651 lines
82 KiB
XML
651 lines
82 KiB
XML
<document id="00338D2F090CC26CF55AD532776BCD16" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard" docDate="2024" docId="038E87DAFFFBE427FC33F8A4FE65FAFC" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pterynotus (Pterymarchia) gaasensis Merle, Pacaud, Ledon & Goret, 2024, n. sp." docType="treatment" docUuid="B1E15CF2-7B4F-4D02-8F34-1D5848D596F5" docUuidSource="ZooBank" docVersion="7" lastPageNumber="526" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="522" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="E769EFA1F76E3283C4C70349B6B72BEA">
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<mods:title id="CCA8827559C7E3C6F4973B180995AD59">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:name id="9D83286F658CF927B1BCAAB87C485B87" type="personal">
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<mods:namePart id="B852BD89C6247ADC80971A1E2DF0407C">Merle, Didier</mods:namePart>
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<mods:namePart id="4B2025138CBB9AD47AE27603FBF410E9">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="A13BA925066F7AA85BF74993D6E01ED6">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="CFD11097B34E6EE6EDAA4C69C2D2B477">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:nameIdentifier id="D0782A33AB37A78C692D4365BA13FB42" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="32EEBEDAD17758A337D2F105D1AF32BD">Goret, Bernard</mods:namePart>
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<mods:affiliation id="F142A70B0AA63F7EE2A0D4CF47F4BED7">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:nameIdentifier id="92FC24EF025187ACAE4235277281C3E2" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="D8894C79316834C20429A6B94DB43E39">Geodiversitas</mods:title>
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<mods:part id="4496DB49D293E1B082F4943CBB95AD4E">
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<mods:date id="EEC2032213B5985423171B44E6657508">2024</mods:date>
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<mods:number id="26958FBF576FDD322DE4C7E6710B7F76">2024-10-24</mods:number>
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<mods:number id="D7D4E3A1932A201A17A6F8019F4A3010">46</mods:number>
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<mods:number id="DF0B0D3D867B7D06029085EBA692B376">15</mods:number>
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<mods:url id="C99A2B9FF4AB96DDDE00C764348F705C">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf</mods:url>
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<mods:classification id="6C174CB063042FAC44B26477DE4E9716">journal article</mods:classification>
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<mods:identifier id="2EF63061A698A1317B2B8503341C0C3A" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="F18670886A5C51247AE85DBADCDA7C01" type="Zenodo-Dep">13985760</mods:identifier>
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<treatment id="038E87DAFFFBE427FC33F8A4FE65FAFC" ID-DOI="http://doi.org/10.5281/zenodo.13991579" ID-GBIF-Taxon="244735017" ID-Zenodo-Dep="13991579" LSID="urn:lsid:zoobank.org:act:B1E15CF2-7B4F-4D02-8F34-1D5848D596F5" httpUri="http://treatment.plazi.org/id/038E87DAFFFBE427FC33F8A4FE65FAFC" lastPageId="33" lastPageNumber="526" pageId="29" pageNumber="522" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae" scope_subGenus="Pterymarchia">
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<subSubSection id="C33D6547FFFBE41BFC33F8A4FAD3F83F" box="[907,1362,1905,1933]" pageId="29" pageNumber="522" type="nomenclature">
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<paragraph id="8B9836CCFFFBE41BFC33F8A4FAD3F83F" blockId="29.[907,1362,1905,1964]" box="[907,1362,1905,1933]" pageId="29" pageNumber="522">
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<taxonomicName id="4C274D4FFFFBE41BFC33F8A4FA91F838" ID-CoL="CN5MR" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[907,1296,1905,1933]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="species" species="gaasensis" status="sp. nov." subGenus="Pterymarchia">
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<emphasis id="B953EADEFFFBE41BFC33F8A4FC7FF83F" bold="true" box="[907,1022,1906,1932]" italics="true" pageId="29" pageNumber="522">Pterynotus</emphasis>
|
||
(
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||
<emphasis id="B953EADEFFFBE41BFBB6F8A5FB20F83E" bold="true" box="[1038,1185,1907,1933]" italics="true" pageId="29" pageNumber="522">Pterymarchia</emphasis>
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||
)
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<emphasis id="B953EADEFFFBE41BFB09F8A7FA91F838" bold="true" box="[1201,1296,1905,1931]" italics="true" pageId="29" pageNumber="522">gaasensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26057A5FFFBE41BFAAFF8A4FAD3F83F" box="[1303,1362,1906,1932]" pageId="29" pageNumber="522" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6547FFFBE41BFC42F847FA31F852" pageId="29" pageNumber="522" type="description">
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||
<paragraph id="8B9836CCFFFBE41BFC42F847FB63F818" blockId="29.[907,1362,1905,1964]" box="[1018,1250,1937,1964]" pageId="29" pageNumber="522">
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||
(
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||
<figureCitation id="131C2A49FFFBE41BFBBBF847FBD0F818" box="[1027,1105,1937,1963]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="29" pageNumber="522">Figs 18</figureCitation>
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A-D; 27E, F)
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</paragraph>
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<paragraph id="8B9836CCFFFBE41BFC95F806FA31F852" blockId="29.[813,1456,2000,2017]" box="[813,1456,2000,2017]" pageId="29" pageNumber="522">
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<uri id="FFB63ACEFFFBE41BFC95F806FA31F852" box="[813,1456,2000,2017]" pageId="29" pageNumber="522">
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urn:lsid:zoobank.org:act:
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<uuid id="FF810C19FFFBE41BFBA8F806FA31F852" box="[1040,1456,2000,2017]" pageId="29" pageNumber="522">B1E15CF2-7B4F-4D02-8F34-1D5848D596F5</uuid>
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</uri>
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</paragraph>
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</subSubSection>
|
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<caption id="DF586644FFF8E418FF3CF886FD7CF85A" ID-DOI="http://doi.org/10.5281/zenodo.13985800" ID-Zenodo-Dep="13985800" httpUri="https://zenodo.org/record/13985800/files/figure.png" pageId="30" pageNumber="523" startId="30.[132,143,1872,1889]" targetBox="[147,1440,260,1687]" targetPageId="30" targetType="figure">
|
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<paragraph id="8B9836CCFFF8E418FF3CF886FD7CF85A" blockId="30.[132,1457,1872,2025]" pageId="30" pageNumber="523">
|
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<smallCapsWord id="8D7EA010FFF8E418FF3CF886FF1CF8D3" baselines="1885,1885" box="[132,157,1872,1889]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="30" pageNumber="523">FIG</smallCapsWord>
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||
. 16. — Spiral cords and internal denticles of the outer lip in
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<taxonomicName id="4C274D4FFFF8E418FD33F886FCE0F8D2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[651,865,1872,1889]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="30" pageNumber="523" phylum="Mollusca" rank="species" species="submicropterus" status="sp. nov.">
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||
<emphasis id="B953EADEFFF8E418FD33F886FCE0F8D2" box="[651,865,1872,1889]" italics="true" pageId="30" pageNumber="523">Timbellus submicropterus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26057A5FFF8E418FCDDF886FC14F8D2" box="[869,917,1872,1889]" pageId="30" pageNumber="523" rank="species">n. sp.</taxonomicNameLabel>
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||
(
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<emphasis id="B953EADEFFF8E418FC25F886FC3EF8D2" bold="true" box="[925,959,1872,1889]" pageId="30" pageNumber="523">A -H</emphasis>
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||
,
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<emphasis id="B953EADEFFF8E418FC71F886FC57F8D2" bold="true" box="[969,982,1872,1889]" pageId="30" pageNumber="523">K</emphasis>
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||
) and
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||
<taxonomicName id="4C274D4FFFF8E418FBBCF886FA8EF8D2" authority="(Deshayes, 1835)" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[1028,1295,1872,1889]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="30" pageNumber="523" phylum="Mollusca" rank="species" species="micropterus">
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<emphasis id="B953EADEFFF8E418FBBCF886FBFDF8D2" box="[1028,1148,1872,1889]" italics="true" pageId="30" pageNumber="523">T. micropterus</emphasis>
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||
(
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<bibRefCitation id="EFB64B3DFFF8E418FB3DF886FA8AF8D2" author="DESHAYES G. - P." box="[1157,1291,1872,1889]" pageId="30" pageNumber="523" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
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||
)
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</taxonomicName>
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||
(
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<emphasis id="B953EADEFFF8E418FAA1F886FA9EF8D2" bold="true" box="[1305,1311,1872,1889]" pageId="30" pageNumber="523">I</emphasis>
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||
,
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<emphasis id="B953EADEFFF8E418FA90F886FAB3F8D2" bold="true" box="[1320,1330,1872,1889]" pageId="30" pageNumber="523">J</emphasis>
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||
,
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<emphasis id="B953EADEFFF8E418FA84F886FAC6F8D2" bold="true" box="[1340,1351,1872,1889]" pageId="30" pageNumber="523">L</emphasis>
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||
):
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||
<emphasis id="B953EADEFFF8E418FAEEF886FAE2F8D2" bold="true" box="[1366,1379,1872,1889]" pageId="30" pageNumber="523">A</emphasis>
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||
,
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||
<emphasis id="B953EADEFFF8E418FAD5F886FAFBF8D2" bold="true" box="[1389,1402,1872,1889]" pageId="30" pageNumber="523">B</emphasis>
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||
, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, Manche, Normandy, France, middle Eocene (Lutetian);
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<emphasis id="B953EADEFFF8E418FC6EF8B1FC65F8CB" bold="true" box="[982,996,1895,1912]" pageId="30" pageNumber="523">C</emphasis>
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||
,
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||
<emphasis id="B953EADEFFF8E418FC57F8B1FC7CF8CB" bold="true" box="[1007,1021,1895,1912]" pageId="30" pageNumber="523">D</emphasis>
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||
, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, Manche, middle Eocene (Lutetian);
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||
<emphasis id="B953EADEFFF8E418FDA4F8ABFDA9F83D" bold="true" box="[540,552,1917,1934]" pageId="30" pageNumber="523">E</emphasis>
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||
,
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||
<emphasis id="B953EADEFFF8E418FD8AF8ABFDBCF83D" bold="true" box="[562,573,1917,1934]" pageId="30" pageNumber="523">F</emphasis>
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||
, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville;
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||
<emphasis id="B953EADEFFF8E418FC44F8ABFB8BF83D" bold="true" box="[1020,1034,1917,1934]" pageId="30" pageNumber="523">G</emphasis>
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||
,
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||
<emphasis id="B953EADEFFF8E418FBADF8ABFBA2F83D" bold="true" box="[1045,1059,1917,1934]" pageId="30" pageNumber="523">H</emphasis>
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||
, MNHN.F.A90557 (Brébion coll.), Saint-AignanGrand-Lieu (Pierre-Aiguë), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian);
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||
<emphasis id="B953EADEFFF8E418FC38F842FC07F816" bold="true" box="[896,902,1940,1957]" pageId="30" pageNumber="523">I</emphasis>
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||
,
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||
<emphasis id="B953EADEFFF8E418FC28F842FC1BF816" bold="true" box="[912,922,1940,1957]" pageId="30" pageNumber="523">J</emphasis>
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||
, MNHN.F.A32522 (Faullummel coll.), Le Limon, Seine-et-Marne, Paris Basin, France,middle Eocene (Bartonian);
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||
<emphasis id="B953EADEFFF8E418FDAAF87DFD9EF80F" bold="true" box="[530,543,1963,1980]" pageId="30" pageNumber="523">K</emphasis>
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||
, paratype (protoconch), MNHN.F.A90556 (MNHN coll.), Fresville;
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||
<emphasis id="B953EADEFFF8E418FB87F87DFBCBF80F" bold="true" box="[1087,1098,1963,1980]" pageId="30" pageNumber="523">L</emphasis>
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||
, MNHN.F.A90558 (Faullummel coll., protoconch), Chavençon (Carrière Tuyttens), Oise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: A-J, 10 mm; K, L, 0.5 mm. Credits: A-D, I, J, P. Loubry (MNHN/CNRS); E, F, E. Robert (UCBL); G, H, K, L, L. Cazes (MNHN/CNRS).
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</paragraph>
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</caption>
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<caption id="DF586644FFF9E419FF3BFCADFC5AFC11" ID-DOI="http://doi.org/10.5281/zenodo.13988488" ID-Zenodo-Dep="13988488" httpUri="https://zenodo.org/record/13988488/files/figure.png" pageId="31" pageNumber="524" startId="31.[131,142,891,908]" targetBox="[269,1278,178,858]" targetPageId="31" targetType="figure">
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<paragraph id="8B9836CCFFF9E419FF3BFCADFC5AFC11" blockId="31.[131,1455,891,930]" pageId="31" pageNumber="524">
|
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<smallCapsWord id="8D7EA010FFF9E419FF3BFCADFF1CFC38" baselines="904,904" box="[131,157,891,908]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="31" pageNumber="524">FIG</smallCapsWord>
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. 17. — Bivariate plot of total length against width comparing
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<taxonomicName id="4C274D4FFFF9E419FD2FFCADFC23FC3F" authority="(Deshayes, 1835)" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[663,930,891,908]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="micropterus">
|
||
<emphasis id="B953EADEFFF9E419FD2FFCADFC91FC3F" box="[663,784,891,908]" italics="true" pageId="31" pageNumber="524">T. micropterus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF9E419FCA1FCADFC1EFC3F" author="DESHAYES G. - P." box="[793,927,891,908]" pageId="31" pageNumber="524" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the lower Bartonian of the Paris Basin (50 complete specimens) and
|
||
<taxonomicName id="4C274D4FFFF9E419FF5DFC47FEFDFC11" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[229,380,913,930]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="submicropterus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF9E419FF5DFC47FEFDFC11" box="[229,380,913,930]" italics="true" pageId="31" pageNumber="524">T. submicropterus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF9E419FE39FC47FE30FC11" box="[385,433,913,930]" pageId="31" pageNumber="524" rank="species">n. sp.</taxonomicNameLabel>
|
||
from the uppper Lutetian of Normandy (five complete specimens).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D6547FFF9E419FF3CFC03FE84FB28" pageId="31" pageNumber="524" type="reference_group">
|
||
<paragraph id="8B9836CCFFF9E419FF3CFC03FE5CFBB5" blockId="31.[132,775,980,1030]" pageId="31" pageNumber="524">
|
||
<taxonomicName id="4C274D4FFFF9E419FF3CFC03FEB6FC5F" ID-CoL="44MDK" authorityName="Lamarck" authorityYear="1822" box="[132,311,980,1004]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="quadrifrons">
|
||
<emphasis id="B953EADEFFF9E419FF3CFC03FEB6FC5F" box="[132,311,980,1004]" italics="true" pageId="31" pageNumber="524">Murex quadrifrons</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EFB64B3DFFF9E419FEECFC02FE72FC5F" author="GRATELOUP J. - P. S. & DE" box="[340,499,980,1004]" pageId="31" pageNumber="524" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup 1845</bibRefCitation>
|
||
: pl. 3 [pl. 31], fig. 10 [
|
||
<emphasis id="B953EADEFFF9E419FD5BFC03FC86FC5F" box="[739,775,981,1004]" italics="true" pageId="31" pageNumber="524">non</emphasis>
|
||
<taxonomicName id="4C274D4FFFF9E419FF3CFC39FE4EFBB5" ID-CoL="44MDK" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[132,463,1006,1030]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="quadrifrons">
|
||
<emphasis id="B953EADEFFF9E419FF3CFC39FEB5FBB5" box="[132,308,1006,1030]" italics="true" pageId="31" pageNumber="524">Murex quadrifrons</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF9E419FE82FC38FE4EFBB5" author="LAMARCK J. - B." box="[314,463,1006,1030]" pageId="31" pageNumber="524" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E419FF3CFBC9FE32FBE3" blockId="31.[132,775,1054,1105]" pageId="31" pageNumber="524">
|
||
<taxonomicName id="4C274D4FFFF9E419FF3CFBC9FDBAFB85" ID-CoL="BM3KY" baseAuthorityName="A. d'Orbigny" baseAuthorityYear="1852" box="[132,571,1054,1078]" class="Gastropoda" family="Muricidae" genus="Pterynotus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="subquadrifrons" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFF9E419FF3CFBC9FF69FB85" box="[132,232,1055,1078]" italics="true" pageId="31" pageNumber="524">Pterynotus</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFF9E419FF41FBC8FEF8FB85" box="[249,377,1054,1078]" italics="true" pageId="31" pageNumber="524">Pterymarchia</emphasis>
|
||
) cf.
|
||
<emphasis id="B953EADEFFF9E419FE14FBC8FDBAFB85" box="[428,571,1054,1078]" italics="true" pageId="31" pageNumber="524">subquadrifrons</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EFB64B3DFFF9E419FDEFFBC8FC86FB85" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[599,775,1054,1078]" pageId="31" pageNumber="524" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
Merle
|
||
<emphasis id="B953EADEFFF9E419FD22FBC9FD4DFB85" box="[666,716,1054,1078]" italics="true" pageId="31" pageNumber="524">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
(
|
||
<emphasis id="B953EADEFFF9E419FF34FBEFFF4DFBE3" box="[140,204,1081,1104]" italics="true" pageId="31" pageNumber="524">partim</emphasis>
|
||
): pl. 90, figs 3-74 only.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E419FF3CFBBFFE84FB28" blockId="31.[132,775,1128,1179]" pageId="31" pageNumber="524">
|
||
<treatmentCitationGroup id="AB3711E2FFF9E419FF3CFBBFFE84FB28" pageId="31" pageNumber="524">
|
||
<taxonomicName id="4C274D4FFFF9E419FF3CFBBFFE16FB33" ID-CoL="BM3KY" baseAuthorityName="A. d'Orbigny" baseAuthorityYear="1852" box="[132,407,1128,1152]" class="Gastropoda" family="Muricidae" genus="Pterynotus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="subquadrifrons">
|
||
<emphasis id="B953EADEFFF9E419FF3CFBBFFF64FB33" box="[132,229,1129,1152]" italics="true" pageId="31" pageNumber="524">Pterynotus</emphasis>
|
||
cf.
|
||
<emphasis id="B953EADEFFF9E419FEB4FBBEFE16FB33" box="[268,407,1128,1152]" italics="true" pageId="31" pageNumber="524">subquadrifrons</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0A8610DDFFF9E419FE0BFBBFFDE2FB33" author="LOZOUET P." box="[435,611,1128,1152]" page="21" pageId="31" pageNumber="524" year="2023">
|
||
<bibRefCitation id="EFB64B3DFFF9E419FE0BFBBFFDE2FB33" author="LOZOUET P." box="[435,611,1128,1152]" pageId="31" pageNumber="524" pagination="3 - 162" refId="ref46112" refString="LOZOUET P. 2023. - Muricidae (Gastropoda, Neogastropoda) de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana 24: 3 - 162." type="journal article" year="2023">Lozouet 2023: 21</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 17, figs 1-9, text-fig. 10E.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF9E419FF3CFB65FD85FA5B" pageId="31" pageNumber="524" type="materials_examined">
|
||
<paragraph id="8B9836CCFFF9E419FF3CFB65FE58FA30" blockId="31.[131,777,1202,1411]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CFB65FF34FB7B" baselines="1220,1220" box="[132,181,1203,1226]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="31" pageNumber="524">TYPE</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF9E419FF05FB6EFE9AFB7A" baselines="1221" box="[189,283,1208,1225]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="31" pageNumber="524">MATERIAL</smallCapsWord>
|
||
. —
|
||
<materialsCitation id="3B4F3C91FFF9E419FEF3FB64FD78FAA9" ID-GBIF-Occurrence="4958671388" pageId="31" pageNumber="524">
|
||
<emphasis id="B953EADEFFF9E419FEF3FB64FE7BFB79" bold="true" box="[331,506,1202,1226]" pageId="31" pageNumber="524">
|
||
<typeStatus id="549C886EFFF9E419FEF3FB64FE2DFB79" box="[331,428,1202,1226]" pageId="31" pageNumber="524" type="holotype">Holotype</typeStatus>
|
||
.
|
||
<collectingCountry id="F330765CFFF9E419FE00FB64FE7BFB79" box="[440,506,1202,1226]" name="France" pageId="31" pageNumber="524">France</collectingCountry>
|
||
</emphasis>
|
||
•
|
||
<collectingRegion id="49E3F82EFFF9E419FDABFB64FDF2FB79" box="[531,627,1202,1226]" country="France" name="Aquitaine" pageId="31" pageNumber="524">Aquitaine</collectingRegion>
|
||
Basin, Landes, Gaas (Espibos); Gaas Formation (biozone NP23); early Oligocene (Rupelian);
|
||
<collectionCode id="ED36AE09FFF9E419FF43FB3EFED1FB4C" box="[251,336,1256,1279]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="524">MNHN</collectionCode>
|
||
.F.A32492 (
|
||
<taxonomicName id="4C274D4FFFF9E419FE7AFB31FE7CFB4C" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[450,509,1255,1279]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.), figured specimen in
|
||
<bibRefCitation id="EFB64B3DFFF9E419FF3EFAD7FEB2FAA9" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[134,307,1281,1306]" pageId="31" pageNumber="524" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF9E419FF3EFAD7FF3EFAAA" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[134,191,1281,1305]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF9E419FF7CFAD4FF72FAAA" box="[196,243,1281,1305]" italics="true" pageId="31" pageNumber="524">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
: pl. 90, fig. 3), H:
|
||
<quantity id="4CDF9B29FFF9E419FE55FAD4FDB4FAAA" box="[493,565,1282,1306]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="31" pageNumber="524" unit="mm" value="27.0">27 mm</quantity>
|
||
(
|
||
<figureCitation id="131C2A49FFF9E419FDFBFAD7FD36FAAA" box="[579,695,1281,1306]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="31" pageNumber="524">Figs 18A, B</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFF9E419FD7AFAD4FD6CFAA9" box="[706,749,1282,1306]" captionStart="FIG" captionStartId="46.[132,143,1606,1623]" captionTargetBox="[132,1455,215,1546]" captionTargetId="figure-366@46.[699,1260,794,1359]" captionTargetPageId="46" captionText="FIG. 27. — New species of Timbellus De Gregorio, 1885, Pterynotus Swainson, 1833 (Pterymarchia Houart, 1995), Textiliomurex Merle, 2011, Ponderia Houart, 1986 and Flexopteron Shuto, 1969: A-C, Timbellus submicropterus n. sp.: A, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; B, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; C, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; D, T. cf. submicropterus n. sp., MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; E, F, Pterynotus (Pterymarchia) gaasensis n. sp.: E, holotype, MNHN.F.A32492 (Merle coll.), Gaas (Espibos), France, Rupelian; F, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos); G, H, P. (P.) pelouatensis n. sp.: G, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, Textiliomurex chodmonensis n. sp.: I, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; J, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en- Vexin (Carrière Darcy); K, L, Ponderia remyi n. sp.: K, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; L, paratype, MNHN.F.A90555, Fresville (Remy coll.); M, N, Flexopteron constantinense n. sp.: M, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; N, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985831" httpUri="https://zenodo.org/record/13985831/files/figure.png" pageId="31" pageNumber="524">27E</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4F3C91FFF9E419FF3CFACAFE83FAFD" ID-GBIF-Occurrence="4958671355" pageId="31" pageNumber="524">
|
||
<emphasis id="B953EADEFFF9E419FF3CFACAFEA7FA87" bold="true" box="[132,294,1308,1332]" pageId="31" pageNumber="524">
|
||
<typeStatus id="549C886EFFF9E419FF3CFACAFF61FA87" box="[132,224,1308,1332]" pageId="31" pageNumber="524" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<collectingCountry id="F330765CFFF9E419FF50FACAFEA7FA87" box="[232,294,1308,1332]" name="France" pageId="31" pageNumber="524">France</collectingCountry>
|
||
</emphasis>
|
||
• 6 spm; same as for the holotype;
|
||
<collectionCode id="ED36AE09FFF9E419FDEDFACAFD27FA80" box="[597,678,1308,1331]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="524">MNHN</collectionCode>
|
||
.F.A77749 (
|
||
<taxonomicName id="4C274D4FFFF9E419FF32FAE0FF43FAFD" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[138,194,1334,1358]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.) •
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4F3C91FFF9E419FEBDFAE0FF63FADB" ID-GBIF-Occurrence="4958671310" pageId="31" pageNumber="524">
|
||
1 spm; idem;
|
||
<collectionCode id="ED36AE09FFF9E419FEC5FAE1FE4FFAFD" box="[381,462,1335,1358]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="524">MNHN</collectionCode>
|
||
.F.A90554 (
|
||
<taxonomicName id="4C274D4FFFF9E419FD82FAE0FDF3FAFD" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[570,626,1334,1358]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.) (
|
||
<figureCitation id="131C2A49FFF9E419FD17FAE0FF1BFADB" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="31" pageNumber="524">Figs 18C, D</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFF9E419FF1CFA87FF4DFADA" box="[164,204,1361,1385]" captionStart="FIG" captionStartId="46.[132,143,1606,1623]" captionTargetBox="[132,1455,215,1546]" captionTargetId="figure-366@46.[699,1260,794,1359]" captionTargetPageId="46" captionText="FIG. 27. — New species of Timbellus De Gregorio, 1885, Pterynotus Swainson, 1833 (Pterymarchia Houart, 1995), Textiliomurex Merle, 2011, Ponderia Houart, 1986 and Flexopteron Shuto, 1969: A-C, Timbellus submicropterus n. sp.: A, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; B, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; C, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; D, T. cf. submicropterus n. sp., MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; E, F, Pterynotus (Pterymarchia) gaasensis n. sp.: E, holotype, MNHN.F.A32492 (Merle coll.), Gaas (Espibos), France, Rupelian; F, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos); G, H, P. (P.) pelouatensis n. sp.: G, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, Textiliomurex chodmonensis n. sp.: I, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; J, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en- Vexin (Carrière Darcy); K, L, Ponderia remyi n. sp.: K, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; L, paratype, MNHN.F.A90555, Fresville (Remy coll.); M, N, Flexopteron constantinense n. sp.: M, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; N, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985831" httpUri="https://zenodo.org/record/13985831/files/figure.png" pageId="31" pageNumber="524">27F</figureCitation>
|
||
) •
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4F3C91FFF9E419FF5FFA87FE55FA30" ID-GBIF-Occurrence="4958671429" pageId="31" pageNumber="524">
|
||
1 spm; idem;
|
||
<collectionCode id="ED36AE09FFF9E419FED2FA87FE3EFADB" box="[362,447,1361,1384]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="524">MNHN</collectionCode>
|
||
.F.A86913 (Lozouet coll.), figured in
|
||
<bibRefCitation id="EFB64B3DFFF9E419FF26FABAFEAEFA30" author="LOZOUET P." box="[158,303,1387,1411]" pageId="31" pageNumber="524" pagination="3 - 162" refId="ref46112" refString="LOZOUET P. 2023. - Muricidae (Gastropoda, Neogastropoda) de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana 24: 3 - 162." type="journal article" year="2023">Lozouet (2023</bibRefCitation>
|
||
: text-fig. 10a, b)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E419FF3CFA4DFD85FA5B" blockId="31.[131,775,1435,1512]" pageId="31" pageNumber="524">
|
||
<typeStatus id="549C886EFFF9E419FF3CFA4DFEDFFA02" box="[132,350,1435,1459]" isOtherTypeStatus="true" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CFA4DFF7CFA02" baselines="1453,1453" box="[132,253,1435,1459]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Additional" pageId="31" pageNumber="524">ADDITIONAL</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF9E419FEBAFA76FEDFFA02" baselines="1453" box="[258,350,1440,1457]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="31" pageNumber="524">MATERIAL</smallCapsWord>
|
||
</typeStatus>
|
||
.
|
||
<materialsCitation id="3B4F3C91FFF9E419FED0FA4DFD85FA5B" ID-GBIF-Occurrence="4958671422" collectorName="Gaas & La Gouarde" country="France" location="France" pageId="31" pageNumber="524" specimenCount="1" stateProvince="Aquitaine">
|
||
—
|
||
<emphasis id="B953EADEFFF9E419FE3CFA4DFE44FA00" bold="true" box="[388,453,1435,1459]" pageId="31" pageNumber="524">
|
||
<collectingCountry id="F330765CFFF9E419FE3CFA4DFE44FA00" box="[388,453,1435,1459]" name="France" pageId="31" pageNumber="524">France</collectingCountry>
|
||
</emphasis>
|
||
• 1 spm;
|
||
<collectingRegion id="49E3F82EFFF9E419FDA5FA4DFDFAFA00" box="[541,635,1435,1459]" country="France" name="Aquitaine" pageId="31" pageNumber="524">Aquitaine</collectingRegion>
|
||
<location id="8EF86017FFF9E419FDC6FA4DFD34FA00" LSID="urn:lsid:plazi:treatment:038E87DAFFFBE427FC33F8A4FE65FAFC:8EF86017FFF9E419FDC6FA4DFD34FA00" box="[638,693,1435,1459]" country="France" name="Basin" pageId="31" pageNumber="524" stateProvince="Aquitaine">Basin</location>
|
||
, Landes,
|
||
<collectorName id="26D2531AFFF9E419FF3BFA63FF33FA7E" box="[131,178,1461,1485]" pageId="31" pageNumber="524">Gaas</collectorName>
|
||
(
|
||
<collectorName id="26D2531AFFF9E419FF06FA63FEB6FA7E" box="[190,311,1461,1485]" pageId="31" pageNumber="524">La Gouarde</collectorName>
|
||
); same as for the
|
||
<typeStatus id="549C886EFFF9E419FE65FA63FDB4FA7E" box="[477,565,1461,1485]" pageId="31" pageNumber="524" type="holotype">holotype</typeStatus>
|
||
; Aucoin coll., figured
|
||
<emphasis id="B953EADEFFF9E419FF3CFA07FF19FA5B" box="[132,152,1489,1512]" italics="true" pageId="31" pageNumber="524">in</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF9E419FF26FA06FECCFA5B" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[158,333,1488,1512]" pageId="31" pageNumber="524" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF9E419FF26FA06FF56FA5B" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[158,215,1488,1512]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF9E419FF66FA07FE8CFA5B" box="[222,269,1488,1512]" italics="true" pageId="31" pageNumber="524">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
: pl. 90, fig. 4a, b).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF9E419FF3CF9D6FD6DF9A4" box="[132,748,1535,1560]" pageId="31" pageNumber="524" type="etymology">
|
||
<paragraph id="8B9836CCFFF9E419FF3CF9D6FD6DF9A4" blockId="31.[132,748,1535,1560]" box="[132,748,1535,1560]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CF9D6FF7FF9A5" baselines="1553,1554" box="[132,254,1536,1559]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="31" pageNumber="524">ETYMOLOGY</smallCapsWord>
|
||
. — From the
|
||
<typeStatus id="549C886EFFF9E419FE33F9D6FE34F9AB" box="[395,437,1536,1560]" pageId="31" pageNumber="524">type</typeStatus>
|
||
locality, Gaas (Landes),
|
||
<collectingCountry id="F330765CFFF9E419FD1BF9D6FD66F9A4" box="[675,743,1536,1559]" name="France" pageId="31" pageNumber="524">France</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF9E419FF3CF9E6FE7AF918" pageId="31" pageNumber="524" type="materials_examined">
|
||
<paragraph id="8B9836CCFFF9E419FF3CF9E6FDFBF9D2" blockId="31.[132,775,1583,1634]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CF9E6FF34F9F6" baselines="1601,1601" box="[132,181,1584,1607]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="31" pageNumber="524">
|
||
<typeStatus id="549C886EFFF9E419FF3CF9E6FF34F9F6" box="[132,181,1584,1607]" pageId="31" pageNumber="524">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF9E419FF01F9E3FE92F9F5" baselines="1602" box="[185,275,1589,1606]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="31" pageNumber="524">HORIZON</smallCapsWord>
|
||
. — Gaas Formation (biozone NP23), see
|
||
<bibRefCitation id="EFB64B3DFFF9E419FD24F9F9FEE0F9D2" author="SZTRAKOS K. & STEURBAUT E." pageId="31" pageNumber="524" pagination="741 - 781" refId="ref48742" refString="SZTRAKOS K. & STEURBAUT E. 2017. - Revision lithostratigraphique et biostratigraphique de l'Oligocene d'Aquitaine occidentale (France). Geodiversitas 39 (4): 741 - 781. https: // doi. org / 10.5252 / g 2017 n 4 a 6" type="journal article" year="2017">Sztrákos & Steurbaut (2017: 752)</bibRefCitation>
|
||
, early Oligocene (Rupelian).
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E419FF3CF9ACFE7AF918" blockId="31.[131,776,1657,1708]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CF9ACFF34F93C" baselines="1675,1675" box="[132,181,1658,1681]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="31" pageNumber="524">
|
||
<typeStatus id="549C886EFFF9E419FF3CF9ACFF34F93C" box="[132,181,1658,1681]" pageId="31" pageNumber="524">TYPE</typeStatus>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF9E419FF01F9A9FE93F923" baselines="1676" box="[185,274,1663,1680]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="31" pageNumber="524">LOCALITY</smallCapsWord>
|
||
. —
|
||
<collectingCountry id="F330765CFFF9E419FE83F9ACFEFCF922" box="[315,381,1658,1681]" name="France" pageId="31" pageNumber="524">France</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E3F82EFFF9E419FE3DF9AFFE62F922" box="[389,483,1657,1681]" country="France" name="Aquitaine" pageId="31" pageNumber="524">Aquitaine</collectingRegion>
|
||
Basin, Landes, Gaas (Espibos), see
|
||
<bibRefCitation id="EFB64B3DFFF9E419FF1FF942FE76F918" author="SZTRAKOS K. & STEURBAUT E." box="[167,503,1683,1708]" pageId="31" pageNumber="524" pagination="741 - 781" refId="ref48742" refString="SZTRAKOS K. & STEURBAUT E. 2017. - Revision lithostratigraphique et biostratigraphique de l'Oligocene d'Aquitaine occidentale (France). Geodiversitas 39 (4): 741 - 781. https: // doi. org / 10.5252 / g 2017 n 4 a 6" type="journal article" year="2017">Sztrákos & Steurbaut (2017: 752)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF9E419FF3CF912FF4EF945" pageId="31" pageNumber="524" type="distribution">
|
||
<paragraph id="8B9836CCFFF9E419FF3CF912FF4EF945" blockId="31.[132,774,1731,1782]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CF912FE93F969" baselines="1749,1750" box="[132,274,1732,1755]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="31" pageNumber="524">DISTRIBUTION</smallCapsWord>
|
||
. —
|
||
<collectingRegion id="49E3F82EFFF9E419FE85F915FE1CF968" box="[317,413,1731,1755]" country="France" name="Aquitaine" pageId="31" pageNumber="524">Aquitaine</collectingRegion>
|
||
Basin (
|
||
<collectingCountry id="F330765CFFF9E419FE5DF915FDAAF968" box="[485,555,1731,1755]" name="France" pageId="31" pageNumber="524">France</collectingCountry>
|
||
), early Oligocene (Rupelian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF9E427FF3CF8E4FE65FAFC" lastPageId="33" lastPageNumber="526" pageId="31" pageNumber="524" type="description">
|
||
<paragraph id="8B9836CCFFF9E419FF3CF8E4FE94F8F9" blockId="31.[131,777,1842,2028]" box="[132,277,1842,1868]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FF3CF8E4FE94F8F9" baselines="1861,1861" box="[132,277,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="31" pageNumber="524">DESCRIPTION</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E419FF3CF887FBC3F93F" blockId="31.[131,777,1842,2028]" lastBlockId="31.[811,1458,980,1676]" pageId="31" pageNumber="524">
|
||
Paucispiral protoconch (
|
||
<emphasis id="B953EADEFFF9E419FE2EF887FE3DF8D8" box="[406,444,1873,1899]" italics="true" pageId="31" pageNumber="524">fide</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF9E419FE7DF884FDE1F8DF" author="LOZOUET P." box="[453,608,1874,1900]" pageId="31" pageNumber="524" pagination="3 - 162" refId="ref46112" refString="LOZOUET P. 2023. - Muricidae (Gastropoda, Neogastropoda) de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana 24: 3 - 162." type="journal article" year="2023">Lozouet 2023</bibRefCitation>
|
||
: text-fig. 10e). Teleoconch up to
|
||
<quantity id="4CDF9B29FFF9E419FEF1F8A7FE19F83F" box="[329,408,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="31" pageNumber="524" unit="mm" value="27.0">27 mm</quantity>
|
||
in height, up to
|
||
<quantity id="4CDF9B29FFF9E419FDE8F8A7FD1EF83F" box="[592,671,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="524" unit="mm" value="12.0">12 mm</quantity>
|
||
in width, biconic in profile, composed of six whorls. Moderately high spire, first two whorls subcarinate; next whorls rounded. Last whorl up to 75% of total length. Apical angle 56°. Spiral sculpture with marked primary cords. First whorl: appearance of primary cords P1 and P2; second whorl: P1 and P2; third whorl: appearance of s1. Fourth whorl: appearance of abis, adis and s2. Fifth whorl: no change or appearance of threads. Sixth whorl: IP, adis and abis, P1 to P6, s1 to s6; s2 and s3 well developed; P6 on base on convex part of whorl; ADP, MP and ABP slightly more developed than ads and ms. Surface of shell with numerous threads appearing on fourth whorl. Axial sculpture with thick varices. On first whorl: nine lamellose varices. From the second to fifth whorl: 9-10 thick varices. On sixth: appearance of three or four varices and one intervarices intercalated. Varices and intervarices thick. From the third to last whorl, numerous, fine microlamellae covering entire surface forming scabrous microsculpture. Ovate aperture up to 25% of diameter and up to 66% of length of last whorl (including siphonal canal). Columellar lip, adherent posteriorly, becoming progressively more erect anteriorly. One or two small columellar tubercles. Parietal lip adherent. Outer lip bearing strong internal denticles: ID, D2 to D6 present, D1 missing. Pseudoumbilicus moderately large. Siphonal canal open, up to 48% of apertural length, slightly dorsally recurved.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E419FC95F964FC46F978" blockId="31.[811,1456,1714,2028]" box="[813,967,1714,1740]" pageId="31" pageNumber="524">
|
||
<smallCapsWord id="8D7EA010FFF9E419FC95F964FC46F978" baselines="1733,1734" box="[813,967,1714,1740]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="31" pageNumber="524">COMPARISONS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF9E427FC95F904FC86FE42" blockId="31.[811,1456,1714,2028]" lastBlockId="33.[130,777,215,1360]" lastPageId="33" lastPageNumber="526" pageId="31" pageNumber="524">
|
||
<bibRefCitation id="EFB64B3DFFF9E419FC95F904FC68F95F" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[813,1001,1746,1772]" pageId="31" pageNumber="524" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF9E419FC95F904FCEDF95F" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[813,876,1746,1772]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF9E419FCC9F905FC25F95F" box="[881,932,1746,1772]" italics="true" pageId="31" pageNumber="524">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
: pl. 90, figs 3-4) attributed with hesitation
|
||
<specimenCount id="9D21FD45FFF9E419FC93F924FC45F8BF" box="[811,964,1778,1804]" count="2" pageId="31" pageNumber="524" type="generic">two specimens</specimenCount>
|
||
from Gaas to
|
||
<taxonomicName id="4C274D4FFFF9E419FBEEF925FCE4F89F" ID-CoL="BM2ZD" authority="d'Orbigny, 1852" authorityName="d'Orbigny" authorityYear="1852" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="subquadrifrons">
|
||
<emphasis id="B953EADEFFF9E419FBEEF925FAB4F8BF" box="[1110,1333,1778,1804]" italics="true" pageId="31" pageNumber="524">Murex subquadrifrons</emphasis>
|
||
d’Orbigny, 1852
|
||
</taxonomicName>
|
||
, originally described as
|
||
<taxonomicName id="4C274D4FFFF9E419FBE5F8C4FA9DF89F" ID-CoL="44MDK" authorityName="Lamarck" authorityYear="1822" box="[1117,1308,1810,1836]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="quadrifrons">
|
||
<emphasis id="B953EADEFFF9E419FBE5F8C4FA9DF89F" box="[1117,1308,1810,1836]" italics="true" pageId="31" pageNumber="524">Murex quadrifrons</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFB64B3DFFF9E419FAF8F8C4FCF7F8FF" author="GRATELOUP J. - P. S. & DE" pageId="31" pageNumber="524" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup (1845)</bibRefCitation>
|
||
from this same Rupelian locality. The Grateloup’s collection is housed in the University of Talence, but the
|
||
<typeStatus id="549C886EFFF9E419FA38F884FA2FF8DF" box="[1408,1454,1874,1900]" pageId="31" pageNumber="524">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C274D4FFFF9E419FC15F8A4FBCFF838" ID-CoL="44MDK" authorityName="Lamarck" authorityYear="1822" box="[941,1102,1905,1931]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="524" phylum="Mollusca" rank="species" species="quadrifrons">
|
||
<emphasis id="B953EADEFFF9E419FC15F8A4FBCFF838" box="[941,1102,1905,1931]" italics="true" pageId="31" pageNumber="524">M. quadrifrons</emphasis>
|
||
</taxonomicName>
|
||
was not found and is likely lost (personal observation of the first author). Therefore, only the drawing of
|
||
<bibRefCitation id="EFB64B3DFFF9E419FC1DF867FBDDF87F" author="GRATELOUP J. - P. S. & DE" box="[933,1116,1969,1996]" pageId="31" pageNumber="524" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup (1845</bibRefCitation>
|
||
: pl. 3 [pl. 31], fig. 10, refigured in
|
||
<bibRefCitation id="EFB64B3DFFF9E419FCF0F807FC7DF858" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[840,1020,2001,2027]" pageId="31" pageNumber="524" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF9E419FCF0F807FC09F858" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[840,904,2001,2027]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF9E419FC35F804FC3EF858" box="[909,959,2001,2027]" italics="true" pageId="31" pageNumber="524">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
: pl. 90, fig. 2) can be used to identify the species. According to
|
||
<bibRefCitation id="EFB64B3DFFC6E426FED2F847FDB2F818" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[362,563,1937,1963]" pageId="32" pageNumber="525" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFC6E426FED2F847FE2BF818" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[362,426,1937,1963]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFC6E426FE09F844FE65F818" box="[433,484,1937,1963]" italics="true" pageId="32" pageNumber="525">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
, the drawing shows a shell with a textiliose surface and a biconic shape, as in the Gaas material at hand. Moreover, to our knowledge, no other muricid from Gaas except
|
||
<emphasis id="B953EADEFFC6E426FBE0F844FBEEF818" box="[1112,1135,1938,1963]" italics="true" pageId="32" pageNumber="525">P.</emphasis>
|
||
(
|
||
<taxonomicName id="4C274D4FFFC6E426FBC1F847FA84F818" ID-CoL="BFGQT" authorityName="Houart" authorityYear="1995" box="[1145,1285,1937,1963]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="525" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFC6E426FBC1F847FA84F818" box="[1145,1285,1937,1963]" italics="true" pageId="32" pageNumber="525">Pterymarchia</emphasis>
|
||
</taxonomicName>
|
||
) is comparable to the specimen illustrated by
|
||
<bibRefCitation id="EFB64B3DFFC6E426FBC0F867FAB8F87F" author="GRATELOUP J. - P. S. & DE" box="[1144,1337,1969,1996]" pageId="32" pageNumber="525" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup (1845)</bibRefCitation>
|
||
. However, the shell illustrated by
|
||
<bibRefCitation id="EFB64B3DFFC6E426FB98F807FB61F85F" author="GRATELOUP J. - P. S. & DE" box="[1056,1248,2001,2028]" pageId="32" pageNumber="525" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup (1845)</bibRefCitation>
|
||
lacks internal denticles within the outer lip, lacks columellar denticles, shows more numerous varices (six on the ventral face vs four in the specimens at hand), has a narrower siphonal canal, and is more likely to represent a specimen of
|
||
<taxonomicName id="4C274D4FFFC7E427FDA9FEEEFDE9FEE2" ID-CoL="7P2BV" authorityName="Jousseaume" authorityYear="1880" box="[529,616,312,337]" class="Gastropoda" family="Muricidae" genus="Favartia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFC7E427FDA9FEEEFDE9FEE2" box="[529,616,312,337]" italics="true" pageId="33" pageNumber="526">Favartia</emphasis>
|
||
</taxonomicName>
|
||
. These numerous differences between
|
||
<bibRefCitation id="EFB64B3DFFC7E427FE2DFE81FDE5FEC1" author="GRATELOUP J. - P. S. & DE" box="[405,612,343,370]" pageId="33" pageNumber="526" pagination="5 - 10" refId="ref44730" refString="GRATELOUP J. - P. S. (DE.) 1845. - Conchyliologie fossile des terrains tertiaires du bassin de l'Adour (environ de Dax). Tome 1. Univalves, Lafargue, Bordeaux: pls 1, 3, 5 - 10, 12 - 48. https: // www. biodiversitylibrary. org / page / 55305392" type="journal article" year="1845">Grateloup’s (1845)</bibRefCitation>
|
||
figure and the studied material suggest that it obviously corresponds to a species other than
|
||
<taxonomicName id="4C274D4FFFC7E427FEFCFE41FD83FE03" ID-CoL="BM2ZD" authorityName="d'Orbigny" authorityYear="1852" box="[324,514,406,432]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="subquadrifrons">
|
||
<emphasis id="B953EADEFFC7E427FEFCFE41FD83FE03" box="[324,514,406,432]" italics="true" pageId="33" pageNumber="526">M. subquadrifrons</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C274D4FFFC7E427FDA8FE41FD70FE03" ID-CoL="BM2ZD" authorityName="d'Orbigny" authorityYear="1852" box="[528,753,406,432]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="subquadrifrons">
|
||
<emphasis id="B953EADEFFC7E427FDA8FE41FD70FE03" box="[528,753,406,432]" italics="true" pageId="33" pageNumber="526">Murex subquadrifrons</emphasis>
|
||
</taxonomicName>
|
||
is therefore not recognizable using the figure of Grateloup, lacks
|
||
<typeStatus id="549C886EFFC7E427FF3AFE01FF30FE42" box="[130,177,471,497]" pageId="33" pageNumber="526">type</typeStatus>
|
||
material and it should be considered a
|
||
<emphasis id="B953EADEFFC7E427FDE7FE01FD7EFE43" box="[607,767,470,496]" italics="true" pageId="33" pageNumber="526">nomen dubium</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF586644FFC6E426FF3CF93CFA30F8E6" ID-DOI="http://doi.org/10.5281/zenodo.13985802" ID-Zenodo-Dep="13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="32" pageNumber="525" startId="32.[132,143,1770,1787]" targetBox="[138,1450,251,1691]" targetPageId="32" targetType="figure">
|
||
<paragraph id="8B9836CCFFC6E426FF3CF93CFA30F8E6" blockId="32.[132,1457,1770,1877]" pageId="32" pageNumber="525">
|
||
<smallCapsWord id="8D7EA010FFC6E426FF3CF93CFF1CF949" baselines="1783,1783" box="[132,157,1770,1787]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="32" pageNumber="525">FIG</smallCapsWord>
|
||
. 18. — Spiral cords and internal denticles of the outer lip in
|
||
<taxonomicName id="4C274D4FFFC6E426FD34F93CFC39F948" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[652,952,1770,1787]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="525" phylum="Mollusca" rank="species" species="gaasensis" status="sp. nov." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC6E426FD34F93CFD67F948" box="[652,742,1770,1787]" italics="true" pageId="32" pageNumber="525">Pterynotus</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC6E426FD48F93CFCDCF948" box="[752,861,1770,1787]" italics="true" pageId="32" pageNumber="525">Pterymarchia</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC6E426FCDEF93CFC39F948" box="[870,952,1770,1787]" italics="true" pageId="32" pageNumber="525">gaasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC6E426FC04F93CFC6DF948" box="[956,1004,1770,1787]" pageId="32" pageNumber="525" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B953EADEFFC6E426FC4CF93CFB96F948" bold="true" box="[1012,1047,1770,1787]" pageId="32" pageNumber="525">A -D</emphasis>
|
||
),
|
||
<taxonomicName id="4C274D4FFFC6E426FB9DF93CFB42F948" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1061,1219,1770,1787]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="525" phylum="Mollusca" rank="species" species="pelouatensis" status="n. sp." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC6E426FB9DF93CFBB9F948" box="[1061,1080,1770,1787]" italics="true" pageId="32" pageNumber="525">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC6E426FBF8F93CFBD2F948" box="[1088,1107,1770,1787]" italics="true" pageId="32" pageNumber="525">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC6E426FBE3F93CFB42F948" box="[1115,1219,1770,1787]" italics="true" pageId="32" pageNumber="525">pelouatensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC6E426FB70F93CFB79F948" box="[1224,1272,1770,1787]" pageId="32" pageNumber="525" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B953EADEFFC6E426FAB8F93CFAA0F948" bold="true" box="[1280,1313,1770,1787]" pageId="32" pageNumber="525">E -H</emphasis>
|
||
) and
|
||
<emphasis id="B953EADEFFC6E426FAF7F93CFAE3F948" box="[1359,1378,1770,1787]" italics="true" pageId="32" pageNumber="525">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC6E426FAD2F93CFAF7F948" box="[1386,1398,1770,1787]" italics="true" pageId="32" pageNumber="525">P</emphasis>
|
||
)
|
||
<taxonomicName id="4C274D4FFFC6E426FA38F93CFEBBF8A2" pageId="32" pageNumber="525">
|
||
<emphasis id="B953EADEFFC6E426FA38F93CFF3FF8A2" italics="true" pageId="32" pageNumber="525">sandbergeri</emphasis>
|
||
(HÖrnes,1856)
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B953EADEFFC6E426FEFBF8D6FEDAF8A2" bold="true" box="[323,347,1792,1809]" pageId="32" pageNumber="525">I -L</emphasis>
|
||
):
|
||
<emphasis id="B953EADEFFC6E426FED1F8D6FEF7F8A2" bold="true" box="[361,374,1792,1809]" pageId="32" pageNumber="525">A</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFC6E426FEC7F8D6FE0DF8A2" bold="true" box="[383,396,1792,1809]" pageId="32" pageNumber="525">B</emphasis>
|
||
, holotype,MNHN.F.A32492 (
|
||
<taxonomicName id="4C274D4FFFC6E426FDC4F8D6FD2AF8A2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[636,683,1792,1809]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian);
|
||
<emphasis id="B953EADEFFC6E426FAEDF8D6FAE2F8A2" bold="true" box="[1365,1379,1792,1809]" pageId="32" pageNumber="525">C</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFC6E426FAD4F8D6FAFBF8A2" bold="true" box="[1388,1402,1792,1809]" pageId="32" pageNumber="525">D</emphasis>
|
||
, paratype, MNHN.F.A90554 (
|
||
<taxonomicName id="4C274D4FFFC6E426FEF1F8C1FEF9F89B" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[329,376,1815,1832]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.), Gaas (Espibos);
|
||
<emphasis id="B953EADEFFC6E426FD8BF8C1FDBEF89B" bold="true" box="[563,575,1815,1832]" pageId="32" pageNumber="525">E</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFC6E426FDF1F8C1FDD5F89B" bold="true" box="[585,596,1815,1832]" pageId="32" pageNumber="525">F</emphasis>
|
||
, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian);
|
||
<emphasis id="B953EADEFFC6E426FA24F8C1FA2BF89B" bold="true" box="[1436,1450,1815,1832]" pageId="32" pageNumber="525">G</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFC6E426FF3CF8F8FF13F88C" bold="true" box="[132,146,1838,1855]" pageId="32" pageNumber="525">H</emphasis>
|
||
, paratype, MNHN.F.J17632, Saucats (Peloua);
|
||
<emphasis id="B953EADEFFC6E426FDA0F8F8FD9FF88C" bold="true" box="[536,542,1838,1855]" pageId="32" pageNumber="525">I</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFC6E426FD9FF8F8FDB0F88C" bold="true" box="[551,561,1838,1855]" pageId="32" pageNumber="525">J</emphasis>
|
||
, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian);
|
||
<emphasis id="B953EADEFFC6E426FF3CF892FF10F8E6" bold="true" box="[132,145,1860,1877]" pageId="32" pageNumber="525">K</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFC6E426FF24F892FF26F8E6" bold="true" box="[156,167,1860,1877]" pageId="32" pageNumber="525">L</emphasis>
|
||
, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9836CCFFC7E427FF23FE2FFE65FAFC" blockId="33.[130,777,215,1360]" pageId="33" pageNumber="526">
|
||
Eocene
|
||
<taxonomicName id="4C274D4FFFC7E427FF49FE2FFD84FDA0" authorityName="Houart" authorityYear="1995" box="[241,517,504,531]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="subGenus" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FF49FE2FFEDFFDA1" box="[241,350,505,530]" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FED7FE2EFE7AFDA1" box="[367,507,504,530]" italics="true" pageId="33" pageNumber="526">Pterymarchia</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="4C274D4FFFC7E427FDDFFE2FFEA3FD81" authority="(Merle 1990)" baseAuthorityName="Merle" baseAuthorityYear="1990" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="antiquus" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FDDFFE2FFDFFFDA1" box="[615,638,505,530]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FD31FE2FFD1DFDA1" box="[649,668,505,530]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FD13FE2FFC86FDA1" box="[683,775,505,530]" italics="true" pageId="33" pageNumber="526">antiquus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFC7E427FF36FDCEFE9BFD81" author="MERLE D." box="[142,282,536,562]" pageId="33" pageNumber="526" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle 1990</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFC7E427FE8EFDCFFD53FD80" ID-CoL="9FYYF" authority="(Deshayes, 1835)" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[310,722,536,563]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="denudatus" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FE8EFDCFFECFFD81" box="[310,334,537,562]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FEE5FDCFFEF0FD81" box="[349,369,537,562]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FE3DFDCEFE79FD81" box="[389,504,536,562]" italics="true" pageId="33" pageNumber="526">denudatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFC7E427FDB3FDCEFD49FD80" author="DESHAYES G. - P." box="[523,712,536,563]" pageId="33" pageNumber="526" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFC7E427FF3CFDEFFE5EFDE1" ID-CoL="BM3FH" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[132,479,568,594]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="defensus" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FF3CFDEFFF1DFDE1" box="[132,156,569,594]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FF14FDEFFF41FDE1" box="[172,192,569,594]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FF6CFDEEFEAFFDE1" box="[212,302,568,594]" italics="true" pageId="33" pageNumber="526">defensus</emphasis>
|
||
(Fuchs, 1870)
|
||
</taxonomicName>
|
||
strongly differ from the
|
||
<taxonomicName id="4C274D4FFFC7E427FF3CFD8FFE9FFDC1" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[132,286,600,626]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="gaasensis" status="n. sp." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FF3CFD8FFF19FDC1" box="[132,152,601,626]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FF1CFD8FFF36FDC1" box="[164,183,601,626]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FF7CFD8FFE9FFDC1" box="[196,286,601,626]" italics="true" pageId="33" pageNumber="526">gaasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC7E427FE9CFD8FFEDFFDC0" box="[292,350,601,627]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
|
||
in having trivaricate sculpture.
|
||
<taxonomicName id="4C274D4FFFC7E427FD15FD8FFD9CFD21" authority="(von Koenen, 1889)" baseAuthorityName="von Koenen" baseAuthorityYear="1889" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="fascistria" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FD15FD8FFF23FD21" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FF0CFDAFFF46FD21" box="[180,199,633,658]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FF60FDAEFEB2FD21" box="[216,307,632,658]" italics="true" pageId="33" pageNumber="526">fascistria</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFC7E427FEFBFDAEFD94FD21" author="VON KOENEN A." box="[323,533,632,658]" pageId="33" pageNumber="526" pagination="1 - 280" refId="ref49077" refString="VON KOENEN A. 1889. - Das Norddeutsche Unter-Oligocan und seine Mollusken-Fauna. Abhandlungen zur geologischen Specialkarte von Preussen und den Thuringischen Staaten 10 (1): 1 - 280." type="journal article" year="1889">von Koenen, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Priabonian of North
|
||
<collectingCountry id="F330765CFFC7E427FF56FD4EFED5FD01" box="[238,340,664,690]" name="Germany" pageId="33" pageNumber="526">Germany</collectingCountry>
|
||
is more similar shape, as it lacks trivaricate sculpture and because their primary cords are poorly distinguishable from the secondary cords on the last whorls.
|
||
<taxonomicName id="4C274D4FFFC7E427FF3BFD2EFEFDFCA1" baseAuthorityName="von Koenen" baseAuthorityYear="1889" box="[131,380,760,786]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="fascistria" subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FF3BFD2EFF71FCA2" box="[131,240,760,785]" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FEB8FD2EFE92FCA2" box="[256,275,760,785]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FE99FD2EFEFDFCA1" box="[289,380,760,786]" italics="true" pageId="33" pageNumber="526">fascistria</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C274D4FFFC7E427FDD2FD2EFC87FCA2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[618,774,760,786]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="gaasensis" status="n. sp." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FDD2FD2EFD00FCA2" box="[618,641,760,785]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FD34FD2EFD1EFCA2" box="[652,671,760,785]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FD14FD2EFC87FCA2" box="[684,774,760,785]" italics="true" pageId="33" pageNumber="526">gaasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC7E427FF3CFCCEFF43FC81" box="[132,194,792,818]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
|
||
in having a lower spire and by lacking intervarices, D1 and ID denticles.
|
||
<taxonomicName id="4C274D4FFFC7E427FEC1FCEEFDF2FCE2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[377,627,823,849]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="gaasensis" status="sp. nov." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FEC1FCEEFE67FCE2" box="[377,486,824,849]" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FE4EFCEEFD88FCE2" box="[502,521,824,849]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FDA1FCEEFDF2FCE2" box="[537,627,824,849]" italics="true" pageId="33" pageNumber="526">gaasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC7E427FDC5FCEEFD38FCE1" box="[637,697,824,850]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
|
||
can be compared to two other Neogene species:
|
||
<taxonomicName id="4C274D4FFFC7E427FDF0FC8EFC86FCC2" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[584,775,855,881]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="pelouatensis" status="n. sp." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FDF0FC8EFDDEFCC2" box="[584,607,856,881]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FDD2FC8EFDFCFCC2" box="[618,637,856,881]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FD34FC81FC86FCC2" box="[652,775,855,881]" italics="true" pageId="33" pageNumber="526">pelouatensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC7E427FF3CFCAEFF43FC21" box="[132,194,888,914]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
|
||
from the Burdigalian of Aquitaine Basin (
|
||
<figureCitation id="131C2A49FFC7E427FD25FCA1FD6CFC22" box="[669,749,887,913]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="33" pageNumber="526">Figs 18</figureCitation>
|
||
EH; 27G, H) and
|
||
<taxonomicName id="4C274D4FFFC7E427FEF5FC4EFD46FC01" authority="(Hornes, 1856)" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" baseAuthorityName="Hornes" baseAuthorityYear="1856" box="[333,711,919,946]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
|
||
<emphasis id="B953EADEFFC7E427FEF5FC4EFEE4FC02" box="[333,357,920,945]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FEC8FC4EFE02FC02" box="[368,387,920,945]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FE2DFC41FD8EFC02" box="[405,527,919,945]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
|
||
(Hörnes, 1856)
|
||
</taxonomicName>
|
||
from the Langhian of the Paratethys. Both species differs from
|
||
<taxonomicName id="4C274D4FFFC7E427FF3EFC0EFEA3FC42" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[134,290,983,1009]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="gaasensis" status="n. sp." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FF3EFC0EFF1CFC42" box="[134,157,984,1009]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FF1EFC0EFF38FC42" box="[166,185,984,1009]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FF70FC0EFEA3FC42" box="[200,290,984,1009]" italics="true" pageId="33" pageNumber="526">gaasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC7E427FE92FC01FEE7FC42" box="[298,358,983,1009]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
|
||
in having more angular whorls with a shoulder delimited by a subspiny P1 and by other primary cords more distinguishable from the secondary cords on the last whorl.
|
||
<bibRefCitation id="EFB64B3DFFC7E427FE9EFBE1FE51FBE2" author="LOZOUET P." box="[294,464,1079,1105]" pageId="33" pageNumber="526" pagination="3 - 162" refId="ref46112" refString="LOZOUET P. 2023. - Muricidae (Gastropoda, Neogastropoda) de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana 24: 3 - 162." type="journal article" year="2023">Lozouet (2023)</bibRefCitation>
|
||
followed
|
||
<bibRefCitation id="EFB64B3DFFC7E427FD83FBE0FC86FBE2" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[571,775,1078,1105]" pageId="33" pageNumber="526" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFC7E427FD83FBE0FDFAFBE3" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[571,635,1078,1104]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFC7E427FD3AFBE1FD37FBE3" box="[642,694,1078,1104]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
and adopted the name
|
||
<taxonomicName id="4C274D4FFFC7E427FE3BFB81FD37FBC3" ID-CoL="BM3KY" baseAuthorityName="A. d'Orbigny" baseAuthorityYear="1852" box="[387,694,1110,1136]" class="Gastropoda" family="Muricidae" genus="Pterynotus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="subquadrifrons">
|
||
<emphasis id="B953EADEFFC7E427FE3BFB81FE71FBC3" box="[387,496,1111,1136]" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
|
||
cf.
|
||
<emphasis id="B953EADEFFC7E427FDA2FB80FD37FBC3" box="[538,694,1110,1136]" italics="true" pageId="33" pageNumber="526">subquadrifrons</emphasis>
|
||
</taxonomicName>
|
||
for the Rupelian material from Gaas, but he did not include Chattian and Aquitanian specimens figured in
|
||
<bibRefCitation id="EFB64B3DFFC7E427FDF8FB40FD7EFB03" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[576,767,1174,1200]" pageId="33" pageNumber="526" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFC7E427FDF8FB40FD01FB03" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[576,640,1174,1200]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFC7E427FD3FFB41FD38FB03" box="[647,697,1174,1200]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
: pl. 90, figs 5, 6), because Rupelian shells have a paucispiral protoconch, whereas the Chattian/Aquitanian species has a multispiral protoconch, and is therefore a separate species. Thus, we consider for instance that
|
||
<taxonomicName id="4C274D4FFFC7E427FDB1FAC0FD26FA9C" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[521,679,1302,1328]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="gaasensis" status="n. sp." subGenus="Pterymarchia">
|
||
<emphasis id="B953EADEFFC7E427FDB1FAC0FDA1FA9C" box="[521,544,1302,1327]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EADEFFC7E427FD93FAC0FDBFFA9C" box="[555,574,1302,1327]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EADEFFC7E427FDF5FAC0FD26FA9C" box="[589,679,1302,1327]" italics="true" pageId="33" pageNumber="526">gaasensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFC7E427FD16FAC0FD6BFA83" box="[686,746,1302,1328]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
|
||
to be present only in the Rupelian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |