251 lines
36 KiB
XML
251 lines
36 KiB
XML
<document id="A1E1DDDCF5F19C19487CC5C2D10BF932" ID-DOI="10.5281/zenodo.206964" ID-GBIF-Dataset="8be3c4b0-eca9-4fa2-b40f-102609149a82" ID-ISSN="1175-5326" ID-Zenodo-Dep="206964" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227623218" checkinUser="plazi" docAuthor="Molineri, Carlos, Grillet, Maria-Eugenia, Nieto, Carolina, Dominguez, Eduardo & Guerrero, Edmundo" docDate="2011" docId="038E879AFFFED12E1DB2F8D5FB52E102" docLanguage="en" docName="zt02750p050.pdf" docOrigin="Zootaxa 2750" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Caenis teipunensis Molineri, Grillet, Nieto, Dominguez & Guerrero, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFB7FFE2FFFFD12A1D25FF8CFF92E33D" masterDocTitle="New species and records for the mayfly families Caenidae, Leptohyphidae and Coryphoridae (Ephemeroptera, Pannota) from Venezuelan Guayana’s Uplands" masterLastPageNumber="50" masterPageNumber="39" pageNumber="40" updateTime="1698238702261" updateUser="plazi">
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<mods:title id="025CB51C6DF640D83A81CF58478882CA">New species and records for the mayfly families Caenidae, Leptohyphidae and Coryphoridae (Ephemeroptera, Pannota) from Venezuelan Guayana’s Uplands</mods:title>
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<mods:namePart id="3D13F59B968F0AD70E31E0B8D3842783">Molineri, Carlos</mods:namePart>
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<mods:namePart id="CBFC28A6520FAA75381AA349F6451526">Grillet, Maria-Eugenia</mods:namePart>
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<mods:namePart id="558B7DDE9934C3D39DF6EDBAC929B69B">Nieto, Carolina</mods:namePart>
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<mods:namePart id="1BB50F8C983A3A84B24DA6C6963CF6DD">Dominguez, Eduardo</mods:namePart>
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<mods:namePart id="18856C306E08527480AC15E620044DFB">Guerrero, Edmundo</mods:namePart>
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<mods:title id="6FBB5115C853E9C55EDF75009643771B">Zootaxa</mods:title>
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<mods:date id="F300D5DB0658ADFADBEBEC415172A679">2011</mods:date>
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<mods:number id="B56015E4DCD04BB3C32B431AB7CB4F79">2750</mods:number>
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<mods:identifier id="17678402752DDF9A90FDFB0BF658BEB4" type="DOI">10.5281/zenodo.206964</mods:identifier>
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<treatment id="038E879AFFFED12E1DB2F8D5FB52E102" ID-DOI="http://doi.org/10.5281/zenodo.3504133" ID-GBIF-Taxon="119386799" ID-Zenodo-Dep="3504133" LSID="urn:lsid:plazi:treatment:038E879AFFFED12E1DB2F8D5FB52E102" httpUri="http://treatment.plazi.org/id/038E879AFFFED12E1DB2F8D5FB52E102" lastPageId="4" lastPageNumber="43" pageId="1" pageNumber="40">
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<subSubSection id="C33D6507FFFED12B1DB2F8D5FE80E4A9" pageId="1" pageNumber="40" type="nomenclature">
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<paragraph id="8B98368CFFFED12B1DB2F8D5FE4DE44E" blockId="1.[151,479,1881,1940]" box="[151,479,1881,1907]" pageId="1" pageNumber="40">
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<heading id="D0D081E0FFFED12B1DB2F8D5FE4DE44E" bold="true" box="[151,479,1881,1907]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
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<emphasis id="B953EA9EFFFED12B1DB2F8D5FE4DE44E" bold="true" box="[151,479,1881,1907]" pageId="1" pageNumber="40">
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<taxonomicName id="4C274D0FFFFED12B1DB2F8D5FEEBE44E" ID-CoL="5WT6W" box="[151,377,1881,1907]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="teipunensis" status="sp. nov.">
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<emphasis id="B953EA9EFFFED12B1DB2F8D5FEEBE44E" bold="true" box="[151,377,1881,1907]" italics="true" pageId="1" pageNumber="40">Caenis teipunensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26057E5FFFED12B1CA4F8D5FE4DE44E" box="[385,479,1881,1907]" pageId="1" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8B98368CFFFED12B1DB2F8F0FE80E4A9" blockId="1.[151,479,1881,1940]" box="[151,274,1916,1940]" pageId="1" pageNumber="40">
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(
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<figureCitation id="131C2A09FFFED12B1DBAF8F0FE9BE4A9" box="[159,265,1916,1940]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 9" captionStartId-0="2.[151,264,1390,1413]" captionStartId-1="3.[151,264,1608,1631]" captionTargetBox-0="[192,1395,256,1366]" captionTargetBox-1="[227,1351,343,1581]" captionTargetId-0="figure@2.[192,1395,256,1366]" captionTargetId-1="figure@3.[225,1362,337,1587]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." captionText-1="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri-0="https://zenodo.org/record/206965/files/figure.png" httpUri-1="https://zenodo.org/record/206966/files/figure.png" pageId="1" pageNumber="40">Figs. 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6507FFFED1291DB2F835FE0FE221" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40" type="materials_examined">
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<paragraph id="8B98368CFFFED1281DB2F835FAAEE3E9" blockId="1.[151,1436,1977,2038]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
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<emphasis id="B953EA9EFFFED12B1DB2F835FE94E4EF" bold="true" box="[151,262,1977,2002]" pageId="1" pageNumber="40">Material.</emphasis>
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<collectingCountry id="F330761CFFFED12B1C28F835FE11E4EC" box="[269,387,1977,2001]" name="Venezuela" pageId="1" pageNumber="40">Venezuela</collectingCountry>
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, Estado Bolívar: 15 3 (slide 518) and 1 Ƥ imagos from Kavanayén stream,
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<geoCoordinate id="EE13504BFFFED12B19F8F836FAFBE4EC" box="[1245,1385,1977,2001]" direction="north" orientation="latitude" pageId="1" pageNumber="40" precision="15" value="5.624167">N 5° 37' 27"</geoCoordinate>
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-
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<geoCoordinate id="EE13504BFFFED12B18A7F836FE84E4C8" direction="west" orientation="longitude" pageId="1" pageNumber="40" precision="15" value="-61.74361">W 61° 44' 37"</geoCoordinate>
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,
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<quantity id="4CDF9B69FFFED12B1C07F851FEEAE4C8" box="[290,376,2013,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="1" pageNumber="40" unit="m" value="1355.0">1355 m</quantity>
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,
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<date id="FF99104CFFFED12B1CA1F851FD95E4C8" box="[388,519,2013,2037]" pageId="1" pageNumber="40" value="2005-11-21">21/XI/2005</date>
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, EG, AMO, CA and MEG cols.; 23 3 (slides 510, 516, 517) and 3 Ƥ imagos from Kako-Parú stream,
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<geoCoordinate id="EE13504BFFFDD1281C51FF14FE6AE392" box="[372,504,151,175]" direction="north" orientation="latitude" pageId="2" pageNumber="41" precision="15" value="4.9">N 4° 54' 0"</geoCoordinate>
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–
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<geoCoordinate id="EE13504BFFFDD1281F3CFF14FD20E392" box="[537,690,151,175]" direction="west" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="-61.090275">W 61° 5' 25"</geoCoordinate>
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,
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<quantity id="4CDF9B69FFFDD1281FE5FF1BFC9EE392" box="[704,780,151,175]" metricMagnitude="2" metricUnit="m" metricValue="9.129999999999999" pageId="2" pageNumber="41" unit="m" value="913.0">913 m</quantity>
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,
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<date id="FF99104CFFFDD1281E3FFF1BFC0CE392" box="[794,926,151,175]" pageId="2" pageNumber="41" value="2007-06-28">28/VI/2007</date>
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, EG, AMO, CA and MEG cols.
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<typeStatus id="549C882EFFFDD1281833FF1BFAECE392" box="[1302,1406,151,175]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
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3 imago, allotype Ƥ imago and 10
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<typeStatus id="549C882EFFFDD1281F22FF31FDF5E3E8" box="[519,615,189,213]" pageId="2" pageNumber="41" type="paratype">paratype</typeStatus>
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3 imagos at
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<collectionCode id="ED36AE49FFFDD1281FDBFF31FCC2E3E9" box="[766,848,189,212]" pageId="2" pageNumber="41">MLBV</collectionCode>
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; 5
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<typeStatus id="549C882EFFFDD1281E55FF31FC43E3E8" box="[880,977,189,213]" pageId="2" pageNumber="41" type="paratype">paratype</typeStatus>
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3 imagos at
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<collectionCode id="ED36AE49FFFDD1281942FF30FB3BE3E9" box="[1127,1193,188,212]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="41">INPA</collectionCode>
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; rest in
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<collectionCode id="ED36AE49FFFDD1281824FF31FAAAE3E9" box="[1281,1336,189,212]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="2" pageNumber="41">IML</collectionCode>
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.
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</paragraph>
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<caption id="DF586604FFFDD1281DB2FAE2FC14E69E" httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="2" pageNumber="41" targetBox="[192,1395,256,1366]" targetPageId="2">
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<paragraph id="8B98368CFFFDD1281DB2FAE2FC14E69E" blockId="2.[151,1437,1390,2036]" pageId="2" pageNumber="41">
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<emphasis id="B953EA9EFFFDD1281DB2FAE2FEA8E6B9" bold="true" box="[151,314,1390,1413]" pageId="2" pageNumber="41">FIGURES 1–4.</emphasis>
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<taxonomicName id="4C274D0FFFFDD1281C65FAE3FE6CE6B8" box="[320,510,1391,1413]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="teipunensis" status="sp. nov.">
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<emphasis id="B953EA9EFFFDD1281C65FAE3FE6CE6B8" box="[320,510,1391,1413]" italics="true" pageId="2" pageNumber="41">Caenis teipunensis</emphasis>
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</taxonomicName>
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<emphasis id="B953EA9EFFFDD1281F21FAE3FDC7E6B9" bold="true" box="[516,597,1391,1412]" pageId="2" pageNumber="41">
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<taxonomicNameLabel id="A26057E5FFFDD1281F21FAE3FDC7E6B9" box="[516,597,1391,1412]" pageId="2" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v.v. (arrows indicate apophyses of styliger sclerite); 4, general view of female.
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</paragraph>
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</caption>
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<paragraph id="8B98368CFFFDD1281DE0FA4CFC7AE4C9" blockId="2.[151,1437,1390,2036]" pageId="2" pageNumber="41">
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Male imago. Length (mm): body, 3.0–3.6; forewings, 2.5–2.9; foreleg (from base of trochanter to apex of claw), 1.8–2.3; caudal filaments, 6.5–9.0. General coloration yellowish light brown (
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<figureCitation id="131C2A09FFFDD1281968FA68FB1DE6C1" box="[1101,1167,1508,1532]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="2" pageNumber="41">Fig. 1</figureCitation>
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). Head yellowish white shaded with gray widely except on a transversal pale band between anterior part of lateral ocelli; venter of head pale. Antennae: scape yellowish gray, pedicel yellowish translucent, flagellum hyaline. Thorax. Prothorax yellowish translucent shaded with gray dorsally, darker on a pair of submedian spots; presternal triangle anteriorly pointed (
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<figureCitation id="131C2A09FFFDD1281DBAF9F8FF7AE5B1" box="[159,232,1652,1676]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="2" pageNumber="41">Fig. 2</figureCitation>
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). Mesonotum and metanotum yellowish orange shaded with gray on carinae, pleurae and sterna paler. Wings hyaline, veins translucent except C, Sc and Rs grayish. Legs. All coxae, fore femora and base of fore tibiae yellowish; rest of foreleg and middle and hind legs whitish; median and subapical gray marks present on all femora, submedian gray mark also on middle and hind tibiae. Abdomen. Segments 1–8 whitish-translucent, 9–10 yellowish, without lateral or dorsal filaments; all terga shaded widely with gray, darker on terga 1–2 and lateral spots of terga 1–9; sterna translucent white with light gray marks on lateral margins of sterna 8–9. Genitalia (
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<figureCitation id="131C2A09FFFDD1281806F8A4FAE0E47D" box="[1315,1394,1832,1856]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="2" pageNumber="41">Figs. 3</figureCitation>
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,
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<figureCitation id="131C2A09FFFDD12818A5F8A4FF34E459" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="2" pageNumber="41">5– 6</figureCitation>
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) with whitish membranes and yellowish sclerites; forceps orangeish, long and slender, apically pointed and sclerotized, generally sharpening abruptly at the apex, few setae and small tubercles present but variable (
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<figureCitation id="131C2A09FFFDD128183CF8FCFAF5E4B5" box="[1305,1383,1904,1928]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="2" pageNumber="41">Figs. 6</figureCitation>
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a–f); penes (
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<figureCitation id="131C2A09FFFDD1281DC0F818FEA1E491" box="[229,307,1940,1964]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="2" pageNumber="41">Figs. 3</figureCitation>
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,
|
||
<figureCitation id="131C2A09FFFDD1281C1AF818FEDFE491" box="[319,333,1940,1964]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="2" pageNumber="41">5</figureCitation>
|
||
) whitish with a yellowish V-shaped mark, ventral surface with small tubercles and shallow groove, lobes of the penes rounded laterally; styliger plate with elongated and oval central sclerite, apophyses strongly developed, wide and long (
|
||
<figureCitation id="131C2A09FFFDD1281CE3F850FD86E4C9" box="[454,532,2012,2036]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="2" pageNumber="41">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="131C2A09FFFDD1281F04F850FDA3E4C9" box="[545,561,2012,2036]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="2" pageNumber="41">5</figureCitation>
|
||
). Caudal filaments whitish translucent.
|
||
</paragraph>
|
||
<paragraph id="8B98368CFFFCD1291DE0FF1BFE21E3CA" blockId="3.[151,1436,151,284]" pageId="3" pageNumber="42">
|
||
Female imago (
|
||
<figureCitation id="131C2A09FFFCD1291C52FF1BFE2CE392" box="[375,446,151,175]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="3" pageNumber="42">Fig. 4</figureCitation>
|
||
). Length (mm): body, 3.0–3.5; forewings, 2.8–3.2; fore leg (from base of trochanter to apex of claw), 1.5; caudal filaments, 2.1–2.5. Color pattern as in male. Abdominal sternum 9 rounded, not projected. Eggs light yellow.
|
||
</paragraph>
|
||
<paragraph id="8B98368CFFFCD1291DE0FE88FE0FE221" blockId="3.[151,1436,151,284]" box="[197,413,260,284]" pageId="3" pageNumber="42">Nymph. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF586604FFFCD1291DB2F9C4FF72E587" httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="3" pageNumber="42" targetBox="[227,1351,343,1581]" targetPageId="3">
|
||
<paragraph id="8B98368CFFFCD1291DB2F9C4FF72E587" blockId="3.[151,1436,1608,1722]" pageId="3" pageNumber="42">
|
||
<emphasis id="B953EA9EFFFCD1291DB2F9C4FEADE563" bold="true" box="[151,319,1608,1631]" pageId="3" pageNumber="42">FIGURES 5–9.</emphasis>
|
||
<taxonomicName id="4C274D0FFFFCD1291C6CF9C6FD9EE562" box="[329,524,1610,1631]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="teipunensis" status="sp. nov.">
|
||
<emphasis id="B953EA9EFFFCD1291C6CF9C6FD9EE562" box="[329,524,1610,1631]" italics="true" pageId="3" pageNumber="42">Caenis teipunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B953EA9EFFFCD1291F32F9C5FDF9E563" bold="true" box="[535,619,1609,1630]" pageId="3" pageNumber="42">
|
||
<taxonomicNameLabel id="A26057E5FFFCD1291F32F9C5FDF9E563" box="[535,619,1609,1630]" pageId="3" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 5, male genitalia, v.v. (a=apophysis, cs=central sclerite, sc=styliger sclerite, st=small tubercles, vg=ventral groove); 6a-f, variations in forceps.
|
||
<taxonomicName id="4C274D0FFFFCD1291E67F9EBFBCAE543" box="[834,1112,1639,1662]" class="Insecta" family="Leptohyphidae" genus="Macunahyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="pemonensis" status="sp. nov.">
|
||
<emphasis id="B953EA9EFFFCD1291E67F9EBFBCAE543" box="[834,1112,1639,1662]" italics="true" pageId="3" pageNumber="42">Macunahyphes pemonensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B953EA9EFFFCD129197AF9E4FB23E540" bold="true" box="[1119,1201,1640,1661]" pageId="3" pageNumber="42">
|
||
<taxonomicNameLabel id="A26057E5FFFCD129197AF9E4FB23E540" box="[1119,1201,1640,1661]" pageId="3" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 7, male genitalia, l.v.; 8, same, v.v.
|
||
<taxonomicName id="4C274D0FFFFCD1291C04F90AFDBEE5A1" box="[289,556,1670,1692]" class="Insecta" family="Leptohyphidae" genus="Macunahyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="incognitus">
|
||
<emphasis id="B953EA9EFFFCD1291C04F90AFDBEE5A1" box="[289,556,1670,1692]" italics="true" pageId="3" pageNumber="42">Macunahyphes incognitus</emphasis>
|
||
</taxonomicName>
|
||
: 9, male genitalia, v.v. (pps= posterior projection of styliger, vs=ventral structure of penes).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D6507FFFCD1291DE0F964FF6EE47A" pageId="3" pageNumber="42" type="etymology">
|
||
<paragraph id="8B98368CFFFCD1291DE0F964FF6EE47A" blockId="3.[151,1436,1767,2008]" pageId="3" pageNumber="42">
|
||
<emphasis id="B953EA9EFFFCD1291DE0F964FED9E43D" bold="true" box="[197,331,1768,1792]" pageId="3" pageNumber="42">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C274D0FFFFCD1291F10F964FD21E5C2" box="[565,691,1768,1791]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="teipunensis">
|
||
<emphasis id="B953EA9EFFFCD1291F10F964FD21E5C2" box="[565,691,1768,1791]" italics="true" pageId="3" pageNumber="42">teipunensis</emphasis>
|
||
</taxonomicName>
|
||
is a combination of "tei-pun" (meaning “sabana”, according to the Pemon indigenous group from "La Gran Sabana", our study area) and the Latin suffix "-ensis" (denoting place or locality).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6507FFFCD12E1DE0F8DFFBCEE221" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42" type="diagnosis">
|
||
<paragraph id="8B98368CFFFCD12E1DE0F8DFFBCEE221" blockId="3.[151,1436,1767,2008]" lastBlockId="4.[151,1437,151,575]" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42">
|
||
<emphasis id="B953EA9EFFFCD1291DE0F8DFFEADE451" bold="true" box="[197,319,1875,1900]" pageId="3" pageNumber="42">Diagnosis.</emphasis>
|
||
Male imagos of
|
||
<taxonomicName id="4C274D0FFFFCD1291CDEF8D8FD5FE451" box="[507,717,1876,1900]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="teipunensis" status="sp. nov.">
|
||
<emphasis id="B953EA9EFFFCD1291CDEF8D8FD5FE451" box="[507,717,1876,1900]" italics="true" pageId="3" pageNumber="42">Caenis teipunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B953EA9EFFFCD1291FF1F8D8FCBFE451" bold="true" box="[724,813,1876,1900]" pageId="3" pageNumber="42">
|
||
<taxonomicNameLabel id="A26057E5FFFCD1291FF1F8D8FCBFE451" box="[724,813,1876,1900]" pageId="3" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from all other species in the genus by the following combination of characters: 1) length of body, 3.0–
|
||
<quantity id="4CDF9B69FFFCD1291EAFF8FBFC71E4B2" box="[906,995,1911,1935]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="42" unit="mm" value="3.6">3.6 mm</quantity>
|
||
, and of forewings,
|
||
<quantity id="4CDF9B69FFFCD129199FF8FBFAD7E4B2" box="[1210,1349,1911,1935]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="3" pageNumber="42" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
|
||
; 2) presternal triangle anteriorly pointed (
|
||
<figureCitation id="131C2A09FFFCD1291F05F817FDF4E48E" box="[544,614,1947,1971]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
|
||
); 3) fore legs 0.6–0.7 times the length of body; 4) finger-like process on abdominal tergum 2 and lateral processes on other abdominal segments absent; 5) cerci 2.1–2.5 times the length of body; 6) styliger plate with elongated and oval central sclerite, apophyses strongly developed, wide and long (
|
||
<figureCitation id="131C2A09FFFBD12E1846FF1BFF35E3E9" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="4" pageNumber="43">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="131C2A09FFFBD12E1D91FF30FF56E3E9" box="[180,196,188,212]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="4" pageNumber="43">5</figureCitation>
|
||
); 7) penes (
|
||
<figureCitation id="131C2A09FFFBD12E1C6EFF30FE09E3E9" box="[331,411,188,212]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1390,1413]" captionTargetBox="[192,1395,256,1366]" captionTargetId="figure@2.[192,1395,256,1366]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Caenis teipunensis sp. nov.: 1, general view of male; 2, male presternal triangle (arrow); 3, male genitalia, v. v. (arrows indicate apophyses of styliger sclerite); 4, general view of female." httpUri="https://zenodo.org/record/206965/files/figure.png" pageId="4" pageNumber="43">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="131C2A09FFFBD12E1C8CFF30FE25E3E9" box="[425,439,188,212]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="4" pageNumber="43">5</figureCitation>
|
||
) completely fused, laterally constricted, lateral lobes rounded, small tubercles and shallow grooves present on ventral surface; 8) forceps sclerotized, sharp, generally with small tubercles and setae on outer basal margin (
|
||
<figureCitation id="131C2A09FFFBD12E1C53FE88FE51E221" box="[374,451,260,284]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1608,1631]" captionTargetBox="[227,1351,343,1581]" captionTargetId="figure@3.[225,1362,337,1587]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Caenis teipunensis sp. nov.: 5, male genitalia, v. v. (a = apophysis, cs = central sclerite, sc = styliger sclerite, st = small tubercles, vg = ventral groove); 6 a-f, variations in forceps. Macunahyphes pemonensis sp. nov.; 7, male genitalia, l. v.; 8, same, v. v. Macunahyphes incognitus: 9, male genitalia, v. v. (pps = posterior projection of styliger, vs = ventral structure of penes)." httpUri="https://zenodo.org/record/206966/files/figure.png" pageId="4" pageNumber="43">Figs. 6</figureCitation>
|
||
a–f), forceps 7.3–9.5 times longer than width at the middle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6507FFFBD12E1DE0FEABFB52E102" pageId="4" pageNumber="43" type="discussion">
|
||
<paragraph id="8B98368CFFFBD12E1DE0FEABFDCAE2BA" blockId="4.[151,1437,151,575]" pageId="4" pageNumber="43">
|
||
<emphasis id="B953EA9EFFFBD12E1DE0FEABFED8E27D" bold="true" box="[197,330,295,320]" pageId="4" pageNumber="43">Discussion.</emphasis>
|
||
Except for the extremely long apophyses of the styliger sclerite and shape of the forceps,
|
||
<taxonomicName id="4C274D0FFFFBD12E186BFEA4FE87E259" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="teipunensis" status="sp. nov.">
|
||
<emphasis id="B953EA9EFFFBD12E186BFEA4FE87E259" italics="true" pageId="4" pageNumber="43">Caenis teipunensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B953EA9EFFFBD12E1C3EFEC0FEE6E259" bold="true" box="[283,372,332,356]" pageId="4" pageNumber="43">
|
||
<taxonomicNameLabel id="A26057E5FFFBD12E1C3EFEC0FEE6E259" box="[283,372,332,356]" pageId="4" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C274D0FFFFBD12E1F21FEC1FD60E259" authority="Malzacher" authorityName="Malzacher" box="[516,754,332,356]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="fittkaui">
|
||
<emphasis id="B953EA9EFFFBD12E1F21FEC1FDE1E259" box="[516,627,332,356]" italics="true" pageId="4" pageNumber="43">C. fittkaui</emphasis>
|
||
Malzacher
|
||
</taxonomicName>
|
||
, and belongs to the same group of species defined by the apically pointed forceps (
|
||
<bibRefCitation id="EFB64B7DFFFBD12E1CB5FEE3FDD9E2BA" author="Malzacher" box="[400,587,367,391]" pageId="4" pageNumber="49" refString="Malzacher, P. (2001) South and Central American Caenis species with rounded forceps tips (Insecta: Ephemeroptera: Caenidae). Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 626, 1 - 20." type="journal article" year="2001">Malzacher 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B98368CFFFBD12E1DE0FE18FB52E102" blockId="4.[151,1437,151,575]" pageId="4" pageNumber="43">
|
||
<emphasis id="B953EA9EFFFBD12E1DE0FE18FEBBE291" bold="true" box="[197,297,404,428]" pageId="4" pageNumber="43">Ecology.</emphasis>
|
||
Imagos were collected along the stream margins, at night, and during the rainy season (
|
||
<date id="FF99104CFFFBD12E1801FE18FF5DE2F2" pageId="4" pageNumber="43" value="2005-11">November 2005</date>
|
||
and
|
||
<date id="FF99104CFFFBD12E1C2CFE34FE13E2F2" box="[265,385,439,463]" pageId="4" pageNumber="43" value="2007-06">June 2007</date>
|
||
), by using light traps. Streams flow through an upland undulating savanna ecosystem (
|
||
<quantity id="4CDF9B69FFFBD12E1846FE3BFF7FE2C9" metricMagnitude="3" metricUnit="m" metricValue="1.134" metricValueMax="1.355" metricValueMin="0.9129999999999999" pageId="4" pageNumber="43" unit="m" value="1134.0" valueMax="1355.0" valueMin="913.0">913– 1355 m</quantity>
|
||
) and they were partially shaded (Kaku-Parú,
|
||
<figureCitation id="131C2A09FFFBD12E1FCEFE50FCA9E2C9" box="[747,827,476,500]" captionStart="FIGURES 23 – 26" captionStartId="9.[151,264,1742,1765]" captionTargetBox="[209,1371,181,1723]" captionTargetId="figure@9.[209,1372,181,1723]" captionTargetPageId="9" captionText="FIGURES 23 – 26. Study sites: 23, Kaku Parú; 24, Kavanayén; 25, Mareman Parú; 26, Aponwao." httpUri="https://zenodo.org/record/206969/files/figure.png" pageId="4" pageNumber="43">Fig. 23</figureCitation>
|
||
) or not shaded at all (Kavanayén,
|
||
<figureCitation id="131C2A09FFFBD12E199BFE50FA83E2C9" box="[1214,1297,476,500]" captionStart="FIGURES 23 – 26" captionStartId="9.[151,264,1742,1765]" captionTargetBox="[209,1371,181,1723]" captionTargetId="figure@9.[209,1372,181,1723]" captionTargetPageId="9" captionText="FIGURES 23 – 26. Study sites: 23, Kaku Parú; 24, Kavanayén; 25, Mareman Parú; 26, Aponwao." httpUri="https://zenodo.org/record/206969/files/figure.png" pageId="4" pageNumber="43">Fig. 24</figureCitation>
|
||
). They were small to medium size streams (
|
||
<quantity id="4CDF9B69FFFBD12E1CD0FD8EFDC9E127" box="[501,603,514,538]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="4" pageNumber="43" unit="m" value="19.0" valueMax="20.0" valueMin="18.0">18–20 m</quantity>
|
||
wide), of low order (1st–2nd), with shallow (
|
||
<quantity id="4CDF9B69FFFBD12E196FFD8EFB3DE127" box="[1098,1199,514,538]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.2" metricValueMin="0.8" pageId="4" pageNumber="43" unit="cm" value="20.0" valueMax="32.0" valueMin="8.0">8–32 cm</quantity>
|
||
) and acidic (pH = 5) waters, and with bottoms mainly composed of bed-rock (Kavanayén) and gravels (Kaku-Parú).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |