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<document id="C3DD57C6A64EFC74D2163C3800BBB3D2" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFC9E429FC09FD60FA31F9BE" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pronucellopsis Merle &amp; Pacaud &amp; Ledon &amp; Goret 2024, n. gen." docType="treatment" docUuid="57351378-2B58-4F05-8CC1-C8DE453B321B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="540" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="540" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="88887A7D267E2223205F765238DAE665">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="DFD5C64EEC3A92D70303DAA15C6420F2">Merle, Didier</mods:namePart>
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<mods:namePart id="9FDC6D0780EFC91B64171F224196B170">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="08E4EAF10B2268741A4DF3AF90620CE0">Ledon, Daniel</mods:namePart>
<mods:affiliation id="0A7DFE2E9029FC4219D4767873DD3774">Muséum national dhistoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
<mods:nameIdentifier id="153C77988EC03899977437A8BC65920C" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="8406E3922C7A76D52BEBD2899166DB9F">Goret, Bernard</mods:namePart>
<mods:affiliation id="7D0332080434950AB176C6258EE74071">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
<mods:nameIdentifier id="CC282A83509FA45E34BD04937E2DC5D1" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="3A1DC527C28C597043ABC41EFE1702F9">Geodiversitas</mods:title>
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<mods:date id="D6460DB37E194190738D750410B2109C">2024</mods:date>
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<treatment id="038E87DAFFC9E429FC09FD60FA31F9BE" ID-DOI="http://doi.org/10.5281/zenodo.13985855" ID-GBIF-Taxon="244735034" ID-Zenodo-Dep="13985855" LSID="urn:lsid:zoobank.org:act:57351378-2B58-4F05-8CC1-C8DE453B321B" httpUri="http://treatment.plazi.org/id/038E87DAFFC9E429FC09FD60FA31F9BE" lastPageNumber="540" pageId="47" pageNumber="540" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae">
<subSubSection id="C33D6547FFC9E429FC09FD60FA2EFCB8" pageId="47" pageNumber="540" type="nomenclature">
<paragraph id="8B9836CCFFC9E429FC09FD60FAAAFD63" blockId="47.[945,1323,694,720]" box="[945,1323,694,720]" pageId="47" pageNumber="540">
<heading id="D0D081A0FFC9E429FC09FD60FAAAFD63" box="[945,1323,694,720]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="540" reason="2">
Genus
<taxonomicName id="4C274D4FFFC9E429FC46FD60FB57FD63" ID-CoL="CN5N5" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1022,1238,694,720]" class="Gastropoda" family="Muricidae" genus="Pronucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="genus" status="n. gen.">
<emphasis id="B953EADEFFC9E429FC46FD60FB0CFD63" box="[1022,1165,694,720]" italics="true" pageId="47" pageNumber="540">Pronucellopsis</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFC9E429FB59FD60FAAAFD63" box="[1249,1323,694,720]" pageId="47" pageNumber="540" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C33D6547FFC9E429FC95FCEAFADEFC0B" pageId="47" pageNumber="540" type="materials_examined">
<paragraph id="8B9836CCFFC9E429FC95FCEAFBCDFCDD" blockId="47.[813,1455,828,878]" pageId="47" pageNumber="540">
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<typeStatus id="549C886EFFC9E429FC95FCEAFCDEFCE1" box="[813,863,828,851]" pageId="47" pageNumber="540">TYPE</typeStatus>
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<smallCapsWord id="8D7EA010FFC9E429FCDFFC97FC2EFCE1" baselines="846" box="[871,943,833,850]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="47" pageNumber="540">SPECIES</smallCapsWord>
. —
<taxonomicName id="4C274D4FFFC9E429FC5AFCEAFA2DFCE7" authority="Merle, Ledon &amp; Goret" authorityName="Merle, Ledon &amp; Goret" authorityYear="2024" box="[994,1452,828,852]" class="Gastropoda" family="Muricidae" genus="Pronucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="pacaudi" status="sp. nov.">
<emphasis id="B953EADEFFC9E429FC5AFCEAFB3BFCE7" box="[994,1210,828,852]" italics="true" pageId="47" pageNumber="540">Pronucellopsis pacaudi</emphasis>
Merle, Ledon &amp; Goret
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFC9E429FC96FC81FCE5FCDD" box="[814,868,855,878]" pageId="47" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
by present designation.
</paragraph>
<paragraph id="8B9836CCFFC9E429FC95FC51FADEFC0B" blockId="47.[811,1456,902,953]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FC95FC51FC10FC2E" baselines="920,921" box="[813,913,903,926]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Included" pageId="47" pageNumber="540">INCLUDED</smallCapsWord>
<smallCapsWord id="8D7EA010FFC9E429FC2DFC5AFC5BFC2E" baselines="921" box="[917,986,908,925]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="47" pageNumber="540">SPECIES</smallCapsWord>
. —
<taxonomicName id="4C274D4FFFC9E429FBB9FC50FA2DFC2D" authority="Merle, Ledon &amp; Goret" authorityName="Merle, Ledon &amp; Goret" authorityYear="2024" box="[1025,1452,902,926]" class="Gastropoda" family="Muricidae" genus="Pronucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="species" species="pacaudi" status="n. gen., n. sp.">
<emphasis id="B953EADEFFC9E429FBB9FC50FB4CFC2D" box="[1025,1229,902,926]" italics="true" pageId="47" pageNumber="540">Pronucellopsis pacaudi</emphasis>
Merle, Ledon &amp; Goret
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFC9E429FC93FC77FC33FC0B" box="[811,946,929,953]" pageId="47" pageNumber="540" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, Paris Basin,
<collectingCountry id="F330765CFFC9E429FB8AFC77FBF7FC0B" box="[1074,1142,929,952]" name="France" pageId="47" pageNumber="540">France</collectingCountry>
; Paleocene (Thanetian).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC9E429FC95FC07FABEFB65" pageId="47" pageNumber="540" type="diagnosis">
<paragraph id="8B9836CCFFC9E429FC95FC07FABEFB65" blockId="47.[811,1457,976,1238]" pageId="47" pageNumber="540">
<smallCapsWord id="8D7EA010FFC9E429FC95FC07FC14FC54" baselines="994,995" box="[813,917,977,1000]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="47" pageNumber="540">DIAGNOSIS</smallCapsWord>
. — Subfusiform teleoconch with rounded whorls. Poorly marked cords. No IP. P6 weak, not atrophied. ADP, MP weak. Secondary cords (s1, s2, s3, s4) appearing late in ontogeny. Primary cords P3 to P6 appearing after third whorl. Varices formed by one or two abaxial lamellae on early whorls, occasionally reinforced by several lamellae on succeeding whorls. Number of varices: between 13 and 15. Weak, rounded shoulder sinus. Posterior sinus on sutural ramp deep. Internal denticles usually weak: ID missing. D1 stronger than D2. D6 present. Columellar lip smooth. Siphonal canal narrow. Microstructure of three aragonite layers.
</paragraph>
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<subSubSection id="C33D6547FFC9E429FC95FB22FA31F9BE" pageId="47" pageNumber="540" type="discussion">
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<smallCapsWord id="8D7EA010FFC9E429FC95FB22FC2DFABE" baselines="1287,1288" box="[813,940,1268,1294]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="47" pageNumber="540">COMMENTS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFC9E429FC95FAC2FA31F9BE" blockId="47.[811,1456,1268,1550]" pageId="47" pageNumber="540">
<taxonomicName id="4C274D4FFFC9E429FC95FAC2FB85FA9D" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[813,1028,1300,1326]" class="Gastropoda" family="Muricidae" genus="Pronucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFC9E429FC95FAC2FC3DFA9D" box="[813,956,1300,1326]" italics="true" pageId="47" pageNumber="540">Pronucellopsis</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFC9E429FBA8FAC3FBDBFA9C" box="[1040,1114,1301,1327]" pageId="47" pageNumber="540" rank="genus">n. gen.</taxonomicNameLabel>
is the sister taxon of
<taxonomicName id="4C274D4FFFC9E429FA86FAC2FA2EFA9D" authorityName="Merle" authorityYear="2005" box="[1342,1455,1300,1326]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC9E429FA86FAC2FA2EFA9D" box="[1342,1455,1300,1326]" italics="true" pageId="47" pageNumber="540">Nucellopsis</emphasis>
</taxonomicName>
(clade E,
<figureCitation id="131C2A49FFC9E429FC28FAE2FC4CFAFD" box="[912,973,1332,1358]" captionStart="FIG" captionStartId="6.[134,143,1784,1801]" captionTargetBox="[134,618,235,1734]" captionTargetId="graphics-161@6.[134,618,235,1734]" captionTargetPageId="6" captionText="FIG. 1. — Strict Consensus tree of 7 equally parsimonious trees (length 107, ci 0.53, ri 0.78) generated by Winclada and computed using the option Heuristic Analysis (mult* max*), all characters non additive." figureDoi="http://doi.org/10.5281/zenodo.13985762" httpUri="https://zenodo.org/record/13985762/files/figure.png" pageId="47" pageNumber="540">Fig. 1</figureCitation>
). Both taxa share a weak or missing shoulder sinus.
<taxonomicName id="4C274D4FFFC9E429FCCAFA82FBCBFADD" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[882,1098,1364,1390]" class="Gastropoda" family="Muricidae" genus="Pronucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFC9E429FCCAFA82FB80FADD" box="[882,1025,1364,1390]" italics="true" pageId="47" pageNumber="540">Pronucellopsis</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFC9E429FBEEFA83FB20FADC" box="[1110,1185,1365,1391]" pageId="47" pageNumber="540" rank="genus">n. gen.</taxonomicNameLabel>
is easily differentiated in having a more elongated shape and poorly developed internal denticles. It also displays poor development of the spiral cords which always remain weak and appear late in ontogeny. This feature is present in early muricids from the Paleocene (e.g.,
<taxonomicName id="4C274D4FFFC9E429FC94FA22FC04F9BE" authorityName="Jousseaume" authorityYear="1880" box="[812,901,1524,1549]" class="Gastropoda" family="Muricidae" genus="Poirieria" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC9E429FC94FA22FC04F9BE" box="[812,901,1524,1549]" italics="true" pageId="47" pageNumber="540">Poirieria</emphasis>
</taxonomicName>
or
<taxonomicName id="4C274D4FFFC9E429FC1DFA25FB9AF9BE" authorityName="Shuto" authorityYear="1969" box="[933,1051,1523,1549]" class="Gastropoda" family="Muricidae" genus="Flexopteron" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="540" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC9E429FC1DFA25FB9AF9BE" box="[933,1051,1523,1549]" italics="true" pageId="47" pageNumber="540">Flexopteron</emphasis>
</taxonomicName>
) and seems a plesiomorphic character.
</paragraph>
</subSubSection>
</treatment>
</document>