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<document id="F68DAAA166257625ACDDFF066B175477" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFC0E42FFCFFF936FD46FD63" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Crassimurex (Pliocrassimurex) hirtus Merle, Pacaud, Ledon &amp; Goret, 2024, n. subgen., n. sp." docType="treatment" docUuid="79134847-C7B3-4A32-BFA1-CD185BD5A1ED" docUuidSource="ZooBank" docVersion="6" lastPageNumber="532" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="531" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="14A484B089D8E486BF9067D2892BDB67">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="DB642B93C54FB3A597B32D178D36452E">Merle, Didier</mods:namePart>
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<mods:namePart id="F80D578A681938F527E04EC34E8E70AD">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="3D5D3EC394F12FF250A0CC22479ECC26">Ledon, Daniel</mods:namePart>
<mods:affiliation id="B3414D13BD63DA357D3F7FC2A525D42F">Muséum national dhistoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
<mods:nameIdentifier id="6EFCBAB3EC380338C27A049C0E074E5B" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="E73BA4C5DE6B615252A3FE84E11742B0">Goret, Bernard</mods:namePart>
<mods:affiliation id="5FE998687D1ACB10A323FB8BB7FB8043">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:title id="5A69D4102969C14C2EFCDA9FA98CFE6D">Geodiversitas</mods:title>
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<mods:date id="FAD5A6774C4C0F0295C682A179E67C14">2024</mods:date>
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<treatment id="038E87DAFFC0E42FFCFFF936FD46FD63" ID-DOI="http://doi.org/10.5281/zenodo.13991589" ID-GBIF-Taxon="244735086" ID-Zenodo-Dep="13991589" LSID="urn:lsid:zoobank.org:act:79134847-C7B3-4A32-BFA1-CD185BD5A1ED" httpUri="http://treatment.plazi.org/id/038E87DAFFC0E42FFCFFF936FD46FD63" lastPageId="41" lastPageNumber="532" pageId="38" pageNumber="531" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae" scope_subGenus="Pliocrassimurex">
<subSubSection id="C33D6547FFC0E420FCFFF936FA17F948" box="[839,1430,1760,1788]" pageId="38" pageNumber="531" type="nomenclature">
<paragraph id="8B9836CCFFC0E420FCFFF936FA17F948" blockId="38.[839,1430,1760,1819]" box="[839,1430,1760,1788]" pageId="38" pageNumber="531">
<taxonomicName id="4C274D4FFFC0E420FCFFF936FB57F94F" ID-CoL="CN5MV" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[839,1238,1760,1788]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="38" pageNumber="531" phylum="Mollusca" rank="species" species="hirtus" status="n. subgen., n. sp." subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFC0E420FCFFF936FC4CF949" bold="true" box="[839,973,1760,1786]" italics="true" pageId="38" pageNumber="531">Crassimurex</emphasis>
(
<emphasis id="B953EADEFFC0E420FC65F934FB06F94F" bold="true" box="[989,1159,1762,1788]" italics="true" pageId="38" pageNumber="531">Pliocrassimurex</emphasis>
)
<emphasis id="B953EADEFFC0E420FB2EF934FB57F94F" bold="true" box="[1174,1238,1762,1788]" italics="true" pageId="38" pageNumber="531">hirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC0E420FB65F937FA17F948" box="[1245,1430,1760,1787]" pageId="38" pageNumber="531" rank="species">n. subgen., n. sp.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C33D6547FFC0E420FBB0F8D7FA2EF8EB" pageId="38" pageNumber="531" type="description">
<paragraph id="8B9836CCFFC0E420FBB0F8D7FB55F8A9" blockId="38.[839,1430,1760,1819]" box="[1032,1236,1793,1819]" pageId="38" pageNumber="531">
(
<figureCitation id="131C2A49FFC0E420FBA9F8D4FBD6F8A9" box="[1041,1111,1794,1818]" captionStart="FIG" captionStartId="39.[132,143,1192,1209]" captionTargetBox="[150,1438,242,1128]" captionTargetId="figure-538@39.[609,1070,505,769]" captionTargetPageId="39" captionText="FIG. 21. — Spiral cords and internal denticles of the outer lip in Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp. (A-F) and Crassimurex (s.s.) calcitrapa (Lamarck, 1803) (G, H) for comparison: A, B, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean); C, D, paratype, MNHN.F.A77752, Molins del Rei; E, F, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei; G, H, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de lOrme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985810" httpUri="https://zenodo.org/record/13985810/files/figure.png" pageId="38" pageNumber="531">Figs 21</figureCitation>
A-F; 28C-E)
</paragraph>
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urn:lsid:zoobank.org:act:
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<smallCapsWord id="8D7EA010FFC0E420FC95F849FCDDF807" baselines="1968,1968" box="[813,860,1951,1974]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="38" pageNumber="531">TYPE</smallCapsWord>
<smallCapsWord id="8D7EA010FFC0E420FCD8F872FC38F806" baselines="1969" box="[864,953,1956,1973]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="38" pageNumber="531">MATERIAL</smallCapsWord>
. —
<materialsCitation id="3B4F3C91FFC0E420FC67F849FA2BF805" ID-GBIF-Occurrence="4958671423" box="[991,1450,1950,1976]" country="Spain" location="Llobregat Basin" municipality="Molins del Rei" pageId="38" pageNumber="531" specimenCount="1" typeStatus="holotype">
<emphasis id="B953EADEFFC0E420FC67F849FBF8F80B" bold="true" box="[991,1145,1951,1976]" pageId="38" pageNumber="531">
<typeStatus id="549C886EFFC0E420FC67F849FBBAF804" box="[991,1083,1951,1975]" pageId="38" pageNumber="531" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F330765CFFC0E420FBFBF876FBF8F80B" box="[1091,1145,1952,1976]" name="Spain" pageId="38" pageNumber="531">Spain</collectingCountry>
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<location id="8EF86017FFC0E420FB32F848FA9AF805" LSID="urn:lsid:plazi:treatment:038E87DAFFC0E42FFCFFF936FD46FD63:8EF86017FFC0E420FB32F848FA9AF805" box="[1162,1307,1950,1974]" country="Spain" municipality="Molins del Rei" name="Llobregat Basin" pageId="38" pageNumber="531">Llobregat Basin</location>
,
<collectingMunicipality id="6BFCACB6FFC0E420FA9BF848FA2BF805" box="[1315,1450,1950,1974]" pageId="38" pageNumber="531">Molins del Rei</collectingMunicipality>
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;
<materialsCitation id="3B4F3C91FFC0E420FC93F86FFA31F85F" ID-GBIF-Occurrence="4958671321" collectionCode="MNHN" collectorName="Yellow" country="Spain" location="Molins del Rei" municipality="Anna" pageId="38" pageNumber="531" specimenCount="1" typeStatus="holotype">
<collectorName id="26D2531AFFC0E420FC93F86FFCE6F862" box="[811,871,1977,2001]" pageId="38" pageNumber="531">Yellow</collectorName>
and grey clays of
<location id="8EF86017FFC0E420FBB2F86FFB10F862" LSID="urn:lsid:plazi:treatment:038E87DAFFC0E42FFCFFF936FD46FD63:8EF86017FFC0E420FBB2F86FFB10F862" box="[1034,1169,1977,2001]" country="Spain" municipality="Anna" name="Molins del Rei" pageId="38" pageNumber="531">Molins del Rei</location>
(quarry
<collectingMunicipality id="6BFCACB6FFC0E420FB58F86FFA97F862" box="[1248,1302,1977,2001]" pageId="38" pageNumber="531">Anna</collectingMunicipality>
); Pliocene (Zanclean);
<collectionCode id="ED36AE09FFC0E420FCD2F802FC38F858" box="[874,953,2004,2027]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="531">MNHN</collectionCode>
.F.A77751 (
<collectionCode id="ED36AE09FFC0E420FB98F805FBF1F858" box="[1056,1136,2003,2027]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="531">MNHN</collectionCode>
coll.),H:
<quantity id="4CDF9B29FFC0E420FB7CF805FA84F858" box="[1220,1285,2003,2027]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="38" pageNumber="531" unit="mm" value="28.0">28 mm</quantity>
(
<figureCitation id="131C2A49FFC0E420FAB7F805FAF4F858" box="[1295,1397,2003,2028]" captionStart="FIG" captionStartId="39.[132,143,1192,1209]" captionTargetBox="[150,1438,242,1128]" captionTargetId="figure-538@39.[609,1070,505,769]" captionTargetPageId="39" captionText="FIG. 21. — Spiral cords and internal denticles of the outer lip in Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp. (A-F) and Crassimurex (s.s.) calcitrapa (Lamarck, 1803) (G, H) for comparison: A, B, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean); C, D, paratype, MNHN.F.A77752, Molins del Rei; E, F, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei; G, H, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de lOrme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985810" httpUri="https://zenodo.org/record/13985810/files/figure.png" pageId="38" pageNumber="531">Figs 21A,B</figureCitation>
;
<figureCitation id="131C2A49FFC0E420FAC2F805FA24F85F" box="[1402,1445,2003,2028]" captionStart="FIG" captionStartId="48.[132,143,1301,1318]" captionTargetBox="[145,1443,235,1238]" captionTargetId="figure-535@48.[700,1258,235,783]" captionTargetPageId="48" captionText="FIG. 28. — New species of Flexopteron Shuto, 1969, Crassimurex (Pliocrassimurex) n. subgen., Nucellopsis Merle, 2005 and Pronucellopsis Merle, n. gen.: A, B, Flexopteron liancurtense n. sp.: A, holotype, MNHN.F.J02578 (Cossmann coll.), Liancourt-Saint-Pierre, France, Ypresian; B, paratype: MNHN.F.A77758 (Merle coll.), Liancourt-Saint-Pierre; C-E, Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp.: C, paratype (juvenile), MNHN.F.A77753, Molins del Rei, Spain, Zanclean; D, paratype, MNHN.F.A77752, Molins del Rei; E, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei; F, G, Nucellopsis parisiensis n. sp.: F, holotype, MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle), France, Bartonian; G, paratype, MNHN.F.A90963 (Faullummel coll.), Saint-Witz (Le Guépelle); H, I, Pronucellopsis pacaudi Merle, Ledon &amp; Goret, n. gen., n. sp.: H, holotype, MNHN.F.J02587 (Cossmann coll.), Châlons-sur-Vesle, France, Thanetian; I, paratype, MNHN.F.A90829 (Pacaud coll.), Jonchery-sur-Vesle, France, Thanetian. Scale bars: 5 mm. Scale bar: A, B, 3 mm; C, D, F-I, 5 mm; E, 10 mm. Credits: P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985833" httpUri="https://zenodo.org/record/13985833/files/figure.png" pageId="38" pageNumber="531">28E</figureCitation>
).
</materialsCitation>
<materialsCitation id="3B4F3C91FFC1E421FF3CFA85FEFBFA13" ID-GBIF-Occurrence="4958671402" collectionCode="MNHN" collectorName="Yellow" country="Spain" location="Molins del Rei" municipality="Anna" pageId="39" pageNumber="532" specimenCount="1" typeStatus="holotype">
<emphasis id="B953EADEFFC1E421FF3CFA85FE9EFADE" bold="true" box="[132,287,1363,1389]" pageId="39" pageNumber="532">
<typeStatus id="549C886EFFC1E421FF3CFA85FF60FAD8" box="[132,225,1363,1387]" pageId="39" pageNumber="532" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F330765CFFC1E421FF51FA83FE9EFADE" box="[233,287,1365,1389]" name="Spain" pageId="39" pageNumber="532">Spain</collectingCountry>
</emphasis>
• 2 spm; same as for the holotype;
<collectionCode id="ED36AE09FFC1E421FDEBFA82FD24FAD8" box="[595,677,1364,1387]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="39" pageNumber="532">MNHN</collectionCode>
.F.A77752 (
<collectionCode id="ED36AE09FFC1E421FF33FAB8FF5EFA35" box="[139,223,1390,1414]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="39" pageNumber="532">MNHN</collectionCode>
coll.) (
<figureCitation id="131C2A49FFC1E421FE90FAB8FE24FA36" box="[296,421,1390,1414]" captionStart="FIG" captionStartId="39.[132,143,1192,1209]" captionTargetBox="[150,1438,242,1128]" captionTargetId="figure-538@39.[609,1070,505,769]" captionTargetPageId="39" captionText="FIG. 21. — Spiral cords and internal denticles of the outer lip in Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp. (A-F) and Crassimurex (s.s.) calcitrapa (Lamarck, 1803) (G, H) for comparison: A, B, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean); C, D, paratype, MNHN.F.A77752, Molins del Rei; E, F, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei; G, H, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de lOrme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985810" httpUri="https://zenodo.org/record/13985810/files/figure.png" pageId="39" pageNumber="532">Figs 21C, D</figureCitation>
;
<figureCitation id="131C2A49FFC1E421FE09FAB8FE61FA36" box="[433,480,1390,1414]" captionStart="FIG" captionStartId="48.[132,143,1301,1318]" captionTargetBox="[145,1443,235,1238]" captionTargetId="figure-535@48.[700,1258,235,783]" captionTargetPageId="48" captionText="FIG. 28. — New species of Flexopteron Shuto, 1969, Crassimurex (Pliocrassimurex) n. subgen., Nucellopsis Merle, 2005 and Pronucellopsis Merle, n. gen.: A, B, Flexopteron liancurtense n. sp.: A, holotype, MNHN.F.J02578 (Cossmann coll.), Liancourt-Saint-Pierre, France, Ypresian; B, paratype: MNHN.F.A77758 (Merle coll.), Liancourt-Saint-Pierre; C-E, Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp.: C, paratype (juvenile), MNHN.F.A77753, Molins del Rei, Spain, Zanclean; D, paratype, MNHN.F.A77752, Molins del Rei; E, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei; F, G, Nucellopsis parisiensis n. sp.: F, holotype, MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle), France, Bartonian; G, paratype, MNHN.F.A90963 (Faullummel coll.), Saint-Witz (Le Guépelle); H, I, Pronucellopsis pacaudi Merle, Ledon &amp; Goret, n. gen., n. sp.: H, holotype, MNHN.F.J02587 (Cossmann coll.), Châlons-sur-Vesle, France, Thanetian; I, paratype, MNHN.F.A90829 (Pacaud coll.), Jonchery-sur-Vesle, France, Thanetian. Scale bars: 5 mm. Scale bar: A, B, 3 mm; C, D, F-I, 5 mm; E, 10 mm. Credits: P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985833" httpUri="https://zenodo.org/record/13985833/files/figure.png" pageId="39" pageNumber="532">28D</figureCitation>
,
<collectionCode id="ED36AE09FFC1E421FE53FAB8FDC1FA36" box="[491,576,1390,1413]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="39" pageNumber="532">MNHN</collectionCode>
.F.A77753 (
<collectionCode id="ED36AE09FFC1E421FD0BFAB8FC86FA35" box="[691,775,1390,1414]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="39" pageNumber="532">MNHN</collectionCode>
coll.) (
<figureCitation id="131C2A49FFC1E421FF7CFA5EFEB7FA13" box="[196,310,1416,1440]" captionStart="FIG" captionStartId="39.[132,143,1192,1209]" captionTargetBox="[150,1438,242,1128]" captionTargetId="figure-538@39.[609,1070,505,769]" captionTargetPageId="39" captionText="FIG. 21. — Spiral cords and internal denticles of the outer lip in Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp. (A-F) and Crassimurex (s.s.) calcitrapa (Lamarck, 1803) (G, H) for comparison: A, B, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean); C, D, paratype, MNHN.F.A77752, Molins del Rei; E, F, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei; G, H, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de lOrme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985810" httpUri="https://zenodo.org/record/13985810/files/figure.png" pageId="39" pageNumber="532">Figs 21E, F</figureCitation>
;
<figureCitation id="131C2A49FFC1E421FEF8FA5EFEEFFA13" box="[320,366,1416,1440]" captionStart="FIG" captionStartId="48.[132,143,1301,1318]" captionTargetBox="[145,1443,235,1238]" captionTargetId="figure-535@48.[700,1258,235,783]" captionTargetPageId="48" captionText="FIG. 28. — New species of Flexopteron Shuto, 1969, Crassimurex (Pliocrassimurex) n. subgen., Nucellopsis Merle, 2005 and Pronucellopsis Merle, n. gen.: A, B, Flexopteron liancurtense n. sp.: A, holotype, MNHN.F.J02578 (Cossmann coll.), Liancourt-Saint-Pierre, France, Ypresian; B, paratype: MNHN.F.A77758 (Merle coll.), Liancourt-Saint-Pierre; C-E, Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp.: C, paratype (juvenile), MNHN.F.A77753, Molins del Rei, Spain, Zanclean; D, paratype, MNHN.F.A77752, Molins del Rei; E, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei; F, G, Nucellopsis parisiensis n. sp.: F, holotype, MNHN.F.R64955 (Merle coll.), Saint-Witz (Le Guépelle), France, Bartonian; G, paratype, MNHN.F.A90963 (Faullummel coll.), Saint-Witz (Le Guépelle); H, I, Pronucellopsis pacaudi Merle, Ledon &amp; Goret, n. gen., n. sp.: H, holotype, MNHN.F.J02587 (Cossmann coll.), Châlons-sur-Vesle, France, Thanetian; I, paratype, MNHN.F.A90829 (Pacaud coll.), Jonchery-sur-Vesle, France, Thanetian. Scale bars: 5 mm. Scale bar: A, B, 3 mm; C, D, F-I, 5 mm; E, 10 mm. Credits: P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985833" httpUri="https://zenodo.org/record/13985833/files/figure.png" pageId="39" pageNumber="532">28C</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF586644FFC1E421FF3CFB7EFEEEFAA7" ID-DOI="http://doi.org/10.5281/zenodo.13985810" ID-Zenodo-Dep="13985810" httpUri="https://zenodo.org/record/13985810/files/figure.png" pageId="39" pageNumber="532" startId="39.[132,143,1192,1209]" targetBox="[150,1438,242,1128]" targetPageId="39" targetType="figure">
<paragraph id="8B9836CCFFC1E421FF3CFB7EFEEEFAA7" blockId="39.[132,1457,1192,1300]" pageId="39" pageNumber="532">
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. 21. — Spiral cords and internal denticles of the outer lip in
<taxonomicName id="4C274D4FFFC1E421FD18FB7EFC4FFB0A" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[672,974,1192,1209]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="hirtus" status="n. subgen., n. sp." subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFC1E421FD18FB7EFC86FB0A" box="[672,775,1192,1209]" italics="true" pageId="39" pageNumber="532">Crassimurex</emphasis>
(
<emphasis id="B953EADEFFC1E421FCAAFB7EFC14FB0A" box="[786,917,1192,1209]" italics="true" pageId="39" pageNumber="532">Pliocrassimurex</emphasis>
)
<emphasis id="B953EADEFFC1E421FC18FB7EFC4FFB0A" box="[928,974,1192,1209]" italics="true" pageId="39" pageNumber="532">hirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC1E421FC6DFB7EFBEAFB0A" box="[981,1131,1192,1209]" pageId="39" pageNumber="532" rank="species">n. subgen., n. sp.</taxonomicNameLabel>
(
<emphasis id="B953EADEFFC1E421FBCCFB7EFB12FB0A" bold="true" box="[1140,1171,1192,1209]" pageId="39" pageNumber="532">A -F</emphasis>
) and
<taxonomicName id="4C274D4FFFC1E421FB7DFB7EFAADFB0A" authorityName="Merle" authorityYear="1990" box="[1221,1324,1192,1209]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC1E421FB7DFB7EFAADFB0A" box="[1221,1324,1192,1209]" italics="true" pageId="39" pageNumber="532">Crassimurex</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFC1E421FA8FFB7EFAD5FB0A" box="[1335,1364,1192,1209]" italics="true" pageId="39" pageNumber="532">s.s.</emphasis>
)
<taxonomicName id="4C274D4FFFC1E421FAE7FB7EFE8BFB63" authority="(Lamarck, 1803)" baseAuthorityName="Lamarck" baseAuthorityYear="1803" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="calcitrapa">
<emphasis id="B953EADEFFC1E421FAE7FB7EFA31FB0A" box="[1375,1456,1192,1209]" italics="true" pageId="39" pageNumber="532">calcitrapa</emphasis>
(
<bibRefCitation id="EFB64B3DFFC1E421FF31FB69FE87FB63" author="LAMARCK J. - B." box="[137,262,1215,1232]" pageId="39" pageNumber="532" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="B953EADEFFC1E421FEADFB69FEA2FB63" bold="true" box="[277,291,1215,1232]" pageId="39" pageNumber="532">G</emphasis>
,
<emphasis id="B953EADEFFC1E421FE96FB69FEBDFB63" bold="true" box="[302,316,1215,1232]" pageId="39" pageNumber="532">H</emphasis>
) for comparison:
<emphasis id="B953EADEFFC1E421FE68FB69FE5CFB63" bold="true" box="[464,477,1215,1232]" pageId="39" pageNumber="532">A</emphasis>
,
<emphasis id="B953EADEFFC1E421FE50FB69FE74FB63" bold="true" box="[488,501,1215,1232]" pageId="39" pageNumber="532">B</emphasis>
, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean);
<emphasis id="B953EADEFFC1E421FA3BFB69FA10FB63" bold="true" box="[1411,1425,1215,1232]" pageId="39" pageNumber="532">C</emphasis>
,
<emphasis id="B953EADEFFC1E421FA24FB69FA2BFB63" bold="true" box="[1436,1450,1215,1232]" pageId="39" pageNumber="532">D</emphasis>
, paratype, MNHN.F.A77752, Molins del Rei;
<emphasis id="B953EADEFFC1E421FE56FB03FE7BFB55" bold="true" box="[494,506,1237,1254]" pageId="39" pageNumber="532">E</emphasis>
,
<emphasis id="B953EADEFFC1E421FDBCFB03FD8EFB55" bold="true" box="[516,527,1237,1254]" pageId="39" pageNumber="532">F</emphasis>
, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei;
<emphasis id="B953EADEFFC1E421FB87FB03FBCCFB55" bold="true" box="[1087,1101,1237,1254]" pageId="39" pageNumber="532">G</emphasis>
,
<emphasis id="B953EADEFFC1E421FBEFFB03FBE4FB55" bold="true" box="[1111,1125,1237,1254]" pageId="39" pageNumber="532">H</emphasis>
, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de lOrme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS).
</paragraph>
</caption>
<subSubSection id="C33D6547FFC1E421FF3CFA6EFD49FA59" pageId="39" pageNumber="532" type="etymology">
<paragraph id="8B9836CCFFC1E421FF3CFA6EFD49FA59" blockId="39.[132,775,1464,1514]" pageId="39" pageNumber="532">
<smallCapsWord id="8D7EA010FFC1E421FF3CFA6EFF7FFA7D" baselines="1481,1482" box="[132,254,1464,1487]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="39" pageNumber="532">ETYMOLOGY</smallCapsWord>
. — From the Latin adjective
<taxonomicName id="4C274D4FFFC1E421FD94FA6EFDE3FA63" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[556,610,1464,1488]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="hirtus">
<emphasis id="B953EADEFFC1E421FD94FA6EFDE3FA63" box="[556,610,1464,1488]" italics="true" pageId="39" pageNumber="532">hirtus</emphasis>
</taxonomicName>
meaning bristly, because of the cord spines of the species; gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC1E421FF3CF9D4FD2EF9D7" pageId="39" pageNumber="532" type="materials_examined">
<paragraph id="8B9836CCFFC1E421FF3CF9D4FC89F9A9" blockId="39.[132,776,1538,1562]" box="[132,776,1538,1562]" pageId="39" pageNumber="532">
<smallCapsWord id="8D7EA010FFC1E421FF3CF9D4FF33F9AB" baselines="1555,1556" box="[132,178,1538,1561]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="39" pageNumber="532">
<typeStatus id="549C886EFFC1E421FF3CF9D4FF33F9AB" box="[132,178,1538,1561]" pageId="39" pageNumber="532">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFC1E421FF0EF9D1FE8AF9AB" baselines="1556" box="[182,267,1543,1560]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="39" pageNumber="532">HORIZON</smallCapsWord>
. — Yellow and grey clays of Molins del Rei (quarry Anna).
</paragraph>
<paragraph id="8B9836CCFFC1E421FF3CF9E4FD2EF9D7" blockId="39.[131,775,1585,1636]" pageId="39" pageNumber="532">
<smallCapsWord id="8D7EA010FFC1E421FF3CF9E4FF34F9F4" baselines="1603,1603" box="[132,181,1586,1609]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="39" pageNumber="532">
<typeStatus id="549C886EFFC1E421FF3CF9E4FF34F9F4" box="[132,181,1586,1609]" pageId="39" pageNumber="532">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFC1E421FF03F9E1FE97F9FB" baselines="1604" box="[187,278,1591,1608]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="39" pageNumber="532">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F330765CFFC1E421FEF8F9E7FEF8F9FA" box="[320,377,1585,1609]" name="Spain" pageId="39" pageNumber="532">Spain</collectingCountry>
,
<location id="8EF86017FFC1E421FE3BF9E7FD9CF9FA" LSID="urn:lsid:plazi:treatment:038E87DAFFC0E42FFCFFF936FD46FD63:8EF86017FFC1E421FE3BF9E7FD9CF9FA" box="[387,541,1585,1609]" country="Spain" county="Molins del Rei" municipality="Pliocene" name="Llobregat Basin" pageId="39" pageNumber="532">Llobregat Basin</location>
,
<collectingMunicipality id="6BFCACB6FFC1E421FD9EF9E7FDF9F9FA" box="[550,632,1585,1609]" pageId="39" pageNumber="532">Pliocene</collectingMunicipality>
(Zanclean; see
<bibRefCitation id="EFB64B3DFFC1E421FF3BF99AFEF0F9D7" author="SALVANY J. M. &amp; AGUIRRE J." box="[131,369,1612,1636]" pageId="39" pageNumber="532" pagination="1 - 19" refId="ref48536" refString="SALVANY J. M. &amp; AGUIRRE J. 2020. - The Neogene and Quaternary deposits of the Barcelona city through the high-speed train line. Geologica Acta 18 (10): 1 - 19. https: // doi. org / 10.1344 / GeologicaActa 2020.18.10" type="journal article" year="2020">Salvany &amp; Aguirre 2020</bibRefCitation>
),
<collectingCounty id="62F94E40FFC1E421FE3FF99AFD96F9D7" box="[391,535,1612,1636]" pageId="39" pageNumber="532">Molins del Rei</collectingCounty>
(quarry Anna).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC1E421FF3CF9AAFD19F920" box="[132,664,1659,1684]" pageId="39" pageNumber="532" type="distribution">
<paragraph id="8B9836CCFFC1E421FF3CF9AAFD19F920" blockId="39.[132,664,1659,1684]" box="[132,664,1659,1684]" pageId="39" pageNumber="532">
<smallCapsWord id="8D7EA010FFC1E421FF3CF9AAFE93F921" baselines="1677,1678" box="[132,274,1660,1683]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="39" pageNumber="532">DISTRIBUTION</smallCapsWord>
. — Only known from the
<typeStatus id="549C886EFFC1E421FDA4F9AAFDC7F927" box="[540,582,1660,1684]" pageId="39" pageNumber="532">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC1E421FF3CF965FBD8F8BF" pageId="39" pageNumber="532" type="description">
<paragraph id="8B9836CCFFC1E421FF3CF965FE94F978" blockId="39.[131,776,1715,2028]" box="[132,277,1715,1741]" pageId="39" pageNumber="532">
<smallCapsWord id="8D7EA010FFC1E421FF3CF965FE94F978" baselines="1734,1734" box="[132,277,1715,1741]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="39" pageNumber="532">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFC1E421FF3CF904FBD8F8BF" blockId="39.[131,776,1715,2028]" lastBlockId="39.[811,1457,1363,1804]" pageId="39" pageNumber="532">
Protoconch not preserved. Teleoconch up to
<quantity id="4CDF9B29FFC1E421FD2AF904FD67F95F" box="[658,742,1746,1772]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="39" pageNumber="532" unit="mm" value="28.0">28 mm</quantity>
in height (
<typeStatus id="549C886EFFC1E421FF59F924FECFF8BF" box="[225,334,1778,1804]" pageId="39" pageNumber="532" type="holotype">holotype</typeStatus>
), up to
<quantity id="4CDF9B29FFC1E421FE08F924FD8CF8BF" box="[432,525,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="39" pageNumber="532" unit="mm" value="7.6">7.6 mm</quantity>
in width, biconic in profile, composed of six whorls. Moderately low spire with carinate whorls. Last whorl up to 76% of total length. Apical angle 68°. Spiral sculpture with marked primary cords. First whorl: not preserved; from second to fourth whorl: presence of P1 and P2, P2 close to abapical suture; fifth whorl: appearance of IP; sixth whorl: appearance of abis infrasutural ramp. P1 to P5 and s1 to s3 on convex part of whorl; P6, ADP, MP and ABP on siphonal canal. Axial sculpture with high, sublamellose varices. On second whorl: 9-10 varices; on third whorl: ten varices; from fourth whorl to fifth whorl: 8-10 varices; on sixth whorl: eight varices. On spire, short cord spines on P1-P2; P1 delimiting periphery. On last whorl, short cord spines on IP, P1-P6, ADP, MP, ABP, but more developed on P1, P2 and P3; P2 delimiting periphery. Ovate aperture up to 31% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip, smooth, adherent. Parietal lip smooth, adherent. Outer lip lacking internal denticles. Pseudoumbilicus moderately large. Siphonal canal open, up to 41% of aperture length, slightly dorsally recurved. Aragonitic microstructure.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC1E42FFC95F8E4FD46FD63" lastPageId="41" lastPageNumber="534" pageId="39" pageNumber="532" type="discussion">
<paragraph id="8B9836CCFFC1E42FFC95F8E4FD46FD63" blockId="39.[811,1455,1842,2028]" lastBlockId="41.[131,775,215,720]" lastPageId="41" lastPageNumber="534" pageId="39" pageNumber="532">
<smallCapsWord id="8D7EA010FFC1E421FC95F8E4FC2DF8F9" baselines="1861,1861" box="[813,940,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="39" pageNumber="532">COMMENTS</smallCapsWord>
By its shape and its sculpture,
<taxonomicName id="4C274D4FFFC1E421FB34F884FA2EF8D8" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1164,1455,1873,1900]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="species" species="hirtus" status="n. subgen., n. sp." subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFC1E421FB34F884FB27F8DF" box="[1164,1190,1874,1900]" italics="true" pageId="39" pageNumber="532">C.</emphasis>
(
<emphasis id="B953EADEFFC1E421FB0FF887FADFF8D8" box="[1207,1374,1873,1899]" italics="true" pageId="39" pageNumber="532">Pliocrassimurex</emphasis>
)
<emphasis id="B953EADEFFC1E421FAC8F887FA2EF8D8" box="[1392,1455,1873,1899]" italics="true" pageId="39" pageNumber="532">hirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC1E421FC95F8A4FC64F83F" box="[813,997,1905,1932]" pageId="39" pageNumber="532" rank="species">n. subgen., n. sp.</taxonomicNameLabel>
resembles members of
<taxonomicName id="4C274D4FFFC1E421FB67F8A4FCE8F818" authority="Merle, 1990" authorityName="Merle" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC1E421FB67F8A4FAE1F83F" box="[1247,1376,1906,1932]" italics="true" pageId="39" pageNumber="532">Crassimurex</emphasis>
<bibRefCitation id="EFB64B3DFFC1E421FADFF8A7FCE8F818" author="MERLE D." pageId="39" pageNumber="532" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle, 1990</bibRefCitation>
</taxonomicName>
. However, until now,
<taxonomicName id="4C274D4FFFC1E421FBE6F844FB63F81F" authorityName="Merle" authorityYear="1990" box="[1118,1250,1938,1964]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC1E421FBE6F844FB63F81F" box="[1118,1250,1938,1964]" italics="true" pageId="39" pageNumber="532">Crassimurex</emphasis>
</taxonomicName>
was not known in the Pliocene and seemed extinct after the Middle Miocene.
<taxonomicName id="4C274D4FFFC1E421FC93F807FC2BF858" authorityName="Merle" authorityYear="1990" box="[811,938,2001,2027]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC1E421FC93F807FC2BF858" box="[811,938,2001,2027]" italics="true" pageId="39" pageNumber="532">Crassimurex</emphasis>
</taxonomicName>
is currently represented by two subgenera,
<taxonomicName id="4C274D4FFFC1E42FFACEF807FF56FF42" authorityName="Merle" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" lastPageId="41" lastPageNumber="534" order="Neogastropoda" pageId="39" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC1E42FFACEF807FF56FF42" italics="true" lastPageId="41" lastPageNumber="534" pageId="39" pageNumber="532">Crassimurex</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFCFE42FFF50FF0EFE86FF42" box="[232,263,216,241]" italics="true" pageId="41" pageNumber="534">s.s.</emphasis>
): early Eocene to middle Eocene,
<collectingCountry id="F330765CFFCFE42FFD3CFF0EFD53FF41" box="[644,722,216,242]" name="France" pageId="41" pageNumber="534">France</collectingCountry>
, and
<taxonomicName id="4C274D4FFFCFE42FFF3CFF2EFE17FEA2" authorityName="Gurs" authorityYear="2001" box="[132,406,247,274]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="subGenus" subGenus="Eopaziella">
<emphasis id="B953EADEFFCFE42FFF3CFF2EFE89FEA1" box="[132,264,248,274]" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
(
<emphasis id="B953EADEFFCFE42FFEA2FF21FE0BFEA2" box="[282,394,247,273]" italics="true" pageId="41" pageNumber="534">Eopaziella</emphasis>
)
</taxonomicName>
: late Eocene to Middle Miocene (Europe). Therefore, the discovery of a species of
<taxonomicName id="4C274D4FFFCFE42FFD75FEC1FF59FEE2" authorityName="Merle" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFD75FEC1FF59FEE2" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
</taxonomicName>
in the Early Pliocene of Spain is surprising and extends the stratigraphic range of the genus.
<taxonomicName id="4C274D4FFFCFE42FFD3CFE81FEC7FE22" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="subGenus" subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFCFE42FFD3CFE81FC86FEC2" box="[644,775,343,369]" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
(
<emphasis id="B953EADEFFCFE42FFF36FEA1FEBDFE22" box="[142,316,375,401]" italics="true" pageId="41" pageNumber="534">Pliocrassimurex</emphasis>
)
</taxonomicName>
differs from
<taxonomicName id="4C274D4FFFCFE42FFE5EFEA1FDEFFE22" authorityName="Merle" authorityYear="1990" box="[486,622,375,401]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFE5EFEA1FDEFFE22" box="[486,622,375,401]" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFCFE42FFD3BFEA1FD22FE23" box="[643,675,375,400]" italics="true" pageId="41" pageNumber="534">s.s.</emphasis>
) and
<emphasis id="B953EADEFFCFE42FFD55FEA1FC86FE22" box="[749,775,375,401]" italics="true" pageId="41" pageNumber="534">C.</emphasis>
(
<emphasis id="B953EADEFFCFE42FFF36FE40FF7FFE03" box="[142,254,406,432]" italics="true" pageId="41" pageNumber="534">Eopaziella</emphasis>
) by having a more obtuse shoulder angle. In
<emphasis id="B953EADEFFCFE42FFF3FFE61FF21FE62" box="[135,160,439,465]" italics="true" pageId="41" pageNumber="534">C.</emphasis>
(
<taxonomicName id="4C274D4FFFCFE42FFF17FE60FED7FE63" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[175,342,438,464]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="531" phylum="Mollusca" rank="subGenus" status="subgen. nov." subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFCFE42FFF17FE60FED7FE63" box="[175,342,438,464]" italics="true" pageId="41" pageNumber="534">Pliocrassimurex</emphasis>
</taxonomicName>
), P2 delimits the periphery on the last whorl, whereas in
<taxonomicName id="4C274D4FFFCFE42FFEF2FE01FE4DFE42" authorityName="Merle" authorityYear="1990" box="[330,460,471,497]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFEF2FE01FE4DFE42" box="[330,460,471,497]" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFCFE42FFE65FE01FE7DFE43" box="[477,508,471,496]" italics="true" pageId="41" pageNumber="534">s.s.</emphasis>
) and
<emphasis id="B953EADEFFCFE42FFD82FE01FDD5FE42" box="[570,596,471,497]" italics="true" pageId="41" pageNumber="534">C.</emphasis>
(
<emphasis id="B953EADEFFCFE42FFDDBFE00FD53FE43" box="[611,722,470,496]" italics="true" pageId="41" pageNumber="534">Eopaziella</emphasis>
), P1 delimits the periphery.
<taxonomicName id="4C274D4FFFCFE42FFE2BFE21FD96FDA2" authorityName="Merle" authorityYear="1990" box="[403,535,503,529]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFE2BFE21FD96FDA2" box="[403,535,503,529]" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFCFE42FFD90FE21FDC6FDA3" box="[552,583,503,528]" italics="true" pageId="41" pageNumber="534">s.s.</emphasis>
) 1990 can share cord spines with
<emphasis id="B953EADEFFCFE42FFEF1FDC1FEE2FD82" box="[329,355,535,561]" italics="true" pageId="41" pageNumber="534">C.</emphasis>
(
<taxonomicName id="4C274D4FFFCFE42FFECCFDC0FD9AFD83" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[372,539,534,560]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="531" phylum="Mollusca" rank="subGenus" status="subgen. nov." subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFCFE42FFECCFDC0FD9AFD83" box="[372,539,534,560]" italics="true" pageId="41" pageNumber="534">Pliocrassimurex</emphasis>
</taxonomicName>
), as seen in the
<typeStatus id="549C886EFFCFE42FFD6FFDC1FC87FD82" box="[727,774,535,561]" pageId="41" pageNumber="534">type</typeStatus>
species
<emphasis id="B953EADEFFCFE42FFF6FFDE0FF70FDE3" box="[215,241,566,592]" italics="true" pageId="41" pageNumber="534">C.</emphasis>
(
<emphasis id="B953EADEFFCFE42FFEB9FDE1FEA1FDE3" box="[257,288,567,592]" italics="true" pageId="41" pageNumber="534">s.s.</emphasis>
)
<taxonomicName id="4C274D4FFFCFE42FFE88FDE0FDDCFDE3" authority="(Lamarck, 1803)" baseAuthorityName="Lamarck" baseAuthorityYear="1803" box="[304,605,566,592]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="calcitrapa">
<emphasis id="B953EADEFFCFE42FFE88FDE0FE18FDE3" box="[304,409,566,592]" italics="true" pageId="41" pageNumber="534">calcitrapa</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFE10FDE0FDD4FDE3" author="LAMARCK J. - B." box="[424,597,566,592]" pageId="41" pageNumber="534" pagination="217 - 227" refId="ref45566" refString="LAMARCK J. - B. 1803. - Memoires sur les fossiles des environs de Paris (suite 1). Annales du Museum d'Histoire naturelle 2 (9): 217 - 227." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131C2A49FFCFE42FFDD5FDE0FD79FDE3" box="[621,760,566,592]" captionStart="FIG" captionStartId="39.[132,143,1192,1209]" captionTargetBox="[150,1438,242,1128]" captionTargetId="figure-538@39.[609,1070,505,769]" captionTargetPageId="39" captionText="FIG. 21. — Spiral cords and internal denticles of the outer lip in Crassimurex (Pliocrassimurex) hirtus n. subgen., n. sp. (A-F) and Crassimurex (s.s.) calcitrapa (Lamarck, 1803) (G, H) for comparison: A, B, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean); C, D, paratype, MNHN.F.A77752, Molins del Rei; E, F, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei; G, H, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de lOrme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985810" httpUri="https://zenodo.org/record/13985810/files/figure.png" pageId="41" pageNumber="534">Fig. 21G, H</figureCitation>
), but it differs by having varices which are not axially aligned.
<taxonomicName id="4C274D4FFFCFE42FFF3BFDA0FE13FD23" authorityName="Gurs" authorityYear="2001" box="[131,402,630,656]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="subGenus" subGenus="Eopaziella">
<emphasis id="B953EADEFFCFE42FFF3BFDA0FE86FD23" box="[131,263,630,656]" italics="true" pageId="41" pageNumber="534">Crassimurex</emphasis>
(
<emphasis id="B953EADEFFCFE42FFEA0FDA0FE09FD23" box="[280,392,630,656]" italics="true" pageId="41" pageNumber="534">Eopaziella</emphasis>
)
</taxonomicName>
resembles
<emphasis id="B953EADEFFCFE42FFDB2FDA0FDA5FD23" box="[522,548,630,656]" italics="true" pageId="41" pageNumber="534">C.</emphasis>
(
<taxonomicName id="4C274D4FFFCFE42FFD8DFDA0FD5DFD23" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[565,732,630,656]" class="Gastropoda" family="Muricidae" genus="Crassimurex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="531" phylum="Mollusca" rank="subGenus" status="subgen. nov." subGenus="Pliocrassimurex">
<emphasis id="B953EADEFFCFE42FFD8DFDA0FD5DFD23" box="[565,732,630,656]" italics="true" pageId="41" pageNumber="534">Pliocrassimurex</emphasis>
</taxonomicName>
) by having varices axially aligned but differs in lacking cord spines (see
<bibRefCitation id="EFB64B3DFFCFE42FFEBBFD63FD80FD63" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[259,513,693,720]" pageId="41" pageNumber="534" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFCFE42FFEBBFD63FEC5FD7C" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[259,324,693,719]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFCFE42FFEF4FD60FE03FD7C" box="[332,386,693,719]" italics="true" pageId="41" pageNumber="534">et al.</emphasis>
2011: 178
</bibRefCitation>
, fig. 62, pl. 144).
</paragraph>
</subSubSection>
</treatment>
</document>