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<mods:title id="9AE4CCA72A1AE143784FABCAE1B42674">Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)</mods:title>
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<paragraph id="8B98369F771D7F3181AFA3F7FD84FA01" blockId="50.[151,655,1483,1546]" box="[151,539,1483,1510]" pageId="50" pageNumber="51">
<heading id="D0D081F3771D7F3181AFA3F7FD84FA01" bold="true" box="[151,539,1483,1510]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="4">
<taxonomicName id="4C274D1C771D7F3181AFA3F7FD84FA01" ID-CoL="LC5W" authority="Sharp, 1885" authorityName="Sharp" authorityYear="1885" box="[151,539,1483,1510]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="bidens">
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<emphasis id="B953EA8D771D7F3181AFA3F7FEE1FA02" bold="true" box="[151,382,1483,1510]" italics="true" pageId="50" pageNumber="51">Belonuchus bidens</emphasis>
<bibRefCitation id="EFB64B6E771D7F3180BDA3F7FD84FA01" author="Sharp, D." box="[389,539,1483,1509]" pageId="50" pageNumber="51" pagination="393 - 536" refId="ref114488" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor &amp; Francis, London, pp. 393 - 536, pls. 10 - 13." type="book chapter" year="1885">Sharp, 1885</bibRefCitation>
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<subSubSection id="C33D6514771D7F3181AFA3D3FD10F9ED" box="[151,655,1519,1546]" pageId="50" pageNumber="51" type="description">
<paragraph id="8B98369F771D7F3181AFA3D3FD10F9ED" blockId="50.[151,655,1483,1546]" box="[151,655,1519,1546]" pageId="50" pageNumber="51">
<figureCitation id="131C2A1A771D7F3181AFA3D3FE95F9ED" box="[151,266,1519,1545]" captionStart="FIGURE 4" captionStartId="98.[151,250,1720,1744]" captionTargetBox="[170,1410,196,1684]" captionTargetId="figure-25@98.[161,1426,181,1696]" captionTargetPageId="98" captionText="FIGURE 4. Dorsal view of males of: a) Belonuchus tenuistylus (holotype), b) B. tepoztecus (paratype), c) B. trochanterinus, d) B. zunilensis (lectotype), e) B. bidens (syntype, NHM©), f) B. bidens, g) B. erichsoni, h) B. jalappensis. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630889" httpUri="https://zenodo.org/record/6630889/files/figure.png" pageId="50" pageNumber="51">Figs. 4e, f</figureCitation>
, 
<figureCitation id="131C2A1A771D7F31802DA3D3FEA0F9ED" box="[277,319,1519,1545]" captionStart="FIGURE 10" captionStartId="104.[151,250,1839,1863]" captionTargetBox="[176,1418,194,1806]" captionTargetId="figure-25@104.[152,1435,181,1816]" captionTargetPageId="104" captionText="FIGURE 10. Dorsal view of the head of males of: a) Belonuchus julietitae, b) B. singularis, c) B. connexus, d) B. platypterus, e) B. bidens, f) B. nigerrimus, g) B. optatus, h) B. cyanipennis, i) B. godmani. Dorsal view of the head and pronotum of the holotype female of: j) B. optatus." figureDoi="http://doi.org/10.5281/zenodo.6630909" httpUri="https://zenodo.org/record/6630909/files/figure.png" pageId="50" pageNumber="51">10e</figureCitation>
, 
<figureCitation id="131C2A1A771D7F318074A3D3FEEAF9ED" box="[332,373,1519,1545]" captionStart="FIGURE 11" captionStartId="105.[151,250,1833,1857]" captionTargetBox="[175,1420,203,1797]" captionTargetId="figure-19@105.[151,1436,181,1809]" captionTargetPageId="105" captionText="FIGURE 11. Mesoventrite of males of: a) Belonuchus cariniventris, b) B. oxyporinus, c) B. rufisternus, d) B. tepoztecus, e) B. bidens, f) B. jalappensis, g) B. longitarsus, h) B. ephippiatus, i) B. dichrous, j) B. flavipennis, k) B. platypterus, l) B. juanmorronei, m) B. iridescens. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge, green arrow: longitudinal carina in anterior half." figureDoi="http://doi.org/10.5281/zenodo.6630913" httpUri="https://zenodo.org/record/6630913/files/figure.png" pageId="50" pageNumber="51">11e</figureCitation>
, 
<figureCitation id="131C2A1A771D7F3180B9A3D3FE31F9ED" box="[385,430,1519,1545]" captionStart="FIGURE 12" captionStartId="106.[151,250,1870,1894]" captionTargetBox="[191,1398,191,1835]" captionTargetId="figure-25@106.[165,1422,181,1846]" captionTargetPageId="106" captionText="FIGURE 12. Mesoventrite of males of: a) Belonuchus viridipennis, b) Belonuchus sp. Q, c) B. cyanipennis. Ventral view of the left femur of male anterior leg of: d) B. apiciventris, e) B. oxyporimimus, f) B. oxyporinus, g) B. tepoztecus, h) B. bidens, i) B. viridipennis. Ventral view of the left posterior leg of males of: j) B. julietitae, k) B. linuzensis (small specimen), l) B. linuzensis (large specimen), m) B. metafemoralis, n) B. oxyporimimus, o) B. oxyporinus. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge." figureDoi="http://doi.org/10.5281/zenodo.6630915" httpUri="https://zenodo.org/record/6630915/files/figure.png" pageId="50" pageNumber="51">12h</figureCitation>
, 
<figureCitation id="131C2A1A771D7F318081A3D3FE72F9EE" box="[441,493,1519,1546]" captionStart="FIGURE 15" captionStartId="109.[151,250,1821,1845]" captionTargetBox="[175,1417,182,1784]" captionTargetId="figure-19@109.[160,1427,181,1797]" captionTargetPageId="109" captionText="FIGURE 15. Male pregenital sternite of: a) Belonuchus alternans, b) B. oxyporinus, c) B. tepoztecus, d) B. jalappensis, e) B. iridescens, f) B. platypterus, g) B. cyanipennis, h) B. apiciventris, i) B. linuzensis, j) B. metafemoralis, k) B. rufisternus, l) B. tenuistylus, m) B. bidens, n) B. pollens, o) B. longitarsus." figureDoi="http://doi.org/10.5281/zenodo.6630921" httpUri="https://zenodo.org/record/6630921/files/figure.png" pageId="50" pageNumber="51">
<quantity id="4CDF9B7A771D7F318081A3D3FE72F9EE" box="[441,493,1519,1546]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="50" pageNumber="51" unit="m" value="15.0">15m</quantity>
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, 
<figureCitation id="131C2A1A771D7F3180C1A3D3FDBAF9ED" box="[505,549,1519,1545]" captionStart="FIGURE 19" captionStartId="113.[151,250,1823,1847]" captionTargetBox="[165,1426,181,1793]" captionTargetId="figure-19@113.[161,1426,181,1800]" captionTargetPageId="113" captionText="FIGURE 19. Male genital sternite of: a) Belonuchus tepoztecus, b) B. trochanterinus, c) B. zunilensis, d) B. bidens, e) B. erichsoni, f) B. jalappensis, g) B. mixtecus, h) B. pollens, i) B. rufipennis, j) B. basiventris, k) B. connexus, l) B. longitarsus, m) B. rufiventris, n) B. xanthomelas, o) B. ephippiatus, p) B. moroni. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630927" httpUri="https://zenodo.org/record/6630927/files/figure.png" pageId="50" pageNumber="51">19d</figureCitation>
, 
<figureCitation id="131C2A1A771D7F318309A3D3FDC4F9ED" box="[561,603,1519,1545]" captionStart="FIGURE 23" captionStartId="117.[151,250,1878,1902]" captionTargetBox="[158,1426,187,1844]" captionTargetId="figure-19@117.[151,1436,181,1854]" captionTargetPageId="117" captionText="FIGURE 23. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus bidens, b) B. erichsoni, c) B. jalappensis, d) B. mixtecus, e) B. pollens, f) B. rufipennis, g) B. basiventris, h) B. connexus (at center, detail of the apex of the median lobe), i) B. longitarsus (at center, detail of the apex of the median lobe). Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630939" httpUri="https://zenodo.org/record/6630939/files/figure.png" pageId="50" pageNumber="51">23a</figureCitation>
, 
<figureCitation id="131C2A1A771D7F31835FA3D3FD10F9ED" box="[615,655,1519,1545]" captionStart="FIGURE 30" captionStartId="124.[151,250,1685,1709]" captionTargetBox="[160,1424,191,1648]" captionTargetId="figure-25@124.[151,1436,181,1661]" captionTargetPageId="124" captionText="FIGURE 30. Geographical distribution in Mexico and Guatemala of: a) Belonuchus abnormalis, B. bidens and B. platypterus; b) B. basiventris and B. jalappensis." figureDoi="http://doi.org/10.5281/zenodo.6630953" httpUri="https://zenodo.org/record/6630953/files/figure.png" pageId="50" pageNumber="51">30a</figureCitation>
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<paragraph id="8B98369F771D7F3181AFA008FD8EF9A8" blockId="50.[151,529,1587,1612]" box="[151,529,1587,1612]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB3711B1771D7F3181AFA008FD8EF9A8" box="[151,529,1587,1612]" pageId="50" pageNumber="51">
<treatmentCitation id="0A86108E771D7F3181AFA008FD92F9A8" author="Sharp, D." box="[151,525,1587,1612]" page="430" pageId="50" pageNumber="51" year="1885">
<taxonomicName id="4C274D1C771D7F3181AFA008FD92F9A8" ID-CoL="LC5W" authority="Sharp, 1885: 430" authorityName="Sharp" authorityPageNumber="430" authorityYear="1885" box="[151,525,1587,1612]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B953EA8D771D7F3181AFA008FEC9F9A8" box="[151,342,1588,1612]" italics="true" pageId="50" pageNumber="51">Belonuchus bidens</emphasis>
<bibRefCitation id="EFB64B6E771D7F318064A008FD92F9A8" author="Sharp, D." box="[348,525,1587,1612]" pageId="50" pageNumber="51" pagination="393 - 536" refId="ref114488" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor &amp; Francis, London, pp. 393 - 536, pls. 10 - 13." type="book chapter" year="1885">Sharp, 1885: 430</bibRefCitation>
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.
