156 lines
18 KiB
XML
156 lines
18 KiB
XML
<document id="549AB031B5DF0CB8B7BA4E4239C1B03E" ID-CLB-Dataset="21707" ID-DOI="10.11646/zootaxa.4790.2.10" ID-GBIF-Dataset="3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889408" ID-ZooBank="7F4A9B99-C12E-473B-B258-7BE5697F4986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859385935" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="038E87D4FF8BFFAEFF157CF5FBA5777B" docLanguage="en" docName="zootaxa.4790.2.10.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus simplicatus Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="369" masterDocId="FFB7FFACFF81FFA5FF827F44FFB87066" masterDocTitle="Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="374" masterPageNumber="358" pageNumber="368" updateTime="1698842324292" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CDF520DF58B7D3FEC404AA62880AC3CD">Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="3C18A3F39BDD00F618ACC804643DAF48" type="personal">
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<mods:namePart id="BCD23FFAD3006B834D7AF3E7DA31689E">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="A899DE92BC63E237E8E3603FEEFA1D4F">text</mods:typeOfResource>
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<mods:title id="42E0500EDE995DCDAAA8D7D158C1F257">Zootaxa</mods:title>
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<mods:part id="02E0451C3CC0E0D2C819BD742DB4341B">
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<mods:date id="6913CCCE30C1031A0185586260EF5127">2020</mods:date>
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<mods:number id="7D64FB157727E1AB4E4C7C6EAA702311">2020-06-11</mods:number>
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<mods:number id="12EB72405C0BB9D5ED2F4D61A807739B">4790</mods:number>
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<treatment id="038E87D4FF8BFFAEFF157CF5FBA5777B" ID-DOI="http://doi.org/10.5281/zenodo.5918724" ID-GBIF-Taxon="164501155" ID-Zenodo-Dep="5918724" LSID="urn:lsid:plazi:treatment:038E87D4FF8BFFAEFF157CF5FBA5777B" httpUri="http://treatment.plazi.org/id/038E87D4FF8BFFAEFF157CF5FBA5777B" lastPageId="11" lastPageNumber="369" pageId="10" pageNumber="368">
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<subSubSection id="C33D6549FF8BFFAFFF157CF5FDF473AD" box="[151,588,945,972]" pageId="10" pageNumber="368" type="nomenclature">
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<paragraph id="8B9836C2FF8BFFAFFF157CF5FDF473AD" blockId="10.[151,588,945,1007]" box="[151,588,945,972]" pageId="10" pageNumber="368">
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<heading id="D0D081AEFF8BFFAFFF157CF5FDF473AD" bold="true" box="[151,588,945,972]" fontSize="11" level="1" pageId="10" pageNumber="368" reason="1">
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<emphasis id="B953EAD0FF8BFFAFFF157CF5FDF473AD" bold="true" box="[151,588,945,972]" pageId="10" pageNumber="368">
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<taxonomicName id="4C274D41FF8BFFAFFF157CF5FE1573AA" ID-CoL="8Y4RG" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,429,945,972]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="368" phylum="Arthropoda" rank="species" species="simplicatus" status="sp. nov.">
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<emphasis id="B953EAD0FF8BFFAFFF157CF5FE1573AA" bold="true" box="[151,429,945,972]" italics="true" pageId="10" pageNumber="368">Hydrochus simplicatus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26057ABFF8BFFAFFE3E7CF5FDF473AD" box="[444,588,945,971]" pageId="10" pageNumber="368" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF8BFFAFFF157C91FF477389" box="[151,255,981,1007]" pageId="10" pageNumber="368" type="description">
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<paragraph id="8B9836C2FF8BFFAFFF157C91FF477389" blockId="10.[151,588,945,1007]" box="[151,255,981,1007]" pageId="10" pageNumber="368">
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<figureCitation id="131C2A47FF8BFFAFFF157C91FF5B7389" box="[151,227,981,1007]" captionStart="FIGURE 1" captionStartId="2.[151,250,2019,2044]" captionTargetBox="[192,1405,448,1963]" captionTargetId="figure@2.[151,1435,417,1993]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3889410" httpUri="https://zenodo.org/record/3889410/files/figure.png" pageId="10" pageNumber="368">Figs. 1</figureCitation>
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,
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<figureCitation id="131C2A47FF8BFFAFFF737C91FF477389" box="[241,255,981,1007]" captionStart="FIGURE 7" captionStartId="11.[151,250,1688,1713]" captionTargetBox="[165,1423,201,1651]" captionTargetId="figure@11.[155,1433,181,1662]" captionTargetPageId="11" captionText="FIGURE 7. Hydrochus simplicatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889424" httpUri="https://zenodo.org/record/3889424/files/figure.png" pageId="10" pageNumber="368">7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF8BFFAFFF157B05FD667419" pageId="10" pageNumber="368" type="materials_examined">
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<paragraph id="8B9836C2FF8BFFAFFF157B05FD667419" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">
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<emphasis id="B953EAD0FF8BFFAFFF157B05FEF1743D" bold="true" box="[151,329,1088,1115]" pageId="10" pageNumber="368">Type Material.</emphasis>
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<materialsCitation id="3B4F3C9FFF8BFFAFFED07B05FD667419" ID-GBIF-Occurrence="2632886338" collectingDate="1974-09-10" collectionCode="NMNH" collectorName="Boyce A. Drummond" country="Ecuador" elevation="300" location="Rio" municipality="Limoncocha" pageId="10" pageNumber="368" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
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<typeStatus id="549C8860FF8BFFAFFED07B05FE02743D" box="[338,442,1089,1115]" pageId="10" pageNumber="368" type="holotype">Holotype</typeStatus>
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(male): “
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<collectingCountry id="F3307652FF8BFFAFFDA77B05FD31743D" box="[549,649,1089,1115]" name="Ecuador" pageId="10" pageNumber="368">Ecuador</collectingCountry>
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:
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<collectingRegion id="49E3F820FF8BFFAFFD177B05FCF8743D" box="[661,832,1089,1115]" country="Ecuador" name="Napo" pageId="10" pageNumber="368">Napo Province</collectingRegion>
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,
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<collectingMunicipality id="6BFCACB8FF8BFFAFFCCF7B05FC64743D" box="[845,988,1089,1115]" pageId="10" pageNumber="368">Limoncocha</collectingMunicipality>
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, on
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<location id="8EF86019FF8BFFAFFB8F7B05FB8D743D" LSID="urn:lsid:plazi:treatment:038E87D4FF8BFFAEFF157CF5FBA5777B:8EF86019FF8BFFAFFB8F7B05FB8D743D" box="[1037,1077,1089,1115]" country="Ecuador" municipality="Limoncocha" name="Rio" pageId="10" pageNumber="368" stateProvince="Napo">Rio</location>
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<collectingRegion id="49E3F820FF8BFFAFFBBC7B05FBC5743D" box="[1086,1149,1089,1115]" country="Ecuador" name="Napo" pageId="10" pageNumber="368">Napo</collectingRegion>
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, elev.
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<quantity id="4CDF9B27FF8BFFAFFB447B05FAA8743D" box="[1222,1296,1089,1115]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="368" unit="m" value="300.0">
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<elevation id="000AD1F1FF8BFFAFFB447B05FAA8743D" box="[1222,1296,1089,1115]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="368" unit="m" value="300.0">300 m</elevation>
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</quantity>
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,
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<date id="FF991002FF8BFFAFFA9F7B05FA20743D" box="[1309,1432,1089,1115]" pageId="10" pageNumber="368" value="1974-09-10">
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<collectingDate id="EFDDE9EAFF8BFFAFFA9F7B05FA20743D" box="[1309,1432,1089,1115]" pageId="10" pageNumber="368" value="1974-09-10">10.ix.1974</collectingDate>
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</date>
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,
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<collectorName id="26D25314FF8BFFAFFF157B21FE327419" box="[151,394,1125,1151]" pageId="10" pageNumber="368">Boyce A. Drummond</collectorName>
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,
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<collectingMethod id="52664ED5FF8BFFAFFE177B21FDE57419" box="[405,605,1125,1151]" pageId="10" pageNumber="368">III blacklight trap</collectingMethod>
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” (
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<collectionCode id="ED36AE07FF8BFFAFFDF47B21FD6A7419" box="[630,722,1125,1151]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="368" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF8BFFAFFF457BCCFD597569" pageId="10" pageNumber="368" type="diagnosis">
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<paragraph id="8B9836C2FF8BFFAFFF457BCCFD597569" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">
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<emphasis id="B953EAD0FF8BFFAFFF457BCCFE6B74C5" bold="true" box="[199,467,1160,1187]" pageId="10" pageNumber="368">Differential Diagnosis.</emphasis>
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Among Ecuadorian species, recognized by the combination of moderate size (ca.
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<quantity id="4CDF9B27FF8BFFAFFAE97BCCFF7F74A1" metricMagnitude="-3" metricUnit="m" metricValue="3.58" pageId="10" pageNumber="368" unit="mm" value="3.58">3.58 mm</quantity>
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), the moderately low pronotal reliefs, the relatively small, rather sparse pronotal punctures, the numerous dorsal microsetae, and the male genitalia (
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<figureCitation id="131C2A47FF8BFFAFFDA27B95FDDB748D" box="[544,611,1233,1259]" captionStart="FIGURE 7" captionStartId="11.[151,250,1688,1713]" captionTargetBox="[165,1423,201,1651]" captionTargetId="figure@11.[155,1433,181,1662]" captionTargetPageId="11" captionText="FIGURE 7. Hydrochus simplicatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889424" httpUri="https://zenodo.org/record/3889424/files/figure.png" pageId="10" pageNumber="368">Fig. 7</figureCitation>
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; described below). This species has a generalized
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<taxonomicName id="4C274D41FF8BFFAFFB0D7B95FAB1748D" authorityName="Leach" authorityYear="1817" box="[1167,1289,1233,1259]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="368" phylum="Arthropoda" rank="genus">
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<emphasis id="B953EAD0FF8BFFAFFB0D7B95FAB1748D" box="[1167,1289,1233,1259]" italics="true" pageId="10" pageNumber="368">Hydrochus</emphasis>
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</taxonomicName>
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habitus; reliable determinations will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF8BFFAEFF457A5CFB76769F" lastPageId="11" lastPageNumber="369" pageId="10" pageNumber="368" type="description">
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<paragraph id="8B9836C2FF8BFFAFFF457A5CFD547581" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">
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<emphasis id="B953EAD0FF8BFFAFFF457A5CFEE97555" bold="true" box="[199,337,1304,1331]" pageId="10" pageNumber="368">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="131C2A47FF8BFFAFFDDF7A5DFD1A7555" box="[605,674,1305,1331]" captionStart="FIGURE 7" captionStartId="11.[151,250,1688,1713]" captionTargetBox="[165,1423,201,1651]" captionTargetId="figure@11.[155,1433,181,1662]" captionTargetPageId="11" captionText="FIGURE 7. Hydrochus simplicatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889424" httpUri="https://zenodo.org/record/3889424/files/figure.png" pageId="10" pageNumber="368">Fig. 7</figureCitation>
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). Size:
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<typeStatus id="549C8860FF8BFFAFFD757A5DFCE17555" box="[759,857,1305,1331]" pageId="10" pageNumber="368" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.58/1.36; head width 0.88; pronotum l/w 0.82/0.85; PA 0.76; PB 0.70; elytra 2.15/1.36. Body size moderate (ca.
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<quantity id="4CDF9B27FF8BFFAFFF157A24FF46751D" box="[151,254,1376,1403]" metricMagnitude="-3" metricUnit="m" metricValue="3.58" pageId="10" pageNumber="368" unit="mm" value="3.58">3.58 mm</quantity>
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). Dorsum dull grey, with numerous microsetae, without iridescence, elytra with black spots. Punctation on elytra ca. 3–5x that of pronotum. Elytra interstriae ca. 1–2x strial puncture diameter. Interstria 9
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<superScript id="7C529B8AFF8BFFAFFB437AC0FB7675F4" attach="left" box="[1217,1230,1412,1426]" fontSize="6" pageId="10" pageNumber="368">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C529B8AFF8BFFAFFE187AECFE1F75D0" attach="left" box="[410,423,1448,1462]" fontSize="6" pageId="10" pageNumber="368">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C529B8AFF8BFFAFFCB97AECFCF075D0" attach="left" box="[827,840,1448,1462]" fontSize="6" pageId="10" pageNumber="368">th</superScript>
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well raised, moderately elongate, elytra transversely humped in this area, posterior declivity rather abrupt.
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</paragraph>
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<paragraph id="8B9836C2FF8BFFAFFF457AB5FF6C7635" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">Pronotum slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin arcuate or very slightly bisinuate; depressions shallow to moderately deep.</paragraph>
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<paragraph id="8B9836C2FF8BFFAFFF457919FBF276FD" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">Elytra with spaces between strial punctures ca. 1x their diameter; apices conjointly rounded in dorsal view, each with single deep oval puncture, in lateral view outer margin with distinct angulation.</paragraph>
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<paragraph id="8B9836C2FF8BFFAFFF4579E1FCB97685" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">Ventral characters: Mentum densely moderately coarsely punctate, with deep median fovea. Submentum with two foveae. Legs testaceous, femora darker than tibiae.</paragraph>
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<paragraph id="8B9836C2FF8BFFAFFF4579A9FABB77F1" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">
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Male genitalia (
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<figureCitation id="131C2A47FF8BFFAFFEFA79A9FE047761" box="[376,444,1773,1799]" captionStart="FIGURE 7" captionStartId="11.[151,250,1688,1713]" captionTargetBox="[165,1423,201,1651]" captionTargetId="figure@11.[155,1433,181,1662]" captionTargetPageId="11" captionText="FIGURE 7. Hydrochus simplicatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889424" httpUri="https://zenodo.org/record/3889424/files/figure.png" pageId="10" pageNumber="368">Fig. 7</figureCitation>
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) general characters: genitalia wide compared to length; parameres much wider in lateral than dorsal/ventral views, in dorsal/ventral views tips narrow, almost parallel sided, arcuate toward midline, not expanded, without plsa, but with narrow pmsm, in lateral view tapering to very small, weakly recurved distal ends; aedeagus lanceolate, wide, widest at about midlength, tapering from midlength to pointed apex, distal ½ with narrow alsm; basal piece shorter than parameres, ratio ca. 65/40, basal orifice asymmetrical, sclerotized.
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</paragraph>
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<paragraph id="8B9836C2FF8BFFAFFF4578E5FEC677B9" blockId="10.[151,1437,1088,2015]" pageId="10" pageNumber="368">Dorsal surface: adtl at about midlength, membranous area distal to adtl large; adbl small; pdmm overlapping aedeagus in basal ½.</paragraph>
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<caption id="DF58664AFF8AFFAEFF1579DCFC6076D7" ID-DOI="http://doi.org/10.5281/zenodo.3889424" ID-Zenodo-Dep="3889424" box="[151,984,1688,1713]" httpUri="https://zenodo.org/record/3889424/files/figure.png" pageId="11" pageNumber="369" startId="11.[151,250,1688,1713]" targetBox="[165,1423,201,1651]" targetPageId="11">
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||
<paragraph id="8B9836C2FF8AFFAEFF1579DCFC6076D7" blockId="11.[151,984,1688,1713]" box="[151,984,1688,1713]" pageId="11" pageNumber="369">
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<emphasis id="B953EAD0FF8AFFAEFF1579DCFEAB76D7" bold="true" box="[151,275,1688,1713]" pageId="11" pageNumber="369">FIGURE 7.</emphasis>
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<taxonomicName id="4C274D41FF8AFFAEFE9B79DDFDB976D7" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,513,1689,1713]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="369" phylum="Arthropoda" rank="species" species="simplicatus" status="sp. nov.">
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<emphasis id="B953EAD0FF8AFFAEFE9B79DDFDB976D7" box="[281,513,1689,1713]" italics="true" pageId="11" pageNumber="369">Hydrochus simplicatus</emphasis>
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</taxonomicName>
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||
<emphasis id="B953EAD0FF8AFFAEFD8A79DDFDF976D7" bold="true" box="[520,577,1689,1713]" pageId="11" pageNumber="369">
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<taxonomicNameLabel id="A26057ABFF8AFFAEFD8A79DDFDF976D7" box="[520,577,1689,1713]" pageId="11" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, habitus and male genitalia of holotype.
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</paragraph>
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</caption>
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||
<paragraph id="8B9836C2FF8AFFAEFF45799BFB76769F" blockId="11.[199,1230,1758,1821]" box="[199,1230,1758,1785]" pageId="11" pageNumber="369">Ventral surface: avtl at about basal ¼; pvmm slightly overlapping proximal 1/2 of aedeagus.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6549FF8AFFAEFF457847FBA5777B" box="[199,1053,1795,1821]" pageId="11" pageNumber="369" type="etymology">
|
||
<paragraph id="8B9836C2FF8AFFAEFF457847FBA5777B" blockId="11.[199,1230,1758,1821]" box="[199,1053,1795,1821]" pageId="11" pageNumber="369">
|
||
<emphasis id="B953EAD0FF8AFFAEFF457847FEF6777B" bold="true" box="[199,334,1795,1821]" pageId="11" pageNumber="369">Etymology.</emphasis>
|
||
Named in reference to the comparatively simple male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |