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<document id="1BB01077F9629B31A9D1FC0D16765D8D" ID-CLB-Dataset="21707" ID-DOI="10.11646/zootaxa.4790.2.10" ID-GBIF-Dataset="3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889408" ID-ZooBank="7F4A9B99-C12E-473B-B258-7BE5697F4986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859385935" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="038E87D4FF8DFFA9FF157EBDFC7073FD" docLanguage="en" docName="zootaxa.4790.2.10.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus bicarinatus Perkins 2019" docType="treatment" docVersion="8" lastPageNumber="370" masterDocId="FFB7FFACFF81FFA5FF827F44FFB87066" masterDocTitle="Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="374" masterPageNumber="358" pageNumber="370" updateTime="1698842324292" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F9898D774B856089A44B9C9B61B232E3">Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="1EBEEE6179B791B995B1A338DEE8A5B9">Perkins, Philip D.</mods:namePart>
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<mods:date id="EE11BF8B34AA349186CC8E07AEA6C3F7">2020</mods:date>
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<treatment id="038E87D4FF8DFFA9FF157EBDFC7073FD" ID-DOI="http://doi.org/10.5281/zenodo.5918728" ID-GBIF-Taxon="164501153" ID-Zenodo-Dep="5918728" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD" httpUri="http://treatment.plazi.org/id/038E87D4FF8DFFA9FF157EBDFC7073FD" lastPageNumber="370" pageId="12" pageNumber="370">
<subSubSection id="C33D6549FF8DFFA9FF157EBDFDAC7272" box="[151,532,505,532]" pageId="12" pageNumber="370" type="nomenclature">
<paragraph id="8B9836C2FF8DFFA9FF157EBDFDAC7272" blockId="12.[151,532,505,532]" box="[151,532,505,532]" pageId="12" pageNumber="370">
<heading id="D0D081AEFF8DFFA9FF157EBDFDAC7272" bold="true" box="[151,532,505,532]" fontSize="11" level="1" pageId="12" pageNumber="370" reason="1">
<taxonomicName id="4C274D41FF8DFFA9FF157EBDFDAC7272" ID-CoL="8Y4PC" authority="Perkins" authorityName="Perkins" authorityYear="2019" box="[151,532,505,532]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="bicarinatus">
<emphasis id="B953EAD0FF8DFFA9FF157EBDFDAC7272" bold="true" box="[151,532,505,532]" pageId="12" pageNumber="370">
<emphasis id="B953EAD0FF8DFFA9FF157EBDFE157272" bold="true" box="[151,429,505,532]" italics="true" pageId="12" pageNumber="370">Hydrochus bicarinatus</emphasis>
Perkins
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFA9FF157D7AFCB072A5" pageId="12" pageNumber="370" type="reference_group">
<paragraph id="8B9836C2FF8DFFA9FF157D7AFD847230" blockId="12.[151,572,573,598]" box="[151,572,573,598]" pageId="12" pageNumber="370">
<treatmentCitationGroup id="AB3711ECFF8DFFA9FF157D7AFD847230" box="[151,572,573,598]" pageId="12" pageNumber="370">
<treatmentCitation id="0A8610D3FF8DFFA9FF157D7AFD847230" author="Perkins, P. D." box="[151,572,573,598]" httpUri="http://treatment.plazi.org/id/03A587E7A875FFF30CBA10D5FC8393DE" page="23" pageId="12" pageNumber="370" year="2019">
<taxonomicName id="4C274D41FF8DFFA9FF157D7AFD847230" ID-CoL="8Y4PC" authority="Perkins 2019 c: 23" authorityName="Perkins" authorityYear="2019" box="[151,572,573,598]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="370" phylum="Arthropoda" rank="species" species="bicarinatus">