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<paragraph id="8B98369F771D7F3181AFA046FEAAF8AC" blockId="50.[151,1437,1658,2008]" pageId="50" pageNumber="51">
Total body length in males 
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(range 9.5–13.0 mm), in females 
<quantity id="4CDF9B7A771D7F3182A6A046FB9FF970" box="[926,1024,1658,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="50" pageNumber="51" unit="mm" value="11.3">11.3 mm</quantity>
(range 
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). Black on head (except mouthparts), antennomeres 4–10 or 4–11, pronotum, visible abdominal segments 1–4, and anterior half (or slightly more) of fifth. Reddish on mouth parts, antennomeres 1–3 and sometimes the apical segment, elytra, scutellum, sterna, legs, from basal half (or basal third) of fifth visible abdominal segment to genital segment. Prosternum with variation from black to reddish. Apical antennomere reddish, paler than anterior antennomeres and with different texture.
</paragraph>
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<paragraph id="8B98369F771D7F3081FFA16EFCDCFE6D" blockId="50.[151,1437,1658,2008]" lastBlockId="51.[151,1437,150,2013]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="B953EA8D771D7F3181FFA16EFE98F888" bold="true" box="[199,263,1874,1900]" pageId="50" pageNumber="51">Head</emphasis>
: transverse, ratio length/width in males 0.71 (range 0.50–0.82), in females 0.79 (range 0.77–0.82); with posterior corners and postero-dorsal part somewhat convex. Dorsal surface with sulcate longitudinal midline visible in anterior half of head; front foveate near anterior margin, in males this foveate area is placed between the two rather short cephalic projections at internal side of each antennal insertion (
<figureCitation id="131C2A1A771D7F318529A182FBEDF83C" box="[1041,1138,1982,2008]" captionStart="FIGURE 10" captionStartId="104.[151,250,1839,1863]" captionTargetBox="[176,1418,194,1806]" captionTargetId="figure-25@104.[152,1435,181,1816]" captionTargetPageId="104" captionText="FIGURE 10. Dorsal view of the head of males of: a) Belonuchus julietitae, b) B. singularis, c) B. connexus, d) B. platypterus, e) B. bidens, f) B. nigerrimus, g) B. optatus, h) B. cyanipennis, i) B. godmani. Dorsal view of the head and pronotum of the holotype female of: j) B. optatus." figureDoi="http://doi.org/10.5281/zenodo.6630909" httpUri="https://zenodo.org/record/6630909/files/figure.png" pageId="50" pageNumber="51">Fig. 10e</figureCitation>
). Eyes large, slightly more than 0.5 times the cephalic lateral length, not protruding laterally. Antennomere 4 elongate, 5 as long as wide, 6–10 transverse. Mandibles longer than head in males (ratio 1.26, range 1.10–1.48) and shorter than head in females (ratio 0.90, range 0.84–0.97); each with two small, moderately separated teeth (basal and middle); mandibular channel slightly developed, with external margin slightly separated from internal margin, internal margin like impressed line, markedly extending anteriad of middle tooth. Apical palpomere of maxillary and labial palpi 1.25 to 1.48 times longer than preapical palpomere. Head slightly wider than pronotum in males (ratio 1.14, range 1.06–1.18), and almost equally wide in females (ratio 1.03, range 0.95–1.07).
</paragraph>
<paragraph id="8B98369F771C7F3081FFA7AFFBECFCB9" blockId="51.[151,1437,150,2013]" pageId="51" pageNumber="52">
<emphasis id="B953EA8D771C7F3081FFA7AFFEBFFE49" bold="true" box="[199,288,403,429]" pageId="51" pageNumber="52">Thorax</emphasis>
: each dorsal row of pronotum with five punctures; pronotum almost as long as wide (ratio in males 1.04, range 1.00–1.14, in females 1.07, range 1.00–1.11) and almost as wide at anterior corners as at posterior corners (ratio in males 1.08, range 1.06–1.12, in females 1.03, range 1.03–1.05). Scutellum and elytra with fine punctures and setae moderately dense. Prosternum with area near to anterior margin slightly elevated and delimitated backward by a faintly impressed transverse line. Intercoxal process of mesoventrite shield-shaped, rather broad and with obtuse-angled tip; transverse discal ridge almost parallel to margin of intercoxal process, not meeting margin of process laterally (
<figureCitation id="131C2A1A771C7F3080B8A457FE7FFD61" box="[384,480,619,645]" captionStart="FIGURE 11" captionStartId="105.[151,250,1833,1857]" captionTargetBox="[175,1420,203,1797]" captionTargetId="figure-19@105.[151,1436,181,1809]" captionTargetPageId="105" captionText="FIGURE 11. Mesoventrite of males of: a) Belonuchus cariniventris, b) B. oxyporinus, c) B. rufisternus, d) B. tepoztecus, e) B. bidens, f) B. jalappensis, g) B. longitarsus, h) B. ephippiatus, i) B. dichrous, j) B. flavipennis, k) B. platypterus, l) B. juanmorronei, m) B. iridescens. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge, green arrow: longitudinal carina in anterior half." figureDoi="http://doi.org/10.5281/zenodo.6630913" httpUri="https://zenodo.org/record/6630913/files/figure.png" pageId="51" pageNumber="52">Fig. 11e</figureCitation>
). Profemur of males with external and internal margins well differentiated and each with row of spines; external margin curved downward near apex, distinctly narrowed and forming kind of notch where protibia would fit; external row of spines initiating slightly before base and reaching initial part of curvature, with fewer spines than on internal margin; spines on internal margin initiating from base, where there are additional non-aligned spines, but row of spines continuing until apex (
<figureCitation id="131C2A1A771C7F308207A4C7FC3EFCF2" box="[831,929,763,790]" captionStart="FIGURE 12" captionStartId="106.[151,250,1870,1894]" captionTargetBox="[191,1398,191,1835]" captionTargetId="figure-25@106.[165,1422,181,1846]" captionTargetPageId="106" captionText="FIGURE 12. Mesoventrite of males of: a) Belonuchus viridipennis, b) Belonuchus sp. Q, c) B. cyanipennis. Ventral view of the left femur of male anterior leg of: d) B. apiciventris, e) B. oxyporimimus, f) B. oxyporinus, g) B. tepoztecus, h) B. bidens, i) B. viridipennis. Ventral view of the left posterior leg of males of: j) B. julietitae, k) B. linuzensis (small specimen), l) B. linuzensis (large specimen), m) B. metafemoralis, n) B. oxyporimimus, o) B. oxyporinus. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge." figureDoi="http://doi.org/10.5281/zenodo.6630915" httpUri="https://zenodo.org/record/6630915/files/figure.png" pageId="51" pageNumber="52">Fig. 12h</figureCitation>
). Metafemur of males with two rows of large and dark spines on ventral surface, internal row well aligned from base until apical third, external row not clearly aligned because of additional, non-aligned spines at base. Tarsomeres dorsally flattened.
</paragraph>
<paragraph id="8B98369F771C7F3081FFA55AFC0DFB45" blockId="51.[151,1437,150,2013]" pageId="51" pageNumber="52">
<emphasis id="B953EA8D771C7F3081FFA55AFEA3FC64" bold="true" box="[199,316,870,896]" pageId="51" pageNumber="52">Abdomen</emphasis>
: first three visible tergites with posterior basal transverse carina well developed; adjacent area slightly depressed and narrow, with moderately dense wide-superficial punctures that are decreasing toward posterior margin, combined with fine, moderately dense punctures; fourth visible tergite with punctures similar to that of previous tergites, visible tergites 5–6 only with fine, moderately dense punctures. Visible sternites 1–4 with similar punctures as on tergites 1–4, but with denser wide-superficial punctures; sternites 5–6 only with fine punctures. Male pregenital sternite slightly emarginate at posterior margin, with long setae at sides and shorter, finer setae at center (
<figureCitation id="131C2A1A771C7F3081D2A203FECBFBBD" box="[234,340,1086,1113]" captionStart="FIGURE 15" captionStartId="109.[151,250,1821,1845]" captionTargetBox="[175,1417,182,1784]" captionTargetId="figure-19@109.[160,1427,181,1797]" captionTargetPageId="109" captionText="FIGURE 15. Male pregenital sternite of: a) Belonuchus alternans, b) B. oxyporinus, c) B. tepoztecus, d) B. jalappensis, e) B. iridescens, f) B. platypterus, g) B. cyanipennis, h) B. apiciventris, i) B. linuzensis, j) B. metafemoralis, k) B. rufisternus, l) B. tenuistylus, m) B. bidens, n) B. pollens, o) B. longitarsus." figureDoi="http://doi.org/10.5281/zenodo.6630921" httpUri="https://zenodo.org/record/6630921/files/figure.png" pageId="51" pageNumber="52">
Fig. 
<quantity id="4CDF9B7A771C7F308026A202FECBFBBD" box="[286,340,1086,1113]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="m" value="15.0">15m</quantity>
</figureCitation>
). Male genital sternite short (2.26 times longer than wide), very asymmetrical, anterior portion occupying 29% and posterior portion 71% of its length, apical emargination deep (
<figureCitation id="131C2A1A771C7F308528A25FFBEEFB9A" box="[1040,1137,1123,1150]" captionStart="FIGURE 19" captionStartId="113.[151,250,1823,1847]" captionTargetBox="[165,1426,181,1793]" captionTargetId="figure-19@113.[161,1426,181,1800]" captionTargetPageId="113" captionText="FIGURE 19. Male genital sternite of: a) Belonuchus tepoztecus, b) B. trochanterinus, c) B. zunilensis, d) B. bidens, e) B. erichsoni, f) B. jalappensis, g) B. mixtecus, h) B. pollens, i) B. rufipennis, j) B. basiventris, k) B. connexus, l) B. longitarsus, m) B. rufiventris, n) B. xanthomelas, o) B. ephippiatus, p) B. moroni. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630927" httpUri="https://zenodo.org/record/6630927/files/figure.png" pageId="51" pageNumber="52">Fig. 19d</figureCitation>
). Abdominal styli pale, but with apical third darker, slender, with lateral margin slightly curved.
</paragraph>
<paragraph id="8B98369F771C7F3081FFA296FE92FAE9" blockId="51.[151,1437,150,2013]" pageId="51" pageNumber="52">
<emphasis id="B953EA8D771C7F3081FFA296FEA5FB20" bold="true" box="[199,314,1194,1220]" pageId="51" pageNumber="52">Aedeagus</emphasis>
: length 
<quantity id="4CDF9B7A771C7F3080AFA296FE61FB21" box="[407,510,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="51" pageNumber="52" unit="mm" value="1.65">1.65 mm</quantity>
; rounded shape in basal half and elongate in apical half; apex wide at sides, with a minute median projection; basal half notably wider than apical half; internal sac visible, with characteristic shape (
<figureCitation id="131C2A1A771C7F308198A2CFFE9FFAE9" box="[160,256,1267,1293]" captionStart="FIGURE 23" captionStartId="117.[151,250,1878,1902]" captionTargetBox="[158,1426,187,1844]" captionTargetId="figure-19@117.[151,1436,181,1854]" captionTargetPageId="117" captionText="FIGURE 23. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus bidens, b) B. erichsoni, c) B. jalappensis, d) B. mixtecus, e) B. pollens, f) B. rufipennis, g) B. basiventris, h) B. connexus (at center, detail of the apex of the median lobe), i) B. longitarsus (at center, detail of the apex of the median lobe). Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630939" httpUri="https://zenodo.org/record/6630939/files/figure.png" pageId="51" pageNumber="52">Fig. 23a</figureCitation>
).
</paragraph>
<paragraph id="8B98369F771C7F3081FFA32AFDE7FA79" blockId="51.[151,1437,150,2013]" pageId="51" pageNumber="52">
<emphasis id="B953EA8D771C7F3081FFA32AFED8FAD4" bold="true" box="[199,327,1302,1328]" pageId="51" pageNumber="52">Variability</emphasis>
. In addition to the variation in the measurements and ratios indicated in the description, it was observed, in both males and small females, that antennomeres 5–10 may be transverse instead of as long as wide. In some specimens the punctures of scutellum are denser than on elytra. Females do not have the apex of the anterior tibiae widened and curved to form a notch.
</paragraph>
<paragraph id="8B98369F771C7F3081FFA39AFD85F97D" blockId="51.[151,1437,150,2013]" pageId="51" pageNumber="52">
<emphasis id="B953EA8D771C7F3081FFA39AFE54FA25" bold="true" box="[199,459,1446,1473]" pageId="51" pageNumber="52">Taxonomic comments</emphasis>
. 
<taxonomicName id="4C274D1C771C7F3080E1A39BFD33FA25" authorityName="Sharp" authorityYear="1885" box="[473,684,1447,1473]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B953EA8D771C7F3080E1A39BFD33FA25" box="[473,684,1447,1473]" italics="true" pageId="51" pageNumber="52">Belonuchus bidens</emphasis>
</taxonomicName>
is differs from the rest of species of 
<taxonomicName id="4C274D1C771C7F308572A39BFB23FA25" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1098,1212,1447,1473]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="B953EA8D771C7F308572A39BFB23FA25" box="[1098,1212,1447,1473]" italics="true" pageId="51" pageNumber="52">rufipennis</emphasis>
</taxonomicName>
group by its unique coloration pattern, in which the elytra, scutellum, legs and last two visible abdominal segments (fifth only the posterior half) are reddish and the rest of the body is black; additionally, the males have two projections in the form of short horns on the frons, next to the antennal insertions. The aedeagus and genital sternite are also rather distinctive in this species. The 
<typeStatus id="549C883D771C7F308049A00BFE4CF9B5" box="[369,467,1591,1617]" pageId="51" pageNumber="52" type="syntype">syntypes</typeStatus>
of this species were not studied, but we analyzed some photos of these specimens (
<figureCitation id="131C2A1A771C7F308457A00BFF2AF991" captionStart="FIGURE 4" captionStartId="98.[151,250,1720,1744]" captionTargetBox="[170,1410,196,1684]" captionTargetId="figure-25@98.[161,1426,181,1696]" captionTargetPageId="98" captionText="FIGURE 4. Dorsal view of males of: a) Belonuchus tenuistylus (holotype), b) B. tepoztecus (paratype), c) B. trochanterinus, d) B. zunilensis (lectotype), e) B. bidens (syntype, NHM©), f) B. bidens, g) B. erichsoni, h) B. jalappensis. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630889" httpUri="https://zenodo.org/record/6630889/files/figure.png" pageId="51" pageNumber="52">Fig. 4e</figureCitation>
; NHM, shared by M. Chani Posse), which together with the information of the original description (
<bibRefCitation id="EFB64B6E771C7F308429A067FA0BF991" author="Sharp, D." box="[1297,1428,1626,1653]" pageId="51" pageNumber="52" pagination="393 - 536" refId="ref114488" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor &amp; Francis, London, pp. 393 - 536, pls. 10 - 13." type="book chapter" year="1885">Sharp 1885</bibRefCitation>
) allowed us to identify this species.
</paragraph>
</subSubSection>
<subSubSection id="C33D6514771C7F3781FFA09EFAC7FC41" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8B98369F771C7F3781FFA09EFAC7FC41" blockId="51.[151,1437,150,2013]" lastBlockId="52.[151,1437,151,933]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis id="B953EA8D771C7F3081FFA09EFD5EF958" bold="true" box="[199,705,1698,1725]" pageId="51" pageNumber="52">
Material examined (
<specimenCount id="9D21FD16771C7F308081A09EFDBBF959" box="[441,548,1698,1725]" pageId="51" pageNumber="52" type="male">34 males</specimenCount>
, 
<specimenCount id="9D21FD16771C7F308308A09EFD2BF958" box="[560,692,1698,1724]" pageId="51" pageNumber="52" type="female">21 females</specimenCount>
):
</emphasis>
<materialsCitation id="3B4F3CC2771C7F3083F0A09EFEF8F905" country="Mexico" location="Hidalgo" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Hidalgo">
<emphasis id="B953EA8D771C7F3083F0A09EFC07F958" bold="true" box="[712,920,1698,1724]" pageId="51" pageNumber="52">
<collectingCountry id="F330760F771C7F3083F0A09EFCBCF958" box="[712,803,1698,1724]" name="Mexico" pageId="51" pageNumber="52">Mexico</collectingCountry>
: 
<collectingRegion id="49E3F87D771C7F308208A09EFC0CF958" box="[816,915,1698,1724]" country="Mexico" name="Hidalgo" pageId="51" pageNumber="52">Hidalgo</collectingRegion>
:
</emphasis>
“Chapulhuacán Arroyo Blanco, bosque tropical subcaducifolio
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771C7F308049A0FBFE0CF8CD" collectingDate="2009-03-30" collectingDateMax="2009-04-06" collectingDateMin="2009-03-30" collectionCode="CC-UAEH" collectorName="J. Marquez &amp; M. Rivero &amp; M. Torres" country="Mexico" elevation="239" latitude="21.154766" location="Huautla" longLatPrecision="1" longitude="-99.005516" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Hidalgo">
<quantity id="4CDF9B7A771C7F308049A0FBFE25F905" box="[369,442,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.3899999999999997" pageId="51" pageNumber="52" unit="m" value="239.0">
<elevation id="000AD1AC771C7F308049A0FBFE25F905" box="[369,442,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.3899999999999997" pageId="51" pageNumber="52" unit="m" value="239.0">239 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058771C7F3080FEA0FBFDC5F905" box="[454,602,1735,1761]" degrees="21" direction="north" minutes="09.286" orientation="latitude" pageId="51" pageNumber="52" precision="1" value="21.154766">N21°09.286’</geoCoordinate>
, 
<geoCoordinate id="EE135058771C7F30835DA0FBFD60F905" box="[613,767,1735,1761]" degrees="99" direction="west" minutes="00.331" orientation="longitude" pageId="51" pageNumber="52" precision="1" value="-99.005516">W99°00.331’</geoCoordinate>
, NTP-80 #2 (calamar), 
<date id="FF99105F771C7F308530A0FBFB78F905" box="[1032,1255,1735,1761]" pageId="51" pageNumber="52" value="2009-03-30" valueMax="2009-04-06" valueMin="2009-03-30">
<collectingDate id="EFDDE9B7771C7F308530A0FBFB78F905" box="[1032,1255,1735,1761]" pageId="51" pageNumber="52" value="2009-03-30" valueMax="2009-04-06" valueMin="2009-03-30">30-III a 06-IV-2009</collectingDate>
</date>
, 
<collectorName id="26D25349771C7F3085CAA0FBFAEEF905" box="[1266,1393,1735,1761]" pageId="51" pageNumber="52">J. Márquez</collectorName>
, 
<collectorName id="26D25349771C7F308444A0FBFF79F8E1" pageId="51" pageNumber="52">M. Rivero</collectorName>
, 
<collectorName id="26D25349771C7F3081CBA0D7FEFAF8E1" box="[243,357,1771,1797]" pageId="51" pageNumber="52">M. Torres</collectorName>
, M. Vargas y J. Sánchez cols.” (1, 
<collectionCode id="ED36AE5A771C7F3083CEA0D7FCE7F8E1" box="[758,888,1771,1797]" pageId="51" pageNumber="52">CC-UAEH</collectionCode>
). Same data, except: “NTP-80 #3 (calamar)” (3, 
<collectionCode id="ED36AE5A771C7F3081AFA133FE86F8CD" box="[151,281,1807,1833]" pageId="51" pageNumber="52">CC-UAEH</collectionCode>
). “ 
<location id="8EF86044771C7F30800DA133FE0CF8CD" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771C7F30800DA133FE0CF8CD" box="[309,403,1807,1833]" country="Mexico" latitude="21.154766" longLatPrecision="1" longitude="-99.005516" name="Huautla" pageId="51" pageNumber="52" stateProvince="Hidalgo">Huautla</location>
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771C7F3080A6A133FE85F895" collectingDate="2015-04-17" collectingDateMax="2015-04-28" collectingDateMin="2015-04-17" collectionCode="CC-UAEH" collectorName="R. Ramirez &amp; J. Marquez" country="Mexico" latitude="21.123056" location="Rio Blanco" longLatPrecision="21" longitude="-98.28084" municipality="Huehuetla" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Hidalgo">
<locationDeviation id="9A2A2FBD771C7F3080A6A133FC86F8CD" box="[414,793,1807,1833]" location="Tohuaco Amatzintla" pageId="51" pageNumber="52">
<quantity id="4CDF9B7A771C7F3080A6A133FE48F8CD" box="[414,471,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="51" pageNumber="52" unit="km" value="2.0">2 km</quantity>
NW de Tohuaco Amatzintla
</locationDeviation>
, 
<geoCoordinate id="EE135058771C7F30821CA133FC2FF8CD" box="[804,944,1807,1833]" degrees="21" direction="north" minutes="07" orientation="latitude" pageId="51" pageNumber="52" precision="15" seconds="23" value="21.123056">N21°07´23”</geoCoordinate>
, 
<geoCoordinate id="EE135058771C7F308283A132FBD2F8CD" box="[955,1101,1806,1833]" degrees="98" direction="west" minutes="16" orientation="longitude" pageId="51" pageNumber="52" precision="15" seconds="51" value="-98.28084">W98°16´51”</geoCoordinate>
, bosque tropical subcaducifolio, trampa cebada con champiñones, 
<date id="FF99105F771C7F308301A10FFD6CF8A9" box="[569,755,1842,1869]" pageId="51" pageNumber="52" value="2015-04-17" valueMax="2015-04-28" valueMin="2015-04-17">
<collectingDate id="EFDDE9B7771C7F308301A10FFD6CF8A9" box="[569,755,1842,1869]" pageId="51" pageNumber="52" value="2015-04-17" valueMax="2015-04-28" valueMin="2015-04-17">17 a 28-IV-2015</collectingDate>
</date>
, 
<collectorName id="26D25349771C7F3083C5A10FFCE3F8A9" box="[765,892,1843,1869]" pageId="51" pageNumber="52">R. Ramírez</collectorName>
y 
<collectorName id="26D25349771C7F3082AEA10FFB8DF8A9" box="[918,1042,1843,1869]" pageId="51" pageNumber="52">J. Márquez</collectorName>
cols.” (2, 
<collectionCode id="ED36AE5A771C7F3085BBA10FFA9AF8A9" box="[1155,1285,1843,1869]" pageId="51" pageNumber="52">CC-UAEH</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5771C7F308427A10FFA07F8A9" box="[1311,1432,1843,1869]" pageId="51" pageNumber="52">Huehuetla</collectingMunicipality>
, 
<location id="8EF86044771C7F3081AFA16BFE85F895" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771C7F3081AFA16BFE85F895" box="[151,282,1879,1905]" country="Mexico" latitude="21.123056" longLatPrecision="21" longitude="-98.28084" municipality="Huehuetla" name="Rio Blanco" pageId="51" pageNumber="52" stateProvince="Hidalgo">Rio Blanco</location>
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771C7F30801EA16AFB16F85D" collectingDate="2010-05-01" collectingDateMax="2011-09-09" collectingDateMin="2010-05-01" collectionCode="CC-, UAEH, CC-UAEH" collectorName="C. Berriozabal &amp; J. Marquez" country="Mexico" elevation="588" latitude="21.1" location="Lomas del Pericon" longLatPrecision="126" longitude="-99.1" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Hidalgo">
<quantity id="4CDF9B7A771C7F30801EA16AFEEFF895" box="[294,368,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="5.88" pageId="51" pageNumber="52" unit="m" value="588.0">
<elevation id="000AD1AC771C7F30801EA16AFEEFF895" box="[294,368,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="5.88" pageId="51" pageNumber="52" unit="m" value="588.0">588 m</elevation>
</quantity>
, UTM (WG-384), 14 94809, NTP-80 (calamar), 
<date id="FF99105F771C7F3082A5A16BFBDCF895" box="[925,1091,1879,1905]" pageId="51" pageNumber="52" value="2010-05-01" valueMax="2010-05-30" valueMin="2010-05-01">
<collectingDate id="EFDDE9B7771C7F3082A5A16BFBDCF895" box="[925,1091,1879,1905]" pageId="51" pageNumber="52" value="2010-05-01" valueMax="2010-05-30" valueMin="2010-05-01">1 a 30-V-2010</collectingDate>
</date>
, 
<collectorName id="26D25349771C7F308577A16BFB6BF895" box="[1103,1268,1879,1905]" pageId="51" pageNumber="52">C. Berriozábal</collectorName>
col.” (22, 
<collectionCode id="ED36AE5A771C7F308456A16BFF75F871" pageId="51" pageNumber="52">CC- UAEH</collectionCode>
). “La Misión, 
<location id="8EF86044771C7F3080A9A147FDF6F871" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771C7F3080A9A147FDF6F871" box="[401,617,1915,1941]" country="Mexico" latitude="21.1" longLatPrecision="126" longitude="-99.1" name="Lomas del Pericon" pageId="51" pageNumber="52" stateProvince="Hidalgo">Lomas del Pericón</location>
, bosque mesófilo de montaña, 
<quantity id="4CDF9B7A771C7F3082F6A147FBB8F871" box="[974,1063,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.377" pageId="51" pageNumber="52" unit="m" value="1377.0">
<elevation id="000AD1AC771C7F3082F6A147FBB8F871" box="[974,1063,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.377" pageId="51" pageNumber="52" unit="m" value="1377.0">1377 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058771C7F30850DA147FB04F871" box="[1077,1179,1915,1941]" degrees="21" direction="north" minutes="6.0" orientation="latitude" pageId="51" pageNumber="52" precision="92" value="21.1">N21°6.0’</geoCoordinate>
46.0”, 
<geoCoordinate id="EE135058771C7F3085D2A146FAC6F871" box="[1258,1369,1914,1941]" degrees="99" direction="west" minutes="6.0" orientation="longitude" pageId="51" pageNumber="52" precision="92" value="-99.1">W99°6.0’</geoCoordinate>
15.6”, trampa de intercepción de vuelo #1, 
<date id="FF99105F771C7F308312A1A3FC8FF85D" box="[554,784,1951,1977]" pageId="51" pageNumber="52" value="2011-08-26" valueMax="2011-09-09" valueMin="2011-08-26">
<collectingDate id="EFDDE9B7771C7F308312A1A3FC8FF85D" box="[554,784,1951,1977]" pageId="51" pageNumber="52" value="2011-08-26" valueMax="2011-09-09" valueMin="2011-08-26">26-VIII a 9-IX-2011</collectingDate>
</date>
, 
<collectorName id="26D25349771C7F308222A1A3FC09F85D" box="[794,918,1951,1977]" pageId="51" pageNumber="52">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A771C7F3082C4A1A3FBE1F85D" box="[1020,1150,1951,1977]" pageId="51" pageNumber="52">CC-UAEH</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4F3CC2771C7F3785B7A1A3FD62FEA5" collectingDate="1994-09-11" collectingDateMax="2018-10-25" collectingDateMin="1994-09-11" collectionCode="CC-UAEH, MAAS" collectorName="J. Marquez &amp; J. D. Silva &amp; J. D. Silva-Hurtado &amp; O. Perez" country="Mexico" county="rio San Marcos" lastPageId="52" lastPageNumber="53" latitude="20.322918" location="Canada de Patla" longLatPrecision="1" longitude="-98.04317" municipality="Xicotepec de Juarez" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Puebla">
<emphasis id="B953EA8D771C7F3085B7A1A3FB75F85D" bold="true" box="[1167,1258,1951,1977]" pageId="51" pageNumber="52">
<collectingRegion id="49E3F87D771C7F3085B7A1A3FB79F85D" box="[1167,1254,1951,1977]" country="Mexico" name="Puebla" pageId="51" pageNumber="52">Puebla</collectingRegion>
:
</emphasis>
“Tlacuilotepec, cerca del 
<collectingCounty id="62F94E13771C7F30803AA1FFFE30F839" box="[258,431,1987,2013]" pageId="51" pageNumber="52">río San Marcos</collectingCounty>
, 
<geoCoordinate id="EE135058771C7F308082A1FEFDC4F839" box="[442,603,1986,2013]" degrees="20" direction="north" minutes="19" orientation="latitude" pageId="51" pageNumber="52" precision="1" seconds="22.5" value="20.322918">N20°19’22.5”</geoCoordinate>
, 
<geoCoordinate id="EE135058771C7F30835DA1FEFC94F839" box="[613,779,1986,2013]" degrees="98" direction="west" minutes="02" orientation="longitude" pageId="51" pageNumber="52" precision="1" seconds="35.4" value="-98.04317">W98°02’35.4”</geoCoordinate>
, bosque tropical subcaducifolio, trampa con fruta, 
<date id="FF99105F771C7F378479A1FFFF70FF55" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" value="2018-10-13" valueMax="2018-10-25" valueMin="2018-10-13">
<collectingDate id="EFDDE9B7771C7F378479A1FFFF70FF55" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" value="2018-10-13" valueMax="2018-10-25" valueMin="2018-10-13">13 a 25- X-2018</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F3781C3A6ABFEE7FF55" box="[251,376,151,177]" pageId="52" pageNumber="53">J. Márquez</collectorName>
y 
<collectorName id="26D25349771B7F3780ACA6ABFD95FF55" box="[404,522,151,177]" pageId="52" pageNumber="53">J. D. Silva</collectorName>
cols.” (m 1, 
<collectionCode id="ED36AE5A771B7F3783A3A6ABFC82FF55" box="[667,797,151,177]" pageId="52" pageNumber="53">CC-UAEH</collectionCode>
). Same data, except: “NTP-80, 
<date id="FF99105F771B7F378546A6ABFAE0FF55" box="[1150,1407,151,177]" pageId="52" pageNumber="53" value="2018-07-17" valueMax="2018-08-19" valueMin="2018-07-17">
<collectingDate id="EFDDE9B7771B7F378546A6ABFAE0FF55" box="[1150,1407,151,177]" pageId="52" pageNumber="53" value="2018-07-17" valueMax="2018-08-19" valueMin="2018-07-17">17-VII a 19-VIII-2018</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F3784B2A6ABFEC7FF31" pageId="52" pageNumber="53">J. D. Silva-Hurtado</collectorName>
col.” (3, 
<collectionCode id="ED36AE5A771B7F378086A687FDDFFF31" box="[446,576,187,213]" pageId="52" pageNumber="53">CC-UAEH</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5771B7F378363A687FCDFFF31" box="[603,832,187,213]" pageId="52" pageNumber="53">Xicotepec de Juárez</collectingMunicipality>
, 
<location id="8EF86044771B7F378273A687FB9EFF31" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378273A687FB9EFF31" box="[843,1025,187,213]" country="Mexico" county="rio San Marcos" latitude="20.322918" longLatPrecision="1" longitude="-98.04317" municipality="Xicotepec de Juarez" name="Canada de Patla" pageId="52" pageNumber="53" stateProvince="Puebla">Cañada de Patla</location>
, bosque mesófilo de montaña perturbado, 
<date id="FF99105F771B7F3781E2A6E3FEFFFF1D" box="[218,352,223,249]" pageId="52" pageNumber="53" value="1994-09-11">
<collectingDate id="EFDDE9B7771B7F3781E2A6E3FEFFFF1D" box="[218,352,223,249]" pageId="52" pageNumber="53" value="1994-09-11">11-IX-1994</collectingDate>
</date>
, en tronco caído, 
<collectorName id="26D25349771B7F37831AA6E3FD02FF1D" box="[546,669,223,249]" pageId="52" pageNumber="53">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A771B7F37823AA6E3FCC7FF1D" box="[770,856,223,249]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<location id="8EF86044771B7F37824BA6E3FBC7FF1D" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F37824BA6E3FBC7FF1D" box="[883,1112,223,249]" country="Mexico" county="rio San Marcos" latitude="20.322918" longLatPrecision="1" longitude="-98.04317" municipality="Xicotepec de Juarez" name="Xicotepec de Juarez" pageId="52" pageNumber="53" stateProvince="Puebla">Xicotepec de Juárez</location>
, Hidroeléctrica Patla, bosque mesófilo de montaña perturbado, en necrotrampa temporal (calamar), 
<date id="FF99105F771B7F378290A73FFBB2FEF9" box="[936,1069,259,285]" pageId="52" pageNumber="53" value="1998-03-27">
<collectingDate id="EFDDE9B7771B7F378290A73FFBB2FEF9" box="[936,1069,259,285]" pageId="52" pageNumber="53" value="1998-03-27">27-III-1998</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F378501A73FFB06FEF9" box="[1081,1177,259,285]" pageId="52" pageNumber="53">O. Pérez</collectorName>
col.” (2, 
<collectionCode id="ED36AE5A771B7F37843AA73FFAC7FEF9" box="[1282,1368,259,285]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<location id="8EF86044771B7F37844DA73FFEFFFEA5" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F37844DA73FFEFFFEA5" country="Mexico" county="rio San Marcos" latitude="20.322918" longLatPrecision="1" longitude="-98.04317" municipality="Xicotepec de Juarez" name="Xicotepec de Juarez" pageId="52" pageNumber="53" stateProvince="Puebla">Xicotepec de Juárez</location>
, 
<location id="8EF86044771B7F378054A71BFDD0FEA5" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378054A71BFDD0FEA5" box="[364,591,295,321]" country="Mexico" county="rio San Marcos" latitude="20.322918" longLatPrecision="1" longitude="-98.04317" municipality="Xicotepec de Juarez" name="Hidroelectrica Plata" pageId="52" pageNumber="53" stateProvince="Puebla">Hidroeléctrica Plata</location>
, selva mediana
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F378231A71AFD90FE81" collectingDate="2002-10-05" collectionCode="MAAS" collectorName="J. Asiain &amp; J. Marquez" country="Mexico" elevation="457" location="Same" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Puebla">
<quantity id="4CDF9B7A771B7F378231A71AFCCDFEA5" box="[777,850,294,321]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="52" pageNumber="53" unit="m" value="457.0">
<elevation id="000AD1AC771B7F378231A71AFCCDFEA5" box="[777,850,294,321]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="52" pageNumber="53" unit="m" value="457.0">457 m</elevation>
</quantity>
, en plantas, 
<date id="FF99105F771B7F3782E6A71AFBD2FEA5" box="[990,1101,294,321]" pageId="52" pageNumber="53" value="2002-10-05">
<collectingDate id="EFDDE9B7771B7F3782E6A71AFBD2FEA5" box="[990,1101,294,321]" pageId="52" pageNumber="53" value="2002-10-05">5-X-2002</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F378561A71BFB25FEA5" box="[1113,1210,295,321]" pageId="52" pageNumber="53">J. Asiain</collectorName>
y 
<collectorName id="26D25349771B7F3785EFA71BFACAFEA5" box="[1239,1365,295,321]" pageId="52" pageNumber="53">J. Márquez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A771B7F378186A777FE8BFE81" box="[190,276,331,357]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). 
<location id="8EF86044771B7F37801FA777FEFAFE81" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F37801FA777FEFAFE81" box="[295,357,331,357]" country="Mexico" name="Same" pageId="52" pageNumber="53" stateProvince="Puebla">Same</location>
data, except: “
</materialsCitation>
<materialsCitation id="3B4F3CC2771B7F378337A777FB50FE81" box="[527,1231,331,357]" collectingDate="2002-03-06" collectionCode="MAAS" country="Mexico" elevation="487" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Puebla">
<quantity id="4CDF9B7A771B7F378337A777FDC3FE81" box="[527,604,331,357]" metricMagnitude="2" metricUnit="m" metricValue="4.87" pageId="52" pageNumber="53" unit="m" value="487.0">
<elevation id="000AD1AC771B7F378337A777FDC3FE81" box="[527,604,331,357]" metricMagnitude="2" metricUnit="m" metricValue="4.87" pageId="52" pageNumber="53" unit="m" value="487.0">487 m</elevation>
</quantity>
, en inflorescencia de plátano, 
<date id="FF99105F771B7F37828EA777FBAFFE81" box="[950,1072,331,357]" pageId="52" pageNumber="53" value="2002-03-06">
<collectingDate id="EFDDE9B7771B7F37828EA777FBAFFE81" box="[950,1072,331,357]" pageId="52" pageNumber="53" value="2002-03-06">6-III-2002</collectingDate>
</date>
.” (1, 
<collectionCode id="ED36AE5A771B7F378555A777FB5CFE81" box="[1133,1219,331,357]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4F3CC2771B7F3785EFA776FC22FE49" collectingDate="2007-07-16" collectionCode="CC-UAEH" collectorName="J. Asiain &amp; J. Marquez" country="Mexico" elevation="585" latitude="21.398806" location="Las Pozas de James" longLatPrecision="1" longitude="-98.935196" municipality="Xilitla" pageId="52" pageNumber="53" specimenCount="1" stateProvince="San Luis Potosi">
<emphasis id="B953EA8D771B7F3785EFA776FA03FE81" bold="true" box="[1239,1436,330,357]" pageId="52" pageNumber="53">
<collectingRegion id="49E3F87D771B7F3785EFA776FA08FE81" box="[1239,1431,330,357]" country="Mexico" name="San Luis Potosi" pageId="52" pageNumber="53">San Luis Potosí</collectingRegion>
:
</emphasis>
“ 
<collectingMunicipality id="6BFCACE5771B7F378199A753FF71FE6D" box="[161,238,367,393]" pageId="52" pageNumber="53">Xilitla</collectingMunicipality>
, 
<location id="8EF86044771B7F3781C4A753FE7AFE6D" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F3781C4A753FE7AFE6D" box="[252,485,367,393]" country="Mexico" latitude="21.398806" longLatPrecision="1" longitude="-98.935196" municipality="Xilitla" name="Las Pozas de James" pageId="52" pageNumber="53" stateProvince="San Luis Potosi">Las Pozas de James</location>
, 
<geoCoordinate id="EE135058771B7F3780CCA752FD0AFE6D" box="[500,661,366,393]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="55.7" value="21.398806">N21°23’55.7”</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F37839BA752FCA4FE6D" box="[675,827,366,393]" degrees="98" direction="west" minutes="56" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="6.7" value="-98.935196">W98°56’6.7”</geoCoordinate>
, bosque tropical subperenifolio, 
<quantity id="4CDF9B7A771B7F37858EA752FA9DFE6D" box="[1206,1282,366,393]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="52" pageNumber="53" unit="m" value="585.0">
<elevation id="000AD1AC771B7F37858EA752FA9DFE6D" box="[1206,1282,366,393]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="52" pageNumber="53" unit="m" value="585.0">585 m</elevation>
</quantity>
, bajo rocas a orilla del río, 
<date id="FF99105F771B7F378015A7AFFE21FE49" box="[301,446,403,429]" pageId="52" pageNumber="53" value="2007-07-16">
<collectingDate id="EFDDE9B7771B7F378015A7AFFE21FE49" box="[301,446,403,429]" pageId="52" pageNumber="53" value="2007-07-16">16-VII-2007</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F3780F0A7AFFDB7FE49" box="[456,552,403,429]" pageId="52" pageNumber="53">J. Asiain</collectorName>
y 
<collectorName id="26D25349771B7F37837AA7AFFD21FE49" box="[578,702,403,429]" pageId="52" pageNumber="53">J. Márquez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A771B7F378217A7AFFC2EFE49" box="[815,945,403,429]" pageId="52" pageNumber="53">CC-UAEH</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4F3CC2771B7F3782FAA7AFFC52FDFD" collectingDate="1994-02-24" collectionCode="MAAS, IEXA" collectorName="J. Marquez &amp; M. A. Moron &amp; de Agua" country="Mexico" county="Same" latitude="18.45" location="Pipiapan" longLatPrecision="1275" longitude="-95.05" municipality="Catemaco" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<emphasis id="B953EA8D771B7F3782FAA7AFFBA6FE49" bold="true" box="[962,1081,403,429]" pageId="52" pageNumber="53">
<collectingRegion id="49E3F87D771B7F3782FAA7AFFBABFE49" box="[962,1076,403,429]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="52" pageNumber="53">Veracruz</collectingRegion>
:
</emphasis>
“Catemaco, Estación Biológica “Los Tuxtlas”, en frutos podridos, 20 al 
<date id="FF99105F771B7F378362A78BFD49FE35" box="[602,726,439,465]" pageId="52" pageNumber="53" value="1994-02-24">
<collectingDate id="EFDDE9B7771B7F378362A78BFD49FE35" box="[602,726,439,465]" pageId="52" pageNumber="53" value="1994-02-24">24-II-1994</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F3783D9A78BFCC1FE35" box="[737,862,439,465]" pageId="52" pageNumber="53">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A771B7F3782FFA78BFB82FE35" box="[967,1053,439,465]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). 
<collectingCounty id="62F94E13771B7F378517A78BFBF2FE35" box="[1071,1133,439,465]" pageId="52" pageNumber="53">Same</collectingCounty>
data, except: “caminando” (1, 
<collectionCode id="ED36AE5A771B7F378185A7E7FE8CFE11" box="[189,275,475,501]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5771B7F378017A7E7FE3AFE11" box="[303,421,475,501]" pageId="52" pageNumber="53">Catemaco</collectingMunicipality>
, 
<location id="8EF86044771B7F378097A7E7FD8BFE11" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378097A7E7FD8BFE11" box="[431,532,475,501]" country="Mexico" county="Same" latitude="18.45" longLatPrecision="1275" longitude="-95.05" municipality="Catemaco" name="Pipiapan" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Pipiapan</location>
, 
<geoCoordinate id="EE135058771B7F378318A7E7FD1CFE11" box="[544,643,475,501]" degrees="18" direction="north" minutes="27" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="18.45">N18°27´</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F3783B6A7E6FD68FE11" box="[654,759,474,501]" degrees="95" direction="west" minutes="03" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="-95.05">W95°03´</geoCoordinate>
, feb-march, 1991, 
<collectorName id="26D25349771B7F3782F2A7E7FBFEFE11" box="[970,1121,475,501]" pageId="52" pageNumber="53">M. A. Morón</collectorName>
, 
<collectingMethod id="52664E88771B7F378555A7E7FA8DFE11" box="[1133,1298,475,501]" pageId="52" pageNumber="53">ex carrion trap</collectingMethod>
” (1, 
<collectionCode id="ED36AE5A771B7F378471A7E7FA0EFE11" box="[1353,1425,475,501]" pageId="52" pageNumber="53">IEXA</collectionCode>
). “ 
<location id="8EF86044771B7F378199A7C3FE98FDFD" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378199A7C3FE98FDFD" box="[161,263,511,537]" country="Mexico" county="Same" latitude="18.45" longLatPrecision="1275" longitude="-95.05" municipality="Catemaco" name="Cordoba" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Córdoba</location>
, 
<location id="8EF86044771B7F378029A7C3FE7CFDFD" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378029A7C3FE7CFDFD" box="[273,483,511,537]" country="Mexico" county="Same" latitude="18.45" longLatPrecision="1275" longitude="-95.05" municipality="Catemaco" name="San Rafael Caleria" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">San Rafael Calería</location>
, 
<location id="8EF86044771B7F3780D4A7C3FDE6FDFD" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F3780D4A7C3FDE6FDFD" box="[492,633,511,537]" country="Mexico" county="Same" latitude="18.45" longLatPrecision="1275" longitude="-95.05" municipality="Catemaco" name="Ojo de Agua" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">
Ojo 
<collectorName id="26D25349771B7F378324A7C3FDE6FDFD" box="[540,633,511,537]" pageId="52" pageNumber="53">de Agua</collectorName>
</location>
, selva mediana subcaducifolia
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F3782EEA7C3FA89FDD9" collectionCode="MAAS" collectorName="Q. Santiago &amp; J. Marquez" country="Mexico" county="Cordoba" elevation="1127" location="Ojo de Agua" municipality="San Rafael Caleria" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F3782EEA7C3FBB4FDFD" box="[982,1067,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="52" pageNumber="53" unit="m" value="1127.0">
<elevation id="000AD1AC771B7F3782EEA7C3FBB4FDFD" box="[982,1067,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="52" pageNumber="53" unit="m" value="1127.0">1127 m</elevation>
</quantity>
, en lima podrida, 
