531 lines
77 KiB
XML
531 lines
77 KiB
XML
<document id="3906706C9765FE0379CBC64CE6F9E358" ID-DOI="10.11646/zootaxa.4790.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889504" ID-ZooBank="6316903F-9894-40CF-A22C-8593637F1C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859535270" checkinUser="plazi" docAuthor="Moravec, Jiří, Razanajaonarivalona, Elisé Hugo & Hori, Michio" docDate="2020" docId="038E87CBD31ACD01FF21F89D8306B66A" docLanguage="en" docName="zootaxa.4790.2.4.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pogonostoma (Pogonostoma) natsuae Moravec & Razanajaonarivalona & Hori 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="269" masterDocId="FFB7FFB3D31BCD09FFB6FFA88227B471" masterDocTitle="New or rare Madagascar tiger beetles- 19. Description of Pogonostoma (Pogonostoma) natsuae sp. nov. and elaborated redescription of P. (M.) meridionale Fleutiaux (Coleoptera: Cicindelidae)" masterLastPageNumber="276" masterPageNumber="261" pageNumber="262" updateTime="1700276581814" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="8F8BBA1F15A5513AB22A61A9E6C0D252" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CC318DDE71D1A7006564BB345E0F82B7">
|
||
<mods:title id="AE5515EF376D1E79943C413EE24A6377">New or rare Madagascar tiger beetles- 19. Description of Pogonostoma (Pogonostoma) natsuae sp. nov. and elaborated redescription of P. (M.) meridionale Fleutiaux (Coleoptera: Cicindelidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="77DDB75A71021233379D121DF7AFCFF4" type="personal">
|
||
<mods:role id="9B0A4C60C7A7EE7DF09C2FCA6AB254D2">
|
||
<mods:roleTerm id="643A262806FEF39681BAABA29DD8B0C0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B106AE310102C504AD55690BE2FAF59C">Moravec, Jiří</mods:namePart>
|
||
<mods:affiliation id="E5264E321E39BFA0E85E87BF1C448AB7">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, 613 00 Brno, Czech Republic</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="15019B832033F0244BD600FC6E51751F" type="personal">
|
||
<mods:role id="CCDBA143BD5C2C6252AB84E2C2313727">
|
||
<mods:roleTerm id="F2FC8AC1C68E415B3CE72FB53347CC9D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0250DDEA77EE60DC645940DA4A3DC563">Razanajaonarivalona, Elisé Hugo</mods:namePart>
|
||
<mods:affiliation id="4253EBB96FF3F96B6A1F97AF1390444E">Département de Biologie Animale, Faculté des Sciences, Université d’Antananarivo, BP 906, Antananarivo 101, Madagascar relysehugo @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9430 - 6793</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="B0491AFC88C0EB97E7C15CB0B559B49B" type="personal">
|
||
<mods:role id="051237D279089FFB39C2E76F25665C4B">
|
||
<mods:roleTerm id="A22400C5B33ED98A0CD6A46305CD8E32">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3774C06872CD3FF7921D77927B742295">Hori, Michio</mods:namePart>
|
||
<mods:affiliation id="A354E0CF4885863DA21BDB88075409A0">Kyoto University, Yoshida-Honmachi, Sakyo-ku, Kyoto, 606 - 8501</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E49B0C4EC086C1B6A14CC63E085E3F0E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F6909D59ED58FE30AC842FC8914C22CE" type="host">
|
||
<mods:titleInfo id="1578D56FBCF35BEA6E7EF0F7AAD33E7D">
|
||
<mods:title id="39B9916AFDE50C880352924CEFF0CC00">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8EEDF79AE3E0544679A78DDAAF40E599">
|
||
<mods:date id="4B6FEA8CAC5A4D03D4E1790661B6243E">2020</mods:date>
|
||
<mods:detail id="941BE168BD0688F7EEA58536540A9511" type="pubDate">
|
||
<mods:number id="0763252D3755DB273966C8E1D5B65B3E">2020-06-11</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="21EC585585447BDA30DFCC921D65079E" type="volume">
|
||
<mods:number id="EE529C9BD52577BF38A8E9D45824ABDC">4790</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="58CBA0EAD329D874C450240F05B9F313" type="issue">
|
||
<mods:number id="0EBD5FAF30DC3D97E7A645D6D9E91607">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="09925C1DB3D3ADB0D911372FD7A25579" unit="page">
|
||
<mods:start id="C34EBB17963C3E798337B3BF4042C143">261</mods:start>
|
||
<mods:end id="9AB0AACAFD20D466FE47C2083E7B772A">276</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="1BF72A3F2970A176AC9A8F02E98030A2">journal article</mods:classification>
|
||
<mods:identifier id="BA7B6851D2393B9EC1979BD05A87EAED" type="DOI">10.11646/zootaxa.4790.2.4</mods:identifier>
|
||
<mods:identifier id="06C19FE9C79E1B6B2CFC7D5213CEB146" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="93BB8D5F9768139C4000C7D5E5A3151B" type="Zenodo-Dep">3889504</mods:identifier>
|
||
<mods:identifier id="8D5057626D4EDAD2E4F02329D31DAC68" type="ZooBank">6316903F-9894-40CF-A22C-8593637F1C9A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038E87CBD31ACD01FF21F89D8306B66A" LSID="urn:lsid:plazi:treatment:038E87CBD31ACD01FF21F89D8306B66A" httpUri="http://treatment.plazi.org/id/038E87CBD31ACD01FF21F89D8306B66A" lastPageId="8" lastPageNumber="269" pageId="1" pageNumber="262">
|
||
<subSubSection id="C33D6556D31ACD08FF21F89D80F4B33F" box="[151,723,1844,1871]" pageId="1" pageNumber="262" type="nomenclature">
|
||
<paragraph id="8B9836DDD31ACD08FF21F89D80F4B33F" blockId="1.[151,723,1844,1906]" box="[151,723,1844,1871]" pageId="1" pageNumber="262">
|
||
<heading id="D0D081B1D31ACD08FF21F89D80F4B33F" bold="true" box="[151,723,1844,1871]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
|
||
<emphasis id="B953EACFD31ACD08FF21F89D80F4B33F" bold="true" box="[151,723,1844,1871]" pageId="1" pageNumber="262">
|
||
<taxonomicName id="4C274D5ED31ACD08FF21F89D804AB33E" ID-CoL="8M8QT" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[151,621,1844,1871]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD31ACD08FF21F89D8319B33E" bold="true" box="[151,318,1845,1871]" italics="true" pageId="1" pageNumber="262">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD31ACD08FEE6F89D83D0B33E" bold="true" box="[336,503,1845,1871]" italics="true" pageId="1" pageNumber="262">Pogonostoma</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD31ACD08FDBFF89D804AB33E" bold="true" box="[521,621,1845,1871]" italics="true" pageId="1" pageNumber="262">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057B4D31ACD08FDC2F89C80F4B33F" box="[628,723,1844,1870]" pageId="1" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6556D31ACD08FF21F8F08363B303" box="[151,324,1879,1906]" pageId="1" pageNumber="262" type="description">
|
||
<paragraph id="8B9836DDD31ACD08FF21F8F08363B303" blockId="1.[151,723,1844,1906]" box="[151,324,1879,1906]" pageId="1" pageNumber="262">
|
||
(
|
||
<figureCitation id="131C2A58D31ACD08FF29F8F08334B303" box="[159,275,1880,1906]" captionStart-0="FIGURES 1–9" captionStart-1="FIGURES 10–15" captionStart-2="FIGURES 16–18" captionStart-3="FIGURES 19–32" captionStartId-0="3.[151,264,1869,1894]" captionStartId-1="4.[151,264,1869,1894]" captionStartId-2="5.[151,264,1861,1886]" captionStartId-3="6.[151,264,1800,1825]" captionTargetBox-0="[162,1427,191,1829]" captionTargetBox-1="[156,1416,193,1828]" captionTargetBox-2="[164,1424,194,1823]" captionTargetBox-3="[151,1435,192,1767]" captionTargetId-0="figure@3.[151,1435,179,1842]" captionTargetId-1="figure@4.[151,1435,181,1844]" captionTargetId-2="figure@5.[151,1435,181,1836]" captionTargetId-3="figure@6.[151,1435,181,1774]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." captionText-1="FIGURES 10–15. Pogonostoma (P.) natsuae sp. nov. from type locality. 10–12—elytral apices: 10—♂, HT (OSAKA); 11—♂, PT (CCJM); 12—♀, AT (OSAKA); 13–14—elytron: 14—♂, HT; 15—♀, AT; 15—sculpture on elytral disc, ♂, HT. Bars = 1 mm." captionText-2="FIGURES 16–18. Pogonostoma (P.) natsuae sp. nov. from type locality, pronotum. 16—♂, HT (OSAKA); 17—♂, PT (CCJM) in lateral view; 18—♀, AT (OSAKA). Bars = 1 mm." captionText-3="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3889508" figureDoi-1="http://doi.org/10.5281/zenodo.3889510" figureDoi-2="http://doi.org/10.5281/zenodo.3889512" figureDoi-3="http://doi.org/10.5281/zenodo.3889514" httpUri-0="https://zenodo.org/record/3889508/files/figure.png" httpUri-1="https://zenodo.org/record/3889510/files/figure.png" httpUri-2="https://zenodo.org/record/3889512/files/figure.png" httpUri-3="https://zenodo.org/record/3889514/files/figure.png" pageId="1" pageNumber="262">Figs 1–26</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A58D31ACD08FEA9F8FF831BB303" box="[287,316,1879,1906]" captionStart="FIGURE 52" captionStartId="13.[151,251,1945,1970]" captionTargetBox="[324,1268,196,1911]" captionTargetId="figure@13.[313,1275,181,1919]" captionTargetPageId="13" captionText="FIGURE 52. Map of distribution of Pogonostoma (P.) natsuae sp. nov. and P. (P.) meridionale Fleutiaux." figureDoi="http://doi.org/10.5281/zenodo.3889524" httpUri="https://zenodo.org/record/3889524/files/figure.png" pageId="1" pageNumber="262">52</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6556D31ACD0BFF21F8088013B516" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262" type="materials_examined">