<emphasis id="B953EAD0FF8DFFA9FF157D7AFEC77230" box="[151,383,574,598]" italics="true" pageId="12" pageNumber="370">Hydrochus bicarinatus</emphasis>
<bibRefCitation id="EFB64B33FF8DFFA9FE077D79FD847230" author="Perkins, P. D." box="[389,572,573,598]" pageId="12" pageNumber="370" pagination="1 - 59" refId="ref7294" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins 2019c: 23</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9836C2FF8DFFA9FF157DC0FCB072A5" blockId="12.[151,1436,644,707]" pageId="12" pageNumber="370">
<materialsCitation id="3B4F3C9FFF8DFFA9FF157DC0FCBC72A5" ID-GBIF-Occurrence="2632886304" country="Venezuela" location="Originally" municipality="Images" pageId="12" pageNumber="370" specimenCount="1" typeStatus="holotype">
<location id="8EF86019FF8DFFA9FF157DC0FEB272F8" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FF157DC0FEB272F8" box="[151,266,644,670]" country="Venezuela" municipality="Images" name="Originally" pageId="12" pageNumber="370">Originally</location>
described from
<collectingCountry id="F3307652FF8DFFA9FE4A7DC0FD8672F8" box="[456,574,644,670]" name="Venezuela" pageId="12" pageNumber="370">Venezuela</collectingCountry>
, the following specimen records are from
<collectingCountry id="F3307652FF8DFFA9FBA57DC0FB3F72F8" box="[1063,1159,644,670]" name="Ecuador" pageId="12" pageNumber="370">Ecuador</collectingCountry>
.
<collectingMunicipality id="6BFCACB8FF8DFFA9FB167DC1FB5D72F9" box="[1172,1253,645,671]" pageId="12" pageNumber="370">Images</collectingMunicipality>
of the
<typeStatus id="549C8860FF8DFFA9FAB87DC0FA2472F8" box="[1338,1436,644,670]" pageId="12" pageNumber="370" type="holotype">holotype</typeStatus>
habitus and male genitalia are given in
<bibRefCitation id="EFB64B33FF8DFFA9FDCD7DECFCBC72A5" author="Perkins, P. D." box="[591,772,680,707]" pageId="12" pageNumber="370" pagination="1 - 59" refId="ref7294" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins (2019c)</bibRefCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF8DFFA9FF157DB4FC7073FD" pageId="12" pageNumber="370" type="materials_examined">
<paragraph id="8B9836C2FF8DFFA9FF157DB4FC7073FD" blockId="12.[151,1436,752,923]" pageId="12" pageNumber="370">
<emphasis id="B953EAD0FF8DFFA9FF157DB4FE02736D" bold="true" box="[151,442,752,779]" pageId="12" pageNumber="370">Material examined (80):</emphasis>
<materialsCitation id="3B4F3C9FFF8DFFA9FE437DB5FE3D7349" ID-GBIF-Occurrence="2632886320" collectingDate="1975-06-21" collectionCode="NMNH" collectorName="Cohen &amp; Langley &amp; Monnig" country="Ecuador" latitude="-1.8" location="Babahoyo" longLatPrecision="1308" longitude="-79.53333" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Los Rios">
<emphasis id="B953EAD0FF8DFFA9FE437DB5FD90736D" bold="true" box="[449,552,753,779]" pageId="12" pageNumber="370">
<collectingCountry id="F3307652FF8DFFA9FE437DB5FD90736D" box="[449,552,753,779]" name="Ecuador" pageId="12" pageNumber="370">Ecuador</collectingCountry>
</emphasis>
:
<collectingRegion id="49E3F820FF8DFFA9FDB57DB5FD27736C" box="[567,671,752,779]" country="Ecuador" name="Los Rios" pageId="12" pageNumber="370">Los Rios</collectingRegion>
,
<location id="8EF86019FF8DFFA9FD287DB4FCA6736C" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FD287DB4FCA6736C" box="[682,798,752,778]" country="Ecuador" latitude="-1.8" longLatPrecision="1308" longitude="-79.53333" name="Babahoyo" pageId="12" pageNumber="370" stateProvince="Los Rios">Babahoyo</location>
, (
<geoCoordinate id="EE135005FF8DFFA9FCB67DB5FC3C736D" box="[820,900,753,779]" degrees="1" direction="south" minutes="48" orientation="latitude" pageId="12" pageNumber="370" precision="925" value="-1.8">1°48S</geoCoordinate>
,
<geoCoordinate id="EE135005FF8DFFA9FC127DB5FC43736D" box="[912,1019,753,779]" degrees="79" direction="west" minutes="32" orientation="longitude" pageId="12" pageNumber="370" precision="925" value="-79.53333">79°32W</geoCoordinate>
),
<date id="FF991002FF8DFFA9FB8D7DB4FB31736D" box="[1039,1161,752,779]" pageId="12" pageNumber="370" value="1975-06-21">
<collectingDate id="EFDDE9EAFF8DFFA9FB8D7DB4FB31736D" box="[1039,1161,752,779]" pageId="12" pageNumber="370" value="1975-06-21">21.vi.1975</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF8DFFA9FB447DB4FAAA736C" box="[1222,1298,752,778]" pageId="12" pageNumber="370">Cohen</collectorName>
,
<collectorName id="26D25314FF8DFFA9FA9C7DB4FAC3736C" box="[1310,1403,752,778]" pageId="12" pageNumber="370">Langley</collectorName>
, &amp;
<collectorName id="26D25314FF8DFFA9FF157C50FF487348" box="[151,240,788,814]" pageId="12" pageNumber="370">Monnig</collectorName>
(60
<collectionCode id="ED36AE07FF8DFFA9FEA07C51FEC57349" box="[290,381,789,815]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF8DFFA9FE127C50FB097349" ID-GBIF-Occurrence="2632886342" box="[400,1201,788,815]" collectingDate="1975-05-11" collectionCode="NMNH" collectorName="Spangler" country="Ecuador" latitude="-1.0333333" location="Quevedo" longLatPrecision="1308" longitude="-79.46667" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Los Rios">
<location id="8EF86019FF8DFFA9FE127C50FE407348" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FE127C50FE407348" box="[400,504,788,814]" country="Ecuador" latitude="-1.0333333" longLatPrecision="1308" longitude="-79.46667" name="Quevedo" pageId="12" pageNumber="370" stateProvince="Los Rios">Quevedo</location>
, (
<geoCoordinate id="EE135005FF8DFFA9FD8F7C51FDF57349" box="[525,589,789,815]" degrees="1" direction="south" minutes="2" orientation="latitude" pageId="12" pageNumber="370" precision="925" value="-1.0333333">1°2S</geoCoordinate>
,
<geoCoordinate id="EE135005FF8DFFA9FDDB7C51FD7C7349" box="[601,708,789,815]" degrees="79" direction="west" minutes="28" orientation="longitude" pageId="12" pageNumber="370" precision="925" value="-79.46667">79°28W</geoCoordinate>
),
<date id="FF991002FF8DFFA9FD557C50FCFF7349" box="[727,839,788,815]" pageId="12" pageNumber="370" value="1975-05-11">
<collectingDate id="EFDDE9EAFF8DFFA9FD557C50FCFF7349" box="[727,839,788,815]" pageId="12" pageNumber="370" value="1975-05-11">11.v.1975</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF8DFFA9FC007C50FC5F7348" box="[898,999,788,814]" pageId="12" pageNumber="370">Spangler</collectorName>
,
<emphasis id="B953EAD0FF8DFFA9FC707C51FB9B7348" box="[1010,1059,788,814]" italics="true" pageId="12" pageNumber="370">et al</emphasis>
. (2
<collectionCode id="ED36AE07FF8DFFA9FBCC7C51FB117349" box="[1102,1193,789,815]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF8DFFA9FB397C50FD257335" ID-GBIF-Occurrence="2632886322" collectingDate="1975-07-03" collectionCode="NMNH" collectorName="Langley &amp; Cohen" country="Ecuador" location="Quevedo" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Los Rios">
<location id="8EF86019FF8DFFA9FB397C50FA987348" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FB397C50FA987348" box="[1211,1312,788,814]" country="Ecuador" name="Quevedo" pageId="12" pageNumber="370" stateProvince="Los Rios">Quevedo</location>
(
<quantity id="4CDF9B27FF8DFFA9FAB27C51FACE7348" box="[1328,1398,788,815]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="370" unit="km" value="11.0">11 km</quantity>
S),
<date id="FF991002FF8DFFA9FF157C7CFEB37335" box="[151,267,824,851]" pageId="12" pageNumber="370" value="1975-07-03">
<collectingDate id="EFDDE9EAFF8DFFA9FF157C7CFEB37335" box="[151,267,824,851]" pageId="12" pageNumber="370" value="1975-07-03">3.vii.1975</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF8DFFA9FECB7C7CFE1D7334" box="[329,421,824,850]" pageId="12" pageNumber="370">Langley</collectorName>
&amp;
<collectorName id="26D25314FF8DFFA9FE487C7CFDAB7334" box="[458,531,824,850]" pageId="12" pageNumber="370">Cohen</collectorName>
(2
<collectionCode id="ED36AE07FF8DFFA9FDB87C7DFD2D7335" box="[570,661,825,851]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF8DFFA9FD2B7C7CFDD07311" ID-GBIF-Occurrence="2632886339" collectingDate="1975-05-10" collectionCode="NMNH" collectorName="P. J. Spangler" country="Ecuador" latitude="-0.67222226" location="Bahia de Caraquez" longLatPrecision="21" longitude="-80.08584" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Manabi">
<collectingRegion id="49E3F820FF8DFFA9FD2B7C7CFCB97334" box="[681,769,824,850]" country="Ecuador" name="Manabi" pageId="12" pageNumber="370">Manabi</collectingRegion>
,
<location id="8EF86019FF8DFFA9FC8F7C7CFC5B7335" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FC8F7C7CFC5B7335" box="[781,995,824,851]" country="Ecuador" latitude="-0.67222226" longLatPrecision="21" longitude="-80.08584" name="Bahia de Caraquez" pageId="12" pageNumber="370" stateProvince="Manabi">Bahia de Caraquez</location>
(
<quantity id="4CDF9B27FF8DFFA9FC707C7CFBE87334" box="[1010,1104,824,851]" metricMagnitude="4" metricUnit="m" metricValue="3.56" pageId="12" pageNumber="370" unit="km" value="35.6">35.6 km</quantity>
E), (
<geoCoordinate id="EE135005FF8DFFA9FB087C7DFAB07335" box="[1162,1288,825,851]" degrees="0" direction="south" minutes="40" orientation="latitude" pageId="12" pageNumber="370" precision="15" seconds="20" value="-0.67222226">0°4020S</geoCoordinate>
,
<geoCoordinate id="EE135005FF8DFFA9FA967C7CFA367335" box="[1300,1422,824,851]" degrees="80" direction="west" minutes="5" orientation="longitude" pageId="12" pageNumber="370" precision="15" seconds="9" value="-80.08584">80°59W</geoCoordinate>
),
<date id="FF991002FF8DFFA9FF157C18FEB07311" box="[151,264,860,887]" pageId="12" pageNumber="370" value="1975-05-10">
<collectingDate id="EFDDE9EAFF8DFFA9FF157C18FEB07311" box="[151,264,860,887]" pageId="12" pageNumber="370" value="1975-05-10">10.v.1975</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF8DFFA9FEC77C19FE677310" box="[325,479,860,887]" pageId="12" pageNumber="370">P. J. Spangler</collectorName>
(3
<collectionCode id="ED36AE07FF8DFFA9FD847C19FDD97311" box="[518,609,861,887]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF8DFFA9FDF67C19FB797311" ID-GBIF-Occurrence="2632886318" box="[628,1217,860,887]" collectingDate="1978-03-05" collectionCode="NMNH" collectorName="J. J. Anderson" country="Ecuador" location="Tosagua" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Manabi">
<location id="8EF86019FF8DFFA9FDF67C19FD6C7311" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FDF67C19FD6C7311" box="[628,724,861,887]" country="Ecuador" name="Tosagua" pageId="12" pageNumber="370" stateProvince="Manabi">Tosagua</location>
,
<date id="FF991002FF8DFFA9FD627C18FCF57311" box="[736,845,860,887]" pageId="12" pageNumber="370" value="1978-03-05">
<collectingDate id="EFDDE9EAFF8DFFA9FD627C18FCF57311" box="[736,845,860,887]" pageId="12" pageNumber="370" value="1978-03-05">5.iii.1978</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF8DFFA9FC097C19FB927310" box="[907,1066,860,887]" pageId="12" pageNumber="370">J. J. Anderson</collectorName>
(11
<collectionCode id="ED36AE07FF8DFFA9FBDC7C19FB017311" box="[1118,1209,861,887]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF8DFFA9FB4F7C19FC7D73FD" ID-GBIF-Occurrence="2632886330" collectingDate="1975-09-21" collectionCode="NMNH" collectorName="Langley &amp; Cohen" country="Ecuador" latitude="-0.3" location="Shushufindi" longLatPrecision="1308" longitude="-76.416664" pageId="12" pageNumber="370" specimenCount="1" stateProvince="Napo">
<collectingRegion id="49E3F820FF8DFFA9FB4F7C19FAB47311" box="[1229,1292,861,887]" country="Ecuador" name="Napo" pageId="12" pageNumber="370">Napo</collectingRegion>
, ca.
<location id="8EF86019FF8DFFA9FAC27C18FF7773FC" LSID="urn:lsid:plazi:treatment:038E87D4FF8DFFA9FF157EBDFC7073FD:8EF86019FF8DFFA9FAC27C18FF7773FC" country="Ecuador" latitude="-0.3" longLatPrecision="1308" longitude="-76.416664" name="Shushufindi" pageId="12" pageNumber="370" stateProvince="Napo">Shushufindi</location>
, (
<geoCoordinate id="EE135005FF8DFFA9FF667CC5FE8C73FD" box="[228,308,897,923]" degrees="0" direction="south" minutes="18" orientation="latitude" pageId="12" pageNumber="370" precision="925" value="-0.3">0°18S</geoCoordinate>
,
<geoCoordinate id="EE135005FF8DFFA9FEBD7CC4FE1273FD" box="[319,426,896,923]" degrees="76" direction="west" minutes="25" orientation="longitude" pageId="12" pageNumber="370" precision="925" value="-76.416664">76°25W</geoCoordinate>
),
<date id="FF991002FF8DFFA9FE3C7CC4FD8073FD" box="[446,568,896,923]" pageId="12" pageNumber="370" value="1975-09-21">
<collectingDate id="EFDDE9EAFF8DFFA9FE3C7CC4FD8073FD" box="[446,568,896,923]" pageId="12" pageNumber="370" value="1975-09-21">21.ix.1975</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF8DFFA9FDF67CC4FD6873FC" box="[628,720,896,922]" pageId="12" pageNumber="370">Langley</collectorName>
&amp;
<collectorName id="26D25314FF8DFFA9FD717CC4FC8473FC" box="[755,828,896,922]" pageId="12" pageNumber="370">Cohen</collectorName>
(2
<collectionCode id="ED36AE07FF8DFFA9FCE07CC5FC0573FD" box="[866,957,897,923]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="370" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>