<collectorName id="26D25349771B7F3785D6A7C3FAEEFDFD" box="[1262,1393,511,537]" pageId="52" pageNumber="53">Q. Santiago</collectorName>
y 
<collectorName id="26D25349771B7F3784B2A7C3FF64FDD9" pageId="52" pageNumber="53">J. Márquez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A771B7F378056A41FFE5BFDD9" box="[366,452,547,573]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<collectingCounty id="62F94E13771B7F3780E7A41FFDDAFDD9" box="[479,581,547,573]" pageId="52" pageNumber="53">Córdoba</collectingCounty>
, 
<collectingMunicipality id="6BFCACE5771B7F378368A41FFCBAFDD9" box="[592,805,547,573]" pageId="52" pageNumber="53">San Rafael Calería</collectingMunicipality>
, 
<location id="8EF86044771B7F378217A41FFC21FDD9" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378217A41FFC21FDD9" box="[815,958,547,573]" country="Mexico" county="Cordoba" municipality="San Rafael Caleria" name="Ojo de Agua" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Ojo de Agua</location>
, selva mediana subcaducifolia
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F378419A41FFD0FFD61" collectionCode="MAAS" collectorName="Q. Santiago &amp; J. Marquez" country="Mexico" county="Cordoba" elevation="1127" latitude="18.954168" location="Ojo de Agua" longLatPrecision="0" longitude="-96.0" municipality="San Rafael Caleria" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F378419A41FFAE9FDD9" box="[1313,1398,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="52" pageNumber="53" unit="m" value="1127.0">
<elevation id="000AD1AC771B7F378419A41FFAE9FDD9" box="[1313,1398,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="52" pageNumber="53" unit="m" value="1127.0">1127 m</elevation>
</quantity>
, en lima podrida, 
<collectorName id="26D25349771B7F378009A47BFE2BFD85" box="[305,436,583,609]" pageId="52" pageNumber="53">Q. Santiago</collectorName>
y 
<collectorName id="26D25349771B7F3780F5A47BFDD6FD85" box="[461,585,583,609]" pageId="52" pageNumber="53">J. Márquez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A771B7F378380A47BFC91FD85" box="[696,782,583,609]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<collectingCounty id="62F94E13771B7F378211A47BFC10FD85" box="[809,911,583,609]" pageId="52" pageNumber="53">Córdoba</collectingCounty>
, 
<collectingMunicipality id="6BFCACE5771B7F3782A1A47BFBF4FD85" box="[921,1131,583,609]" pageId="52" pageNumber="53">San Rafael Calería</collectingMunicipality>
, 
<location id="8EF86044771B7F37854CA47BFA9DFD85" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F37854CA47BFA9DFD85" box="[1140,1282,583,609]" country="Mexico" county="Cordoba" latitude="18.954168" longLatPrecision="0" longitude="-96.0" municipality="San Rafael Caleria" name="Ojo de Agua" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Ojo de Agua</location>
, 
<geoCoordinate id="EE135058771B7F378434A47AFA08FD85" box="[1292,1431,582,609]" degrees="18" direction="north" minutes="57" orientation="latitude" pageId="52" pageNumber="53" precision="15" seconds="15" value="18.954168">N18°57’15”</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F3781AFA456FF41FD61" box="[151,222,618,645]" degrees="96" direction="west" orientation="longitude" pageId="52" pageNumber="53" precision="55555" value="-96.0">W96°</geoCoordinate>
,56’24”, selva mediana subcaducifolia
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F3783A6A457FCFAFD4D" collectingDate="1999-03-20" collectingDateMax="1999-04-17" collectingDateMin="1999-03-20" collectionCode="MAAS" collectorName="Q. J. Santiago &amp; J. Marquez &amp; Asiain" country="Mexico" elevation="1127" location="Cuauhtemoc" municipality="Cordoba" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F3783A6A457FD69FD61" box="[670,758,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="52" pageNumber="53" unit="m" value="1127.0">
<elevation id="000AD1AC771B7F3783A6A457FD69FD61" box="[670,758,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="52" pageNumber="53" unit="m" value="1127.0">1127 m</elevation>
</quantity>
, NTP-80 (calamar), 
<date id="FF99105F771B7F3782DCA457FB55FD61" box="[996,1226,619,645]" pageId="52" pageNumber="53" value="1999-03-20" valueMax="1999-04-17" valueMin="1999-03-20">
<collectingDate id="EFDDE9B7771B7F3782DCA457FB55FD61" box="[996,1226,619,645]" pageId="52" pageNumber="53" value="1999-03-20" valueMax="1999-04-17" valueMin="1999-03-20">20-III a 17-IV-1999</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F3785EFA457FAE2FD61" box="[1239,1405,619,645]" pageId="52" pageNumber="53">Q. J. Santiago</collectorName>
, 
<collectorName id="26D25349771B7F3784B2A457FF64FD4D" pageId="52" pageNumber="53">J. Márquez</collectorName>
y J. 
<collectorName id="26D25349771B7F378009A4B3FEE5FD4D" box="[305,378,655,681]" pageId="52" pageNumber="53">Asiain</collectorName>
cols.” (2, 
<collectionCode id="ED36AE5A771B7F3780C8A4B3FDD9FD4D" box="[496,582,655,681]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5771B7F37835BA4B3FD56FD4D" box="[611,713,655,681]" pageId="52" pageNumber="53">Córdoba</collectingMunicipality>
, 
<location id="8EF86044771B7F3783EDA4B3FCFAFD4D" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F3783EDA4B3FCFAFD4D" box="[725,869,655,681]" country="Mexico" municipality="Cordoba" name="Cuauhtemoc" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Cuauhtémoc</location>
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F378249A4B3FCE4FD29" collectingDate="1992-06-24" collectionCode="IEXA" collectorName="R. Hernandez" country="Mexico" elevation="940" location="Xococapa" municipality="Ilamatlan" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F378249A4B3FC24FD4D" box="[881,955,655,681]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="52" pageNumber="53" unit="m" value="940.0">
<elevation id="000AD1AC771B7F378249A4B3FC24FD4D" box="[881,955,655,681]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="52" pageNumber="53" unit="m" value="940.0">940 m</elevation>
</quantity>
, 
<date id="FF99105F771B7F3782F0A4B3FBD1FD4D" box="[968,1102,655,681]" pageId="52" pageNumber="53" value="1992-06-24">
<collectingDate id="EFDDE9B7771B7F3782F0A4B3FBD1FD4D" box="[968,1102,655,681]" pageId="52" pageNumber="53" value="1992-06-24">24-VI-1992</collectingDate>
</date>
, NTP-80, 
<collectorName id="26D25349771B7F3785FCA4B3FAC1FD4D" box="[1220,1374,655,681]" pageId="52" pageNumber="53">R. Hernández</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A771B7F378186A48FFE99FD29" box="[190,262,691,717]" pageId="52" pageNumber="53">IEXA</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5771B7F37801CA48FFE0CFD29" box="[292,403,691,717]" pageId="52" pageNumber="53">Ilamatlán</collectingMunicipality>
, 
<location id="8EF86044771B7F378098A48FFD8CFD29" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378098A48FFD8CFD29" box="[416,531,691,717]" country="Mexico" municipality="Ilamatlan" name="Xococapa" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Xococapa</location>
, bosque tropical subperenifolio
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F3782B0A48FFB01FD15" collectingDate="2006-04-10" collectingDateMax="2006-08-13" collectingDateMin="2006-04-10" collectionCode="CC-UAEH" collectorName="F. Ramirez" country="Mexico" elevation="388" latitude="20.806915" location="Same" longLatPrecision="1" longitude="-98.340195" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F3782B0A48FFC4CFD29" box="[904,979,691,717]" metricMagnitude="2" metricUnit="m" metricValue="3.88" pageId="52" pageNumber="53" unit="m" value="388.0">
<elevation id="000AD1AC771B7F3782B0A48FFC4CFD29" box="[904,979,691,717]" metricMagnitude="2" metricUnit="m" metricValue="3.88" pageId="52" pageNumber="53" unit="m" value="388.0">388 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058771B7F3782D8A48FFB1EFD29" box="[992,1153,691,717]" degrees="20" direction="north" minutes="48" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="24.9" value="20.806915">N20°48’24.9”</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F3785B6A48FFAABFD29" box="[1166,1332,691,717]" degrees="98" direction="west" minutes="20" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="24.7" value="-98.340195">W98°20’24.7”</geoCoordinate>
, NTP-80 (calamar) # 3, 
<date id="FF99105F771B7F378005A4EBFDADFD15" box="[317,562,727,753]" pageId="52" pageNumber="53" value="2006-08-13" valueMax="2006-04-10" valueMin="2006-08-13">
<collectingDate id="EFDDE9B7771B7F378005A4EBFDADFD15" box="[317,562,727,753]" pageId="52" pageNumber="53" value="2006-08-13" valueMax="2006-04-10" valueMin="2006-08-13">13-VIII a 10-IV-2006</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F378306A4EBFD26FD15" box="[574,697,727,753]" pageId="52" pageNumber="53">F. Ramírez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A771B7F37821CA4EBFC39FD15" box="[804,934,727,753]" pageId="52" pageNumber="53">CC-UAEH</collectionCode>
). 
<location id="8EF86044771B7F378281A4EBFC68FD15" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378281A4EBFC68FD15" box="[953,1015,727,753]" country="Mexico" latitude="20.806915" longLatPrecision="1" longitude="-98.340195" name="Same" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Same</location>
data, except: “
</materialsCitation>
<materialsCitation id="3B4F3CC2771B7F3785A7A4EBFB06FCF1" collectingDate="2006-03-05" collectingDateMax="2006-04-01" collectingDateMin="2006-03-05" collectionCode="CC-UAEH" country="Mexico" elevation="397" latitude="20.806665" location="Same" longLatPrecision="1" longitude="-98.34017" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F3785A7A4EBFB74FD15" box="[1183,1259,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="52" pageNumber="53" unit="m" value="397.0">
<elevation id="000AD1AC771B7F3785A7A4EBFB74FD15" box="[1183,1259,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="52" pageNumber="53" unit="m" value="397.0">397 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058771B7F3785CFA4EBFA08FD15" box="[1271,1431,727,753]" degrees="20" direction="north" minutes="48" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="24.0" value="20.806665">N20°48’24.0”</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F3781AFA4C7FED7FCF1" box="[151,328,763,789]" degrees="98" direction="west" minutes="20" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="24.6" value="-98.34017">W 98°20’24.6”</geoCoordinate>
, NTP-80 #4, 
<date id="FF99105F771B7F3780D1A4C6FD4CFCF1" box="[489,723,762,789]" pageId="52" pageNumber="53" value="2006-03-05" valueMax="2006-04-01" valueMin="2006-03-05">
<collectingDate id="EFDDE9B7771B7F3780D1A4C6FD4CFCF1" box="[489,723,762,789]" pageId="52" pageNumber="53" value="2006-03-05" valueMax="2006-04-01" valueMin="2006-03-05">05-III a 01-IV-2006</collectingDate>
</date>
.” (1, 
<collectionCode id="ED36AE5A771B7F37822CA4C7FC09FCF1" box="[788,918,763,789]" pageId="52" pageNumber="53">CC-UAEH</collectionCode>
). 
<location id="8EF86044771B7F378293A4C7FC76FCF1" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378293A4C7FC76FCF1" box="[939,1001,763,789]" country="Mexico" latitude="20.806665" longLatPrecision="1" longitude="-98.34017" name="Same" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Same</location>
data, except: “
</materialsCitation>
<materialsCitation id="3B4F3CC2771B7F3785A1A4C7FA08FCDD" collectionCode="CC-UAEH" country="Mexico" county="Totutla" elevation="362" latitude="19.20639" location="Rancho Zacuapan" longLatPrecision="21" longitude="-96.842224" municipality="Mata Oscura" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F3785A1A4C7FB77FCF1" box="[1177,1256,763,789]" metricMagnitude="2" metricUnit="m" metricValue="3.62" pageId="52" pageNumber="53" unit="m" value="362.0">
<elevation id="000AD1AC771B7F3785A1A4C7FB77FCF1" box="[1177,1256,763,789]" metricMagnitude="2" metricUnit="m" metricValue="3.62" pageId="52" pageNumber="53" unit="m" value="362.0">362 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058771B7F3785CFA4C7FA08FCF1" box="[1271,1431,763,789]" degrees="20" direction="north" minutes="48" orientation="latitude" pageId="52" pageNumber="53" precision="1" seconds="20.3" value="20.805637">N20°48’20.3”</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F3781AFA522FEA1FCDD" box="[151,318,798,825]" degrees="98" direction="west" minutes="20" orientation="longitude" pageId="52" pageNumber="53" precision="1" seconds="31.5" value="-98.34209">W98°20’31.5”</geoCoordinate>
, NTP-80 #2” (1, 
<collectionCode id="ED36AE5A771B7F3780C7A523FD1EFCDD" box="[511,641,799,825]" pageId="52" pageNumber="53">CC-UAEH</collectionCode>
). “ 
<collectingCounty id="62F94E13771B7F3783A4A523FD6EFCDD" box="[668,753,799,825]" pageId="52" pageNumber="53">Totutla</collectingCounty>
, 
<collectingMunicipality id="6BFCACE5771B7F3783C3A523FC12FCDD" box="[763,909,799,825]" pageId="52" pageNumber="53">Mata Oscura</collectingMunicipality>
, 
<location id="8EF86044771B7F3782A1A523FBFAFCDD" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F3782A1A523FBFAFCDD" box="[921,1125,799,825]" country="Mexico" county="Totutla" latitude="19.20639" longLatPrecision="21" longitude="-96.842224" municipality="Mata Oscura" name="Rancho Zacuapan" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Rancho Zacuapan</location>
, 
<geoCoordinate id="EE135058771B7F378557A523FB64FCDD" box="[1135,1275,799,825]" degrees="19" direction="north" minutes="12" orientation="latitude" pageId="52" pageNumber="53" precision="15" seconds="23" value="19.20639">N19°12’23”</geoCoordinate>
, 
<geoCoordinate id="EE135058771B7F37843EA522FA08FCDD" box="[1286,1431,798,825]" degrees="96" direction="west" minutes="50" orientation="longitude" pageId="52" pageNumber="53" precision="15" seconds="32" value="-96.842224">W96°50’32”</geoCoordinate>
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2771B7F3781AFA57FFACBFC41" collectingDate="1994-02-24" collectingDateMax="1999-05-18" collectingDateMin="1994-02-24" collectionCode="MAAS, QJSJ" collectorName="J. Asiain &amp; Q. Santiago &amp; J. Marquez" country="Mexico" county="Same" elevation="869" location="Xico" municipality="Same" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A771B7F3781AFA57FFF7FFCB9" box="[151,224,835,861]" metricMagnitude="2" metricUnit="m" metricValue="8.69" pageId="52" pageNumber="53" unit="m" value="869.0">
<elevation id="000AD1AC771B7F3781AFA57FFF7FFCB9" box="[151,224,835,861]" metricMagnitude="2" metricUnit="m" metricValue="8.69" pageId="52" pageNumber="53" unit="m" value="869.0">869 m</elevation>
</quantity>
, naranja podrida, 
<date id="FF99105F771B7F378090A57FFDBCFCB9" box="[424,547,835,861]" pageId="52" pageNumber="53" value="1999-05-18">
<collectingDate id="EFDDE9B7771B7F378090A57FFDBCFCB9" box="[424,547,835,861]" pageId="52" pageNumber="53" value="1999-05-18">18-V-1999</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F378316A57FFD0EFCB9" box="[558,657,835,861]" pageId="52" pageNumber="53">J. Asiain</collectorName>
, 
<collectorName id="26D25349771B7F3783A3A57FFC80FCB9" box="[667,799,835,861]" pageId="52" pageNumber="53">Q. Santiago</collectorName>
y 
<collectorName id="26D25349771B7F378203A57FFC28FCB9" box="[827,951,835,861]" pageId="52" pageNumber="53">J. Márquez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A771B7F378512A57FFB1FFCB9" box="[1066,1152,835,861]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). 
<collectingCounty id="62F94E13771B7F3785AAA57FFB4FFCB9" box="[1170,1232,835,861]" pageId="52" pageNumber="53">Same</collectingCounty>
data, except: “ 
<date id="FF99105F771B7F37844DA57FFF71FC65" pageId="52" pageNumber="53" value="1994-02-24">
<collectingDate id="EFDDE9B7771B7F37844DA57FFF71FC65" pageId="52" pageNumber="53" value="1994-02-24">24- II-1994</collectingDate>
</date>
” (1, 
<collectionCode id="ED36AE5A771B7F37801CA55BFEE5FC65" box="[292,378,871,897]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). 
<collectingMunicipality id="6BFCACE5771B7F3780B5A55BFE54FC65" box="[397,459,871,897]" pageId="52" pageNumber="53">Same</collectingMunicipality>
data, except: “en mandarina podrida, 
<date id="FF99105F771B7F37824EA55BFC6EFC65" box="[886,1009,871,897]" pageId="52" pageNumber="53" value="1999-05-18">
<collectingDate id="EFDDE9B7771B7F37824EA55BFC6EFC65" box="[886,1009,871,897]" pageId="52" pageNumber="53" value="1999-05-18">18-V-1999</collectingDate>
</date>
” (2, 
<collectionCode id="ED36AE5A771B7F378510A55BFBE1FC65" box="[1064,1150,871,897]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="52" pageNumber="53" type="Herbarium">MAAS</collectionCode>
). “ 
<location id="8EF86044771B7F3785A2A55BFB4AFC65" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F3785A2A55BFB4AFC65" box="[1178,1237,871,897]" country="Mexico" county="Same" municipality="Same" name="Xico" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Xico</location>
, km 1.2 camino a cascadas de 
<location id="8EF86044771B7F378019A5B7FEEEFC41" LSID="urn:lsid:plazi:treatment:038E8789771D7F3781AFA3F7FAC7FC41:8EF86044771B7F378019A5B7FEEEFC41" box="[289,369,907,933]" country="Mexico" county="Same" municipality="Same" name="Texolo" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Texolo</location>
, cultivo de café, en plátano en descomposición, 
<date id="FF99105F771B7F3782B6A5B7FB8AFC41" box="[910,1045,907,933]" pageId="52" pageNumber="53" value="1997-11-12">
<collectingDate id="EFDDE9B7771B7F3782B6A5B7FB8AFC41" box="[910,1045,907,933]" pageId="52" pageNumber="53" value="1997-11-12">12-XI-1997</collectingDate>
</date>
, 
<collectorName id="26D25349771B7F378518A5B7FB3BFC41" box="[1056,1188,907,933]" pageId="52" pageNumber="53">Q. Santiago</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A771B7F378435A5B7FAD4FC41" box="[1293,1355,907,933]" pageId="52" pageNumber="53">QJSJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>