbellus crenulatus tricarinatus (Lamarck, 1803) (A-C) and T. curvispina n. sp. (D-H): A, original figure of Lamarck (1816: pl. 418, fig. 5a), Thiverval-Grignon, Yvelines, Paris Basin, France, middle Eocene (Lutetian); B, C, neotype designated herein, MNHN.F.A90547 (MNHN coll.), Thiverval-Grignon (Falunière); D, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin (Bois-du-But), Eure, Paris Basin, France,middle Eocene (Lutetian); E, F, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, Oise, middle Eocene (Lutetian);G, H, paratype,MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin. Scale bars: B-H, 10 mm. Credits: P. Loubry." figureDoi="http://doi.org/10.5281/zenodo.13985794" httpUri="https://zenodo.org/record/13985794/files/figure.png" pageId="27" pageNumber="520">Figs 14G, H</figureCitation>
|
||
,
|
||
<figureCitation id="131C2A49FFFDE41DFDFDFF01FDF1FF5C" box="[581,624,215,239]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="27" pageNumber="520">26K</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4F3C91FFFDE41DFD3FFF01FE83FE97" ID-GBIF-Occurrence="4958671316" collectorName="Lutetian" country="France" location="Eocene" pageId="27" pageNumber="520" specimenCount="1" typeStatus="paratype">
|
||
1 spm; idem;
|
||
<collectionCode id="ED36AE09FFFDE41DFF3CFF24FF59FEBA" box="[132,216,242,265]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A32514 (
|
||
<collectorName id="26D2531AFFFDE41DFEFDFF27FE0AFEBA" box="[325,395,241,265]" pageId="27" pageNumber="520">Pacaud</collectorName>
|
||
coll.), figured
|
||
<emphasis id="B953EADEFFFDE41DFDA8FF24FDA2FEBA" box="[528,547,242,265]" italics="true" pageId="27" pageNumber="520">in</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFD92FF27FC82FEBA" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[554,771,241,265]" pageId="27" pageNumber="520" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFFDE41DFD92FF27FDE3FEBA" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[554,610,241,265]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFFDE41DFDDFFF24FD12FEBA" box="[615,659,241,265]" italics="true" pageId="27" pageNumber="520">et al.</emphasis>
|
||
(2011: 440
|
||
</bibRefCitation>
|
||
, pl. 98, fig. 7)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4F3C91FFFDE41DFEA1FEDAFEB7FEC0" ID-GBIF-Occurrence="4958671301" collectorName="Butte Saint-Leonard" country="France" location="Paris Basin" pageId="27" pageNumber="520" specimenCount="1" typeStatus="paratype">
|
||
1 spm;
|
||
<location id="8EF86017FFFDE41DFED9FEDAFE4CFE97" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFED9FEDAFE4CFE97" box="[353,461,268,292]" country="France" name="Paris Basin" pageId="27" pageNumber="520">Paris Basin</location>
|
||
, Yvelines, Villiers-Saint-Frédéric (
|
||
<collectorName id="26D2531AFFFDE41DFF33FEF0FED7FE8D" box="[139,342,294,318]" pageId="27" pageNumber="520">Butte Saint-Léonard</collectorName>
|
||
);
|
||
<location id="8EF86017FFFDE41DFEDEFEF0FE7EFE8D" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFEDEFEF0FE7EFE8D" box="[358,511,294,318]" country="France" name="Falun de Parnes" pageId="27" pageNumber="520">Falun de Parnes</location>
|
||
(biozone NP15);
|
||
<location id="8EF86017FFFDE41DFD16FEF0FC82FE8D" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFD16FEF0FC82FE8D" box="[686,771,294,318]" country="France" name="Lutetian" pageId="27" pageNumber="520">Lutetian</location>
|
||
;
|
||
<collectionCode id="ED36AE09FFFDE41DFF3CFE97FF58FEEB" box="[132,217,321,344]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A32516 (
|
||
<collectorName id="26D2531AFFFDE41DFEF3FE97FE22FEEA" box="[331,419,321,345]" pageId="27" pageNumber="520">Schtrock</collectorName>
|
||
coll.), figured
|
||
<emphasis id="B953EADEFFFDE41DFD8DFE97FDC8FEEB" box="[565,585,321,344]" italics="true" pageId="27" pageNumber="520">in</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFDF7FE97FF2BFEC0" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." pageId="27" pageNumber="520" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFFDE41DFDF7FE97FD09FEEA" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[591,648,321,345]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFFDE41DFD28FE97FD41FEEA" box="[656,704,321,345]" italics="true" pageId="27" pageNumber="520">et al.</emphasis>
|
||
(2011: 440
|
||
</bibRefCitation>
|
||
pl. 98, fig. 6)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4F3C91FFFDE41DFEFEFE8DFEB4FE1B" ID-GBIF-Occurrence="4958671425" collectionCode="MNHN" collectorName="Yvelines & Parnes" country="France" location="Paris Basin" pageId="27" pageNumber="520" specimenCount="1" typeStatus="paratype">
|
||
1 spm;
|
||
<location id="8EF86017FFFDE41DFE29FE8DFE7EFEC0" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFE29FE8DFE7EFEC0" box="[401,511,347,371]" country="France" name="Paris Basin" pageId="27" pageNumber="520">Paris Basin</location>
|
||
,
|
||
<collectorName id="26D2531AFFFDE41DFDB2FE8DFDDAFEC0" box="[522,603,347,371]" pageId="27" pageNumber="520">Yvelines</collectorName>
|
||
,
|
||
<collectorName id="26D2531AFFFDE41DFDDEFE8AFD29FEC0" box="[614,680,348,371]" pageId="27" pageNumber="520">Parnes</collectorName>
|
||
;
|
||
<location id="8EF86017FFFDE41DFD0AFE8DFF45FE3E" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFD0AFE8DFF45FE3E" country="France" name="Falun de Parnes" pageId="27" pageNumber="520">Falun de Parnes</location>
|
||
(biozone NP15);
|
||
<location id="8EF86017FFFDE41DFEC2FEA0FE4EFE3D" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFEC2FEA0FE4EFE3D" box="[378,463,374,398]" country="France" name="Lutetian" pageId="27" pageNumber="520">Lutetian</location>
|
||
;
|
||
<collectionCode id="ED36AE09FFFDE41DFE62FEA0FDAEFE3E" box="[474,559,374,397]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="520">MNHN</collectionCode>
|
||
.F.A90544 (ex
|
||
<location id="8EF86017FFFDE41DFD78FEA0FF7BFE1B" LSID="urn:lsid:plazi:treatment:038E87DAFFFCE41DFC63FA22FB82FAA3:8EF86017FFFDE41DFD78FEA0FF7BFE1B" country="France" name="Galerie de Zoologie" pageId="27" pageNumber="520">Galerie de Zoologie</location>
|
||
coll.)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFF3CFE16FE93FDBE" pageId="27" pageNumber="520" type="etymology">
|
||
<paragraph id="8B9836CCFFFDE41DFF3CFE16FE93FDBE" blockId="27.[131,775,448,525]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF3CFE16FF7FFE65" baselines="465,466" box="[132,254,448,471]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="27" pageNumber="520">ETYMOLOGY</smallCapsWord>
|
||
. — Name derived from Latin
|
||
<emphasis id="B953EADEFFFDE41DFD9BFE16FDCDFE64" box="[547,588,448,471]" italics="true" pageId="27" pageNumber="520">curv</emphasis>
|
||
- (meaning curved) and
|
||
<emphasis id="B953EADEFFFDE41DFF15FE0DFF5CFE41" box="[173,221,475,498]" italics="true" pageId="27" pageNumber="520">spina</emphasis>
|
||
(meaning spine) and refers to the P1 spine dorsally curved; gender female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFF3CFDF3FEDCFD12" pageId="27" pageNumber="520" type="materials_examined">
|
||
<paragraph id="8B9836CCFFFDE41DFF3CFDF3FD22FDE4" blockId="27.[132,776,548,599]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF3CFDF3FF34FD89" baselines="566,566" box="[132,181,549,572]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="520">TYPE</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF05FDFCFE99FD88" baselines="567" box="[189,280,554,571]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="27" pageNumber="520">HORIZON</smallCapsWord>
|
||
. — Falun de Parnes (biozone NP15), see
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFD06FDF2FE89FDE4" author="GELY J. - P. & LORENZ C." pageId="27" pageNumber="520" pagination="713 - 747" refId="ref44561" refString="GELY J. - P. & LORENZ C. 1991. - Analyse sequentielle de l'Eocene et de l'Oligocene du Bassin parisien (France). Revue de l'institut francais du petrole 46 (6): 713 - 747." type="journal article" year="1991">Gély & Lorenz (1991</bibRefCitation>
|
||
: pl. 1), middle Eocene (middle Lutetian).
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFDE41DFF3CFDB9FEDCFD12" blockId="27.[132,775,622,673]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF3CFDB9FF34FD37" baselines="640,640" box="[132,181,623,646]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="520">TYPE</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF04FDA2FE96FD36" baselines="641" box="[188,279,628,645]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="520">LOCALITY</smallCapsWord>
|
||
. —
|
||
<collectingCountry id="F330765CFFFDE41DFEF4FDB9FE11FD35" box="[332,400,623,646]" name="France" pageId="27" pageNumber="520">France</collectingCountry>
|
||
, Paris Basin, Oise, Parnes (Les Bôves), see
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFF1FFD5FFEDBFD12" author="FRITEL P. H." box="[167,346,649,673]" pageId="27" pageNumber="520" refId="ref44321" refString="FRITEL P. H. 1910. - Guide geologique et paleontologique de la region parisienne dans un rayon de 100 kilometres. Deyrolle, Paris, 356 p." type="book" year="1910">Fritel (1910: 221)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFF3CFD6FFDEDFD58" pageId="27" pageNumber="520" type="distribution">
|
||
<paragraph id="8B9836CCFFFDE41DFF3CFD6FFDEDFD58" blockId="27.[132,775,696,747]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF3CFD6FFE93FD7C" baselines="714,715" box="[132,274,697,720]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="27" pageNumber="520">DISTRIBUTION</smallCapsWord>
|
||
. — Calcaire grossier moyen, biozone NP15, Paris Basin (
|
||
<collectingCountry id="F330765CFFFDE41DFF70FD05FE8FFD58" box="[200,270,723,747]" name="France" pageId="27" pageNumber="520">France</collectingCountry>
|
||
), middle Eocene (middle Lutetian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFFDE41DFF3CFCC0FB82FAA3" pageId="27" pageNumber="520" type="description">
|
||
<paragraph id="8B9836CCFFFDE41DFF3CFCC0FE94FC9D" blockId="27.[130,777,790,1645]" box="[132,277,790,816]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF3CFCC0FE94FC9D" baselines="809,809" box="[132,277,790,816]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="27" pageNumber="520">DESCRIPTION</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFDE41DFF3CFCE3FF53F9DE" blockId="27.[130,777,790,1645]" pageId="27" pageNumber="520">
|
||
Protoconch not preserved. Teleoconch up to
|
||
<quantity id="4CDF9B29FFFDE41DFDC7FCE3FD66FCE3" box="[639,743,821,848]" metricMagnitude="-2" metricUnit="m" metricValue="5.18" pageId="27" pageNumber="520" unit="mm" value="51.8">51.8 mm</quantity>
|
||
in height, up to
|
||
<quantity id="4CDF9B29FFFDE41DFEA1FC83FEE9FCC3" box="[281,360,853,880]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="520" unit="mm" value="30.0">30 mm</quantity>
|
||
in width, subfusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 85% of total length. Apical angle 63°. Spiral sculpture with marked primary cords on varices, but weaker on intervarices. First and second whorls eroded. From third: appearance of weak P1 and P2. Third word: no change. Fourth whorl: appearance of s1. Fifth whorl: occasional appearance of s2. Sixth whorl: P1 to P6, s1, s2, P6 not atrophied, s1 well developed; ADP developed, MP, ABP weak or missing on siphonal canal. Axial sculpture eroded on two early whorls. On second whorl, eight protovarices. Third whorl: appearance of varices; three varices and one intervarix between. From fourth to last whorl, no change. Higher relief of intervarices on P2. Varices sublamellose. Appearance of short straight P1 spine on fourth whorl. On last whorl, axial ornamentation forming very short spinelets, dorsally recurved on P1 to P6. P1 spine more strongly developed than other cord spines. Rounded aperture up to 34% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with very weak denticles including D1 to D6. Small crenulations at edge of outer lip. Siphonal canal narrow, open, bent, slightly dorsally recurved, up to 81% of apertural length. Pseudoumbilicus narrow
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFDE41DFF3CF944FE9EF918" blockId="27.[130,777,1682,2027]" box="[132,287,1682,1708]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFF3CF944FE9EF918" baselines="1701,1702" box="[132,287,1682,1708]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="27" pageNumber="520">COMPARISONS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFDE41DFF3CF964FA2FFD7C" blockId="27.[130,777,1682,2027]" lastBlockId="27.[811,1457,215,720]" pageId="27" pageNumber="520">
|
||
This species can be included in the group of
|
||
<taxonomicName id="4C274D4FFFFDE41DFDC7F965FC88F97F" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[639,777,1714,1740]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="crenulatus">
|
||
<emphasis id="B953EADEFFFDE41DFDC7F965FC88F97F" box="[639,777,1714,1740]" italics="true" pageId="27" pageNumber="520">T. crenulatus</emphasis>
|
||
</taxonomicName>
|
||
defined by
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFF42F904FDB4F95F" author="HARASEWYCH M. G. & JENSEN R. H." box="[250,565,1746,1772]" pageId="27" pageNumber="520" pagination="1 - 16" refId="ref44794" refString="HARASEWYCH M. G. & JENSEN R. H. 1979. - Review of the subgenus Pterynotus (Gastropoda: Muricidae) in the Western Atlantic. Nemouria 22: 1 - 16. https: // www. biodiversitylibrary. org / page / 45974443" type="journal article" year="1979">Harasewych & Jensen (1979)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFDFAF904FD53F95F" author="VOKES E. H." box="[578,722,1746,1772]" pageId="27" pageNumber="520" pagination="1 - 108" refId="ref48996" refString="VOKES E. H. 1992. - Cenozoic Muricidae of the Western Atlantic region, Part IX - Pterynotus, Poirieria, Aspella, Dermomurex, Calotrophon, Acantolabia, and Attiliosa; additions and corrections. Tulane Studies in Geology and Paleontology 25 (1 - 3): 1 - 108." type="journal article" year="1992">Vokes (1992)</bibRefCitation>
|
||
, and updated by
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFF44F924FE66F8BE" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[252,487,1778,1805]" pageId="27" pageNumber="520" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFFDE41DFF44F924FEBBF8BF" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[252,314,1778,1804]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFFDE41DFEF9F925FEF0F8BF" box="[321,369,1778,1804]" italics="true" pageId="27" pageNumber="520">et al.</emphasis>
|
||
(2011: 15)
|
||
</bibRefCitation>
|
||
. In this group the last whorl is generally wider than in species of the
|
||
<taxonomicName id="4C274D4FFFFDE41DFD93F8C4FD38F89F" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[555,697,1810,1836]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFFDE41DFD93F8C4FD38F89F" box="[555,697,1810,1836]" italics="true" pageId="27" pageNumber="520">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
group, the shoulder spine (P1) is more delineated and the internal denticles within the outer lip are weaker. The spiral sculpture corresponds to fine primary cords, as in the
|
||
<taxonomicName id="4C274D4FFFFDE41DFD8FF8A4FD3DF838" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[567,700,1905,1931]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFFDE41DFD8FF8A4FD3DF838" box="[567,700,1905,1931]" italics="true" pageId="27" pageNumber="520">T.tripteroides</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<taxonomicName id="4C274D4FFFFDE41DFF3AF847FED1F818" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[130,336,1937,1963]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="curvispina" status="sp. nov.">
|
||
<emphasis id="B953EADEFFFDE41DFF3AF847FED1F818" box="[130,336,1937,1963]" italics="true" pageId="27" pageNumber="520">Timbellus curvispina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFEEEF844FE0CF81F" box="[342,397,1938,1964]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
was mistaken for the Lutetian species
|
||
<taxonomicName id="4C274D4FFFFDE41DFF3AF864FDBAF878" authority="(Lamarck, 1803)" baseAuthorityName="Lamarck" baseAuthorityYear="1803" box="[130,571,1969,1995]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
|
||
<emphasis id="B953EADEFFFDE41DFF3AF864FE01F878" box="[130,384,1969,1995]" italics="true" pageId="27" pageNumber="520">T. crenulatus tricarinatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFE37F867FDB2F878" author="LAMARCK J. - B." box="[399,563,1969,1995]" pageId="27" pageNumber="520" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but it differs by the construction of the P1 spine. This spine is straight, long and oriented abaperturally in
|
||
<taxonomicName id="4C274D4FFFFDE41DFB8FFF0EFAB9FF42" baseAuthorityName="Lamarck" baseAuthorityYear="1803" box="[1079,1336,215,241]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
|
||
<emphasis id="B953EADEFFFDE41DFB8FFF0EFAB9FF42" box="[1079,1336,215,241]" italics="true" pageId="27" pageNumber="520">T. crenulatus tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is shorter, curved and dorsally oriented in
|
||
<taxonomicName id="4C274D4FFFFDE41DFB51FF2EFAEEFEA2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1257,1391,248,273]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="curvispina" status="n. sp.">
|
||
<emphasis id="B953EADEFFFDE41DFB51FF2EFAEEFEA2" box="[1257,1391,248,273]" italics="true" pageId="27" pageNumber="520">T. curvispina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFACDFF2EFA2FFEA1" box="[1397,1454,248,274]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
(see
|
||
<figureCitation id="131C2A49FFFDE41DFCE2FEC1FC22FE82" box="[858,931,279,305]" captionStart="FIG" captionStartId="26.[132,143,1378,1395]" captionTargetBox="[132,1455,215,1334]" captionTargetId="figure-411@26.[698,1267,213,775]" captionTargetPageId="26" captionText="FIG. 14. — Spiral cords and internal denticles of the outer lip in Timbellus crenulatus tricarinatus (Lamarck, 1803) (A-C) and T. curvispina n. sp. (D-H): A, original figure of Lamarck (1816: pl. 418, fig. 5a), Thiverval-Grignon, Yvelines, Paris Basin, France, middle Eocene (Lutetian); B, C, neotype designated herein, MNHN.F.A90547 (MNHN coll.), Thiverval-Grignon (Falunière); D, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin (Bois-du-But), Eure, Paris Basin, France,middle Eocene (Lutetian); E, F, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, Oise, middle Eocene (Lutetian);G, H, paratype,MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin. Scale bars: B-H, 10 mm. Credits: P. Loubry." figureDoi="http://doi.org/10.5281/zenodo.13985794" httpUri="https://zenodo.org/record/13985794/files/figure.png" pageId="27" pageNumber="520">Fig. 14</figureCitation>
|
||
).
|
||
<taxonomicName id="4C274D4FFFFDE41DFC00FEC1FB0FFE82" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[952,1166,279,305]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="curvispina" status="sp. nov.">
|
||
<emphasis id="B953EADEFFFDE41DFC00FEC1FB0FFE82" box="[952,1166,279,305]" italics="true" pageId="27" pageNumber="520">Timbellus curvispina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFB29FECEFB4BFE81" box="[1169,1226,280,306]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
also bears cord spines that are always more developed than in
|
||
<taxonomicName id="4C274D4FFFFDE41DFB58FEEEFCEDFEC2" baseAuthorityName="Lamarck" baseAuthorityYear="1803" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
|
||
<emphasis id="B953EADEFFFDE41DFB58FEEEFCEDFEC2" italics="true" pageId="27" pageNumber="520">T. crenulatus tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the spiral cords of
|
||
<taxonomicName id="4C274D4FFFFDE41DFB04FE8EFAC4FEC2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1212,1349,344,369]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="curvispina" status="n. sp.">
|
||
<emphasis id="B953EADEFFFDE41DFB04FE8EFAC4FEC2" box="[1212,1349,344,369]" italics="true" pageId="27" pageNumber="520">T. curvispina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFAF4FE81FA09FEC2" box="[1356,1416,343,369]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
are weak in the intervarical spaces, whereas those of
|
||
<taxonomicName id="4C274D4FFFFDE41DFA92FEA1FC24FE03" baseAuthorityName="Lamarck" baseAuthorityYear="1803" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
|
||
<emphasis id="B953EADEFFFDE41DFA92FEA1FC24FE03" italics="true" pageId="27" pageNumber="520">T. crenulatus tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
are well marked.
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFBCAFE40FAD3FE02" author="PACAUD J. - M. & LEDON D. & GORET B." box="[1138,1362,406,433]" pageId="27" pageNumber="520" pagination="58 - 61" refId="ref48127" refString="PACAUD J. - M., LEDON D. & GORET B. 2017. - Timbellus capitaneus n. sp., une espece nouvelle de Muricidae (Mollusca, Gastropoda) du Lutetien (Eocene moyen) du bassin de Paris. Fossiles. Revue francaise de paleontologie 32: 58 - 61." type="journal article" year="2017">
|
||
Pacaud
|
||
<emphasis id="B953EADEFFFDE41DFB7EFE41FB7CFE03" box="[1222,1277,406,432]" italics="true" pageId="27" pageNumber="520">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
recently designated the
|
||
<typeStatus id="549C886EFFFDE41DFC61FE60FBBCFE63" box="[985,1085,438,464]" pageId="27" pageNumber="520" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C274D4FFFFDE41DFBDFFE61FCE6FE42" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="tricarinoides">
|
||
<emphasis id="B953EADEFFFDE41DFBDFFE61FABBFE63" box="[1127,1338,438,464]" italics="true" pageId="27" pageNumber="520">Murex tricarinoides</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFAF9FE60FCE6FE42" author="DESHAYES G. - P." pageId="27" pageNumber="520" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
|
||
</taxonomicName>
|
||
demonstrating that authors confused this species with another:
|
||
<taxonomicName id="4C274D4FFFFDE41DFC2BFE21FA0BFDA3" authority="Pacaud, Ledon & Goret, 2017" authorityName="Pacaud, Ledon & Goret" authorityYear="2017" box="[915,1418,502,528]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="capitaneus">
|
||
<emphasis id="B953EADEFFFDE41DFC2BFE21FBA0FDA3" box="[915,1057,503,528]" italics="true" pageId="27" pageNumber="520">T. capitaneus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFB93FE20FA0BFDA3" author="PACAUD J. - M. & LEDON D. & GORET B." box="[1067,1418,502,528]" pageId="27" pageNumber="520" pagination="58 - 61" refId="ref48127" refString="PACAUD J. - M., LEDON D. & GORET B. 2017. - Timbellus capitaneus n. sp., une espece nouvelle de Muricidae (Mollusca, Gastropoda) du Lutetien (Eocene moyen) du bassin de Paris. Fossiles. Revue francaise de paleontologie 32: 58 - 61." type="journal article" year="2017">Pacaud, Ledon & Goret, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
. In addition, they synonymised
|
||
<taxonomicName id="4C274D4FFFFDE41DFBEEFDC1FB6EFD83" authorityName="Deshayes" authorityYear="1835" box="[1110,1263,534,560]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="tricarinoides">
|
||
<emphasis id="B953EADEFFFDE41DFBEEFDC1FB6EFD83" box="[1110,1263,534,560]" italics="true" pageId="27" pageNumber="520">T. tricarinoides</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C274D4FFFFDE41DFA93FDC1FC22FDE3" baseAuthorityName="Lamarck" baseAuthorityYear="1803" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
|
||
<emphasis id="B953EADEFFFDE41DFA93FDC1FC22FDE3" italics="true" pageId="27" pageNumber="520">T. crenulatus tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C274D4FFFFDE41DFB9CFDE1FB43FDE3" authorityName="Deshayes" authorityYear="1835" box="[1060,1218,566,592]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="tricarinoides">
|
||
<emphasis id="B953EADEFFFDE41DFB9CFDE1FB43FDE3" box="[1060,1218,566,592]" italics="true" pageId="27" pageNumber="520">T. tricarinoides</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C274D4FFFFDE41DFAEAFDE1FC5DFDC3" baseAuthorityName="Lamarck" baseAuthorityYear="1803" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="tricarinatus">
|
||
<emphasis id="B953EADEFFFDE41DFAEAFDE1FC5DFDC3" italics="true" pageId="27" pageNumber="520">T. crenulatus tricarinatus</emphasis>
|
||
</taxonomicName>
|
||
in lacking true P1 spines and in displaying a rounded outer lip.
|
||
<taxonomicName id="4C274D4FFFFDE41DFBB3FDA1FA2EFD23" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[1035,1455,630,656]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="crenulatus">
|
||
<emphasis id="B953EADEFFFDE41DFBB3FDA1FA80FD23" box="[1035,1281,630,656]" italics="true" pageId="27" pageNumber="520">T. crenulatus crenulatus</emphasis>
|
||
(Röding, 1798)
|
||
</taxonomicName>
|
||
from the Bartonian is easily distinguishable from
|
||
<taxonomicName id="4C274D4FFFFDE41DFA92FD40FA2FFD1C" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1322,1454,662,687]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="curvispina" status="n. sp.">
|
||
<emphasis id="B953EADEFFFDE41DFA92FD40FA2FFD1C" box="[1322,1454,662,687]" italics="true" pageId="27" pageNumber="520">T. curvispina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFC93FD60FCE7FD63" box="[811,870,694,720]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
by its acute and long P1 spine pointing abaperturally.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFFDE41DFC95FD21FB82FAA3" blockId="27.[811,1457,759,1296]" pageId="27" pageNumber="520">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC95FD21FCFFFCBC" baselines="778,778" box="[813,894,759,785]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Species" pageId="27" pageNumber="520">SPECIES</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC3DFD2AFC4FFCBC" baselines="778" box="[901,974,764,783]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="group" pageId="27" pageNumber="520">GROUP</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFFDE41DFC6DFD2AFC70FCBC" baselines="778" box="[981,1009,764,783]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="27" pageNumber="520">OF</smallCapsWord>
|
||
<emphasis id="B953EADEFFFDE41DFC40FD21FB90FCA3" box="[1016,1041,759,784]" italics="true" pageId="27" pageNumber="520">T.</emphasis>
|
||
<taxonomicName id="4C274D4FFFFDE41DFBA0FD2AFAF7FCA2" authority="(LAMARCK, 1822)" baseAuthorityName="LAMARCK" baseAuthorityYear="1822" box="[1048,1398,759,785]" class="Insecta" family="Tettigoniidae" genus="Phyllopterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="genus">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFBA0FD2AFB31FCBC" baselines="778" box="[1048,1200,764,783]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="phyllopterus" pageId="27" pageNumber="520">
|
||
<emphasis id="B953EADEFFFDE41DFBA0FD2AFB31FCBC" box="[1048,1200,764,783]" italics="true" pageId="27" pageNumber="520">PHYLLOPTERUS</emphasis>
|
||
</smallCapsWord>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFB06FD21FAEFFCA2" author="LAMARCK J. - B." box="[1214,1390,759,785]" pageId="27" pageNumber="520" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">
|
||
<smallCapsWord id="8D7EA010FFFDE41DFB06FD21FAA6FCBC" baselines="778,778" box="[1214,1319,759,785]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Lamarck" pageId="27" pageNumber="520">LAMARCK</smallCapsWord>
|
||
, 1822
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
Members of this group are usually large sized and share a strong development of the internal denticles within the outer lip and strong primary cords; secondary cords are poorly developed. All species lack columellar denticles, except in the
|
||
<typeStatus id="549C886EFFFDE41DFCE0FC41FC36FC02" box="[856,951,919,945]" pageId="27" pageNumber="520" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C274D4FFFFDE41DFC63FC4EFA97FC02" authority="(Harasewych, 1982)" baseAuthorityName="Harasewych" baseAuthorityYear="1982" box="[987,1302,919,946]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="xenos">
|
||
<emphasis id="B953EADEFFFDE41DFC63FC4EFC6DFC02" box="[987,1004,920,945]" italics="true" pageId="27" pageNumber="520">T</emphasis>
|
||
.
|
||
<emphasis id="B953EADEFFFDE41DFC43FC4EFBB0FC02" box="[1019,1073,920,945]" italics="true" pageId="27" pageNumber="520">xenos</emphasis>
|
||
(Harasewych, 1982)
|
||
</taxonomicName>
|
||
, which is presumably a pathological specimen (
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFB1FFC61FAD7FC62" author="VOKES E. H." box="[1191,1366,951,978]" pageId="27" pageNumber="520" pagination="1 - 108" refId="ref48996" refString="VOKES E. H. 1992. - Cenozoic Muricidae of the Western Atlantic region, Part IX - Pterynotus, Poirieria, Aspella, Dermomurex, Calotrophon, Acantolabia, and Attiliosa; additions and corrections. Tulane Studies in Geology and Paleontology 25 (1 - 3): 1 - 108." type="journal article" year="1992">Vokes 1992: 18</bibRefCitation>
|
||
). In the middle Eocene from Europe,
|
||
<taxonomicName id="4C274D4FFFFDE41DFBD0FC01FAEEFC42" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1128,1391,983,1009]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="submicropterus" status="sp. nov.">
|
||
<emphasis id="B953EADEFFFDE41DFBD0FC01FAEEFC42" box="[1128,1391,983,1009]" italics="true" pageId="27" pageNumber="520">Timbellus submicropterus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFACDFC01FA31FC42" box="[1397,1456,983,1009]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
(upper Lutetian) and the Bartonian
|
||
<taxonomicName id="4C274D4FFFFDE41DFB1CFC2EFCEDFB82" authority="(Deshayes, 1835)" baseAuthorityName="Deshayes" baseAuthorityYear="1835" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="micropterus">
|
||
<emphasis id="B953EADEFFFDE41DFB1CFC2EFAB7FBA2" box="[1188,1334,1016,1041]" italics="true" pageId="27" pageNumber="520">T. micropterus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFAFBFC21FCE5FB82" author="DESHAYES G. - P." pageId="27" pageNumber="520" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
represent the oldest members of this species group. It becomes more diversified in the late Eocene/early Oligocene with
|
||
<taxonomicName id="4C274D4FFFFDE41DFCDAFB81FB29FBC2" authority="(Grateloup, 1845)" baseAuthorityName="Grateloup" baseAuthorityYear="1845" box="[866,1192,1110,1137]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="lamarckii">
|
||
<emphasis id="B953EADEFFFDE41DFCDAFB81FC5EFBC3" box="[866,991,1110,1136]" italics="true" pageId="27" pageNumber="520">T. lamarckii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFC54FB80FB20FBC2" author="GRATELOUP J. - P. S. & DE" box="[1004,1185,1110,1137]" pageId="27" pageNumber="520" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup, 1845</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFFDE41DFB09FB81FCEFFB23" authority="(von Koenen, 1890)" baseAuthorityName="von Koenen" baseAuthorityYear="1890" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="detritus">
|
||
<emphasis id="B953EADEFFFDE41DFB09FB81FA99FBC3" box="[1201,1304,1110,1136]" italics="true" pageId="27" pageNumber="520">T. detritus</emphasis>
|
||
(von Koenen, 1890)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFFDE41DFCC3FBA1FB19FB23" authority="(Steuer, 1912)" baseAuthorityName="Steuer" baseAuthorityYear="1912" box="[891,1176,1142,1168]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="arenarius">
|
||
<emphasis id="B953EADEFFFDE41DFCC3FBA1FC77FB23" box="[891,1014,1143,1168]" italics="true" pageId="27" pageNumber="520">T. arenarius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFBBCFBA0FB0EFB23" author="STEUER A." box="[1028,1167,1142,1168]" pageId="27" pageNumber="520" pagination="1 - 65" refId="ref48658" refString="STEUER A. 1912. - Marine Conchylien aus dem Mainzer Becken, I. Abhandlungen der Grossherzoglich Hessischen Geologischen Landeshanstalt zu Darmstadt 6 (1): 1 - 65." type="journal article" year="1912">Steuer, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFFDE41DFB19FBA1FA2EFB23" authority="Pacaud, 2018" authorityName="Pacaud" authorityYear="2018" box="[1185,1455,1142,1168]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="danapris">
|
||
<emphasis id="B953EADEFFFDE41DFB19FBA1FA97FB23" box="[1185,1302,1142,1168]" italics="true" pageId="27" pageNumber="520">T. danapris</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFAA5FBA0FA2EFB23" author="PACAUD J. - M." box="[1309,1455,1142,1168]" pageId="27" pageNumber="520" pagination="107 - 116" refId="ref48075" refString="PACAUD J. - M. 2018. - Timbellus danapris n. sp., une espece nouvelle de Muricidae (Mollusca, Gastropoda) du Priabonien (Eocene superieur) de Dnipro (Oblast de Dnipropetrovsk, Ukraine). Novapex 19 (4): 107 - 116." type="journal article" year="2018">Pacaud, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFFDE41DFCE6FB41FC6EFB03" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[862,1007,1174,1200]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="radulfiensis" status="n. sp.">
|
||
<emphasis id="B953EADEFFFDE41DFCE6FB41FC6EFB03" box="[862,1007,1174,1200]" italics="true" pageId="27" pageNumber="520">T. radulfiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFFDE41DFC40FB41FBB5FB02" box="[1016,1076,1175,1201]" pageId="27" pageNumber="520" rank="species">n. sp.</taxonomicNameLabel>
|
||
It contains three Miocene species:
|
||
<taxonomicName id="4C274D4FFFFDE41DFC96FB61FBE0FB63" authority="(Bellardi, 1872)" baseAuthorityName="Bellardi" baseAuthorityYear="1872" box="[814,1121,1206,1232]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="perlongus">
|
||
<emphasis id="B953EADEFFFDE41DFC96FB61FC2DFB63" box="[814,940,1206,1232]" italics="true" pageId="27" pageNumber="520">T. perlongus</emphasis>
|
||
(Bellardi, 1872)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFFDE41DFBD7FB61FA2EFB63" authority="(Bellardi, 1872)" baseAuthorityName="Bellardi" baseAuthorityYear="1872" box="[1135,1455,1206,1232]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="rosavendae">
|
||
<emphasis id="B953EADEFFFDE41DFBD7FB61FB7AFB63" box="[1135,1275,1206,1232]" italics="true" pageId="27" pageNumber="520">T. rosavendae</emphasis>
|
||
(Bellardi, 1872)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFFDE41DFCE4FB01FB47FB43" authority="(Magne, 1941)" baseAuthorityName="Magne" baseAuthorityYear="1941" box="[860,1222,1238,1264]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="520" phylum="Mollusca" rank="species" species="pseudolamarcki">
|
||
<emphasis id="B953EADEFFFDE41DFCE4FB01FB99FB43" box="[860,1048,1238,1264]" italics="true" pageId="27" pageNumber="520">T. pseudolamarcki</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFFDE41DFB90FB00FB3FFB43" author="MAGNE A." box="[1064,1214,1238,1264]" pageId="27" pageNumber="520" pagination="28 - 36" refId="ref46270" refString="MAGNE A. 1941. - Sur quelques especes nouvelles ou interessantes provenant du Calcaire a Asteries de la Souys. Proces-verbaux de la Societe linneenne de Bordeaux 91: 28 - 36." type="journal article" year="1941">Magne, 1941</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and becomes extinct before the Pliocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |