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<document id="D3F8210B4C31A863EB940A27C8A321CA" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFF1E41FFE8EFA21FEE9FBDC" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Timbellus magnei" docType="treatment" docVersion="8" lastPageNumber="517" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="516" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="EC77AFE8F9837654BB76E24532788695">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="EBD1653731748B77C582FE13212EA3D6">Ledon, Daniel</mods:namePart>
<mods:affiliation id="712A6ABA0EA6C984545C7397DCA335C8">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:affiliation id="2E41DF6C09A66255D37480416E03BBA6">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<taxonomicName id="4C274D4FFFF1E411FE8EFA21FE76F9BC" ID-CoL="CN5MQ" box="[310,503,1525,1553]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFF1E411FE8EFA21FE76F9BC" bold="true" box="[310,503,1525,1553]" italics="true" pageId="23" pageNumber="516">Timbellus magnei</emphasis>
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n. name
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<paragraph id="8B9836CCFFF1E411FEE0F9C0FDB2F983" blockId="23.[310,597,1525,1584]" box="[344,563,1557,1584]" pageId="23" pageNumber="516">
(
<figureCitation id="131C2A49FFF1E411FED9F9C3FE2EF983" box="[353,431,1557,1584]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="23" pageNumber="516">Figs 12</figureCitation>
A-F; 26G-I)
</paragraph>
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<subSubSection id="C33D6547FFF1E411FF3CF998FEF8F858" pageId="23" pageNumber="516" type="reference_group">
<paragraph id="8B9836CCFFF1E411FF3CF998FEEFF9CC" blockId="23.[132,774,1613,1664]" pageId="23" pageNumber="516">
<treatmentCitationGroup id="AB3711E2FFF1E411FF3CF998FD10F9D6" box="[132,657,1613,1637]" pageId="23" pageNumber="516">
<treatmentCitation id="0A8610DDFFF1E411FF3CF998FE5CF9D6" author="ROUAULT A." box="[132,477,1613,1637]" page="204" pageId="23" pageNumber="516" year="1850">
<taxonomicName id="4C274D4FFFF1E411FF3CF998FE5CF9D6" ID-CoL="CM8R7" authority="Rouault, 1850: 204" authorityName="sensu Rouault" authorityPageNumber="204" authorityYear="1850" box="[132,477,1613,1637]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FF3CF998FE92F9D6" box="[132,275,1614,1637]" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FEA2F99BFE5CF9D6" author="ROUAULT A." box="[282,477,1613,1637]" pageId="23" pageNumber="516" pagination="457 - 502" refId="ref48421" refString="ROUAULT A. 1850. - Description des fossiles eocenes des environs de Pau. Memoires de la Societe geologique de France, serie 2, 3: 457 - 502." type="journal article" year="1850">Rouault, 1850: 204</bibRefCitation>
</taxonomicName>
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, pl. XVII, fig. 17a
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[
<emphasis id="B953EADEFFF1E411FD27F998FD42F9D6" box="[671,707,1614,1637]" italics="true" pageId="23" pageNumber="516">non</emphasis>
<taxonomicName id="4C274D4FFFF1E411FD71F998FEE1F933" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FD71F998FF4EF9CC" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FF6DF9B1FEE1F933" author="GMELIN J. F." box="[213,352,1639,1664]" pageId="23" pageNumber="516" pagination="3021 - 3910" refId="ref44669" refString="GMELIN J. F. 1791 (ed.). - Caroli a Linnaei Systema Naturae per Regna Tria Naturae, part Vermes. Ed. 13. Tome 1 (6). G. E. Beer, Leipzig: 3021 - 3910. https: // doi. org / 10.5962 / bhl. title. 36932" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
</taxonomicName>
].
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<paragraph id="8B9836CCFFF1E411FF3CF941FF6DF97B" blockId="23.[132,775,1686,1736]" pageId="23" pageNumber="516">
<treatmentCitationGroup id="AB3711E2FFF1E411FF3CF941FE7AF91D" box="[132,507,1686,1710]" pageId="23" pageNumber="516">
<taxonomicName id="4C274D4FFFF1E411FF3CF941FEB9F91D" box="[132,312,1687,1710]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FF3CF941FEB9F91D" box="[132,312,1687,1710]" italics="true" pageId="23" pageNumber="516">Pterynotus trigonus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8610DDFFF1E411FEEDF940FE7AF91D" author="GLIBERT M." box="[341,507,1686,1710]" page="11" pageId="23" pageNumber="516" year="1963">
<bibRefCitation id="EFB64B3DFFF1E411FEEDF940FE7AF91D" author="GLIBERT M." box="[341,507,1686,1710]" pageId="23" pageNumber="516" pagination="1 - 179" refId="ref44626" refString="GLIBERT M. 1963. - Les Muricacea et les Buccinacea du Cenozoique etranger des collections de l'Institut royal des Sciences naturelles de Belgique. Memoires de l'Institut royal des Sciences naturelles de Belgique serie 2, 74: 1 - 179." type="journal article" year="1963">Glibert 1963: 11</bibRefCitation>
</treatmentCitation>
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[
<emphasis id="B953EADEFFF1E411FDB3F941FDAEF91D" box="[523,559,1687,1710]" italics="true" pageId="23" pageNumber="516">non</emphasis>
<taxonomicName id="4C274D4FFFF1E411FD8FF941FF5FF97B" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FD8FF941FD46F91D" box="[567,711,1687,1710]" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FD76F940FF5FF97B" author="GMELIN J. F." pageId="23" pageNumber="516" pagination="3021 - 3910" refId="ref44669" refString="GMELIN J. F. 1791 (ed.). - Caroli a Linnaei Systema Naturae per Regna Tria Naturae, part Vermes. Ed. 13. Tome 1 (6). G. E. Beer, Leipzig: 3021 - 3910. https: // doi. org / 10.5962 / bhl. title. 36932" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="8B9836CCFFF1E411FF3CF909FD94F8A2" blockId="23.[131,774,1758,1809]" pageId="23" pageNumber="516">
<treatmentCitationGroup id="AB3711E2FFF1E411FF3CF909FE38F945" box="[132,441,1759,1782]" pageId="23" pageNumber="516">
<taxonomicName id="4C274D4FFFF1E411FF3CF909FE29F945" box="[132,424,1759,1782]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="subSpecies" species="tricarinatus" subSpecies="trigonus">
<emphasis id="B953EADEFFF1E411FF3CF909FE29F945" box="[132,424,1759,1782]" italics="true" pageId="23" pageNumber="516">Pterynotus tricarinatus trigonus</emphasis>
</taxonomicName>
–
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<treatmentCitation id="0A8610DDFFF1E411FE05F909FDE2F945" author="MERLE D." box="[445,611,1758,1783]" page="158" pageId="23" pageNumber="516" year="1990">
<bibRefCitation id="EFB64B3DFFF1E411FE05F909FDE2F945" author="MERLE D." box="[445,611,1758,1783]" pageId="23" pageNumber="516" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle 1990: 158</bibRefCitation>
</treatmentCitation>
, pl. 2, figs 13, 14 only [
<emphasis id="B953EADEFFF1E411FF06F92CFF63F8A2" box="[190,226,1786,1809]" italics="true" pageId="23" pageNumber="516">non</emphasis>
<emphasis id="B953EADEFFF1E411FF50F92CFEF6F8A2" box="[232,375,1786,1809]" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FEC5F92FFD86F8A2" author="GMELIN J. F." box="[381,519,1785,1809]" pageId="23" pageNumber="516" pagination="3021 - 3910" refId="ref44669" refString="GMELIN J. F. 1791 (ed.). - Caroli a Linnaei Systema Naturae per Regna Tria Naturae, part Vermes. Ed. 13. Tome 1 (6). G. E. Beer, Leipzig: 3021 - 3910. https: // doi. org / 10.5962 / bhl. title. 36932" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
].
</paragraph>
<paragraph id="8B9836CCFFF1E411FF3CF8FEFEA6F8EA" blockId="23.[132,775,1831,1882]" pageId="23" pageNumber="516">
<treatmentCitationGroup id="AB3711E2FFF1E411FF3CF8FEFEA6F8EA" pageId="23" pageNumber="516">
<taxonomicName id="4C274D4FFFF1E411FF3CF8FEFED9F88C" ID-CoL="8ZHC9" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[132,344,1831,1855]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF1E411FF3CF8FEFED9F88C" box="[132,344,1831,1855]" italics="true" pageId="23" pageNumber="516">Pterynotus tripteroides</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8610DDFFF1E411FECFF8F1FDA3F88C" author="MERLE D." box="[375,546,1831,1855]" page="156" pageId="23" pageNumber="516" year="1990">
<bibRefCitation id="EFB64B3DFFF1E411FECFF8F1FDA3F88C" author="MERLE D." box="[375,546,1831,1855]" pageId="23" pageNumber="516" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle 1990: 156</bibRefCitation>
</treatmentCitation>
, pl. 3, fig. 1 only [
<emphasis id="B953EADEFFF1E411FD5BF8FEFC86F88C" box="[739,775,1832,1855]" italics="true" pageId="23" pageNumber="516">non</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FF3CF894FE98F8E9" author="LAMARCK J. - B." box="[132,281,1858,1882]" pageId="23" pageNumber="516" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
].
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</paragraph>
<paragraph id="8B9836CCFFF1E411FF3CF8A6FED8F811" blockId="23.[132,775,1904,1955]" pageId="23" pageNumber="516">
<treatmentCitationGroup id="AB3711E2FFF1E411FF3CF8A6FED8F811" pageId="23" pageNumber="516">
<taxonomicName id="4C274D4FFFF1E411FF3CF8A6FECFF83B" ID-CoL="8YPF3" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[132,334,1904,1928]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF1E411FF3CF8A6FECFF83B" box="[132,334,1904,1928]" italics="true" pageId="23" pageNumber="516">Timbellus tripteroides</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8610DDFFF1E411FEDFF8A6FDB4F83B" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[359,565,1904,1928]" page="44" pageId="23" pageNumber="516" year="2011">
<bibRefCitation id="EFB64B3DFFF1E411FEDFF8A6FDB4F83B" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[359,565,1904,1928]" pageId="23" pageNumber="516" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
Merle
<emphasis id="B953EADEFFF1E411FE1EF8A7FE54F83B" box="[422,469,1904,1928]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
2011: 44
</bibRefCitation>
</treatmentCitation>
, pl. 100 figs 3, 4 only [
<emphasis id="B953EADEFFF1E411FF34F85DFF31F811" box="[140,176,1931,1954]" italics="true" pageId="23" pageNumber="516">non</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FF0EF85DFECAF810" author="LAMARCK J. - B." box="[182,331,1931,1955]" pageId="23" pageNumber="516" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9836CCFFF1E411FF3CF86FFEF8F858" blockId="23.[132,775,1977,2028]" pageId="23" pageNumber="516">
<treatmentCitationGroup id="AB3711E2FFF1E411FF3CF86FFEF8F858" pageId="23" pageNumber="516">
<taxonomicName id="4C274D4FFFF1E411FF3CF86FFE12F862" box="[132,403,1977,2001]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="subSpecies" species="crenulatus" subSpecies="palensis">
<emphasis id="B953EADEFFF1E411FF3CF86FFE12F862" box="[132,403,1977,2001]" italics="true" pageId="23" pageNumber="516">Timbellus crenulatus palensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8610DDFFF1E411FE13F86FFD07F862" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[427,646,1977,2001]" page="440" pageId="23" pageNumber="516" year="2011">
<bibRefCitation id="EFB64B3DFFF1E411FE13F86FFD07F862" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[427,646,1977,2001]" pageId="23" pageNumber="516" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
Merle
<emphasis id="B953EADEFFF1E411FE53F86CFD98F862" box="[491,537,1977,2001]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
2011: 440
</bibRefCitation>
</treatmentCitation>
, pl. 98, fig. 1 only [
<emphasis id="B953EADEFFF1E411FF06F802FF63F858" box="[190,226,2004,2027]" italics="true" pageId="23" pageNumber="516">non</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FF50F802FEEAF858" author="MAGNE A." box="[232,363,2003,2028]" pageId="23" pageNumber="516" pagination="28 - 36" refId="ref46270" refString="MAGNE A. 1941. - Sur quelques especes nouvelles ou interessantes provenant du Calcaire a Asteries de la Souys. Proces-verbaux de la Societe linneenne de Bordeaux 91: 28 - 36." type="journal article" year="1941">Magne, 1941</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF1E411FC95FF07FB08FCDD" pageId="23" pageNumber="516" type="materials_examined">
<paragraph id="8B9836CCFFF1E411FC95FF07FB9FFEE0" blockId="23.[811,1455,209,339]" pageId="23" pageNumber="516">
<materialsCitation id="3B4F3C91FFF1E411FC95FF07FB9FFEE0" ID-GBIF-Occurrence="4958671311" collectorName="de Gan &amp; Seaman" country="France" location="Marnes de Gan" pageId="23" pageNumber="516" specimenCount="1" stateProvince="Aquitaine" typeStatus="neotype">
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<smallCapsWord id="8D7EA010FFF1E411FCDCFF00FC43FF54" baselines="227" box="[868,962,214,231]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="23" pageNumber="516">MATERIAL</smallCapsWord>
. —
<emphasis id="B953EADEFFF1E411FC48FF07FB12FF5A" bold="true" box="[1008,1171,209,233]" pageId="23" pageNumber="516">
<typeStatus id="549C886EFFF1E411FC48FF07FBC6FF5A" box="[1008,1095,209,233]" pageId="23" pageNumber="516" type="neotype">Neotype</typeStatus>
.
<collectingCountry id="F330765CFFF1E411FBE9FF07FB12FF5A" box="[1105,1171,209,233]" name="France" pageId="23" pageNumber="516">France</collectingCountry>
</emphasis>
•
<collectingRegion id="49E3F82EFFF1E411FB12FF07FA8BFF5A" box="[1194,1290,209,233]" country="France" name="Aquitaine" pageId="23" pageNumber="516">Aquitaine</collectingRegion>
Basin, Pyrénées-Atlantiques, Bos d’Arros; uppermost part of the
<location id="8EF86017FFF1E411FB57FF3AFA00FEB0" LSID="urn:lsid:plazi:treatment:038E87DAFFF1E41FFE8EFA21FEE9FBDC:8EF86017FFF1E411FB57FF3AFA00FEB0" box="[1263,1409,235,259]" country="France" name="Marnes de Gan" pageId="23" pageNumber="516" stateProvince="Aquitaine">
Marnes
<collectorName id="26D2531AFFF1E411FA84FF3DFA00FEB0" box="[1340,1409,235,259]" pageId="23" pageNumber="516">de Gan</collectorName>
</location>
only (top biozone N12/base biozone NP13); Early Eocene (Ypresian); BMNH 69802 (
<collectorName id="26D2531AFFF1E411FC6CFEF6FBA2FE8B" box="[980,1059,288,312]" pageId="23" pageNumber="516">Seaman</collectorName>
coll.) designated by
<bibRefCitation id="EFB64B3DFFF1E411FB4FFEF6FA2AFE8B" author="MERLE D." box="[1271,1451,288,312]" pageId="23" pageNumber="516" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
<taxonomicName id="4C274D4FFFF1E411FB4FFEF6FAB0FE8B" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1271,1329,288,312]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1990: 158
</bibRefCitation>
, pl. 2, fig. 14) (
<figureCitation id="131C2A49FFF1E411FC03FEECFB92FEE0" box="[955,1043,314,339]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="23" pageNumber="516">Fig. 12A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9836CCFFF1E411FC95FEBCFC55FD73" blockId="23.[812,1456,362,704]" pageId="23" pageNumber="516">
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<smallCapsWord id="8D7EA010FFF1E411FC10FEA6FB83FE32" baselines="381" box="[936,1026,368,385]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="23" pageNumber="516">MATERIAL</smallCapsWord>
. —
<materialsCitation id="3B4F3C91FFF1E411FB90FEBCFA2EFE5F" ID-GBIF-Occurrence="4958671378" pageId="23" pageNumber="516">
<emphasis id="B953EADEFFF1E411FB90FEBCFBE9FE31" bold="true" box="[1064,1128,362,386]" pageId="23" pageNumber="516">
<collectingCountry id="F330765CFFF1E411FB90FEBCFBE9FE31" box="[1064,1128,362,386]" name="France" pageId="23" pageNumber="516">France</collectingCountry>
</emphasis>
• 1 spm
<collectingRegion id="49E3F82EFFF1E411FB00FEBCFA95FE31" box="[1208,1300,362,386]" country="France" name="Aquitaine" pageId="23" pageNumber="516">Aquitaine</collectingRegion>
Basin, Pyrénées-Atlantiques, Gan (Tuilerie); uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13); Early Eocene (Ypresian, uppermost part of the Marnes de Gan);
<collectionCode id="ED36AE09FFF1E411FB4FFE6CFACAFE62" box="[1271,1355,442,465]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.A32506 (
<taxonomicName id="4C274D4FFFF1E411FC8BFE02FCEFFE5F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[819,878,468,492]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), figured specimen of
<bibRefCitation id="EFB64B3DFFF1E411FBC8FE02FA9DFE5F" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[1136,1308,468,492]" pageId="23" pageNumber="516" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFF1E411FBC8FE02FB28FE5F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1136,1193,468,492]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFF1E411FB17FE03FB5CFE5F" box="[1199,1245,468,492]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
(2011
</bibRefCitation>
: pl. 100, fig. 3)
</materialsCitation>
•
<materialsCitation id="3B4F3C91FFF1E411FC85FE39FAD7FD92" ID-GBIF-Occurrence="4958671358" pageId="23" pageNumber="516">
1 spm; idem;
<collectionCode id="ED36AE09FFF1E411FC7CFE39FB98FDB5" box="[964,1049,495,518]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.A32508 (
<taxonomicName id="4C274D4FFFF1E411FB32FE38FB44FDB5" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1162,1221,494,518]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), figured specimen of
<bibRefCitation id="EFB64B3DFFF1E411FCFFFDDFFC77FD92" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[839,1014,521,545]" pageId="23" pageNumber="516" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFF1E411FCFFFDDFFC01FD92" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[839,896,521,545]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFF1E411FC3EFDDCFC34FD92" box="[902,949,521,545]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
(2011
</bibRefCitation>
: pl. 100, fig. 1) (
<figureCitation id="131C2A49FFF1E411FB25FDDFFA99FD93" box="[1181,1304,521,545]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="23" pageNumber="516">Figs 12C, D</figureCitation>
;
<figureCitation id="131C2A49FFF1E411FA9AFDDFFACEFD92" box="[1314,1359,521,545]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="23" pageNumber="516">26G</figureCitation>
)
</materialsCitation>
•
<materialsCitation id="3B4F3C91FFF1E411FAD5FDDFFC77FDE5" ID-GBIF-Occurrence="4958671384" pageId="23" pageNumber="516">
1 spm; idem;
<collectionCode id="ED36AE09FFF1E411FCD4FDF2FC40FD88" box="[876,961,548,571]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.R61437,
<taxonomicName id="4C274D4FFFF1E411FB8CFDF5FBECFD88" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1076,1133,547,571]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll., figured specimen of
<bibRefCitation id="EFB64B3DFFF1E411FACFFDF5FCEBFDE5" author="MERLE D." pageId="23" pageNumber="516" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
<taxonomicName id="4C274D4FFFF1E411FACFFDF5FA31FD88" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1399,1456,547,571]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1990
</bibRefCitation>
: pl. 3, fig. 1)
</materialsCitation>
•
<materialsCitation id="3B4F3C91FFF1E411FBAAFDE8FCE1FDC3" ID-GBIF-Occurrence="4958671302" pageId="23" pageNumber="516">
1 spm; idem;
<collectionCode id="ED36AE09FFF1E411FB1DFDE8FB7AFDE6" box="[1189,1275,574,597]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.A24115 (
<taxonomicName id="4C274D4FFFF1E411FACAFDE8FA2EFDE5" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1394,1455,574,598]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.)
</materialsCitation>
•
<materialsCitation id="3B4F3C91FFF1E411FCC0FD8EFCF3FD38" ID-GBIF-Occurrence="4958671391" pageId="23" pageNumber="516">
2 spm; idem;
<collectionCode id="ED36AE09FFF1E411FC47FD8FFBD2FDC3" box="[1023,1107,601,624]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.A72443 (Varone coll.) (
<figureCitation id="131C2A49FFF1E411FAEBFD8EFCBCFD39" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="23" pageNumber="516">Figs 12E, F</figureCitation>
;
<figureCitation id="131C2A49FFF1E411FCF0FDA5FCEBFD38" box="[840,874,627,651]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="23" pageNumber="516">26I</figureCitation>
)
</materialsCitation>
•
<materialsCitation id="3B4F3C91FFF1E411FC32FDA5FADBFD38" ID-GBIF-Occurrence="4958671381" box="[906,1370,627,651]" pageId="23" pageNumber="516">
35 spm; idem;
<collectionCode id="ED36AE09FFF1E411FB99FDA5FBF4FD39" box="[1057,1141,627,650]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.A24114 (
<taxonomicName id="4C274D4FFFF1E411FB5FFDA5FAA3FD38" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1255,1314,627,651]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.)
</materialsCitation>
•
<materialsCitation id="3B4F3C91FFF1E411FACAFDA5FC55FD73" ID-GBIF-Occurrence="4958671334" pageId="23" pageNumber="516">
1 spm (juvenile with protoconch); idem;
<collectionCode id="ED36AE09FFF1E411FBC0FD58FB4CFD16" box="[1144,1229,654,677]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="516">MNHN</collectionCode>
.F.A90545 (
<taxonomicName id="4C274D4FFFF1E411FA86FD5BFAF8FD16" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1342,1401,653,677]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.) (
<figureCitation id="131C2A49FFF1E411FC8DFD7EFC0DFD0C" box="[821,908,680,704]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="23" pageNumber="516">Figs 12B</figureCitation>
;
<figureCitation id="131C2A49FFF1E411FC2FFD7EFC46FD73" box="[919,967,680,704]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="23" pageNumber="516">26H</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9836CCFFF1E411FC95FD0EFBE5FC97" blockId="23.[812,1455,727,804]" pageId="23" pageNumber="516">
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<typeStatus id="549C886EFFF1E411FC95FD0EFCDEFD5E" box="[813,863,728,751]" pageId="23" pageNumber="516">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFF1E411FCDFFD0BFC42FD5D" baselines="746" box="[871,963,733,750]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="23" pageNumber="516">HORIZON</smallCapsWord>
. — Uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13), see
<bibRefCitation id="EFB64B3DFFF1E411FB6DFD24FA2AFCB9" author="NOLF D. &amp; STEURBAUT E. &amp; CAHUZAC B." box="[1237,1451,754,778]" pageId="23" pageNumber="516" pagination="164 - 194" refId="ref47746" refString="NOLF D., STEURBAUT E. &amp; CAHUZAC B. 2002. - Progres recents dans la connaissance des gisements cenozoiques en Aquitaine meridionale (Chalosse, Bearn et Bas-Adour: SW France. Revue de Micropaleontologie 45 (3): 164 - 194. https: // doi. org / 10.1016 / S 0035 - 1598 (02) 90017 - 6" type="journal article" year="2002">
Nolf
<emphasis id="B953EADEFFF1E411FAB1FD24FAB6FCB9" box="[1289,1335,754,778]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
(2002: 173
</bibRefCitation>
, fig. 4), Early Eocene (Ypresian).
</paragraph>
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<typeStatus id="549C886EFFF1E411FC95FCEAFCDCFCE1" box="[813,861,828,851]" pageId="23" pageNumber="516">TYPE</typeStatus>
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. —
<collectingCountry id="F330765CFFF1E411FC51FCEAFBACFCE0" box="[1001,1069,828,851]" name="France" pageId="23" pageNumber="516">France</collectingCountry>
,
<collectingRegion id="49E3F82EFFF1E411FB8EFCEAFB17FCE7" box="[1078,1174,828,852]" country="France" name="Aquitaine" pageId="23" pageNumber="516">Aquitaine</collectingRegion>
Basin, Pyrénées-Atlantiques, Bos d’Arros (
<bibRefCitation id="EFB64B3DFFF1E411FC15FC80FBFCFCDD" author="NOLF D. &amp; STEURBAUT E. &amp; CAHUZAC B." box="[941,1149,854,878]" pageId="23" pageNumber="516" pagination="164 - 194" refId="ref47746" refString="NOLF D., STEURBAUT E. &amp; CAHUZAC B. 2002. - Progres recents dans la connaissance des gisements cenozoiques en Aquitaine meridionale (Chalosse, Bearn et Bas-Adour: SW France. Revue de Micropaleontologie 45 (3): 164 - 194. https: // doi. org / 10.1016 / S 0035 - 1598 (02) 90017 - 6" type="journal article" year="2002">
Nolf
<emphasis id="B953EADEFFF1E411FC58FC81FB8EFCDD" box="[992,1039,854,878]" italics="true" pageId="23" pageNumber="516">et al.</emphasis>
2002: 185
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF1E411FC95FC50FA96FC0B" pageId="23" pageNumber="516" type="distribution">
<paragraph id="8B9836CCFFF1E411FC95FC50FA96FC0B" blockId="23.[811,1455,902,952]" pageId="23" pageNumber="516">
<smallCapsWord id="8D7EA010FFF1E411FC95FC50FC39FC2F" baselines="919,920" box="[813,952,902,925]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="23" pageNumber="516">DISTRIBUTION</smallCapsWord>
. — Known from the
<typeStatus id="549C886EFFF1E411FB3DFC50FB2FFC2D" box="[1157,1198,902,926]" pageId="23" pageNumber="516">type</typeStatus>
locality and Gan (tuilerie),
<collectingRegion id="49E3F82EFFF1E411FC93FC76FC0AFC0B" box="[811,907,928,952]" country="France" name="Aquitaine" pageId="23" pageNumber="516">Aquitaine</collectingRegion>
Basin (
<collectingCountry id="F330765CFFF1E411FC6BFC76FB98FC0B" box="[979,1049,928,952]" name="France" pageId="23" pageNumber="516">France</collectingCountry>
), early Eocene (Ypresian).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF1E41FFC95FC23FEE9FBDC" lastPageId="25" lastPageNumber="518" pageId="23" pageNumber="516" type="description">
<paragraph id="8B9836CCFFF1E411FC95FC23FC3CFBBE" blockId="23.[810,1457,1013,1836]" box="[813,957,1013,1039]" pageId="23" pageNumber="516">
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</paragraph>
<paragraph id="8B9836CCFFF1E411FC95FBC2FAEDF89F" blockId="23.[810,1457,1013,1836]" pageId="23" pageNumber="516">
Smooth, multispiral protoconch of 3.5 whorls (
<figureCitation id="131C2A49FFF1E411FAFDFBC2FA2AFB9C" box="[1349,1451,1044,1071]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="23" pageNumber="516">Figs 12B</figureCitation>
;
<figureCitation id="131C2A49FFF1E411FC95FBE2FCE0FBFD" box="[813,865,1076,1102]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="23" pageNumber="516">26H</figureCitation>
). Sinusigeral scar well marked.Teleoconch up to
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in height, up to
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in width, subfusiform in profile, composed of eight whorls. Moderately high spire. Last whorl up to 70% of total length. Apical angle 53° excluding wings, 70° including wings. Spiral sculpture with marked primary cords on varices and intevarices. First whorl: no cord. Second whorl: appearance of weak P1, P2 and s1. Third whorl: P1, P2, s1 well marked; appearance of s2 weak. Fourth whorl: s2 well marked. From fifth to seventh whorl: no change or appearance of threads. Eighth whorl: P1 to P6, s1, s2, s3 (well developed), P6; ADP, MP weak or missing on siphonal canal. First whorl:around ten protovarices.Second whorl: appearance of varices, three varices and two intervarices between. From third to last whorl, three to five low intervarices. Higher relief of intervarices on P2. Varices lamellose and strongly winged. Appearance of short P1 spine on second whorl.P1 spine short, almost included in wing. No other cord spines. Ovate aperture up to 22% of diameter and up to 71% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with well developed internal denticles including ID, D1 to D6. D1 and D2 bifid. Outer lip not crenulate. Siphonal canal narrow, open, straight, up to 57% of apertural length. Pseudoumbilicus narrow.
</paragraph>
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</paragraph>
<paragraph id="8B9836CCFFF1E41EFC95F8A7FE3BF858" blockId="23.[813,1457,1874,2028]" lastBlockId="24.[130,777,1618,2028]" lastPageId="24" lastPageNumber="517" pageId="23" pageNumber="516">
<bibRefCitation id="EFB64B3DFFF1E411FC95F8A7FC57F83F" author="ROUAULT A." box="[813,982,1905,1932]" pageId="23" pageNumber="516" pagination="457 - 502" refId="ref48421" refString="ROUAULT A. 1850. - Description des fossiles eocenes des environs de Pau. Memoires de la Societe geologique de France, serie 2, 3: 457 - 502." type="journal article" year="1850">Rouault (1850)</bibRefCitation>
described from Bos d’Arros (a locality very close to Gan),
<taxonomicName id="4C274D4FFFF1E411FC76F844FBEFF818" authorityName="Gmelin" authorityYear="1791" box="[974,1134,1938,1963]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FC76F844FBEFF818" box="[974,1134,1938,1963]" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
</taxonomicName>
based on juvenile specimens. Unfortunately,
<taxonomicName id="4C274D4FFFF1E411FC64F864FAB0F87F" authority="Rouault, 1850" authorityName="sensu Rouault" authorityYear="1850" box="[988,1329,1969,1996]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FC64F864FB03F878" box="[988,1154,1970,1995]" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FB34F867FAB0F87F" author="ROUAULT A." box="[1164,1329,1969,1996]" pageId="23" pageNumber="516" pagination="457 - 502" refId="ref48421" refString="ROUAULT A. 1850. - Description des fossiles eocenes des environs de Pau. Memoires de la Societe geologique de France, serie 2, 3: 457 - 502." type="journal article" year="1850">Rouault, 1850</bibRefCitation>
</taxonomicName>
is a junior homonym of
<taxonomicName id="4C274D4FFFF1E411FC77F804FAA2F858" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[975,1315,2001,2027]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="516" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFF1E411FC77F804FBF9F858" box="[975,1144,2002,2027]" italics="true" pageId="23" pageNumber="516">Murex trigonus</emphasis>
<bibRefCitation id="EFB64B3DFFF1E411FB38F807FAA2F858" author="GMELIN J. F." box="[1152,1315,2001,2027]" pageId="23" pageNumber="516" pagination="3021 - 3910" refId="ref44669" refString="GMELIN J. F. 1791 (ed.). - Caroli a Linnaei Systema Naturae per Regna Tria Naturae, part Vermes. Ed. 13. Tome 1 (6). G. E. Beer, Leipzig: 3021 - 3910. https: // doi. org / 10.5962 / bhl. title. 36932" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
</taxonomicName>
and
<bibRefCitation id="EFB64B3DFFF1E41EFADBF804FF53F9DF" author="MAGNE A." lastPageId="24" lastPageNumber="517" pageId="23" pageNumber="516" pagination="28 - 36" refId="ref46270" refString="MAGNE A. 1941. - Sur quelques especes nouvelles ou interessantes provenant du Calcaire a Asteries de la Souys. Proces-verbaux de la Societe linneenne de Bordeaux 91: 28 - 36." type="journal article" year="1941">Magne (1941)</bibRefCitation>
, a homonymy as yet uncorrected. Therefore, we propose the new name
<taxonomicName id="4C274D4FFFFEE41EFE37F9A4FDCFF93F" box="[399,590,1650,1676]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFFEE41EFE37F9A4FDCFF93F" box="[399,590,1650,1676]" italics="true" pageId="24" pageNumber="517">Timbellus magnei</emphasis>
</taxonomicName>
n. name herein. The juvenile specimens illustrated by
<bibRefCitation id="EFB64B3DFFFEE41EFD89F944FD5EF91E" author="ROUAULT A." box="[561,735,1682,1709]" pageId="24" pageNumber="517" pagination="457 - 502" refId="ref48421" refString="ROUAULT A. 1850. - Description des fossiles eocenes des environs de Pau. Memoires de la Societe geologique de France, serie 2, 3: 457 - 502." type="journal article" year="1850">Rouault (1850)</bibRefCitation>
are characterized by low intervarices varying from three to five (compare
<figureCitation id="131C2A49FFFEE41EFF4AF904FE46F95F" box="[242,455,1746,1772]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="24" pageNumber="517">Fig. 12C, D and G</figureCitation>
). This character is present on the early whorls of the adults of
<taxonomicName id="4C274D4FFFFEE41EFE48F925FF44F89F" authority="sensu Merle (1990" authorityName="sensu Merle" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFFEE41EFE48F925FD3FF8BF" box="[496,702,1778,1804]" italics="true" pageId="24" pageNumber="517">T. tripteroides sensu</emphasis>
<bibRefCitation id="EFB64B3DFFFEE41EFD7FF924FF44F89F" author="MERLE D." pageId="24" pageNumber="517" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle (1990</bibRefCitation>
</taxonomicName>
: pl. 2, figs 13, 14 only). Thus, one can accept that these large
<taxonomicName id="4C274D4FFFFEE41EFF78F8E4FDA9F8FF" authority="sensu Merle (1990)" authorityName="sensu Merle" authorityYear="1990" box="[192,552,1841,1868]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFFEE41EFF78F8E4FE0FF8F8" box="[192,398,1841,1867]" italics="true" pageId="24" pageNumber="517">T. tripteroides sensu</emphasis>
<bibRefCitation id="EFB64B3DFFFEE41EFE2DF8E7FDA9F8FF" author="MERLE D." box="[405,552,1841,1868]" pageId="24" pageNumber="517" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle (1990)</bibRefCitation>
</taxonomicName>
correspond to adult specimens of
<taxonomicName id="4C274D4FFFFEE41EFEAFF884FEFEF8D8" box="[279,383,1874,1899]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFFEE41EFEAFF884FEFEF8D8" box="[279,383,1874,1899]" italics="true" pageId="24" pageNumber="517">T. magnei</emphasis>
</taxonomicName>
n. name.
<taxonomicName id="4C274D4FFFFEE41EFE55F887FD29F8D8" box="[493,680,1873,1899]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFFEE41EFE55F887FD29F8D8" box="[493,680,1873,1899]" italics="true" pageId="24" pageNumber="517">Timbellus magnei</emphasis>
</taxonomicName>
n. name is easily separated from typical
<taxonomicName id="4C274D4FFFFEE41EFE50F8A4FDFCF838" authorityName="sensu Merle" authorityYear="1990" box="[488,637,1905,1931]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFFEE41EFE50F8A4FDFCF838" box="[488,637,1905,1931]" italics="true" pageId="24" pageNumber="517">T. tripteroides</emphasis>
</taxonomicName>
by having a multispiral protoconch (
<figureCitation id="131C2A49FFFEE41EFE34F847FE6AF818" box="[396,491,1937,1963]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="24" pageNumber="517">Fig. 12B</figureCitation>
), whilst typical
<taxonomicName id="4C274D4FFFFEE41EFD29F847FC84F818" authorityName="sensu Merle" authorityYear="1990" box="[657,773,1937,1963]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFFEE41EFD29F847FC84F818" box="[657,773,1937,1963]" italics="true" pageId="24" pageNumber="517">tripteroides</emphasis>
</taxonomicName>
has a paucipiral protoconch (see
<figureCitation id="131C2A49FFFEE41EFE51F867FDDFF878" box="[489,606,1969,1996]" captionStart="FIG" captionStartId="19.[132,143,711,728]" captionTargetBox="[132,1455,215,669]" captionTargetId="figure-804@19.[160,986,229,644]" captionTargetPageId="19" captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de l’Orme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985776" httpUri="https://zenodo.org/record/13985776/files/figure.png" pageId="24" pageNumber="517">Fig. 8D, E</figureCitation>
) and by having more and lower intervarices.
</paragraph>
<caption id="DF586644FFFEE41EFF3CFA6DFBAAF994" ID-DOI="http://doi.org/10.5281/zenodo.13985786" ID-Zenodo-Dep="13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="24" pageNumber="517" startId="24.[132,143,1467,1484]" targetBox="[132,1455,215,1424]" targetPageId="24" targetType="figure">
<paragraph id="8B9836CCFFFEE41EFF3CFA6DFBAAF994" blockId="24.[132,1457,1467,1575]" pageId="24" pageNumber="517">
<smallCapsWord id="8D7EA010FFFEE41EFF3CFA6DFF1CFA78" baselines="1480,1480" box="[132,157,1467,1484]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="24" pageNumber="517">FIG</smallCapsWord>
. 12. — Spiral cords and internal denticles of the outer lip in
<taxonomicName id="4C274D4FFFFEE41EFD2FFA6DFCA8FA7F" box="[663,809,1467,1484]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFFEE41EFD2FFA6DFCA8FA7F" box="[663,809,1467,1484]" italics="true" pageId="24" pageNumber="517">Timbellus magnei</emphasis>
</taxonomicName>
n. name (
<emphasis id="B953EADEFFFEE41EFCC5FA6DFC1DFA7F" bold="true" box="[893,924,1467,1484]" pageId="24" pageNumber="517">A -F</emphasis>
) and
<taxonomicName id="4C274D4FFFFEE41EFC74FA6DFB23FA7F" authority="(Magne, 1941)" baseAuthorityName="Magne" baseAuthorityYear="1941" box="[972,1186,1467,1484]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="palensis">
<emphasis id="B953EADEFFFEE41EFC74FA6DFBA4FA7F" box="[972,1061,1467,1484]" italics="true" pageId="24" pageNumber="517">T. palensis</emphasis>
(
<bibRefCitation id="EFB64B3DFFFEE41EFB97FA6DFB1FFA7F" author="MAGNE A." box="[1071,1182,1467,1484]" pageId="24" pageNumber="517" pagination="28 - 36" refId="ref46270" refString="MAGNE A. 1941. - Sur quelques especes nouvelles ou interessantes provenant du Calcaire a Asteries de la Souys. Proces-verbaux de la Societe linneenne de Bordeaux 91: 28 - 36." type="journal article" year="1941">Magne, 1941</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="B953EADEFFFEE41EFB15FA6DFB49FA7F" bold="true" box="[1197,1224,1467,1484]" pageId="24" pageNumber="517">G -I</emphasis>
):
<emphasis id="B953EADEFFFEE41EFB6FFA6DFB65FA7F" bold="true" box="[1239,1252,1467,1484]" pageId="24" pageNumber="517">A</emphasis>
, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);
<emphasis id="B953EADEFFFEE41EFC78FA04FC4CFA50" bold="true" box="[960,973,1490,1507]" pageId="24" pageNumber="517">B</emphasis>
, MNHN.F.A90545 (
<taxonomicName id="4C274D4FFFFEE41EFBD5FA04FB1DFA50" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1133,1180,1490,1507]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);
<emphasis id="B953EADEFFFEE41EFD7EFA3EFD55FA4A" bold="true" box="[710,724,1512,1529]" pageId="24" pageNumber="517">C</emphasis>
,
<emphasis id="B953EADEFFFEE41EFD65FA3EFD6AFA4A" bold="true" box="[733,747,1512,1529]" pageId="24" pageNumber="517">D</emphasis>
, MNHN.F.A32508 (
<taxonomicName id="4C274D4FFFFEE41EFC30FA3EFC36FA4A" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[904,951,1512,1529]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), juvenile specimen,Gan (Tuilerie);
<emphasis id="B953EADEFFFEE41EFB41FA3EFA84FA4A" bold="true" box="[1273,1285,1512,1529]" pageId="24" pageNumber="517">E</emphasis>
,
<emphasis id="B953EADEFFFEE41EFAB6FA3EFA98FA4A" bold="true" box="[1294,1305,1512,1529]" pageId="24" pageNumber="517">F</emphasis>
, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie);
<emphasis id="B953EADEFFFEE41EFECAFA29FE01F9A3" bold="true" box="[370,384,1535,1552]" pageId="24" pageNumber="517">G</emphasis>
, MNHN.F.A90553 (
<taxonomicName id="4C274D4FFFFEE41EFD9AFA29FDD0F9A3" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[546,593,1535,1552]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), juvenile specimen, Gan (Tuilerie);
<emphasis id="B953EADEFFFEE41EFC26FA29FC2DF9A3" bold="true" box="[926,940,1535,1552]" pageId="24" pageNumber="517">H</emphasis>
,
<emphasis id="B953EADEFFFEE41EFC0FFA29FC3CF9A3" bold="true" box="[951,957,1535,1552]" pageId="24" pageNumber="517">I</emphasis>
, MNHN.F.A32509 (
<taxonomicName id="4C274D4FFFFEE41EFBE6FA29FB0CF9A3" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1118,1165,1535,1552]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS).
</paragraph>
</caption>
<paragraph id="8B9836CCFFFEE41FFCFBF984FEE9FBDC" blockId="24.[810,1457,1618,2028]" lastBlockId="25.[131,778,949,1135]" lastPageId="25" lastPageNumber="518" pageId="24" pageNumber="517">
<bibRefCitation id="EFB64B3DFFFEE41EFCFBF984FB32F9DF" author="COSSMANN M. &amp; O'GORMAN G." box="[835,1203,1618,1644]" pageId="24" pageNumber="517" refId="ref43730" refString="COSSMANN M. &amp; O'GORMAN G. 1923. - Description des mollusques, in O'GORMAN G. (ed.), Le gisement cuisien de Gan (Basses Pyrenees). Cossmann, Pau, 188 p." type="book" year="1923">Cossmann &amp; O’Gorman (1923)</bibRefCitation>
misinterpreted
<taxonomicName id="4C274D4FFFFEE41EFAD3F985FB9AF93F" authority="sensu Rouault" authorityName="sensu Rouault" authorityYear="1850" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFFEE41EFAD3F985FC3DF93F" italics="true" pageId="24" pageNumber="517">Murex trigonus sensu</emphasis>
Rouault
</taxonomicName>
and illustrated a specimen typical of the genus
<taxonomicName id="4C274D4FFFFEE41EFC26F944FC78F91F" authorityName="Houart" authorityYear="1986" box="[926,1017,1682,1708]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFFEE41EFC26F944FC78F91F" box="[926,1017,1682,1708]" italics="true" pageId="24" pageNumber="517">Ponderia</emphasis>
</taxonomicName>
:
<taxonomicName id="4C274D4FFFFEE41EFBB0F945FAA3F91F" authority="(Magne, 1941)" baseAuthorityName="Magne" baseAuthorityYear="1941" box="[1032,1314,1682,1708]" class="Gastropoda" family="Muricidae" genus="Ponderia" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="daguini">
<emphasis id="B953EADEFFFEE41EFBB0F945FBF3F91F" box="[1032,1138,1682,1708]" italics="true" pageId="24" pageNumber="517">P. daguini</emphasis>
(
<bibRefCitation id="EFB64B3DFFFEE41EFB39F944FA98F91F" author="MAGNE A." box="[1153,1305,1682,1708]" pageId="24" pageNumber="517" pagination="28 - 36" refId="ref46270" refString="MAGNE A. 1941. - Sur quelques especes nouvelles ou interessantes provenant du Calcaire a Asteries de la Souys. Proces-verbaux de la Societe linneenne de Bordeaux 91: 28 - 36." type="journal article" year="1941">Magne, 1941</bibRefCitation>
)
</taxonomicName>
. In addition,
<bibRefCitation id="EFB64B3DFFFEE41EFC96F964FB17F97F" author="COSSMANN M. &amp; O'GORMAN G." box="[814,1174,1714,1740]" pageId="24" pageNumber="517" refId="ref43730" refString="COSSMANN M. &amp; O'GORMAN G. 1923. - Description des mollusques, in O'GORMAN G. (ed.), Le gisement cuisien de Gan (Basses Pyrenees). Cossmann, Pau, 188 p." type="book" year="1923">Cossmann &amp; O’Gorman (1923)</bibRefCitation>
described and figured an adult specimen of
<taxonomicName id="4C274D4FFFFEE41EFC47F904FBE9F95F" box="[1023,1128,1746,1772]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFFEE41EFC47F904FBE9F95F" box="[1023,1128,1746,1772]" italics="true" pageId="24" pageNumber="517">Timbellus</emphasis>
</taxonomicName>
that Cosmann named
<taxonomicName id="4C274D4FFFFEE41EFAD4F905FBEEF8BF" authority="Lamarck, 1803" authorityName="Lamarck" authorityYear="1803" class="Gastropoda" family="Muricidae" genus="Murex" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="tricarinatus">
<emphasis id="B953EADEFFFEE41EFAD4F905FA30F95F" box="[1388,1457,1747,1772]" italics="true" pageId="24" pageNumber="517">Murex</emphasis>
cf.
<emphasis id="B953EADEFFFEE41EFCF4F925FC45F8BF" box="[844,964,1779,1804]" italics="true" pageId="24" pageNumber="517">tricarinatus</emphasis>
<bibRefCitation id="EFB64B3DFFFEE41EFC72F924FBEEF8BF" author="LAMARCK J. - B." box="[970,1135,1778,1804]" pageId="24" pageNumber="517" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFB64B3DFFFEE41EFBCFF924FA90F8BF" author="MAGNE A." box="[1143,1297,1778,1804]" pageId="24" pageNumber="517" pagination="28 - 36" refId="ref46270" refString="MAGNE A. 1941. - Sur quelques especes nouvelles ou interessantes provenant du Calcaire a Asteries de la Souys. Proces-verbaux de la Societe linneenne de Bordeaux 91: 28 - 36." type="journal article" year="1941">Magne (1941)</bibRefCitation>
gave the name
<taxonomicName id="4C274D4FFFFEE41EFC93F8C4FBD4F89F" box="[811,1109,1810,1836]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="palensis" subGenus="Pteropurpura">
<emphasis id="B953EADEFFFEE41EFC93F8C4FCECF898" box="[811,877,1810,1835]" italics="true" pageId="24" pageNumber="517">Murex</emphasis>
(
<emphasis id="B953EADEFFFEE41EFCC3F8C4FC7FF898" box="[891,1022,1810,1835]" italics="true" pageId="24" pageNumber="517">Pteropurpura</emphasis>
)
<emphasis id="B953EADEFFFEE41EFBB3F8C4FBD4F89F" box="[1035,1109,1810,1836]" italics="true" pageId="24" pageNumber="517">palensis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C274D4FFFFEE41EFBCAF8C4FCE2F8FF" authority="Lamarck, 1803" authorityName="Lamarck" authorityYear="1803" class="Gastropoda" family="Muricidae" genus="Murex" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="tricarinatus">
<emphasis id="B953EADEFFFEE41EFBCAF8C4FB35F898" box="[1138,1204,1810,1835]" italics="true" pageId="24" pageNumber="517">Murex</emphasis>
cf.
<emphasis id="B953EADEFFFEE41EFB6FF8C4FACBF898" box="[1239,1354,1810,1835]" italics="true" pageId="24" pageNumber="517">tricarinatus</emphasis>
<bibRefCitation id="EFB64B3DFFFEE41EFAF6F8C4FCE2F8FF" author="LAMARCK J. - B." pageId="24" pageNumber="517" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
</taxonomicName>
(
<emphasis id="B953EADEFFFEE41EFCC1F8E4FC2EF8F8" box="[889,943,1842,1867]" italics="true" pageId="24" pageNumber="517">sensu</emphasis>
<bibRefCitation id="EFB64B3DFFFEE41EFC00F8E4FBF0F8FF" author="COSSMANN M. &amp; O'GORMAN G." box="[952,1137,1842,1868]" pageId="24" pageNumber="517" refId="ref43730" refString="COSSMANN M. &amp; O'GORMAN G. 1923. - Description des mollusques, in O'GORMAN G. (ed.), Le gisement cuisien de Gan (Basses Pyrenees). Cossmann, Pau, 188 p." type="book" year="1923">Cossmann 1923</bibRefCitation>
) and demonstrated that the juvenile specimens illustrated by
<bibRefCitation id="EFB64B3DFFFEE41EFB24F887FAC7F8DF" author="ROUAULT A." box="[1180,1350,1873,1900]" pageId="24" pageNumber="517" pagination="457 - 502" refId="ref48421" refString="ROUAULT A. 1850. - Description des fossiles eocenes des environs de Pau. Memoires de la Societe geologique de France, serie 2, 3: 457 - 502." type="journal article" year="1850">Rouault (1850)</bibRefCitation>
as
<taxonomicName id="4C274D4FFFFEE41EFAD5F884FC01F838" authorityName="sensu Rouault" authorityYear="1850" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="trigonus">
<emphasis id="B953EADEFFFEE41EFAD5F884FC01F838" italics="true" pageId="24" pageNumber="517">Murex trigonus</emphasis>
</taxonomicName>
were not juveniles of
<taxonomicName id="4C274D4FFFFEE41EFBD5F8A4FA29F838" box="[1133,1448,1905,1931]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="palensis" subGenus="Pteropurpura">
<emphasis id="B953EADEFFFEE41EFBD5F8A4FB30F838" box="[1133,1201,1906,1931]" italics="true" pageId="24" pageNumber="517">Murex</emphasis>
(
<emphasis id="B953EADEFFFEE41EFB79F8A4FACBF838" box="[1217,1354,1906,1931]" italics="true" pageId="24" pageNumber="517">Pteropurpura</emphasis>
)
<emphasis id="B953EADEFFFEE41EFAE2F8A7FA29F838" box="[1370,1448,1905,1931]" italics="true" pageId="24" pageNumber="517">palensis</emphasis>
</taxonomicName>
. We discovered a young specimen from Gan differing from the juveniles of
<taxonomicName id="4C274D4FFFFEE41EFC69F864FBB7F878" box="[977,1078,1970,1995]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="517" phylum="Mollusca" rank="species" species="magnei">
<emphasis id="B953EADEFFFEE41EFC69F864FBB7F878" box="[977,1078,1970,1995]" italics="true" pageId="24" pageNumber="517">T. magnei</emphasis>
</taxonomicName>
n. name. It displays one high intervarix between each varix (
<figureCitation id="131C2A49FFFEE41EFBF9F807FB22F858" box="[1089,1187,2001,2027]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="24" pageNumber="517">Fig. 12G</figureCitation>
), a character also present on the early teleoconch whorls of well preserved adults of
<taxonomicName id="4C274D4FFFFFE41FFD49FC63FF53FC5D" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="518" phylum="Mollusca" rank="species" species="palensis">
<emphasis id="B953EADEFFFFE41FFD49FC63FF53FC5D" italics="true" pageId="25" pageNumber="518">T. palensis</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFFFE41FFF5AFC02FEDDFC5C" box="[226,348,980,1007]" captionStart="FIG" captionStartId="24.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-413@24.[698,1267,772,1332]" captionTargetPageId="24" captionText="FIG. 12. — Spiral cords and internal denticles of the outer lip in Timbellus magnei n. name (A-F) and T. palensis (Magne, 1941) (G-I): A, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian);B, MNHN.F.A90545 (Merle coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian);C, D, MNHN.F.A32508 (Merle coll.), juvenile specimen,Gan (Tuilerie);E, F, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); G, MNHN.F.A90553 (Merle coll.), juvenile specimen, Gan (Tuilerie); H, I, MNHN.F.A32509 (Merle coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985786" httpUri="https://zenodo.org/record/13985786/files/figure.png" pageId="25" pageNumber="518">Fig. 12H, I</figureCitation>
).
<bibRefCitation id="EFB64B3DFFFFE41FFEC9FC02FD32FC5C" author="DOUVILLE H. &amp; O'GORMAN G." box="[369,691,980,1007]" pageId="25" pageNumber="518" pagination="351 - 383" refId="ref44174" refString="DOUVILLE H. &amp; O'GORMAN G. 1929. - L'Eocene du Bearn. Description de la faune de Biron. Bulletin de la Societe Geologique de France serie 4, 29: 351 - 383." type="journal article" year="1929">Douvillé &amp; O’Gorman (1929</bibRefCitation>
: pl. 29, fig. 1) figured a specimen from the early Eocene of Biron (near Orthez) and called it
<taxonomicName id="4C274D4FFFFFE41FFE19FBC3FDDCFB9D" authorityName="Lamarck" authorityYear="1822" box="[417,605,1044,1070]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="518" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFFFE41FFE19FBC3FDDCFB9D" box="[417,605,1044,1070]" italics="true" pageId="25" pageNumber="518">Murex tripteroides</emphasis>
</taxonomicName>
. That specimen bears one high intervarix between each varix and possibly represents
<taxonomicName id="4C274D4FFFFFE41FFF4EFB83FEE0FBDD" box="[246,353,1108,1134]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="518" phylum="Mollusca" rank="species" species="palensis">
<emphasis id="B953EADEFFFFE41FFF4EFB83FEE0FBDD" box="[246,353,1108,1134]" italics="true" pageId="25" pageNumber="518">T. palensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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