|
||
<paragraph id="8B9836DDD31ACD08FF21F80881AEB3AF" blockId="1.[151,1436,1951,2015]" pageId="1" pageNumber="262">
|
||
<materialsCitation id="3B4F3C80D31ACD08FF21F80881A3B3AF" country="Madagascar" county="Sakaraha" latitude="-22.685" location="Southwestern" longLatPrecision="21" longitude="44.15861" municipality="Analavelona Ankokony Forest" pageId="1" pageNumber="262" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B953EACFD31ACD08FF21F8088308B3CB" bold="true" box="[151,303,1951,1978]" pageId="1" pageNumber="262">
|
||
<typeStatus id="549C887FD31ACD08FF21F80882F5B3CB" box="[151,210,1952,1978]" pageId="1" pageNumber="262">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<location id="8EF86006D31ACD08FE8AF80883F3B3CB" LSID="urn:lsid:plazi:treatment:038E87CBD31ACD01FF21F89D8306B66A:8EF86006D31ACD08FE8AF80883F3B3CB" box="[316,468,1952,1978]" country="Madagascar" county="Sakaraha" latitude="-22.685" longLatPrecision="21" longitude="44.15861" municipality="Analavelona Ankokony Forest" name="Southwestern" pageId="1" pageNumber="262">Southwestern</location>
|
||
<collectingCountry id="F330764DD31ACD08FE6DF8088045B3CB" box="[475,610,1952,1978]" name="Madagascar" pageId="1" pageNumber="262">Madagascar</collectingCountry>
|
||
(district of Atsimo-andrefana): isolated subhumid
|
||
<collectingMunicipality id="6BFCACA7D31ACD08FB22F80882FAB3AE" pageId="1" pageNumber="262">Analavelona Ankokony Forest</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="EE13501AD31ACD08FF5BF86C835CB3AF" box="[237,379,1988,2014]" degrees="22" direction="south" minutes="41" orientation="latitude" pageId="1" pageNumber="262" precision="15" seconds="06" value="-22.685">22°41´06´´S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE13501AD31ACD08FE31F86C8003B3AF" box="[391,548,1988,2014]" degrees="044" direction="east" minutes="09" orientation="longitude" pageId="1" pageNumber="262" precision="15" seconds="31" value="44.15861">044°09´31´´E</geoCoordinate>
|
||
),
|
||
<quantity id="4CDF9B38D31ACD08FD8FF86C80A7B3AF" box="[569,640,1988,2014]" metricMagnitude="4" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="262" unit="km" value="46.0">46 km</quantity>
|
||
northwest of
|
||
<collectingCounty id="62F94E51D31ACD08FCACF86C81A3B3AF" box="[794,900,1988,2014]" pageId="1" pageNumber="262">Sakaraha</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FF318013B516" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
<emphasis id="B953EACFD319CD0BFF71FF31834AB4C2" bold="true" box="[199,365,152,179]" pageId="2" pageNumber="263">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B4F3C80D319CD0BFECDFF308676B4A6" collectionCode="OSAKA" collectorName="Elise Razanajaonarivalona" country="Madagascar" location="Analavelona Ankokony Forest" pageId="2" pageNumber="263" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus id="549C887FD319CD0BFECDFF3083C4B4C3" box="[379,483,152,178]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D21FD54D319CD0BFE5CFF3183D8B4C2" box="[490,511,153,179]" pageId="2" pageNumber="263" type="male">♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFD97FF3180A3B4C2" box="[545,644,153,179]" pageId="2" pageNumber="263">OSAKA</collectionCode>
|
||
, labelled: “
|
||
<location id="8EF86006D319CD0BFCB6FF30867FB4C2" LSID="urn:lsid:plazi:treatment:038E87CBD31ACD01FF21F89D8306B66A:8EF86006D319CD0BFCB6FF30867FB4C2" box="[768,1112,152,179]" country="Madagascar" name="Analavelona Ankokony Forest" pageId="2" pageNumber="263">Analavelona Ankokony Forest</location>
|
||
/ Sakaraha (Atsimo-andrefa- na) / Region
|
||
<collectingCountry id="F330764DD319CD0BFE9FFF148397B4A7" box="[297,432,188,214]" name="Madagascar" pageId="2" pageNumber="263">Madagascar</collectingCountry>
|
||
/ Dec. 1–4.2016 /
|
||
<collectorName id="26D2530BD319CD0BFD36FF14818CB4A7" box="[640,939,188,214]" pageId="2" pageNumber="263">Elisé Razanajaonarivalona</collectorName>
|
||
leg.” [printed]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4F3C80D319CD0BFBECFF148356B48B" collectionCode="OSAKA" collectorName="Elise Razanajaonarivalona" country="Madagascar" location="Analavelona Ankokony Forest" pageId="2" pageNumber="263" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
|
||
<typeStatus id="549C887FD319CD0BFBECFF14869BB4A7" box="[1114,1212,188,214]" pageId="2" pageNumber="263" type="allotype">Allotype</typeStatus>
|
||
<specimenCount id="9D21FD54D319CD0BFB75FF1586FFB4A6" box="[1219,1240,189,215]" pageId="2" pageNumber="263" type="female">♀</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFB4AFF158778B4A6" box="[1276,1375,189,215]" pageId="2" pageNumber="263">OSAKA</collectionCode>
|
||
, with same locality label
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4F3C80D319CD0BFECBFF498082B48A" box="[381,677,224,251]" collectionCode="MNHN" country="Madagascar" pageId="2" pageNumber="263" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus id="549C887FD319CD0BFECBFF4983C9B48A" box="[381,494,225,251]" pageId="2" pageNumber="263" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<specimenCount id="9D21FD54D319CD0BFE4CFF498001B48A" box="[506,550,225,251]" pageId="2" pageNumber="263" type="male">1 ♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFDFBFF498082B48A" box="[589,677,225,251]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="263">MNHN</collectionCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4F3C80D319CD0BFD05FF498160B48A" box="[691,839,224,251]" collectionCode="SDEI" country="Madagascar" pageId="2" pageNumber="263" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D21FD54D319CD0BFD05FF4980F8B48A" box="[691,735,225,251]" pageId="2" pageNumber="263" type="male">1 ♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFCB0FF498160B48A" box="[774,839,225,251]" pageId="2" pageNumber="263">SDEI</collectionCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4F3C80D319CD0BFCE2FF498614B48A" box="[852,1075,224,251]" collectionCode="ZUAC" country="Madagascar" pageId="2" pageNumber="263" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D21FD54D319CD0BFCE2FF4981A5B48A" box="[852,898,225,251]" pageId="2" pageNumber="263" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D21FD54D319CD0BFC26FF49819BB48A" box="[912,956,225,251]" pageId="2" pageNumber="263" type="female">1 ♀</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFC52FF498614B48A" box="[996,1075,225,251]" country="Madagascar" httpUri="http://grbio.org/cool/dprk-p82z" name="University of Antananarivo" pageId="2" pageNumber="263">ZUAC</collectionCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4F3C80D319CD0BFBF6FF498717B48A" box="[1088,1328,224,251]" collectionCode="CCJM" country="Madagascar" pageId="2" pageNumber="263" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount id="9D21FD54D319CD0BFBF6FF4986A4B48A" box="[1088,1155,225,251]" pageId="2" pageNumber="263" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D21FD54D319CD0BFB27FF49869AB48A" box="[1169,1213,225,251]" pageId="2" pageNumber="263" type="female">1 ♀</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFB52FF498717B48A" box="[1252,1328,225,251]" pageId="2" pageNumber="263">CCJM</collectionCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4F3C80D319CD0BFA8BFF498317B56E" collectionCode="CJVB" country="Madagascar" pageId="2" pageNumber="263" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
|
||
<specimenCount id="9D21FD54D319CD0BFA8BFF4987A7B48A" box="[1341,1408,225,251]" pageId="2" pageNumber="263" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D21FD54D319CD0BFA38FF4982E6B56E" pageId="2" pageNumber="263" type="female">2 ♀♀</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFF5FFEAD8317B56E" box="[233,304,261,287]" pageId="2" pageNumber="263">CJVB</collectionCode>
|
||
</materialsCitation>
|
||
, other
|
||
<materialsCitation id="3B4F3C80D319CD0BFE37FEAC80B0B56E" box="[385,663,260,287]" collectionCode="MHCW" country="Madagascar" pageId="2" pageNumber="263" specimenCount="12" specimenCount-female="6" specimenCount-male="6" typeStatus="paratype">
|
||
<specimenCount id="9D21FD54D319CD0BFE37FEAC83E3B56E" box="[385,452,260,287]" pageId="2" pageNumber="263" type="male">6 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D21FD54D319CD0BFE64FEAC8034B56E" box="[466,531,260,287]" pageId="2" pageNumber="263" type="female">6 ♀♀</specimenCount>
|
||
in
|
||
<collectionCode id="ED36AE18D319CD0BFD8AFEAD80B0B56E" box="[572,663,261,287]" pageId="2" pageNumber="263">MHCW</collectionCode>
|
||
</materialsCitation>
|
||
. The type specimens labelled: “
|
||
<typeStatus id="549C887FD319CD0BFBB5FEAC8654B56F" box="[1027,1139,260,286]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
|
||
(“
|
||
<typeStatus id="549C887FD319CD0BFB39FEAC86DCB56F" box="[1167,1275,260,286]" pageId="2" pageNumber="263" type="allotype">Allotype</typeStatus>
|
||
or
|
||
<typeStatus id="549C887FD319CD0BFA9EFEAD87B5B56E" box="[1320,1426,261,287]" pageId="2" pageNumber="263" type="paratype">Paratype</typeStatus>
|
||
” respectively) /
|
||
<taxonomicName id="4C274D5ED319CD0BFEFFFE8183CEB532" authorityName="Klug" authorityYear="1835" box="[329,489,297,323]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">Pogonostoma</taxonomicName>
|
||
(s. str.) / natsuae
|
||
<taxonomicNameLabel id="A26057B4D319CD0BFD0AFE81813AB532" box="[700,797,297,323]" pageId="2" pageNumber="263" rank="species">
|
||
<emphasis id="B953EACFD319CD0BFD0AFE818133B532" bold="true" box="[700,788,297,323]" pageId="2" pageNumber="263">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
/ det. Jiří Moravec, / Elisé Hugo Razanajaonarivalona / & Michio Hori, 2019” [red, printed].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6556D319CD0BFF71FED88105B7F2" pageId="2" pageNumber="263" type="diagnosis">
|
||
<paragraph id="8B9836DDD319CD0BFF71FED8811DB6F6" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
<emphasis id="B953EACFD319CD0BFF71FED883E9B5FA" bold="true" box="[199,462,368,395]" pageId="2" pageNumber="263">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C274D5ED319CD0BFE63FED98123B5FB" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[469,772,369,395]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFE63FED98056B5FB" box="[469,625,369,394]" italics="true" pageId="2" pageNumber="263">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFD34FED980B0B5FB" box="[642,663,369,394]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFD1EFED98123B5FB" box="[680,772,369,394]" italics="true" pageId="2" pageNumber="263">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B953EACFD319CD0BFCBDFED98141B5FA" bold="true" box="[779,870,369,395]" pageId="2" pageNumber="263">
|
||
<taxonomicNameLabel id="A26057B4D319CD0BFCBDFED98141B5FA" box="[779,870,369,395]" pageId="2" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is considered here a species of the
|
||
<taxonomicName id="4C274D5ED319CD0BFB4CFED987BBB5FB" authorityName=", in Audouin & Brullé" authorityYear="2013" baseAuthorityName="Brullé" baseAuthorityYear="1834" box="[1274,1436,368,395]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="elegans" subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFB4CFED98728B5FB" box="[1274,1295,369,394]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFA96FED98712B5FB" box="[1312,1333,369,394]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFAF3FED887BBB5FB" box="[1349,1436,368,394]" italics="true" pageId="2" pageNumber="263">elegans</emphasis>
|
||
</taxonomicName>
|
||
species-group (sensu
|
||
<bibRefCitation id="EFB64B2CD319CD0BFE3DFE3D8013B5DE" author="Moravec, J." box="[395,564,405,431]" pageId="2" pageNumber="263" refId="ref7327" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstv Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
). The new species is externally similar to
|
||
<taxonomicName id="4C274D5ED319CD0BFBBCFE3D87B0B5DE" authority="Fleutiaux, 1899" authorityName="Fleutiaux" authorityYear="1899" box="[1034,1431,404,431]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFBBCFE3D8638B5DF" box="[1034,1055,405,430]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFB98FE3D8663B5DF" box="[1070,1092,405,430]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFBE5FE3C86FAB5DF" box="[1107,1245,404,430]" italics="true" pageId="2" pageNumber="263">meridionale</emphasis>
|
||
<bibRefCitation id="EFB64B2CD319CD0BFB55FE3C87B0B5DE" author="Fleutiaux, E." box="[1251,1431,404,431]" pageId="2" pageNumber="263" pagination="384" refId="ref6999" refString="Fleutiaux, E. (1899) Notes recificatives sur deux Cicindelidae (Col.) et descriptions d'uen espece nouvelle. Buletin de la Societ entomologique de France, 1899, 384." type="journal article" year="1899">Fleutiaux, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
, but can be distinguished by its more glossy elytra due to sparser and paler, blonde to straw-coloured ornamental setae, and elytral surface with generally larger and more regular punctures (
|
||
<figureCitation id="131C2A58D319CD0BFC4BFE7486A4B586" box="[1021,1155,476,503]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1869,1894]" captionTargetBox="[156,1416,193,1828]" captionTargetId="figure@4.[151,1435,181,1844]" captionTargetPageId="4" captionText="FIGURES 10–15. Pogonostoma (P.) natsuae sp. nov. from type locality. 10–12—elytral apices: 10—♂, HT (OSAKA); 11—♂, PT (CCJM); 12—♀, AT (OSAKA); 13–14—elytron: 14—♂, HT; 15—♀, AT; 15—sculpture on elytral disc, ♂, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889510" httpUri="https://zenodo.org/record/3889510/files/figure.png" pageId="2" pageNumber="263">Figs 10–15</figureCitation>
|
||
). Moreover, elytral apex possesses right-angled, rarely acute-angled inner tooth in both sexes (
|
||
<figureCitation id="131C2A58D319CD0BFC1CFDA88608B66A" box="[938,1071,512,539]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1869,1894]" captionTargetBox="[156,1416,193,1828]" captionTargetId="figure@4.[151,1435,181,1844]" captionTargetPageId="4" captionText="FIGURES 10–15. Pogonostoma (P.) natsuae sp. nov. from type locality. 10–12—elytral apices: 10—♂, HT (OSAKA); 11—♂, PT (CCJM); 12—♀, AT (OSAKA); 13–14—elytron: 14—♂, HT; 15—♀, AT; 15—sculpture on elytral disc, ♂, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889510" httpUri="https://zenodo.org/record/3889510/files/figure.png" pageId="2" pageNumber="263">Figs 10–14</figureCitation>
|
||
). In contrast, the elytral punctation in
|
||
<taxonomicName id="4C274D5ED319CD0BFF5FFD8D8398B64F" box="[233,447,548,575]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFF5FFD8D82D9B64F" box="[233,254,549,574]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFEB9FD8D8303B64F" box="[271,292,549,574]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFE83FD8C8398B64F" box="[309,447,548,574]" italics="true" pageId="2" pageNumber="263">meridionale</emphasis>
|
||
</taxonomicName>
|
||
is generally finer and more irregular, densest and with multiform intervals, particularly irregular along sutures, with more distinctly aciculate anterior margins on posterior elytral area, and the elytral apices are mostly sexually dimorphic in shape (
|
||
<figureCitation id="131C2A58D319CD0BFD1CFDC48109B6F6" box="[682,814,620,647]" captionStart="FIGURES 42–47" captionStartId="10.[151,264,1850,1875]" captionTargetBox="[170,1418,192,1817]" captionTargetId="figure@10.[155,1432,181,1825]" captionTargetPageId="10" captionText="FIGURES 42–47. Pogonostoma (P.) meridionale Fleutiaux. 42–44— elytral apices: 42—♂, Tolagnaro (CCJM); 43—♂, Be- hara, CCJM; 44—♀, Bekily (CCJM ex APCA); 45–46—elytron: 45—♂, “Fort Dauphin” = Tolagnaro, LT (MNHN); 46—♀, Bekily (CCJM ex APCA); 47—♂, Tolagnaro (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889520" httpUri="https://zenodo.org/record/3889520/files/figure.png" pageId="2" pageNumber="263">Figs 42–46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FD3880C5B74A" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
The lateral aspect of the aedeagus in both species appears somewhat similar, but the aedeagus of the new species is straighter (
|
||
<figureCitation id="131C2A58D319CD0BFEE9FD1C83C4B6BE" box="[351,483,692,719]" captionStart="FIGURES 19–32" captionStartId="6.[151,264,1800,1825]" captionTargetBox="[151,1435,192,1767]" captionTargetId="figure@6.[151,1435,181,1774]" captionTargetPageId="6" captionText="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="2" pageNumber="263">Figs 19, 22</figureCitation>
|
||
), while it is notably boomerang-bent in
|
||
<taxonomicName id="4C274D5ED319CD0BFC1AFD1D86A4B6BF" box="[940,1155,692,719]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFC1AFD1D81E6B6BF" box="[940,961,693,718]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFC64FD1D81CFB6BF" box="[978,1000,693,718]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFC4FFD1C86A4B6BF" box="[1017,1155,692,718]" italics="true" pageId="2" pageNumber="263">meridionale</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131C2A58D319CD0BFB25FD1C873EB6BE" box="[1171,1305,692,719]" captionStart="FIGURES 19–32" captionStartId="6.[151,264,1800,1825]" captionTargetBox="[151,1435,192,1767]" captionTargetId="figure@6.[151,1435,181,1774]" captionTargetPageId="6" captionText="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="2" pageNumber="263">Figs 28, 30</figureCitation>
|
||
). However, the major difference lies in the shape of the aedeagus apex in its ventral (and dorsal) aspect: while it is conically attenuated and nearly pointed in
|
||
<emphasis id="B953EACFD319CD0BFE49FD558037B767" box="[511,528,765,790]" italics="true" pageId="2" pageNumber="263">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C274D5ED319CD0BFDAAFD558083B767" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[540,676,765,791]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFD90FD558010B767" box="[550,567,765,790]" italics="true" pageId="2" pageNumber="263">P</emphasis>
|
||
.)
|
||
<emphasis id="B953EACFD319CD0BFDFBFD558083B767" box="[589,676,765,790]" italics="true" pageId="2" pageNumber="263">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057B4D319CD0BFD1CFD558124B766" box="[682,771,765,791]" pageId="2" pageNumber="263" rank="species">
|
||
<emphasis id="B953EACFD319CD0BFD1CFD5580DAB766" bold="true" box="[682,765,765,791]" pageId="2" pageNumber="263">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="131C2A58D319CD0BFCA7FD54862DB766" box="[785,1034,764,791]" captionStart="FIGURES 19–32" captionStartId="6.[151,264,1800,1825]" captionTargetBox="[151,1435,192,1767]" captionTargetId="figure@6.[151,1435,181,1774]" captionTargetPageId="6" captionText="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="2" pageNumber="263">Figs 20, 21, 23, 24, 26</figureCitation>
|
||
), it is constricted into thin, elongatecylindric tip in
|
||
<taxonomicName id="4C274D5ED319CD0BFEF5FC89803FB74B" box="[323,536,800,827]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFEF5FC89837EB74B" box="[323,345,801,826]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFEDFFC898359B74B" box="[361,382,801,826]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFE38FC88803FB74B" box="[398,536,800,826]" italics="true" pageId="2" pageNumber="263">meridionale</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131C2A58D319CD0BFD91FC8880F2B74A" box="[551,725,800,827]" captionStart="FIGURES 19–32" captionStartId="6.[151,264,1800,1825]" captionTargetBox="[151,1435,192,1767]" captionTargetId="figure@6.[151,1435,181,1774]" captionTargetPageId="6" captionText="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="2" pageNumber="263">Figs 29, 31, 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FCED8105B7F2" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
For differences from other relevant species of the
|
||
<taxonomicName id="4C274D5ED319CD0BFD4EFCED8158B72F" authorityName=", in Audouin & Brullé" authorityYear="2013" baseAuthorityName="Brullé" baseAuthorityYear="1834" box="[760,895,836,863]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="elegans" subGenus="Pogonostoma">
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFCB4FCED813FB72F" box="[770,792,837,862]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFC9EFCEC8158B72F" box="[808,895,836,862]" italics="true" pageId="2" pageNumber="263">elegans</emphasis>
|
||
</taxonomicName>
|
||
species-group see under
|
||
<taxonomicName id="4C274D5ED319CD0BFB2DFCED8756B72F" box="[1179,1393,836,863]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD319CD0BFB2DFCED8696B72F" box="[1179,1201,837,862]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD319CD0BFB77FCED86F0B72F" box="[1217,1239,837,862]" italics="true" pageId="2" pageNumber="263">P.</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD319CD0BFB51FCEC8756B72F" box="[1255,1393,836,862]" italics="true" pageId="2" pageNumber="263">meridionale</emphasis>
|
||
</taxonomicName>
|
||
below and in the
|
||
<taxonomicName id="4C274D5ED319CD0BFE88FCC183FFB7F3" authorityName="Klug" authorityYear="1835" box="[318,472,873,898]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953EACFD319CD0BFE88FCC183FFB7F3" box="[318,472,873,898]" italics="true" pageId="2" pageNumber="263">Pogonostoma</emphasis>
|
||
</taxonomicName>
|
||
monograph (
|
||
<bibRefCitation id="EFB64B2CD319CD0BFDDBFCC18131B7F2" author="Moravec, J." box="[621,790,873,899]" pageId="2" pageNumber="263" refId="ref7327" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstv Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6556D319CD0EFF71FC2487A6B2ED" lastPageId="7" lastPageNumber="268" pageId="2" pageNumber="263" type="description">
|
||
<paragraph id="8B9836DDD319CD0BFF71FC2482C1B79E" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
<emphasis id="B953EACFD319CD0BFF71FC248376B7D6" bold="true" box="[199,337,908,935]" pageId="2" pageNumber="263">Description</emphasis>
|
||
. Body 10.8–12.5 (HT 10.9, AT 12.3) mm long, 2.70–3.30 (HT 2.80, AT 3.10) mm wide, shiny black, setal vesture whitish-yellow to ochraceous or straw-coloured (the coloration changes depending on lightangle).
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FC50838EB046" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
Head (
|
||
<figureCitation id="131C2A58D319CD0BFEA4FC50835FB062" box="[274,376,1016,1043]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Figs 2–3</figureCitation>
|
||
) notably narrower than body, width
|
||
<quantity id="4CDF9B38D319CD0BFCAFFC5081E7B062" box="[793,960,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="2" pageNumber="263" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.30–2.60 mm</quantity>
|
||
; temples rather short, (slightly less than 2 times shorter than eyes).
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FBE98020B0D3" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">Frons merging with clypeus in middle and not differentiated from vertex, flat or moderately convex; supraantennal keels consisting of elevated, blunt to acute anterior crest and usually less distinct posterior one; surface irregularly scabrous to alveolate.</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FB05803BB143" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">Vertex moderately convex in middle with faint posteromedian impression, surface rather finely irregularly scabrous-alveolate, sculpture coarser on posterior area and passing to tubercles and short transverse-wavy rugae on occipital area; dorsal surface of head covered with erect, very short, whitish to ochre-yellow hairlike setae which are better obvious only in lateral view.</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FA958039B10A" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">Genae finely or rather coarsely rugulose on posterior and juxtaorbital area, with several decumbent and erect, whitish, partly barely visible setae.</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FA2C863BB1EE" blockId="2.[151,1437,152,1979]" box="[199,1052,1412,1439]" pageId="2" pageNumber="263">Clypeus black, rough-coriaceous, with dense, whitish or blonde, short setae.</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71FA008681B206" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
Labrum primarily 6-setose, rarely 7-setose with 4–5 anterior and 2 lateral, ochre to ochre-brown setae, surface black, with moderate central convexity, rough-coriaceous, nearly glabrous or with scattered, barely obvious short microtrichia; lateral margins moderately arcuate towards indistinct lateral indentations, with rounded anterolateral margins (anterolateral teeth effaced), anterior margin irregularly dentate (with five to six small teeth); male labrum (
|
||
<figureCitation id="131C2A58D319CD0BFF29F9908325B222" box="[159,258,1592,1619]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Figs 7–8</figureCitation>
|
||
)
|
||
<quantity id="4CDF9B38D319CD0BFEA7F9918394B222" box="[273,435,1593,1619]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="2" pageNumber="263" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.70–0.80 mm</quantity>
|
||
long,
|
||
<quantity id="4CDF9B38D319CD0BFE4EF99080BCB222" box="[504,667,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.35" metricValueMin="1.2" pageId="2" pageNumber="263" unit="mm" value="1.275" valueMax="1.35" valueMin="1.2">1.20–1.35 mm</quantity>
|
||
wide, dentate anterior margin subtruncate to truncate; female labrum more prolonged anteriad, mostly semicircular, length 0.90–0.1.00 mm, width
|
||
<quantity id="4CDF9B38D319CD0BFC4AF9F48685B206" box="[1020,1186,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="2" pageNumber="263" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.40–1.50 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71F9288392B293" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
Maxillae (
|
||
<figureCitation id="131C2A58D319CD0BFE8CF92883A7B2EA" box="[314,384,1664,1691]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Fig. 6</figureCitation>
|
||
): galea black, sometimes with brownish apex of basal (longest) galeomere; lacinia black with brownish, rarely reddish-brown faint tinge and with brownish or rusty setae; apical portion moderately clavate-dilated,
|
||
<quantity id="4CDF9B38D319CD0BFF6AF9608356B292" box="[220,369,1736,1763]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="2" pageNumber="263" unit="mm" value="0.375" valueMax="0.4" valueMin="0.35">0.35–0.4 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71F94480E5B33F" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
Palpi (
|
||
<figureCitation id="131C2A58D319CD0BFEA7F9448371B376" box="[273,342,1772,1799]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Fig. 2</figureCitation>
|
||
) of usual length and shape, black, except for cinnamon-brown inner apical area of terminal palpomeres, covered with scattered, short whitish or greyish microtrichia, longest palpomeres of labial palpi with long, black-brown setae with mahogany to violet tinge.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD319CD0BFF71F8F087ABB3CB" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="263">
|
||
Mandibles (
|
||
<figureCitation id="131C2A58D319CD0BFEFAF8F08388B302" box="[332,431,1880,1907]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Figs 3–5</figureCitation>
|
||
) with rather short terminal teeth, sexually dimorphic in shape, in male (
|
||
<figureCitation id="131C2A58D319CD0BFB79F8F08708B302" box="[1231,1327,1880,1907]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Fig. 3–4</figureCitation>
|
||
) subsymmetrical (apart from the longer terminal tooth in right mandible as usual) with second tooth in both mandibles notably smaller than third tooth; female mandibles (
|
||
<figureCitation id="131C2A58D319CD0BFD1AF80880C8B3CA" box="[684,751,1952,1979]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1869,1894]" captionTargetBox="[162,1427,191,1829]" captionTargetId="figure@3.[151,1435,179,1842]" captionTargetPageId="3" captionText="FIGURES 1–9. Pogonostoma (P.) natsuae sp. nov. from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="2" pageNumber="263">Fig. 5</figureCitation>
|
||
) with inner teeth of right mandible almost of the same size.
|
||
</paragraph>
|
||
<caption id="DF586655D318CD0AFF21F8E58631B3DC" ID-DOI="http://doi.org/10.5281/zenodo.3889508" ID-Zenodo-Dep="3889508" httpUri="https://zenodo.org/record/3889508/files/figure.png" pageId="3" pageNumber="264" startId="3.[151,264,1869,1894]" targetBox="[162,1427,191,1829]" targetPageId="3">
|
||
<paragraph id="8B9836DDD318CD0AFF21F8E58631B3DC" blockId="3.[151,1437,1869,1966]" pageId="3" pageNumber="264">
|
||
<emphasis id="B953EACFD318CD0AFF21F8E5831CB317" bold="true" box="[151,315,1869,1894]" pageId="3" pageNumber="264">FIGURES 1–9.</emphasis>
|
||
<taxonomicName id="4C274D5ED318CD0AFEF7F8E78074B317" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[321,595,1870,1894]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="263" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD318CD0AFEF7F8E783EBB317" box="[321,460,1871,1894]" italics="true" pageId="3" pageNumber="264">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD318CD0AFE6AF8E783CCB317" box="[476,491,1871,1894]" italics="true" pageId="3" pageNumber="264">P</emphasis>
|
||
.)
|
||
<emphasis id="B953EACFD318CD0AFDB6F8E78074B317" box="[512,595,1871,1894]" italics="true" pageId="3" pageNumber="264">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057B4D318CD0AFDEDF8E68088B314" box="[603,687,1870,1894]" pageId="3" pageNumber="264" rank="species">
|
||
<emphasis id="B953EACFD318CD0AFDEDF8E6808FB317" bold="true" box="[603,680,1870,1894]" pageId="3" pageNumber="264">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from type locality. 1—habitus, ♂, HT (OSAKA); 2–3—head or its part: 2—♂, HT (OSAKA); 3—♂, PT (ZUAC); 4–5—mandibles: 4—♂, PT (ZUAC); 5—♀, AT (OSAKA); 6—galea and lacinia, ♂, PT (CCJM); 7–9—labrum: 7—♂, HT; 8—♂, PT (ZUAC); 9—♀, AT. Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF586655D31FCD0DFF21F8E58303B3DC" ID-DOI="http://doi.org/10.5281/zenodo.3889510" ID-Zenodo-Dep="3889510" httpUri="https://zenodo.org/record/3889510/files/figure.png" pageId="4" pageNumber="265" startId="4.[151,264,1869,1894]" targetBox="[156,1416,193,1828]" targetPageId="4">
|
||
<paragraph id="8B9836DDD31FCD0DFF21F8E58303B3DC" blockId="4.[151,1437,1869,1966]" pageId="4" pageNumber="265">
|
||
<emphasis id="B953EACFD31FCD0DFF21F8E58370B317" bold="true" box="[151,343,1869,1894]" pageId="4" pageNumber="265">FIGURES 10–15.</emphasis>
|
||
<taxonomicName id="4C274D5ED31FCD0DFED7F8E7805EB317" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[353,633,1870,1894]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="263" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD31FCD0DFED7F8E783CBB317" box="[353,492,1871,1894]" italics="true" pageId="4" pageNumber="265">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD31FCD0DFE48F8E7802AB317" box="[510,525,1871,1894]" italics="true" pageId="4" pageNumber="265">P</emphasis>
|
||
.)
|
||
<emphasis id="B953EACFD31FCD0DFD90F8E7805EB317" box="[550,633,1871,1894]" italics="true" pageId="4" pageNumber="265">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057B4D31FCD0DFD35F8E680FDB314" box="[643,730,1870,1894]" pageId="4" pageNumber="265" rank="species">
|
||
<emphasis id="B953EACFD31FCD0DFD35F8E680F4B317" bold="true" box="[643,723,1870,1894]" pageId="4" pageNumber="265">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from type locality. 10–12—elytral apices: 10—♂, HT (OSAKA); 11—♂, PT (CCJM); 12—♀, AT (OSAKA); 13–14—elytron: 14—♂, HT; 15—♀, AT; 15—sculpture on elytral disc, ♂, HT. Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF586655D31ECD0CFF21F8ED8128B3F3" ID-DOI="http://doi.org/10.5281/zenodo.3889512" ID-Zenodo-Dep="3889512" httpUri="https://zenodo.org/record/3889512/files/figure.png" pageId="5" pageNumber="266" startId="5.[151,264,1861,1886]" targetBox="[164,1424,194,1823]" targetPageId="5">
|
||
<paragraph id="8B9836DDD31ECD0CFF21F8ED8128B3F3" blockId="5.[151,1436,1861,1922]" pageId="5" pageNumber="266">
|
||
<emphasis id="B953EACFD31ECD0CFF21F8ED837FB32F" bold="true" box="[151,344,1861,1886]" pageId="5" pageNumber="266">FIGURES 16–18.</emphasis>
|
||
<taxonomicName id="4C274D5ED31ECD0CFED7F8EF805DB32F" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[353,634,1862,1886]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD31ECD0CFED7F8EF83CBB32F" box="[353,492,1863,1886]" italics="true" pageId="5" pageNumber="266">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD31ECD0CFE49F8EF8029B32F" box="[511,526,1863,1886]" italics="true" pageId="5" pageNumber="266">P</emphasis>
|
||
.)
|
||
<emphasis id="B953EACFD31ECD0CFD91F8EF805DB32F" box="[551,634,1863,1886]" italics="true" pageId="5" pageNumber="266">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057B4D31ECD0CFD32F8EE80FBB32F" box="[644,732,1862,1886]" pageId="5" pageNumber="266" rank="species">
|
||
<emphasis id="B953EACFD31ECD0CFD32F8EE80F2B32F" bold="true" box="[644,725,1862,1886]" pageId="5" pageNumber="266">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from type locality, pronotum. 16—♂, HT (OSAKA); 17—♂, PT (CCJM) in lateral view; 18—♀, AT (OSAKA). Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF586655D31DCD0FFF21F8A0805DB3C1" ID-DOI="http://doi.org/10.5281/zenodo.3889514" ID-Zenodo-Dep="3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="6" pageNumber="267" startId="6.[151,264,1800,1825]" targetBox="[151,1435,192,1767]" targetPageId="6">
|
||
<paragraph id="8B9836DDD31DCD0FFF21F8A0805DB3C1" blockId="6.[151,1437,1800,1969]" pageId="6" pageNumber="267">
|
||
<emphasis id="B953EACFD31DCD0FFF21F8A0837FB350" bold="true" box="[151,344,1800,1825]" pageId="6" pageNumber="267">FIGURES 19–32.</emphasis>
|
||
two species of
|
||
<taxonomicName id="4C274D5ED31DCD0FFDB5F8A180B7B351" authorityName="Klug" authorityYear="1835" box="[515,656,1801,1824]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="267" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953EACFD31DCD0FFDB5F8A180B7B351" box="[515,656,1801,1824]" italics="true" pageId="6" pageNumber="267">Pogonostoma</emphasis>
|
||
</taxonomicName>
|
||
, male aedeagus or its apex. 19–26—
|
||
<taxonomicName id="4C274D5ED31DCD0FFB90F8A1871AB351" authority="Moravec & Razanajaonarivalona & Hori, 2020" authorityName="Moravec & Razanajaonarivalona & Hori" authorityYear="2020" box="[1062,1341,1801,1825]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="263" phylum="Arthropoda" rank="species" species="natsuae" status="sp. nov." subGenus="Pogonostoma">
|
||
<emphasis id="B953EACFD31DCD0FFB90F8A18696B351" box="[1062,1201,1801,1824]" italics="true" pageId="6" pageNumber="267">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD31DCD0FFB74F8A186F6B351" box="[1218,1233,1801,1824]" italics="true" pageId="6" pageNumber="267">P</emphasis>
|
||
.)
|
||
<emphasis id="B953EACFD31DCD0FFB5CF8A1871AB351" box="[1258,1341,1801,1824]" italics="true" pageId="6" pageNumber="267">natsuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057B4D31DCD0FFAF0F8A187BBB351" box="[1350,1436,1801,1825]" pageId="6" pageNumber="267" rank="species">
|
||
<emphasis id="B953EACFD31DCD0FFAF0F8A187B1B350" bold="true" box="[1350,1430,1801,1825]" pageId="6" pageNumber="267">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—
|
||
<taxonomicName id="4C274D5ED31DCD0FFDC5F8F980D9B319" authorityName="Klug" authorityYear="1835" box="[627,766,1873,1896]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="267" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953EACFD31DCD0FFDC5F8F980D9B319" box="[627,766,1873,1896]" italics="true" pageId="6" pageNumber="267">Pogonostoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C274D5ED31DCD0FFCBEF8F98639B318" authority="Fleutiaux" authorityName="Fleutiaux" box="[776,1054,1872,1897]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="267" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
|
||
(
|
||
<emphasis id="B953EACFD31DCD0FFCA6F8F98138B319" box="[784,799,1873,1896]" italics="true" pageId="6" pageNumber="267">P</emphasis>
|
||
.)
|
||
<emphasis id="B953EACFD31DCD0FFC81F8F98196B318" box="[823,945,1873,1897]" italics="true" pageId="6" pageNumber="267">meridionale</emphasis>
|
||
Fleutiaux
|
||
</taxonomicName>
|
||
: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9836DDD31CCD0EFF71FF31872CB48A" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="268">Antennae black, in male as long as body or slightly longer, in female somewhat shorter; scape covered with rather dense whitish microsetae and with long, yellow-brown subanterior (easily abraded) setae; antennomeres 2–4 with rather numerous but scattered whitish microsetae, 5–11 with pubescence of greyish microtrichia.</paragraph>
|
||
<paragraph id="8B9836DDD31CCD0EFF71FEAD875BB74A" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="268">
|
||
Thorax: pronotum (
|
||
<figureCitation id="131C2A58D31CCD0EFE10FEAD800FB551" box="[422,552,261,288]" captionStart="FIGURES 16–18" captionStartId="5.[151,264,1861,1886]" captionTargetBox="[164,1424,194,1823]" captionTargetId="figure@5.[151,1435,181,1836]" captionTargetPageId="5" captionText="FIGURES 16–18. Pogonostoma (P.) natsuae sp. nov. from type locality, pronotum. 16—♂, HT (OSAKA); 17—♂, PT (CCJM) in lateral view; 18—♀, AT (OSAKA). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889512" httpUri="https://zenodo.org/record/3889512/files/figure.png" pageId="7" pageNumber="268">Figs 16–18</figureCitation>
|
||
) elongate,
|
||
<quantity id="4CDF9B38D31CCD0EFD1EFEAD816AB56E" box="[680,845,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="7" pageNumber="268" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.70–3.10 mm</quantity>
|
||
long,
|
||
<quantity id="4CDF9B38D31CCD0EFC21FEAC861BB56E" box="[919,1084,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.1" metricValueMin="1.85" pageId="7" pageNumber="268" unit="mm" value="1.975" valueMax="2.1" valueMin="1.85">1.85–2.10 mm</quantity>
|
||
wide; anterior lobe as wide as posterior lobe in female, in male often slightly narrower, its surface finely tuberculate and rugulose; pronotal disc in female mostly subglobose, in male with moderately convex, rarely subparallel lateral margins; in lateral view subgibbose (more distinctly in male,
|
||
<figureCitation id="131C2A58D31CCD0EFD82FED980A2B5FA" box="[564,645,369,395]" captionStart="FIGURES 16–18" captionStartId="5.[151,264,1861,1886]" captionTargetBox="[164,1424,194,1823]" captionTargetId="figure@5.[151,1435,181,1836]" captionTargetPageId="5" captionText="FIGURES 16–18. Pogonostoma (P.) natsuae sp. nov. from type locality, pronotum. 16—♂, HT (OSAKA); 17—♂, PT (CCJM) in lateral view; 18—♀, AT (OSAKA). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889512" httpUri="https://zenodo.org/record/3889512/files/figure.png" pageId="7" pageNumber="268">Fig. 17</figureCitation>
|
||
); notopleural sutures invisible from above or very indistinctly at posterior pronotal angles in male; median line indistinct, merging with irregularly cristate-tuberculate surface sculpture consisting of tubercles commonly transversely connected into short, transverse-elongate crests, which are finer on anterior area and becoming larger on median area; posteromedian area adjacent to posterior lobe covered with fine and shallow, more transverse-continuous stria-like rugae; whole discal surface covered with scattered, short, decumbent or erect, whitish to ochre-yellow or straw-yellow hairlike setae which are barely obvious in dorsal view, but well recognizable in lateral view (
|
||
<figureCitation id="131C2A58D31CCD0EFDA2FDE18040B612" box="[532,615,585,611]" captionStart="FIGURES 16–18" captionStartId="5.[151,264,1861,1886]" captionTargetBox="[164,1424,194,1823]" captionTargetId="figure@5.[151,1435,181,1836]" captionTargetPageId="5" captionText="FIGURES 16–18. Pogonostoma (P.) natsuae sp. nov. from type locality, pronotum. 16—♂, HT (OSAKA); 17—♂, PT (CCJM) in lateral view; 18—♀, AT (OSAKA). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889512" httpUri="https://zenodo.org/record/3889512/files/figure.png" pageId="7" pageNumber="268">Fig. 17</figureCitation>
|
||
); posterior lobe shiny and smooth except for few indistinct shallow wrinkles in middle, entirely glabrous; proepisterna large, polished and smooth, finely and shortly parallel-wrinkled on juxtanotopleural area, nearly glabrous, except for only occasional hairlike setae; mesepisterna large, polished, with only occasional setae, mesepimeron deeply impressed; metepisterna notably long, deeply impressed, smooth and glabrous; prosternum smooth, except for very fine transverse wrinkles, rather densely covered with long and erect, whitish hairlike setae arising from barely visible setigerous micropunctures; mesosternum sparsely punctate-setose with shorter whitish setae; metasternum almost smooth, indistinctly sparsely whitish setose, glabrous in middle.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD31CCD0EFF71FCED8095B142" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="268">
|
||
Elytra (
|
||
<figureCitation id="131C2A58D31CCD0EFEACFCED83BEB72E" box="[282,409,836,863]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1869,1894]" captionTargetBox="[156,1416,193,1828]" captionTargetId="figure@4.[151,1435,181,1844]" captionTargetPageId="4" captionText="FIGURES 10–15. Pogonostoma (P.) natsuae sp. nov. from type locality. 10–12—elytral apices: 10—♂, HT (OSAKA); 11—♂, PT (CCJM); 12—♀, AT (OSAKA); 13–14—elytron: 14—♂, HT; 15—♀, AT; 15—sculpture on elytral disc, ♂, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889510" httpUri="https://zenodo.org/record/3889510/files/figure.png" pageId="7" pageNumber="268">Figs 10–15</figureCitation>
|
||
) elongate, 6.00–
|
||
<quantity id="4CDF9B38D31CCD0EFDE5FCED8091B72E" box="[595,694,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="268" unit="mm" value="7.3">7.30 mm</quantity>
|
||
long; humeri obliquely arcuate; lateral margins almost parallel ex- cept for slightly arched bulge in anterior third of the margin in male, and only indistinctly enlarged towards rounded lateral anteapical angles; elytral apices almost uniform in both sexes, with faint sutural emargination towards indistinct sutural spine, and with right-angled, rarely acute-angled inner tooth between external tooth which is thornlike in male, while blunter and right-angled in female; elytral surface convex with moderate basodiscal convexity delineated by deep discal impression, punctate throughout (except for effaced narrow basal and basohumeral area); punctures large and deep on two elytral thirds, large and almost isolated on lateral areas, deepest, large and irregularly anastomosing by lowered lateral intervals on basodiscal convexity, reduced or partly (never entirely) effaced within discal impression, and again large but only occasionally anastomosing on elytral disc except for much finer and irregular sculpture on limited area along sutures; punctures become smaller, more spaced and with aciculate anterior margins of intervals on posterior declivity and whole anteapical area, but larger and irregular on apices; surface of intervals shiny; setal vesture consisting of nearly erect, copious, moderately long, pale to straw-yellow ornamental setae which are mixed with very sparse, long and erect, whitish hairlike sensory setae, which are very long and numerous on lateral portion of humeri.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD31CCD0EFF71FA958058B10A" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="268">Abdomen. Ventrites black, sometimes with very faint diffusing bluish lustre, covered with short, dense, but scattered and mostly appressed white setae.</paragraph>
|
||
<paragraph id="8B9836DDD31CCD0EFF71FA2D8657B25E" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="268">Legs black; coxae rather densely whitish-setose; profemora covered with rather dense medium-long whitish setae; mesofemora with dense whitish setae on their dorsal surface, much sparser setae on ventral area; metafemora with sparser, ochre and brownish setae; pro- and mesotibiae with whitish setae except for dense, ochre-ferrugineous setae on apices of tibiae; tarsi dark reddish-brown, their surface covered with whitish setae (and as in all species, first three protarsomeres in male dilated and with dense setose pad); claws rusty-brown.</paragraph>
|
||
<paragraph id="8B9836DDD31CCD0EFF71F99187A6B2ED" blockId="7.[151,1437,153,2015]" pageId="7" pageNumber="268">
|
||
Aedeagus
|
||
<quantity id="4CDF9B38D31CCD0EFE89F99083C3B222" box="[319,484,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="7" pageNumber="268" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.50–2.60 mm</quantity>
|
||
long,
|
||
<quantity id="4CDF9B38D31CCD0EFD99F99180B2B222" box="[559,661,1593,1619]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="268" unit="mm" value="0.4">0.40 mm</quantity>
|
||
wide, in its left lateral aspect (
|
||
<figureCitation id="131C2A58D31CCD0EFC4DF991868CB222" box="[1019,1195,1592,1619]" captionStart="FIGURES 19–32" captionStartId="6.[151,264,1800,1825]" captionTargetBox="[151,1435,192,1767]" captionTargetId="figure@6.[151,1435,181,1774]" captionTargetPageId="6" captionText="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="7" pageNumber="268">Figs 19, 22, 25</figureCitation>
|
||
) rather distinctly but gradually bent in middle, apical portion conically tapered towards rather narrow and small, dorsad-bent, rounded apex, which is in its ventral (and dorsal) aspect conically attenuated and nearly pointed (
|
||
<figureCitation id="131C2A58D31CCD0EFBC2F9298753B2ED" box="[1140,1396,1665,1692]" captionStart="FIGURES 19–32" captionStartId="6.[151,264,1800,1825]" captionTargetBox="[151,1435,192,1767]" captionTargetId="figure@6.[151,1435,181,1774]" captionTargetPageId="6" captionText="FIGURES 19–32. two species of Pogonostoma, male aedeagus or its apex. 19–26— Pogonostoma (P.) natsuae sp. nov. from type locality: 19–21—HT (OSAKA); 22–24—PT (CCJM); 25–26—PT (ZUAC) (19, 22, 25—lateral view; 20, 23, 26— ventral view; 21, 24—dorsal view). 27–32—Pogonostoma (P.) meridionale Fleutiaux: 27—“Fort Dauphin”, LT (MNHN); 28— Bekily (CCJM ex APCA); 29—ditto, ventral view; 30—“Madagascar” (MFNB); 31—ditto, dorsal view; 32— Behara (CCJM ex APCA), ventral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889514" httpUri="https://zenodo.org/record/3889514/files/figure.png" pageId="7" pageNumber="268">Figs 20, 21, 23, 24, 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6556D31CCD01FF71F90C8300B5A2" lastPageId="8" lastPageNumber="269" pageId="7" pageNumber="268" type="distribution">
|
||
<paragraph id="8B9836DDD31CCD01FF71F90C870DB516" blockId="7.[151,1437,153,2015]" lastBlockId="8.[151,1437,153,539]" lastPageId="8" lastPageNumber="269" pageId="7" pageNumber="268">
|
||
<emphasis id="B953EACFD31CCD0EFF71F90C83CBB2CE" bold="true" box="[199,492,1700,1727]" pageId="7" pageNumber="268">Distribution and biology</emphasis>
|
||
(
|
||
<figureCitation id="131C2A58D31CCD0EFE4DF90D8077B2CE" box="[507,592,1700,1727]" captionStart="FIGURE 52" captionStartId="13.[151,251,1945,1970]" captionTargetBox="[324,1268,196,1911]" captionTargetId="figure@13.[313,1275,181,1919]" captionTargetPageId="13" captionText="FIGURE 52. Map of distribution of Pogonostoma (P.) natsuae sp. nov. and P. (P.) meridionale Fleutiaux." figureDoi="http://doi.org/10.5281/zenodo.3889524" httpUri="https://zenodo.org/record/3889524/files/figure.png" pageId="7" pageNumber="268">Fig. 52</figureCitation>
|
||
). The
|
||
<typeStatus id="549C887FD31CCD0EFD23F90D80E2B2CE" box="[661,709,1701,1727]" pageId="7" pageNumber="268">type</typeStatus>
|
||
locality in the Analavelona Massif
|
||
<quantity id="4CDF9B38D31CCD0EFBE2F90D86BCB2CE" box="[1108,1179,1701,1727]" metricMagnitude="4" metricUnit="m" metricValue="4.6" pageId="7" pageNumber="268" unit="km" value="46.0">46 km</quantity>
|
||
northwest of Sakaraha is characterized as a subhumid forest on basalt at an elevation of
|
||
<quantity id="4CDF9B38D31CCD0EFCC0F9608621B292" box="[886,1030,1736,1763]" metricMagnitude="3" metricUnit="m" metricValue="1.0125" metricValueMax="1.325" metricValueMin="0.7" pageId="7" pageNumber="268" unit="m" value="1012.5" valueMax="1325.0" valueMin="700.0">700–1325 m</quantity>
|
||
. The adults of the new species were caught by the second author on trunks of high but not very large trees at a higher altitude. This uniquely isolated forest, surrounded by dry succulent forest formations at lower elevations, is maintained owing to a condensation of moisture from ascending air. The unique system of the Analavelona Massif encompasses 77 tree and plant families with high level of endemism (
|
||
<bibRefCitation id="EFB64B2CD31CCD0EFE57F8F180C4B302" author="Randrianarivony, T. N." box="[481,739,1880,1907]" pageId="7" pageNumber="268" refId="ref7747" refString="Randrianarivony, T. N. (2015) Etudes Ethnobotaniques Et Ecologiques Des Plantes Utiles De La Nouvelle Aire Protegee D'analavelona (District De Sakaraha, Region Du Sud-Ouest De Madagascar). Antananarivo, 2015. Ph. D. Thesis, Univer- site D'antananarivo Faculte des Sciences, Antananarivo, 138 pp. [http: // biblio. univ-antananarivo. mg / pdfs / randrianarivonyTabitaN _ SN _ DNR _ 15. pdf]" type="book" year="2015">Randrianarivony 2015</bibRefCitation>
|
||
). As specified by
|
||
<bibRefCitation id="EFB64B2CD31CCD0EFC13F8F18685B302" author="Goodman, S. M. & Raherilalao, M. J. & Wohlhauser, S." box="[933,1186,1881,1907]" pageId="7" pageNumber="268" pagination="1589 - 1599" refId="ref7144" refString="Goodman, S. M., Raherilalao, M. J. & Wohlhauser, S. (Eds.) (2018) Les aires protegees terrestres de Madagascar: Leur histoire, description et biote / The terrestrial protected areas of Madagascar: Their history, description, and biota; part. Association Vahatra, Antananarivo, 11 pp. [pp. 1589 - 1599]" type="book chapter" year="2018">Goodman et al. (2018)</bibRefCitation>
|
||
, the subhumid climate is associated with a local orographic effect on the upper portions of the massif with an average annual rainfall of around
|
||
<quantity id="4CDF9B38D31CCD0EFF5BF808836AB3CA" box="[237,333,1952,1979]" metricMagnitude="-1" metricUnit="m" metricValue="8.559999999999999" pageId="7" pageNumber="268" unit="mm" value="856.0">856 mm</quantity>
|
||
, 93% usually falling between November and April. The basaltic substrate probably contributes to the persistence of a particularly moist
|
||
<typeStatus id="549C887FD31CCD0EFDF4F86D8055B3AE" box="[578,626,1989,2015]" pageId="7" pageNumber="268">type</typeStatus>
|
||
of forest of the region which justified the massif being the western most extension of the phytogeographical zone known as the Central Domain. Given its multi-layered composition with absence of
|
||
<taxonomicName id="4C274D5ED313CD01FEAEFF1683DCB4A6" authority="Baill" authorityName="Baill" authorityYear="1874" box="[280,507,189,215]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="269" phylum="Tracheophyta" rank="species" species="bojeri">
|
||
<emphasis id="B953EACFD313CD01FEAEFF16839BB4A6" box="[280,444,189,215]" italics="true" pageId="8" pageNumber="269">Uapaca bojeri</emphasis>
|
||
Baill
|
||
</taxonomicName>
|
||
trees (
|
||
<taxonomicName id="4C274D5ED313CD01FDFFFF1580DEB4A9" authorityName="Martinov" authorityYear="1820" box="[585,761,189,216]" class="Magnoliopsida" family="Phyllanthaceae" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="269" phylum="Tracheophyta" rank="family">Phyllanthaceae</taxonomicName>
|
||
), this forest is not sclerophylous woodland, despite its geographical placement. In fact, it has characteristics of medium altitude moist evergreen forest, as of the 75 recognized tree species only four are deciduous. For other details and the list of tree species see Goodman et. al. (2018). Fortu- nately, the main parts of the nice and now newly protected forest have remained almost untouched, being a sacred place for the Malagasy local Bara people, who believe that it is the home of the spirits of their ancestors.
|
||
</paragraph>
|
||
<paragraph id="8B9836DDD313CD01FF71FED98300B5A2" blockId="8.[151,1437,153,539]" pageId="8" pageNumber="269">
|
||
Two other tiger beetle species were caught at the same time in the
|
||
<typeStatus id="549C887FD313CD01FC0AFED981CBB5FA" box="[956,1004,369,395]" pageId="8" pageNumber="269">type</typeStatus>
|
||
locality of the new species:
|
||
<taxonomicName id="4C274D5ED313CD01FA87FEDA81A7B5DE" authority="Moravec & Vybiral, 2010" authorityName="Moravec & Vybiral" authorityYear="2010" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="zombitsynense" subGenus="Microstenocera">
|
||
<emphasis id="B953EACFD313CD01FA87FEDA82E8B5DE" italics="true" pageId="8" pageNumber="269">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD313CD01FF52FE3E83B3B5DE" box="[228,404,406,431]" italics="true" pageId="8" pageNumber="269">Microstenocera</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD313CD01FE1EFE3D806DB5DE" box="[424,586,405,431]" italics="true" pageId="8" pageNumber="269">zombitsynense</emphasis>
|
||
<bibRefCitation id="EFB64B2CD313CD01FDE3FE3D81A7B5DE" author="Moravec, J. & Vybiral, J." box="[597,896,405,431]" pageId="8" pageNumber="269" pagination="95 - 102" refId="ref7484" refString="Moravec, J. & Vybiral, J. (2010) New or rare Madagascar tiger beetles- 15. Pogonostoma (Microstenocera) zombitsynense sp. nov., a new species from southwestern Madagascar (Coleoptera: Cicindelidae). Folia Heyrovskyana, Series A, 18 (1 - 3), 95 - 102." type="journal article" year="2010">Moravec & Vybíral, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D5ED313CD01FC09FE3D8304B5A2" authority="(W. Horn, 1934)" baseAuthorityName="W. Horn" baseAuthorityYear="1934" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="tricolorata" subGenus="Diarrhiza">
|
||
<emphasis id="B953EACFD313CD01FC09FE3D867EB5DE" box="[959,1113,405,431]" italics="true" pageId="8" pageNumber="269">Physodeutera</emphasis>
|
||
(
|
||
<emphasis id="B953EACFD313CD01FBDBFE3D86FCB5DE" box="[1133,1243,405,431]" italics="true" pageId="8" pageNumber="269">Diarrhiza</emphasis>
|
||
)
|
||
<emphasis id="B953EACFD313CD01FB46FE3D874EB5DE" box="[1264,1385,405,431]" italics="true" pageId="8" pageNumber="269">tricolorata</emphasis>
|
||
(W.
|
||
<bibRefCitation id="EFB64B2CD313CD01FF21FE11833CB5A2" author="Horn, W." box="[151,283,441,467]" pageId="8" pageNumber="269" pagination="7 - 28" refId="ref7085" refString="Horn, W. (1934) Catalogue Bibliographique et synonymique des Cicindelides de Madagascar (Coleoptera, Fam. Carabidae, sbfam. Cicindelini). In: HORN, W. & OLSOUfiEff, G., Les Cicindelides de Madagascar (premiere partie). Memories de l'Academie Malgache, 1934, 7 - 28." type="journal article" year="1934">Horn, 1934</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6556D313CD01FF71FE758306B66A" pageId="8" pageNumber="269" type="etymology">
|
||
<paragraph id="8B9836DDD313CD01FF71FE758306B66A" blockId="8.[151,1437,153,539]" pageId="8" pageNumber="269">
|
||
<emphasis id="B953EACFD313CD01FF71FE75836FB586" bold="true" box="[199,328,477,503]" pageId="8" pageNumber="269">Etymology</emphasis>
|
||
. This species is cordially dedicated to Madame Natsu Hori, the recently deceased mother of the third author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |