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<document id="E66A1ECBF579850F353755B074E3E43E" ID-CLB-Dataset="21707" ID-DOI="10.11646/zootaxa.4790.2.10" ID-GBIF-Dataset="3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889408" ID-ZooBank="7F4A9B99-C12E-473B-B258-7BE5697F4986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859385935" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="038E87D4FF8DFFAAFF157A37FC9A70B1" docLanguage="en" docName="zootaxa.4790.2.10.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus choennii Makhan 1992" docType="treatment" docVersion="8" lastPageNumber="373" masterDocId="FFB7FFACFF81FFA5FF827F44FFB87066" masterDocTitle="Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="374" masterPageNumber="358" pageNumber="370" updateTime="1698842324292" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E6BCFCD5F656E8372142630BB9EECBFF">Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<treatment id="038E87D4FF8DFFAAFF157A37FC9A70B1" ID-DOI="http://doi.org/10.5281/zenodo.5918732" ID-GBIF-Taxon="164501152" ID-Zenodo-Dep="5918732" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFAAFF157A37FC9A70B1" httpUri="http://treatment.plazi.org/id/038E87D4FF8DFFAAFF157A37FC9A70B1" lastPageId="15" lastPageNumber="373" pageId="12" pageNumber="370">
<subSubSection id="C33D6549FF8DFFA9FF157A37FE4775EB" box="[151,511,1395,1422]" pageId="12" pageNumber="370" type="nomenclature">
<paragraph id="8B9836C2FF8DFFA9FF157A37FE4775EB" blockId="12.[151,511,1395,1457]" box="[151,511,1395,1422]" pageId="12" pageNumber="370">
<heading id="D0D081AEFF8DFFA9FF157A37FE4775EB" bold="true" box="[151,511,1395,1422]" fontSize="11" level="1" pageId="12" pageNumber="370" reason="1">
<taxonomicName id="4C274D41FF8DFFA9FF157A37FE4775EB" ID-CoL="3N66M" authority="Makhan" authorityName="Makhan" authorityYear="1992" box="[151,511,1395,1422]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="choennii">
<emphasis id="B953EAD0FF8DFFA9FF157A37FE4775EB" bold="true" box="[151,511,1395,1422]" pageId="12" pageNumber="370">
<emphasis id="B953EAD0FF8DFFA9FF157A37FE3675E8" bold="true" box="[151,398,1395,1422]" italics="true" pageId="12" pageNumber="370">Hydrochus choennii</emphasis>
Makhan
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFA9FF157AD3FF4775D7" box="[151,255,1431,1457]" pageId="12" pageNumber="370" type="description">
<paragraph id="8B9836C2FF8DFFA9FF157AD3FF4775D7" blockId="12.[151,511,1395,1457]" box="[151,255,1431,1457]" pageId="12" pageNumber="370">
<figureCitation id="131C2A47FF8DFFA9FF157AD3FF5B75D7" box="[151,227,1431,1457]" captionStart="FIGURE 1" captionStartId="2.[151,250,2019,2044]" captionTargetBox="[192,1405,448,1963]" captionTargetId="figure@2.[151,1435,417,1993]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3889410" httpUri="https://zenodo.org/record/3889410/files/figure.png" pageId="12" pageNumber="370">Figs. 1</figureCitation>
,
<figureCitation id="131C2A47FF8DFFA9FF737AD3FF4775D7" box="[241,255,1431,1457]" captionStart="FIGURE 8" captionStartId="13.[151,250,1759,1784]" captionTargetBox="[169,1422,310,1716]" captionTargetId="figure@13.[151,1435,309,1734]" captionTargetPageId="13" captionText="FIGURE 8. Hydrochus choennii Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3889428" httpUri="https://zenodo.org/record/3889428/files/figure.png" pageId="12" pageNumber="370">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFA9FF157A9BFD9376CB" pageId="12" pageNumber="370" type="reference_group">
<paragraph id="8B9836C2FF8DFFA9FF157A9BFDFD759F" blockId="12.[151,581,1503,1529]" box="[151,581,1503,1529]" pageId="12" pageNumber="370">
<treatmentCitationGroup id="AB3711ECFF8DFFA9FF157A9BFDFD759F" box="[151,581,1503,1529]" pageId="12" pageNumber="370">
<treatmentCitation id="0A8610D3FF8DFFA9FF157A9BFDFD759F" author="Makhan, D." box="[151,581,1503,1529]" page="96" pageId="12" pageNumber="370" year="1992">
<taxonomicName id="4C274D41FF8DFFA9FF157A9BFDFD759F" ID-CoL="3N66M" authority="Makhan 1992: 96" authorityName="Makhan" authorityYear="1992" box="[151,581,1503,1529]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="choennii">
<emphasis id="B953EAD0FF8DFFA9FF157A9BFEC0759F" box="[151,376,1503,1529]" italics="true" pageId="12" pageNumber="370">Hydrochus choennii</emphasis>
<bibRefCitation id="EFB64B33FF8DFFA9FEFD7A9BFDFD759F" author="Makhan, D." box="[383,581,1503,1529]" pageId="12" pageNumber="370" pagination="95 - 103" refId="ref6657" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992: 96</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9836C2FF8DFFA9FF157963FD9376CB" blockId="12.[151,1437,1575,1710]" pageId="12" pageNumber="370">
Originally described from
<collectingCountry id="F3307652FF8DFFA9FE427963FD967627" box="[448,558,1575,1601]" name="Suriname" pageId="12" pageNumber="370">Suriname</collectingCountry>
, based on one male. Only one male is now known from
<collectingCountry id="F3307652FF8DFFA9FB237963FAB97627" box="[1185,1281,1575,1601]" name="Ecuador" pageId="12" pageNumber="370">Ecuador</collectingCountry>
. I have examined the
<typeStatus id="549C8860FF8DFFA9FF74790FFEE07603" box="[246,344,1611,1637]" pageId="12" pageNumber="370" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D41FF8DFFA9FEF97908FD187603" ID-CoL="3N66M" authority="Makhan 1992" authorityName="Makhan" authorityYear="1992" box="[379,672,1611,1637]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="choennii">
<emphasis id="B953EAD0FF8DFFA9FEF97908FE457603" box="[379,509,1611,1637]" italics="true" pageId="12" pageNumber="370">H. choennii</emphasis>
<bibRefCitation id="EFB64B33FF8DFFA9FD81790FFD187603" author="Makhan, D." box="[515,672,1611,1637]" pageId="12" pageNumber="370" pagination="95 - 103" refId="ref6657" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992</bibRefCitation>
</taxonomicName>
, kindly loaned by the RBINS. Labels on the
<typeStatus id="549C8860FF8DFFA9FB0F790FFB577603" box="[1165,1263,1611,1637]" pageId="12" pageNumber="370" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D41FF8DFFA9FA917908FA2C7603" ID-CoL="3N66M" authorityName="Makhan" authorityYear="1992" box="[1299,1428,1611,1637]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="choennii">
<emphasis id="B953EAD0FF8DFFA9FA917908FA2C7603" box="[1299,1428,1611,1637]" italics="true" pageId="12" pageNumber="370">H. choennii</emphasis>
</taxonomicName>
: “Coll. R. I. Sc. N. B. //
<collectingCountry id="F3307652FF8DFFA9FE1A792BFDBB76EF" box="[408,515,1647,1673]" name="Suriname" pageId="12" pageNumber="370">Suriname</collectingCountry>
District
<collectingRegion id="49E3F820FF8DFFA9FDE1792BFD4276EF" box="[611,762,1647,1673]" country="Suriname" name="Commewijne" pageId="12" pageNumber="370">Commewijne</collectingRegion>
<date id="FF991002FF8DFFA9FC82792BFCC276EF" box="[768,890,1647,1673]" pageId="12" pageNumber="370" value="1984-08-04">4.viii.1984</date>
leg. D. Makhan //
<taxonomicName id="4C274D41FF8DFFA9FBCA792BFB7B76EF" ID-CoL="8VRRD" authorityName="Leach" authorityYear="1817" box="[1096,1219,1647,1673]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="genus">Hydrochus</taxonomicName>
choenii” [sic] “det. D. Makhan 1990 //
<typeStatus id="549C8860FF8DFFA9FEF079D7FE6276CB" box="[370,474,1683,1709]" pageId="12" pageNumber="370" type="holotype">Holotype</typeStatus>
[red].”
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFA9FF15799EFF46777C" pageId="12" pageNumber="370" type="materials_examined">
<paragraph id="8B9836C2FF8DFFA9FF15799EFF46777C" blockId="12.[151,1437,1754,2034]" pageId="12" pageNumber="370">
<emphasis id="B953EAD0FF8DFFA9FF15799EFE3E7693" bold="true" box="[151,390,1754,1781]" pageId="12" pageNumber="370">Material examined.</emphasis>
<materialsCitation id="3B4F3C9FFF8DFFA9FE10799FFF42777C" ID-GBIF-Occurrence="2632886309" collectingDate="1977-06-11" collectionCode="NMNH" collectorName="P. J. Spangler &amp; D. R. Givens" country="Ecuador" location="Limoncocha" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Napo">
<collectingCountry id="F3307652FF8DFFA9FE10799FFDA47693" box="[402,540,1755,1781]" name="Ecuador" pageId="12" pageNumber="370">ECUADOR</collectingCountry>
,
<collectingRegion id="49E3F820FF8DFFA9FDAF7998FDD27690" box="[557,618,1756,1782]" country="Ecuador" name="Napo" pageId="12" pageNumber="370">Napo</collectingRegion>
<location id="8EF86019FF8DFFA9FDF4799FFCBA7693" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFAAFF157A37FC9A70B1:8EF86019FF8DFFA9FDF4799FFCBA7693" box="[630,770,1755,1781]" country="Ecuador" name="Limoncocha" pageId="12" pageNumber="370" stateProvince="Napo">Limoncocha</location>
<date id="FF991002FF8DFFA9FC8C799FFC087693" box="[782,944,1755,1782]" pageId="12" pageNumber="370" value="1977-06-11">
<collectingDate id="EFDDE9EAFF8DFFA9FC8C799FFC087693" box="[782,944,1755,1782]" pageId="12" pageNumber="370" value="1977-06-11">11 June 1977</collectingDate>
</date>
,
<collectorName id="26D25314FF8DFFA9FC437998FBDB7693" box="[961,1123,1755,1782]" pageId="12" pageNumber="370">P. J. Spangler</collectorName>
&amp;
<collectorName id="26D25314FF8DFFA9FB137998FA897693" box="[1169,1329,1755,1782]" pageId="12" pageNumber="370">D. R. Givens</collectorName>
, #115 (1
<collectionCode id="ED36AE07FF8DFFA9FF157844FF4A777C" box="[151,242,1792,1818]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFA9FF457866FA3877E3" pageId="12" pageNumber="370" type="diagnosis">
<paragraph id="8B9836C2FF8DFFA9FF457866FA3877E3" blockId="12.[151,1437,1754,2034]" pageId="12" pageNumber="370">
<emphasis id="B953EAD0FF8DFFA9FF457866FE6B775B" bold="true" box="[199,467,1826,1853]" pageId="12" pageNumber="370">Differential Diagnosis.</emphasis>
Among Ecuadorian species, recognized by the combination of moderate size (ca.
<quantity id="4CDF9B27FF8DFFA9FAE97867FF7F7704" metricMagnitude="-3" metricUnit="m" metricValue="3.21" pageId="12" pageNumber="370" unit="mm" value="3.21">3.21 mm</quantity>
), the moderately low and relatively impunctate pronotal reliefs, and the male genitalia (
<figureCitation id="131C2A47FF8DFFA9FB327803FB4E7707" box="[1200,1270,1863,1889]" captionStart="FIGURE 8" captionStartId="13.[151,250,1759,1784]" captionTargetBox="[169,1422,310,1716]" captionTargetId="figure@13.[151,1435,309,1734]" captionTargetPageId="13" captionText="FIGURE 8. Hydrochus choennii Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3889428" httpUri="https://zenodo.org/record/3889428/files/figure.png" pageId="12" pageNumber="370">Fig. 8</figureCitation>
; described below). This species has a generalized
<taxonomicName id="4C274D41FF8DFFA9FDB2782FFD1277E3" authorityName="Leach" authorityYear="1817" box="[560,682,1899,1925]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B953EAD0FF8DFFA9FDB2782FFD1277E3" box="[560,682,1899,1925]" italics="true" pageId="12" pageNumber="370">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFAAFF4578CAFC9A70B1" lastPageId="15" lastPageNumber="373" pageId="12" pageNumber="370" type="description">
<paragraph id="8B9836C2FF8DFFA8FF4578CAFCA2709D" blockId="12.[151,1437,1754,2034]" lastBlockId="13.[151,1436,152,251]" lastPageId="13" lastPageNumber="371" pageId="12" pageNumber="370">
<emphasis id="B953EAD0FF8DFFA9FF4578CAFED577CF" bold="true" box="[199,365,1934,1961]" pageId="12" pageNumber="370">Redescription</emphasis>
. Habitus as illustrated (
<figureCitation id="131C2A47FF8DFFA9FDFA78CBFD0577CF" box="[632,701,1935,1961]" captionStart="FIGURE 8" captionStartId="13.[151,250,1759,1784]" captionTargetBox="[169,1422,310,1716]" captionTargetId="figure@13.[151,1435,309,1734]" captionTargetPageId="13" captionText="FIGURE 8. Hydrochus choennii Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3889428" httpUri="https://zenodo.org/record/3889428/files/figure.png" pageId="12" pageNumber="370">Fig. 8</figureCitation>
). Size (specimen from
<collectingCountry id="F3307652FF8DFFA9FC4078CBFB9B77CF" box="[962,1059,1935,1961]" name="Ecuador" pageId="12" pageNumber="370">Ecuador</collectingCountry>
; length/width, mm): body (length to elytral apices) 3.21/1.21; head width 0.77; pronotum l/w 0.69/0.74; PA 0.69; PB 0.62; elytra 2.08/1.21. Body size moderate (ca.
<quantity id="4CDF9B27FF8DFFA9FEB87893FE197794" box="[314,417,2007,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.21" pageId="12" pageNumber="370" unit="mm" value="3.21">3.21 mm</quantity>
, Ecuadorian specimen). Dorsum grey, with slight to moderately strong iridescence, elytra with black spots. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
<superScript id="7C529B8AFF8CFFA8FEC47FF8FEEB70AC" attach="left" box="[326,339,188,202]" fontSize="6" pageId="13" pageNumber="371">th</superScript>
more convex than others, overhanging 10
<superScript id="7C529B8AFF8CFFA8FCA47FF8FC8B70AC" attach="left" box="[806,819,188,202]" fontSize="6" pageId="13" pageNumber="371">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C529B8AFF8CFFA8FB397FF8FB7070AC" attach="left" box="[1211,1224,188,202]" fontSize="6" pageId="13" pageNumber="371">th</superScript>
moderately raised, moderately elongate, ca. 5 punctures in row on each side.
</paragraph>
<caption id="DF58664AFF8CFFA8FF15799BFCE4769E" ID-DOI="http://doi.org/10.5281/zenodo.3889428" ID-Zenodo-Dep="3889428" box="[151,860,1759,1784]" httpUri="https://zenodo.org/record/3889428/files/figure.png" pageId="13" pageNumber="371" startId="13.[151,250,1759,1784]" targetBox="[169,1422,310,1716]" targetPageId="13">
<paragraph id="8B9836C2FF8CFFA8FF15799BFCE4769E" blockId="13.[151,860,1759,1784]" box="[151,860,1759,1784]" pageId="13" pageNumber="371">
<emphasis id="B953EAD0FF8CFFA8FF15799BFEAB769E" bold="true" box="[151,275,1759,1784]" pageId="13" pageNumber="371">FIGURE 8.</emphasis>
<taxonomicName id="4C274D41FF8CFFA8FE9B79A4FDF9769E" authority="Makhan" authorityName="Makhan" authorityYear="1992" box="[281,577,1759,1784]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="371" phylum="Arthropoda" rank="species" species="choennii">
<emphasis id="B953EAD0FF8CFFA8FE9B79A4FE5D769E" box="[281,485,1760,1784]" italics="true" pageId="13" pageNumber="371">Hydrochus choennii</emphasis>
Makhan
</taxonomicName>
, habitus and male genitalia.
</paragraph>
</caption>
<paragraph id="8B9836C2FF8CFFA8FF457863FC5C77EE" blockId="13.[151,1436,1830,2001]" pageId="13" pageNumber="371">Pronotum length greater than width, widest at anterior margin, narrowed at base, sides weakly arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions moderately deep, much more densely punctate than low reliefs between depressions, latter almost entirely impunctate.</paragraph>
<paragraph id="8B9836C2FF8CFFA8FF4578D6FCC377B6" blockId="13.[151,1436,1830,2001]" pageId="13" pageNumber="371">Elytra with spaces between strial punctures ca. 0.5–1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
<caption id="DF58664AFF8FFFABFF1579D5FCEE76CC" ID-DOI="http://doi.org/10.5281/zenodo.3889430" ID-Zenodo-Dep="3889430" box="[151,854,1681,1706]" httpUri="https://zenodo.org/record/3889430/files/figure.png" pageId="14" pageNumber="372" startId="14.[151,250,1681,1706]" targetBox="[164,1427,207,1645]" targetPageId="14">
<paragraph id="8B9836C2FF8FFFABFF1579D5FCEE76CC" blockId="14.[151,854,1681,1706]" box="[151,854,1681,1706]" pageId="14" pageNumber="372">
<emphasis id="B953EAD0FF8FFFABFF1579D5FEAB76CC" bold="true" box="[151,275,1681,1706]" pageId="14" pageNumber="372">FIGURE 9.</emphasis>
<taxonomicName id="4C274D41FF8FFFABFE9B79D6FD8476CC" authority="Perkins" authorityName="Perkins" authorityYear="2019" box="[281,572,1681,1706]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="372" phylum="Arthropoda" rank="species" species="hellmani">
<emphasis id="B953EAD0FF8FFFABFE9B79D6FE5F76CC" box="[281,487,1682,1706]" italics="true" pageId="14" pageNumber="372">Hydrochus hellmani</emphasis>
Perkins
</taxonomicName>
, habitus and male genitalia.
</paragraph>
</caption>
<paragraph id="8B9836C2FF8FFFABFF45799CFDFA7770" blockId="14.[151,1437,1752,2031]" pageId="14" pageNumber="372">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae. Legs testaceous, with tibiofemoral joints darker.</paragraph>
<paragraph id="8B9836C2FF8FFFABFF457864FCD97788" blockId="14.[151,1437,1752,2031]" pageId="14" pageNumber="372">
Male genitalia (
<figureCitation id="131C2A47FF8FFFABFEFA7864FE02775C" box="[376,442,1824,1850]" captionStart="FIGURE 8" captionStartId="13.[151,250,1759,1784]" captionTargetBox="[169,1422,310,1716]" captionTargetId="figure@13.[151,1435,309,1734]" captionTargetPageId="13" captionText="FIGURE 8. Hydrochus choennii Makhan, habitus and male genitalia." figureDoi="http://doi.org/10.5281/zenodo.3889428" httpUri="https://zenodo.org/record/3889428/files/figure.png" pageId="14" pageNumber="372">Fig. 8</figureCitation>
) general characters: Genitalia large, strongly sclerotized, with margins between strongly sclerotized areas and weakly or non-sclerotized areas very distinct; parameres much longer than aedeagus, plsa large, parameres in dorsal/ventral views with middle 1/3 widened and strongly sclerotized, in lateral view parameres much wider, pointed apically, with two small, indistinct pwsa; pseudoparameres large, as long as or slightly longer than aedeagus; aedeagus distal ½ widened, with alsm, apex sharply pointed, agw at about same level as adtl; basal piece ca. same length as parameres, orifice strongly sclerotized.
</paragraph>
<paragraph id="8B9836C2FF8EFFAAFF457FDDFCC270D5" blockId="15.[199,890,153,215]" box="[199,890,153,179]" pageId="15" pageNumber="373">Dorsal surface: adbl moderately large; adtl at about distal 1/3.</paragraph>
<paragraph id="8B9836C2FF8EFFAAFF457FF9FC9A70B1" blockId="15.[199,890,153,215]" box="[199,802,188,215]" pageId="15" pageNumber="373">Ventral surface: pvmm in basal ¼ approximating alm.</paragraph>
</subSubSection>
</treatment>
</document>
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