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<document id="52E160640DA0BB295109833820557654" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFEEE40FFEBFF8A7FC89F97E" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Paleochicoreus Merle 2024, n. gen." docType="treatment" docUuid="3E5850A7-582D-48BF-B126-BA40739DDC00" docUuidSource="ZooBank" docVersion="6" lastPageNumber="502" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="501" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="6E554E23606A7BFDA15A0E2B23F891F1">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="1C220E9C14CDBCE69DAB91973ED86910">Merle, Didier</mods:namePart>
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<mods:namePart id="8FEB5D287BFD71AB8E901C778C993949">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="91BB638456BA3D2F2CBC02EAC2BCC693">Ledon, Daniel</mods:namePart>
<mods:affiliation id="95CA3F0F27E500828A56D3A7DF9D7315">Muséum national dhistoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:namePart id="A0458540F735EAC3C3B93619DC3EB0B2">Goret, Bernard</mods:namePart>
<mods:affiliation id="2F0466B9FA8A98B41D6A35C7B62B3158">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:title id="C632D160AA2192291C6596E95351AD0C">Geodiversitas</mods:title>
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<mods:date id="BBB06B6A94EA97502B30832498F88C0D">2024</mods:date>
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<treatment id="038E87DAFFEEE40FFEBFF8A7FC89F97E" ID-DOI="http://doi.org/10.5281/zenodo.13989314" ID-GBIF-Taxon="244735005" ID-Zenodo-Dep="13989314" LSID="urn:lsid:zoobank.org:act:3E5850A7-582D-48BF-B126-BA40739DDC00" httpUri="http://treatment.plazi.org/id/038E87DAFFEEE40FFEBFF8A7FC89F97E" lastPageId="9" lastPageNumber="502" pageId="8" pageNumber="501" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Ergalataxinae">
<subSubSection id="C33D6547FFEEE40EFEBFF8A7FD04F83F" box="[263,645,1905,1932]" pageId="8" pageNumber="501" type="nomenclature">
<paragraph id="8B9836CCFFEEE40EFEBFF8A7FD04F83F" blockId="8.[263,645,1905,1963]" box="[263,645,1905,1932]" pageId="8" pageNumber="501">
<heading id="D0D081A0FFEEE40EFEBFF8A7FD04F83F" box="[263,645,1905,1932]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="501" reason="2">
Genus
<taxonomicName id="4C274D4FFFEEE40EFEECF8A7FDAEF838" ID-CoL="CN5MX" authority="Merle" authorityName="Merle" authorityYear="2024" box="[340,559,1905,1931]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="genus" status="n. gen.">
<emphasis id="B953EADEFFEEE40EFEECF8A7FE67F838" box="[340,486,1905,1931]" italics="true" pageId="8" pageNumber="501">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEEE40EFD82F8A4FD04F83F" box="[570,645,1906,1932]" pageId="8" pageNumber="501" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
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<subSubSection id="C33D6547FFEEE40EFE26F847FC86F852" pageId="8" pageNumber="501" type="description">
<paragraph id="8B9836CCFFEEE40EFE26F847FE6CF818" blockId="8.[263,645,1905,1963]" box="[414,493,1937,1963]" pageId="8" pageNumber="501">
(
<figureCitation id="131C2A49FFEEE40EFE1FF847FE65F818" box="[423,484,1937,1963]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2</figureCitation>
)
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urn:lsid:zoobank.org:act:
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. —
<taxonomicName id="4C274D4FFFEEE40EFC73F947FAE2F91A" authority="Vokes, 1990" authorityName="Vokes" authorityYear="1990" box="[971,1379,1681,1705]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="initialis" subGenus="Phyllonotus">
<emphasis id="B953EADEFFEEE40EFC73F947FBA2F91A" box="[971,1059,1681,1705]" italics="true" pageId="8" pageNumber="501">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEEE40EFB89F947FB18F91A" box="[1073,1177,1681,1705]" italics="true" pageId="8" pageNumber="501">Phyllonotus</emphasis>
)
<emphasis id="B953EADEFFEEE40EFB1DF947FB6AF91A" box="[1189,1259,1681,1705]" italics="true" pageId="8" pageNumber="501">initialis</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFB55F947FAE2F91A" author="VOKES E. H." box="[1261,1379,1681,1705]" pageId="8" pageNumber="501" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes, 1990</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="F330765CFFEEE40EFAD3F947FCE7F977" name="United States of America" pageId="8" pageNumber="501">United States</collectingCountry>
,
<collectingRegion id="49E3F82EFFEEE40EFCD6F97AFC27F970" box="[878,934,1708,1731]" country="United States of America" name="Texas" pageId="8" pageNumber="501">Texas</collectingRegion>
, middle Eocene (Bartonian) (
<figureCitation id="131C2A49FFEEE40EFB71F97DFAB1F977" box="[1225,1328,1707,1732]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2A, B</figureCitation>
).
</paragraph>
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. —
<taxonomicName id="4C274D4FFFEEE40EFBB8F900FAB2F95E" authority="Dickerson, 1915" authorityName="Dickerson" authorityYear="1915" box="[1024,1331,1749,1773]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="packardi">
<emphasis id="B953EADEFFEEE40EFBB8F900FB0EF95E" box="[1024,1167,1749,1773]" italics="true" pageId="8" pageNumber="501">Murex packardi</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFB2BF903FAB2F95E" author="DICKERSON R. E." box="[1171,1331,1749,1773]" pageId="8" pageNumber="501" pagination="33 - 98" refId="ref44126" refString="DICKERSON R. E. 1915. - Fauna of the type Tejon: its relation to the Cowlitz phase of the Tejon group of Washington. Proceedings of the California Academy of Sciences, 4 th series, 5 (3): 33 - 98." type="journal article" year="1915">Dickerson, 1915</bibRefCitation>
</taxonomicName>
,
<collectingRegion id="49E3F82EFFEEE40EFA83F903FA2FF95E" box="[1339,1454,1749,1773]" country="United States of America" name="Washington" pageId="8" pageNumber="501">Washington</collectingRegion>
(
<collectingCountry id="F330765CFFEEE40EFC89F926FC30F8BB" box="[817,945,1776,1800]" name="United States of America" pageId="8" pageNumber="501">United States</collectingCountry>
), middle Eocene;
<taxonomicName id="4C274D4FFFEEE40EFBEBF926FCE3F891" authority="Vokes, 1990" authorityName="Vokes" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="initialis" subGenus="Phyllonotus">
<emphasis id="B953EADEFFEEE40EFBEBF926FB2BF8BB" box="[1107,1194,1776,1800]" italics="true" pageId="8" pageNumber="501">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEEE40EFB0FF926FA9CF8BB" box="[1207,1309,1776,1800]" italics="true" pageId="8" pageNumber="501">Phyllonotus</emphasis>
)
<emphasis id="B953EADEFFEEE40EFA91F926FAEFF8BB" box="[1321,1390,1776,1800]" italics="true" pageId="8" pageNumber="501">initialis</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFAC8F926FCE3F891" author="VOKES E. H." pageId="8" pageNumber="501" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes, 1990</bibRefCitation>
</taxonomicName>
,
<collectingRegion id="49E3F82EFFEEE40EFCD3F8DDFC21F891" box="[875,928,1803,1826]" country="United States of America" name="Texas" pageId="8" pageNumber="501">Texas</collectingRegion>
(
<collectingCountry id="F330765CFFEEE40EFC15F8DCFBB4F891" box="[941,1077,1802,1826]" name="United States of America" pageId="8" pageNumber="501">United States</collectingCountry>
), middle Eocene (Lutetian);
<emphasis id="B953EADEFFEEE40EFAF4F8DDFA2EF891" box="[1356,1455,1803,1826]" italics="true" pageId="8" pageNumber="501">Pterynoyus</emphasis>
<taxonomicName id="4C274D4FFFEEE40EFC95F8F0FC7AF88E" authority="Eames, 1957" authorityName="Eames" authorityYear="1957" box="[813,1019,1829,1853]" class="Gastropoda" family="Muricidae" genus="Paleoochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="newtoni">
<emphasis id="B953EADEFFEEE40EFC95F8F0FCF9F88E" box="[813,888,1830,1853]" italics="true" pageId="8" pageNumber="501">newtoni</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFCC5F8F3FC7AF88E" author="EAMES F. E." box="[893,1019,1829,1853]" pageId="8" pageNumber="501" pagination="25 - 70" refId="ref44211" refString="EAMES F. E. 1957. - Eocene Mollusca from Nigeria: a revision. Bulletin of the British Museum of the Natural History (Geology) 3 (2): 25 - 70. https: // doi. org / 10.5962 / p. 143489" type="journal article" year="1957">Eames, 1957</bibRefCitation>
</taxonomicName>
[new name for “
<emphasis id="B953EADEFFEEE40EFB23F8F0FA96F88E" box="[1179,1303,1830,1853]" italics="true" pageId="8" pageNumber="501">Pteropurpura</emphasis>
<taxonomicName id="4C274D4FFFEEE40EFA9EF8F3FC3FF8EB" authority="Newton, 1922" authorityName="Newton" authorityYear="1922" class="Gastropoda" family="Muricidae" genus="Murex" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="tricarinatus">
cf.
<emphasis id="B953EADEFFEEE40EFAFAF8F0FA2EF88E" box="[1346,1455,1830,1853]" italics="true" pageId="8" pageNumber="501">tricarinatus</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFC95F896FC3FF8EB" author="NEWTON R. B." box="[813,958,1856,1880]" pageId="8" pageNumber="501" pagination="7 - 114" refId="ref47686" refString="NEWTON R. B. 1922. - Eocene Mollusca from Nigeria. Geological Survey of Nigeria Bulletin 3: 7 - 114. https: // www. biodiversitylibrary. org / page / 22642662" type="journal article" year="1922">Newton, 1922</bibRefCitation>
</taxonomicName>
],
<collectingCountry id="F330765CFFEEE40EFC76F8E9FB98F8E4" box="[974,1049,1855,1879]" name="Nigeria" pageId="8" pageNumber="501">Nigeria</collectingCountry>
, middle Eocene,
<figureCitation id="131C2A49FFEEE40EFB04F8E9FA86F8EB" box="[1212,1287,1855,1880]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2C</figureCitation>
;
<taxonomicName id="4C274D4FFFEEE40EFAA9F896FC32F8C1" authority="Vincent, 1913" authorityName="Vincent" authorityYear="1913" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="diderrichi">
<emphasis id="B953EADEFFEEE40EFAA9F896FA2EF8E4" box="[1297,1455,1855,1879]" italics="true" pageId="8" pageNumber="501">Murex diderrichi</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFC95F88CFC32F8C1" author="VINCENT G." box="[813,947,1882,1906]" pageId="8" pageNumber="501" pagination="1 - 46" refId="ref48789" refString="VINCENT G. 1913. - La faune Paleocene de Landana. Annales du Musee du Congo Belge. Geologie, Paleontologie, Mineralogie serie 3, 1 (1): 1 - 46." type="journal article" year="1913">Vincent, 1913</bibRefCitation>
</taxonomicName>
[?synonym of
<taxonomicName id="4C274D4FFFEEE40EFB8FF88DFA22F8C1" authority="Oppenheim, 1904" authorityName="Oppenheim" authorityYear="1904" box="[1079,1443,1882,1906]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="camerunensis">
<emphasis id="B953EADEFFEEE40EFB8FF88DFB6FF8C1" box="[1079,1262,1883,1906]" italics="true" pageId="8" pageNumber="501">Murex camerunensis</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFB49F88CFA22F8C1" author="OPPENHEIM P." box="[1265,1443,1882,1906]" pageId="8" pageNumber="501" pagination="243 - 283" refId="ref47867" refString="OPPENHEIM P. 1904. - Uber Tertiarfossilien, warscheinlich Eocanen Altes von Kamerun. Beitrage zur Geologie von Kamerun 3: 243 - 283." type="journal article" year="1904">Oppenheim, 1904</bibRefCitation>
</taxonomicName>
],
<collectingCountry id="F330765CFFEEE40EFC93F8A2FCF2F83F" box="[811,883,1908,1932]" name="Angola" pageId="8" pageNumber="501">Angola</collectingCountry>
,
<collectingCountry id="F330765CFFEEE40EFCC6F8A3FC68F83E" box="[894,1001,1909,1933]" name="Cameroon" pageId="8" pageNumber="501">Cameroon</collectingCountry>
, late Paleocene-early Eocene;
<taxonomicName id="4C274D4FFFEEE40EFAAAF8A3FC4BF814" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="bicostatus">
<emphasis id="B953EADEFFEEE40EFAAAF8A3FA31F83F" box="[1298,1456,1908,1932]" italics="true" pageId="8" pageNumber="501">Murex bicostatus</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFC96F859FC4BF814" author="DESHAYES G. - P." box="[814,970,1935,1959]" pageId="8" pageNumber="501" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel &amp; Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="F330765CFFEEE40EFC6CF846FB99F814" box="[980,1048,1936,1959]" name="France" pageId="8" pageNumber="501">France</collectingCountry>
, middle Eocene (Bartonian) (
<figureCitation id="131C2A49FFEEE40EFA80F859FA23F814" box="[1336,1442,1935,1959]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2D, E</figureCitation>
).
<smallCapsWord id="8D7EA010FFEEE40EFC95F86FFC27F87C" baselines="1994,1995" box="[813,934,1977,2000]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="8" pageNumber="501">ETYMOLOGY</smallCapsWord>
. — Name formed by
<emphasis id="B953EADEFFEEE40EFBC7F86FFB3EF862" box="[1151,1215,1977,2001]" italics="true" pageId="8" pageNumber="501">palaiós</emphasis>
(meaning old in ancient Greek) and
<taxonomicName id="4C274D4FFFEEE40EFC19F805FC7AF858" authorityName="Montfort" authorityYear="1810" box="[929,1019,2003,2027]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEEE40EFC19F805FC7AF858" box="[929,1019,2003,2027]" italics="true" pageId="8" pageNumber="501">Chicoreus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFEFE40FFF3CFF07FE57FE64" pageId="9" pageNumber="502" type="diagnosis">
<paragraph id="8B9836CCFFEFE40FFF3CFF07FE57FE64" blockId="9.[131,777,209,471]" pageId="9" pageNumber="502">
<smallCapsWord id="8D7EA010FFEFE40FFF3CFF07FF6EFF54" baselines="226,227" box="[132,239,209,232]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="9" pageNumber="502">DIAGNOSIS</smallCapsWord>
. — Biconic and middle-sized shell with
<taxonomicName id="4C274D4FFFEFE40FFDD3FF07FD45FF5A" authorityName="Montfort" authorityYear="1810" box="[619,708,209,233]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFDD3FF07FD45FF5A" box="[619,708,209,233]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
shape. Well marked primary cords. Spire: IP (infrasutural ramp), P1 to P3 (convex part of whorl). Last whorl: IP, P1 to P6 (convex part of whorl), P6 not atrophied, ADP, MP and ABP (siphonal canal). Nodulose protovarices on early whorls. Three varices with one or two intervarices on succeeding whorls. Varices slightly winged on last whorl. No cord spine even on P1. Shoulder sinus very short. Aperture ovate with strong internal denticles within outer lip: ID, D1 to D6. Columellar lip smooth. Parietal callus and anal notch present. Aragonite microstructure.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFEFE40FFF3CFE21FC89F97E" pageId="9" pageNumber="502" type="discussion">
<paragraph id="8B9836CCFFEFE40FFF3CFE21FF69FDBC" blockId="9.[130,778,503,1744]" box="[132,232,503,529]" pageId="9" pageNumber="502">
<smallCapsWord id="8D7EA010FFEFE40FFF3CFE21FF69FDBC" baselines="522,522" box="[132,232,503,529]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="9" pageNumber="502">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFEFE40FFF3CFDC0FC89F97E" blockId="9.[130,778,503,1744]" pageId="9" pageNumber="502">
During the Eocene several species of
<taxonomicName id="4C274D4FFFEFE40FFD9DFDC0FC82FD83" authority="Merle" authorityName="Merle" authorityYear="2024" box="[549,771,534,560]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFD9DFDC0FD38FD83" box="[549,697,534,560]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFF3EFDE1FF50FDE2" box="[134,209,567,593]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
appeared in Eastern Pacific (North American coast), Western Atlantic (North American coast) and Eastern Atlantic (West Africa and Europe), which have been attributed in the literature to
<taxonomicName id="4C274D4FFFEFE40FFEB9FD41FD97FD03" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" box="[257,534,662,688]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFEB9FD41FEEBFD03" box="[257,362,663,688]" italics="true" pageId="9" pageNumber="502">Pterynotus</emphasis>
Swainson, 1833
</taxonomicName>
or
<taxonomicName id="4C274D4FFFEFE40FFD8EFD40FF3DFD63" authority="Montfort, 1810" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFD8EFD40FD16FD03" box="[566,663,662,688]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
Montfort, 1810
</taxonomicName>
(
<bibRefCitation id="EFB64B3DFFEFE40FFF74FD60FED3FD63" author="EAMES F. E." box="[204,338,693,720]" pageId="9" pageNumber="502" pagination="25 - 70" refId="ref44211" refString="EAMES F. E. 1957. - Eocene Mollusca from Nigeria: a revision. Bulletin of the British Museum of the Natural History (Geology) 3 (2): 25 - 70. https: // doi. org / 10.5962 / p. 143489" type="journal article" year="1957">Eames 1957</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFEFE40FFEE7FD60FE5FFD63" author="VOKES E. H." box="[351,478,694,720]" pageId="9" pageNumber="502" pagination="1 - 141" refId="ref48907" refString="VOKES E. H. 1971. - Catalogue of the genus Murex Linne (Mollusca: Gastropoda); Muricinae, Ocenebrinae. Bulletins of American Paleontology 61 (268): 1 - 141." type="journal article" year="1971">Vokes 1971</bibRefCitation>
,
<bibRefCitation id="EFB64B3DFFEFE40FFE54FD60FDA5FD63" author="VOKES E. H." box="[492,548,694,720]" pageId="9" pageNumber="502" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">1990</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFEFE40FFD8AFD60FD6AFD63" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[562,747,694,720]" pageId="9" pageNumber="502" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFEFE40FFD8AFD60FDF0FD63" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[562,625,694,720]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFEFE40FFDC1FD60FD2CFD63" box="[633,685,694,720]" italics="true" pageId="9" pageNumber="502">et al.</emphasis>
2011
</bibRefCitation>
as?
<taxonomicName id="4C274D4FFFEFE40FFF35FD00FF6CFD43" authorityName="Montfort" authorityYear="1810" box="[141,237,726,752]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFF35FD00FF6CFD43" box="[141,237,726,752]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
). With
<emphasis id="B953EADEFFEFE40FFE82FD00FE23FD43" box="[314,418,726,752]" italics="true" pageId="9" pageNumber="502">
<taxonomicName id="4C274D4FFFEFE40FFE82FD00FE1CFD43" authorityName="Montfort" authorityYear="1810" box="[314,413,726,752]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">Chicoreus</taxonomicName>
,
</emphasis>
they share a biconic shell, medium length of siphonal canal, trivaricate morphology, nodulose protovarices, an anal notch, and a parietal callus. However, these features are also shared with the genera
<taxonomicName id="4C274D4FFFEFE40FFDD0FCE0FE9CFCDC" authority="Jousseaume, 1880" authorityName="Jousseaume" authorityYear="1880" class="Gastropoda" family="Muricidae" genus="Naquetia" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFDD0FCE0FD48FCFC" box="[616,713,822,847]" italics="true" pageId="9" pageNumber="502">Naquetia</emphasis>
<bibRefCitation id="EFB64B3DFFEFE40FFD68FCE0FE9CFCDC" author="JOUSSEAUME F. P." pageId="9" pageNumber="502" pagination="335 - 338" refId="ref45378" refString="JOUSSEAUME F. P. 1880. - Division methodique de la famille des Purpurides. Le Naturaliste 2 (42): 335 - 338. http: // biodiversitylibrary. org / page / 33928744" type="journal article" year="1880">Jousseaume, 1880</bibRefCitation>
</taxonomicName>
,
<emphasis id="B953EADEFFEFE40FFE91FC83FE21FCDC" box="[297,416,853,879]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
Swainson, 1833 and
<taxonomicName id="4C274D4FFFEFE40FFD33FC83FEB1FC3C" authority="Arakawa, 1964" authorityName="Arakawa" authorityYear="1964" class="Gastropoda" family="Muricidae" genus="Chicomurex" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFD33FC83FC89FCDC" box="[651,776,853,879]" italics="true" pageId="9" pageNumber="502">Chicomurex</emphasis>
Arakawa, 1964
</taxonomicName>
.
<taxonomicName id="4C274D4FFFEFE40FFE87FCA3FDA1FC3C" authority="Merle" authorityName="Merle" authorityYear="2024" box="[319,544,885,911]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFE87FCA3FE54FC3C" box="[319,469,885,911]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFD95FCA0FDFAFC23" box="[557,635,886,912]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
differs from these three genera by having a smooth columellar lip. For example, according to
<bibRefCitation id="EFB64B3DFFEFE40FFECEFC63FD87FC7C" author="VOKES E. H." box="[374,518,949,975]" pageId="9" pageNumber="502" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes (1990)</bibRefCitation>
,
<taxonomicName id="4C274D4FFFEFE40FFDACFC63FF55FC5C" authorityName="Vokes" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" subGenus="Phyllonotus">
<emphasis id="B953EADEFFEFE40FFDACFC63FDF9FC7C" box="[532,632,949,975]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEFE40FFD3FFC63FD7FFC7C" box="[647,766,949,975]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
)
<emphasis id="B953EADEFFEFE40FFF3CFC03FF55FC5C" box="[132,212,981,1007]" italics="true" pageId="9" pageNumber="502">initialis</emphasis>
</taxonomicName>
(herein
<taxonomicName id="4C274D4FFFEFE40FFE95FC03FD95FC5C" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Vokes" baseAuthorityYear="1990" box="[301,532,981,1007]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" status="comb. nov.">
<emphasis id="B953EADEFFEFE40FFE95FC03FD95FC5C" box="[301,532,981,1007]" italics="true" pageId="9" pageNumber="502">Paleochicoreus initialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFEFE40FFDA2FC03FDFAFC5C" box="[538,635,981,1007]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
) (
<figureCitation id="131C2A49FFEFE40FFD29FC03FD7EFC5C" box="[657,767,981,1007]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="9" pageNumber="502">Fig. 2A, B</figureCitation>
) is most similar to members of
<emphasis id="B953EADEFFEFE40FFE74FC23FDC3FBBC" box="[460,578,1013,1039]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
, because of developed internal denticles of the outer lip, but it lacks the derived characters of
<emphasis id="B953EADEFFEFE40FFEB3FBE2FE01FBFD" box="[267,384,1076,1102]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
(rugae and expanded parietal shield).
<taxonomicName id="4C274D4FFFEFE40FFF3AFB82FEEAFBDD" baseAuthorityName="Eames" baseAuthorityYear="1957" box="[130,363,1108,1134]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="newtoni">
<emphasis id="B953EADEFFEFE40FFF3AFB82FEEAFBDD" box="[130,363,1108,1134]" italics="true" pageId="9" pageNumber="502">Paleochicoreus newtoni</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFEFE40FFEC1FB82FE49FBDC" box="[377,456,1108,1135]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="9" pageNumber="502">Fig. 2C</figureCitation>
) possesses an expanded varical flange, like that in
<taxonomicName id="4C274D4FFFEFE40FFEEBFBA2FDBDFB3D" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Vokes" baseAuthorityYear="1990" box="[339,572,1140,1166]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" status="comb. nov.">
<emphasis id="B953EADEFFEFE40FFEEBFBA2FDBDFB3D" box="[339,572,1140,1166]" italics="true" pageId="9" pageNumber="502">Paleochicoreus initialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFEFE40FFDFBFBA3FD26FB3D" box="[579,679,1140,1167]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
, but this character is lacking in many
<taxonomicName id="4C274D4FFFEFE40FFE78FB42FDFDFB1D" box="[448,636,1172,1198]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B953EADEFFEFE40FFE78FB42FDA5FB1D" box="[448,548,1172,1198]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
species.
</taxonomicName>
Members of
<taxonomicName id="4C274D4FFFEFE40FFF3EFB62FECAFB7D" authorityName="G. Perry" authorityYear="1810" box="[134,331,1204,1230]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="subGenus" subGenus="Triplex">
<emphasis id="B953EADEFFEFE40FFF3EFB62FF6BFB7D" box="[134,234,1204,1230]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEFE40FFF41FB62FEC3FB7D" box="[249,322,1204,1230]" italics="true" pageId="9" pageNumber="502">Triplex</emphasis>
)
</taxonomicName>
and
<emphasis id="B953EADEFFEFE40FFE3CFB62FE1CFB7D" box="[388,413,1204,1230]" italics="true" pageId="9" pageNumber="502">C.</emphasis>
(
<taxonomicName id="4C274D4FFFEFE40FFE15FB62FDE1FB7D" authorityName="Oyama" authorityYear="1950" box="[429,608,1204,1230]" class="Gastropoda" family="Muricidae" genus="Rhizophorimurex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFE15FB62FDE1FB7D" box="[429,608,1204,1230]" italics="true" pageId="9" pageNumber="502">Rhizophorimurex</emphasis>
</taxonomicName>
) also bear similarities with those of
<taxonomicName id="4C274D4FFFEFE40FFED6FB02FDC8FB5D" authority="Merle" authorityName="Merle" authorityYear="2024" box="[366,585,1236,1262]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFED6FB02FD81FB5D" box="[366,512,1236,1262]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFDECFB03FD20FB5C" box="[596,673,1237,1263]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
, but they lack an expanded varical flange, their internal denticles are reduced or missing, and they often develop spiny processes.
<taxonomicName id="4C274D4FFFEFE40FFF3DFAE2FF68FAFD" authorityName="Montfort" authorityYear="1810" box="[133,233,1332,1358]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFF3DFAE2FF68FAFD" box="[133,233,1332,1358]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFEFE40FFF40FAE2FE94FAFE" box="[248,277,1332,1357]" italics="true" pageId="9" pageNumber="502">s.s.</emphasis>
), which is rather similar to
<emphasis id="B953EADEFFEFE40FFD84FAE2FDD4FAFD" box="[572,597,1332,1358]" italics="true" pageId="9" pageNumber="502">C.</emphasis>
(
<emphasis id="B953EADEFFEFE40FFDDDFAE2FD2FFAFD" box="[613,686,1332,1358]" italics="true" pageId="9" pageNumber="502">Triplex</emphasis>
), differs by having a labral groove spine placed between P4 and P5 (
<bibRefCitation id="EFB64B3DFFEFE40FFF36FAA5FED0FA3D" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[142,337,1395,1422]" pageId="9" pageNumber="502" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFEFE40FFF36FAA5FF4EFA3E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[142,207,1395,1421]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFEFE40FFF60FAA2FE8EFA3E" box="[216,271,1395,1421]" italics="true" pageId="9" pageNumber="502">et al.</emphasis>
2011
</bibRefCitation>
: fig. 37A, B), lacking in
<taxonomicName id="4C274D4FFFEFE40FFDCBFAA5FF47FA1E" authority="Merle" authorityName="Merle" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFDCBFAA5FC88FA3E" box="[627,777,1395,1421]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFF77FA42FE99FA1D" box="[207,280,1428,1454]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
Species such as
<taxonomicName id="4C274D4FFFEFE40FFE04FA45FD22FA1E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Vokes" baseAuthorityYear="1990" box="[444,675,1427,1453]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" status="comb. nov.">
<emphasis id="B953EADEFFEFE40FFE04FA45FD22FA1E" box="[444,675,1427,1453]" italics="true" pageId="9" pageNumber="502">Paleochicoreus initialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFEFE40FFD11FA42FC89FA1E" box="[681,776,1427,1454]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFEFE40FFF09FA62FE9FFA7E" baseAuthorityName="Eames" baseAuthorityYear="1957" box="[177,286,1460,1486]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="newtoni">
<emphasis id="B953EADEFFEFE40FFF09FA62FF41FA7E" box="[177,192,1460,1485]" italics="true" pageId="9" pageNumber="502">P</emphasis>
.
<emphasis id="B953EADEFFEFE40FFF74FA62FE9FFA7E" box="[204,286,1460,1485]" italics="true" pageId="9" pageNumber="502">newtoni</emphasis>
</taxonomicName>
represent ideal ancestral lineages for
<taxonomicName id="4C274D4FFFEFE40FFD23FA65FD7FFA7E" authorityName="Montfort" authorityYear="1810" box="[667,766,1459,1485]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFD23FA65FD7FFA7E" box="[667,766,1459,1485]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFEFE40FFF3CFA05FE80FA5E" authorityName="Arakawa" authorityYear="1964" box="[132,257,1491,1517]" class="Gastropoda" family="Muricidae" genus="Chicomurex" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFF3CFA05FE80FA5E" box="[132,257,1491,1517]" italics="true" pageId="9" pageNumber="502">Chicomurex</emphasis>
</taxonomicName>
or
<emphasis id="B953EADEFFEFE40FFE90FA05FE1EFA5E" box="[296,415,1491,1517]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
(
<bibRefCitation id="EFB64B3DFFEFE40FFE08FA05FDB1FA5E" author="VOKES E. H." box="[432,560,1491,1517]" pageId="9" pageNumber="502" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes 1990</bibRefCitation>
) but cannot readily be assign to one of these branches. This comment is true for other species attributed here to
<taxonomicName id="4C274D4FFFEFE40FFE6FF9C5FD33F99E" authority="Merle" authorityName="Merle" authorityYear="2024" box="[471,690,1555,1581]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFE6FF9C5FDE8F99E" box="[471,617,1555,1581]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFD05F9C2FC86F99D" box="[701,775,1556,1582]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
Here, we figure and give a new description of the
<typeStatus id="549C886EFFEFE40FFD10F9E5FC86F9FE" box="[680,775,1587,1613]" pageId="9" pageNumber="502" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D4FFFEFE40FFF1CF982FE73F9DD" authority="(Deshayes, 1835)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[164,498,1619,1646]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="bicostatus" status="n. comb.">
<emphasis id="B953EADEFFEFE40FFF1CF982FEA5F9DE" box="[164,292,1619,1645]" italics="true" pageId="9" pageNumber="502">P. bicostatus</emphasis>
(
<bibRefCitation id="EFB64B3DFFEFE40FFE8DF985FE6BF9DD" author="DESHAYES G. - P." box="[309,490,1619,1646]" pageId="9" pageNumber="502" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel &amp; Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFDB8F985FDE4F9DE" box="[512,613,1619,1645]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
This very rare species has not been found subsequent to its original description. Information given hereunder is based on the
<typeStatus id="549C886EFFEFE40FFD10F944FC86F91F" box="[680,775,1682,1708]" pageId="9" pageNumber="502" type="holotype">holotype</typeStatus>
retrieved from the collections of the École des Mines, Lyon.
</paragraph>
</subSubSection>
</treatment>
</document>