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<mods:title id="BD5FA08D1B685AB9BCA14E8C7BFEC1D8">Taxonomic notes on the genus Sinodrepanus Simonis, 1985, with description of the male of S. falsus (Sharp, 1875) (Coleoptera: Scarabaeidae: Oniticellini)</mods:title>
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<taxonomicName id="4C27A6DEC77EFFA0FF04DA1D5622BE6B" ID-CoL="4XL34" authority="(Sharp, 1875)" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[151,575,1111,1138]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">
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<emphasis id="B953014FC77EFFA0FF04DA1D5593BE6B" bold="true" box="[151,398,1111,1138]" italics="true" pageId="1" pageNumber="586">Sinodrepanus falsus</emphasis>
(
<bibRefCitation id="EFB6A0ACC77EFFA0FE0DDA12562ABE6B" author="Sharp, D." box="[414,567,1112,1138]" pageId="1" pageNumber="586" pagination="33 - 54" refId="ref5611" refString="Sharp, D. (1875) Description of some new genera and species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 13, 33 - 54." type="journal article" year="1875">Sharp, 1875</bibRefCitation>
)
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(
<figureCitation id="131CC1D8C77EFFA0FF0CDA3654F0BE8F" box="[159,237,1148,1174]" captionStart="FIGURES 1, 2" captionStartId="2.[151,264,1264,1288]" captionTargetBox="[191,1394,188,1229]" captionTargetId="figure-19@2.[177,1411,179,1238]" captionTargetPageId="2" captionText="FIGURES 1, 2. Sinodrepanus spp. 1: habitus of a male specimen of S. falsus (Sharp, 1875) from Thailand (picture by Marco Uliana); 2: habitus of the male holotype of S. exsul (Sharp, 1875), from Thailand (picture by Antoine Mantilleri, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5683453" httpUri="https://zenodo.org/record/5683453/files/figure.png" pageId="1" pageNumber="586">Figs. 1</figureCitation>
,
<figureCitation id="131CC1D8C77EFFA0FF68DA36553BBE8F" box="[251,294,1148,1174]" captionStart="FIGURES 39" captionStartId="4.[151,264,1228,1252]" captionTargetBox="[157,1414,199,1205]" captionTargetId="figure-19@4.[151,1435,166,1205]" captionTargetPageId="4" captionText="FIGURES 39. Sinodrepanus spp.—S. falsus (Sharp, 1875), specimens from Thailand (37) (pictures by Marco Uliana). 3: female forebody; 4: male metasternum; 5: lamella copulatrix; 6: parameres in dorsal view; 7: aedeagus in lateral view.—S. exsul (Sharp, 1875), male holotype from Thailand (8, 9) (pictures by Christophe Rivier, MNHN). 8: parameres in dorsal view; 9: aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5683455" httpUri="https://zenodo.org/record/5683455/files/figure.png" pageId="1" pageNumber="586">37</figureCitation>
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<treatmentCitation id="0A86FB4CC77EFFA0FF04DA8A55EBBEC1" author="Sharp, D." box="[151,502,1216,1240]" page="52" pageId="1" pageNumber="586" year="1875">
<taxonomicName id="4C27A6DEC77EFFA0FF04DA8A55EBBEC1" ID-CoL="6SN8C" authority=": Sharp, 1875: 52" authorityName="Sharp" authorityPageNumber="52" authorityYear="1875" box="[151,502,1216,1240]" class="Insecta" family="Scarabaeidae" genus="Oniticellus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="B953014FC77EFFA0FF04DA8A5555BEC1" box="[151,328,1216,1240]" italics="true" pageId="1" pageNumber="586">Oniticellus falsus</emphasis>
:
<bibRefCitation id="EFB6A0ACC77EFFA0FEC6DA8A55EBBEC1" author="Sharp, D." box="[341,502,1216,1240]" pageId="1" pageNumber="586" pagination="33 - 54" refId="ref5611" refString="Sharp, D. (1875) Description of some new genera and species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 13, 33 - 54." type="journal article" year="1875">Sharp, 1875: 52</bibRefCitation>
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<treatmentCitationGroup id="AB37FA73C77EFFA0FF04DA9556A8BEEE" box="[151,693,1247,1271]" pageId="1" pageNumber="586">
<treatmentCitation id="0A86FB4CC77EFFA0FF04DA9556A8BEEE" author="Boucomont, A. &amp; Gillet, G." box="[151,693,1247,1271]" page="19" pageId="1" pageNumber="586" year="1921">
<taxonomicName id="4C27A6DEC77EFFA0FF04DA9556A8BEEE" authority=": Boucomont &amp; Gillet, 1921: 19" authorityName="Boucomont &amp; Gillet" authorityPageNumber="19" authorityYear="1921" box="[151,693,1247,1271]" class="Insecta" family="Scarabaeidae" genus="Drepanochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="B953014FC77EFFA0FF04DA95556CBEEE" box="[151,369,1247,1271]" italics="true" pageId="1" pageNumber="586">Drepanochirus falsus</emphasis>
:
<bibRefCitation id="EFB6A0ACC77EFFA0FEEDDAAA56A8BEEE" author="Boucomont, A. &amp; Gillet, G." box="[382,693,1247,1271]" pageId="1" pageNumber="586" refId="ref4686" refString="Boucomont, A. &amp; Gillet, G. (1921) Faune Entomologique de l'Indochine Francaise. Fam. Scarabaeidae Laparosticti (Coleopteres). Fascicule n ° 4. Imprimerie Nouvelle Albert Portail, Saigon, 76 pp." type="book" year="1921">Boucomont &amp; Gillet, 1921: 19</bibRefCitation>
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<paragraph id="8B98DD5DC77EFFA0FF04DAB4576CBF0F" blockId="1.[151,1436,1216,1425]" box="[151,881,1278,1302]" pageId="1" pageNumber="586">
<treatmentCitationGroup id="AB37FA73C77EFFA0FF04DAB4576CBF0F" box="[151,881,1278,1302]" pageId="1" pageNumber="586">
<treatmentCitation id="0A86FB4CC77EFFA0FF04DAB456B5BF0F" author="Boucomont, A. &amp; Gillet, J. J. E." box="[151,680,1278,1302]" page="103" pageId="1" pageNumber="586" year="1927">
<taxonomicName id="4C27A6DEC77EFFA0FF04DAB456B5BF0F" authority=": Boucomont &amp; Gillet, 1927: 103" authorityName="Boucomont &amp; Gillet" authorityPageNumber="103" authorityYear="1927" box="[151,680,1278,1302]" class="Insecta" family="Scarabaeidae" genus="Cyptochirus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="B953014FC77EFFA0FF04DAB45549BF0F" box="[151,340,1278,1302]" italics="true" pageId="1" pageNumber="586">Cyptochirus falsus</emphasis>
:
<bibRefCitation id="EFB6A0ACC77EFFA0FEF1DAB456B5BF0F" author="Boucomont, A. &amp; Gillet, J. J. E." box="[354,680,1278,1302]" pageId="1" pageNumber="586" pagination="100 - 264" refId="ref4727" refString="Boucomont, A. &amp; Gillet, J. J. E. (1927) Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 100 - 264." type="book chapter" year="1927">Boucomont &amp; Gillet, 1927: 103</bibRefCitation>
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;
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<bibRefCitation id="EFB6A0ACC77EFFA0FD21DAB4576CBF0F" author="Paulian, R." box="[690,881,1278,1302]" pageId="1" pageNumber="586" refId="ref5261" refString="Paulian, R. (1945) Faune de L'Empire Francais. III. Coleopteres Scarabeides de L'Indochine. Librairie Larose, Paris, 227 pp." type="book" year="1945">Paulian, 1945: 128</bibRefCitation>
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<treatmentCitationGroup id="AB37FA73C77EFFA0FF04DB575560BF4A" pageId="1" pageNumber="586">
<treatmentCitation id="0A86FB4CC77EFFA0FF04DB57565CBF2D" author="Janssens, A." box="[151,577,1308,1332]" page="36" pageId="1" pageNumber="586" year="1953">
<taxonomicName id="4C27A6DEC77EFFA0FF04DB57565CBF2D" authority=": Janssens, 1953: 36" authorityName="Janssens" authorityPageNumber="36" authorityYear="1953" box="[151,577,1308,1332]" class="Insecta" family="Scarabaeidae" genus="Drepanocerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="B953014FC77EFFA0FF04DB575576BF2D" box="[151,363,1308,1332]" italics="true" pageId="1" pageNumber="586">Drepanocerus falsus</emphasis>
:
<bibRefCitation id="EFB6A0ACC77EFFA0FEE8DB57565CBF2D" author="Janssens, A." box="[379,577,1308,1332]" pageId="1" pageNumber="586" pagination="1 - 118" refId="ref4838" refString="Janssens, A. (1953) Exploration du Parc National de l'Upemba. Mission G. F. De Witte (1946 - 1949). Oniticellini (Coleoptera Lamellicornia). Institut des parcs nationaux du Congo Belge, 11, 1 - 118." type="journal article" year="1953">Janssens, 1953: 36</bibRefCitation>
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, 37;
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<bibRefCitation id="EFB6A0ACC77EFFA0FDE4DB565759BF2D" author="Balthasar, V." box="[631,836,1308,1332]" pageId="1" pageNumber="586" refId="ref4466" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae (Onitini, Oniticellini, Onthophagini). Tschechoslowakischen Akademie der Wissenschaften, Prag, 627 pp." type="book" year="1963">Balthasar, 1963: 62</bibRefCitation>
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, 66;
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<bibRefCitation id="EFB6A0ACC77EFFA0FCE9DB5650D1BF2D" author="Biswas, S. &amp; Chatterjee, S. K." box="[890,1228,1308,1333]" pageId="1" pageNumber="586" pagination="147 - 177" refId="ref4650" refString="Biswas, S. &amp; Chatterjee, S. K. (1985) Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the zoological Survey of India, 84, 147 - 177." type="journal article" year="1985">Biswas &amp; Chatterjee, 1985: 151</bibRefCitation>
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<treatmentCitation id="0A86FB4CC77EFFA0FB49DB565185BF2C" author="Sewak, R." box="[1242,1432,1308,1333]" page="183" pageId="1" pageNumber="586" year="2003">
<bibRefCitation id="EFB6A0ACC77EFFA0FB49DB565185BF2C" author="Sewak, R." box="[1242,1432,1308,1333]" pageId="1" pageNumber="586" pagination="249 - 287" refId="ref5334" refString="Sewak, R. (2003) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of India with special reference to Arunachal Pradesh, Uttar Pradesh, Gujarat and Rajasthan. In: Gupta, R. K. (Ed.), Advancements in Insect Biodiversity. Agrobios, Jodhpur, pp. 249 - 287." type="book chapter" year="2003">Sewak, 2003: 183</bibRefCitation>
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;
<treatmentCitation id="0A86FB4CC77EFFA0FF54DB715560BF4A" author="Sewak, R." box="[199,381,1339,1363]" page="219" pageId="1" pageNumber="586" year="2006">
<bibRefCitation id="EFB6A0ACC77EFFA0FF54DB715560BF4A" author="Sewak, R." box="[199,381,1339,1363]" pageId="1" pageNumber="586" pagination="191 - 224" refId="ref5394" refString="Sewak, R. (2006) Insecta: Coleoptera: Scarabaeidae: Coprinae (dung beetles). In: Alfred, J. R. B. (Ed.), Zoological Survey of India. Fauna of Arunachal Pradesh. State Fauna. Series 13. Part 2. New Delhi, pp. 191 - 224." type="book chapter" year="2006">Sewak, 2006: 219</bibRefCitation>
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<treatmentCitationGroup id="AB37FA73C77EFFA0FF04DB1057CCBF88" pageId="1" pageNumber="586">
<treatmentCitation id="0A86FB4CC77EFFA0FF04DB105632BF6B" author="Simonis, A." box="[151,559,1370,1394]" page="98" pageId="1" pageNumber="586" year="1985">
<taxonomicName id="4C27A6DEC77EFFA0FF04DB105632BF6B" ID-CoL="4XL34" authority=": Simonis, 1985: 98" authorityName="Simonis" authorityPageNumber="98" authorityYear="1985" box="[151,559,1370,1394]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="B953014FC77EFFA0FF04DB105579BF6B" box="[151,356,1370,1394]" italics="true" pageId="1" pageNumber="586">Sinodrepanus falsus</emphasis>
:
<bibRefCitation id="EFB6A0ACC77EFFA0FEE1DB105632BF6B" author="Simonis, A." box="[370,559,1370,1394]" pageId="1" pageNumber="586" pagination="93 - 105" refId="ref5642" refString="Simonis, A. (1985) Un nuovo genere e tre nuove specie di Drepanocerina (Coleoptera, Scarabaeidae: Oniticellini). Revue Suisse de Zoologie, 92 (1), 93 - 105. https: // doi. org / 10.5962 / bhl. part. 81602" type="journal article" year="1985">Simonis, 1985: 98</bibRefCitation>
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, 103, 104;
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<bibRefCitation id="EFB6A0ACC77EFFA0FD31DB1057FDBF6B" author="Kabakov, O. N. &amp; Napolov, A." box="[674,992,1370,1394]" pageId="1" pageNumber="586" pagination="58 - 96" refId="ref4885" refString="Kabakov, O. N. &amp; Napolov, A. (1999) Fauna and ecology of Lamellicornia of subfamily Scarabaeinae (Scarabaeidae, Coleoptera) of Vietnam and some parts of adjacent countries: South China, Laos and Thailand. Latvijas Entomologs, 37, 58 - 96." type="journal article" year="1999">Kabakov &amp; Napolov, 1999: 73</bibRefCitation>
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<bibRefCitation id="EFB6A0ACC77EFFA0FC7FDB1050D2BF6B" author="Ochi, T. &amp; Kon, M. &amp; Masumoto, K." box="[1004,1231,1370,1394]" pageId="1" pageNumber="586" pagination="347 - 356" refId="ref5212" refString="Ochi, T., Kon, M. &amp; Masumoto, K. (2004) Two new species of the genus Sinodrepanus (Coleoptera, Scarabaeidae, Oniticellini) from China and Thailand. Elytra, Tokyo, 32 (2), 347 - 356." type="journal article" year="2004">
Ochi
<emphasis id="B953014FC77EFFA0FBB7DB115045BF6B" box="[1060,1112,1370,1394]" italics="true" pageId="1" pageNumber="586">et al.</emphasis>
, 2004: 355
</bibRefCitation>
</treatmentCitation>
; Krikken, 2009: 21;
<treatmentCitation id="0A86FB4CC77EFFA0FF54DB32561DBF89" author="Kabakov, O. N. &amp; Shokhin, I. V." box="[199,512,1400,1425]" page="58" pageId="1" pageNumber="586" year="2014">
<bibRefCitation id="EFB6A0ACC77EFFA0FF54DB32561DBF89" author="Kabakov, O. N. &amp; Shokhin, I. V." box="[199,512,1400,1425]" pageId="1" pageNumber="586" pagination="47 - 59" refId="ref4937" refString="Kabakov, O. N. &amp; Shokhin, I. V. (2014) Contribution to the knowledge of the subfamily Scarabaeinae (Coleoptera) from China with nomenclatural notes. Caucasian Entomological Bulletin, 10 (1), 47 - 59. https: // doi. org / 10.23885 / 1814 - 3326 - 2014 - 10 - 1 - 47 - 59" type="journal article" year="2014">Kabakov &amp; Shokhin, 2014: 58</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A86FB4CC77EFFA0FD98DB325715BF88" author="Mittal, I. C. &amp; Jain, R." box="[523,776,1400,1425]" page="393" pageId="1" pageNumber="586" year="2015">
<bibRefCitation id="EFB6A0ACC77EFFA0FD98DB325715BF88" author="Mittal, I. C. &amp; Jain, R." box="[523,776,1400,1425]" pageId="1" pageNumber="586" pagination="383 - 404" refId="ref5160" refString="Mittal, I. C. &amp; Jain, R. (2015) A checklist of Indian dung beetles (Coleoptera: Scarabaeidae). Indian Journal of Entomology, 77 (4), 383 - 404. https: // doi. org / 10.5958 / 0974 - 8172.2015.00076.0" type="journal article" year="2015">Mittal &amp; Jain, 2015: 393</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A86FB4CC77EFFA0FC81DB3257CCBF88" author="Bezdek, A." box="[786,977,1400,1425]" page="174" pageId="1" pageNumber="586" year="2016">
<bibRefCitation id="EFB6A0ACC77EFFA0FC81DB3257CCBF88" author="Bezdek, A." box="[786,977,1400,1425]" pageId="1" pageNumber="586" pagination="1905" refId="ref4510" refString="Bezdek, A. (2016) Tribe Oniticellini H. J. Kolbe, 1905. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. Brill, Leiden, Boston, pp. 174 - 177. https: // doi. org / 10.1163 / 9789004309142 _ 003" type="book chapter" year="2016">Bezděk, 2016: 174</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33D8ED6C77EFFA0FF04DBF55596BD90" pageId="1" pageNumber="586" type="materials_examined">
<paragraph id="8B98DD5DC77EFFA0FF04DBF55563BFC0" blockId="1.[151,1436,1470,1929]" box="[151,382,1470,1497]" pageId="1" pageNumber="586">
<materialsCitation id="3B4FD700C77EFFA0FF04DBF55567BFC0" ID-GBIF-Occurrence="3403373306" box="[151,378,1470,1497]" country="Laos" location="Laos" pageId="1" pageNumber="586" specimenCount="1" typeStatus="holotype">
<emphasis id="B953014FC77EFFA0FF04DBF55524BFC1" bold="true" box="[151,313,1470,1497]" pageId="1" pageNumber="586">
<typeStatus id="549C63FFC77EFFA0FF04DBF554CFBFC0" box="[151,210,1471,1497]" pageId="1" pageNumber="586">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3309DCDC77EFFA0FED3DBF55567BFC0" box="[320,378,1471,1497]" name="Laos" pageId="1" pageNumber="586">Laos</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B98DD5DC77EFFA0FF54DBA95149BC5C" blockId="1.[151,1436,1470,1929]" pageId="1" pageNumber="586">
<emphasis id="B953014FC77EFFA0FF54DBA955F7BFE5" bold="true" box="[199,490,1506,1533]" pageId="1" pageNumber="586">Type material examined</emphasis>
.
<emphasis id="B953014FC77EFFA0FE6ADBA9567BBFE4" bold="true" box="[505,614,1507,1533]" pageId="1" pageNumber="586">
<typeStatus id="549C63FFC77EFFA0FE6ADBA9567BBFE4" box="[505,614,1507,1533]" pageId="1" pageNumber="586" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D2116D4C77EFFA0FDE9DBA95690BFE4" box="[634,653,1507,1533]" pageId="1" pageNumber="586" type="female"></specimenCount>
):
<collectingCountry id="F3309DCDC77EFFA0FD35DBA956C1BFE4" box="[678,732,1507,1533]" name="Laos" pageId="1" pageNumber="586">Laos</collectingCountry>
/ Mouhot //
<taxonomicName id="4C27A6DEC77EFFA0FCFFDBA957FABFE4" authorityName="Le Peletier &amp; Serville" authorityYear="1828" box="[876,999,1507,1533]" class="Insecta" family="Scarabaeidae" genus="Oniticellus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">Oniticellus</taxonomicName>
/ falsus / Type D.S. // Ex Musaeo / D. Sharp 1890 // MUSÉUM PARIS / 1952 // COLL. R. OBERTHÜR // A. Janssens det. 1953 /
<taxonomicName id="4C27A6DEC77EFFA0FB3AD84D5159BC38" authorityName="Kirby" authorityYear="1828" box="[1193,1348,1543,1569]" class="Insecta" family="Scarabaeidae" genus="Drepanocerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">Drepanocerus</taxonomicName>
/
<taxonomicName id="4C27A6DEC77EFFA0FAC8D84D54C5BC5C" authority="Sharp" authorityName="Sharp" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">falsus Sharp</taxonomicName>
<specimenCount id="9D2116D4C77EFFA0FF73D86154E8BC5C" box="[224,245,1579,1605]" pageId="1" pageNumber="586" type="female"></specimenCount>
/ Type // TYPE //
<typeStatus id="549C63FFC77EFFA0FE51D861564BBC5C" box="[450,598,1579,1605]" pageId="1" pageNumber="586" type="holotype">HOLOTYPE</typeStatus>
//
<typeStatus id="549C63FFC77EFFA0FDE0D861571ABC5C" box="[627,775,1579,1605]" pageId="1" pageNumber="586" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C27A6DEC77EFFA0FC8ED86157A8BC5C" authorityName="Simonis" authorityYear="1985" box="[797,949,1579,1605]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">Sinodrepanus</taxonomicName>
/
<taxonomicName id="4C27A6DEC77EFFA0FC58D8615075BC5F" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[971,1128,1579,1606]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="falsus">falsus (Sharp)</taxonomicName>
//
<collectionCode id="ED364598C77EFFA0FB16D86150C6BC5C" box="[1157,1243,1579,1605]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="586">MNHN</collectionCode>
/ EC8103.
</paragraph>
<paragraph id="8B98DD5DC77EFFA0FF54D8045596BD90" blockId="1.[151,1436,1470,1929]" pageId="1" pageNumber="586">
<emphasis id="B953014FC77EFFA0FF54D8045632BC71" bold="true" box="[199,559,1614,1640]" pageId="1" pageNumber="586">Additional material examined</emphasis>
(
<specimenCount id="9D2116D4C77EFFA0FDD3D8055666BC70" box="[576,635,1615,1641]" pageId="1" pageNumber="586" type="male">7♂♂</specimenCount>
,
<specimenCount id="9D2116D4C77EFFA0FD1BD80556B4BC70" box="[648,681,1615,1641]" pageId="1" pageNumber="586" type="female">1♀</specimenCount>
).
<materialsCitation id="3B4FD700C77EFFA0FD50D80450A8BC94" ID-GBIF-Occurrence="3403373309" collectingDate="2003-01-19" collectingDateMax="2003-01-27" collectingDateMin="2003-01-19" collectionCode="PMOC" collectorName="N Jinghong &amp; S. Murzin" country="China" elevation="1000" location="Guangpin" municipality="Iquanbanna" pageId="1" pageNumber="586" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<emphasis id="B953014FC77EFFA0FD50D8045711BC71" bold="true" box="[707,780,1614,1640]" pageId="1" pageNumber="586">
<collectingCountry id="F3309DCDC77EFFA0FD50D8045711BC71" box="[707,780,1614,1640]" name="China" pageId="1" pageNumber="586">China</collectingCountry>
</emphasis>
: CHINA, S
<collectingRegion id="49E313BFC77EFFA0FC08D805500ABC70" box="[923,1047,1615,1641]" country="China" name="Yunnan" pageId="1" pageNumber="586">YUNNAN</collectingRegion>
,
<collectingMunicipality id="6BFC4727C77EFFA0FBB0D80550BBBC70" box="[1059,1190,1615,1641]" pageId="1" pageNumber="586">Iquanbanna</collectingMunicipality>
[likely Xishuangbanna],
<location id="8EF88B86C77EFFA0FF5BD839552BBC94" LSID="urn:lsid:plazi:treatment:038E6C4BC77EFFA2FF04DA1D55FCBEF0:8EF88B86C77EFFA0FF5BD839552BBC94" box="[200,310,1651,1677]" country="China" municipality="Iquanbanna" name="Guangpin" pageId="1" pageNumber="586" stateProvince="Yunnan">Guangpin</location>
env., 0km [
<emphasis id="B953014FC77EFFA0FE29D83955C4BC95" box="[442,473,1651,1676]" italics="true" pageId="1" pageNumber="586">sic</emphasis>
!]
<collectorName id="26D2B88BC77EFFA0FE61D839566ABC94" box="[498,631,1651,1677]" pageId="1" pageNumber="586">N Jinghong</collectorName>
,
<quantity id="4CDF70B8C77EFFA0FD12D83956CCBC94" box="[641,721,1651,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="586" unit="m" value="1000.0">
<elevation id="000A3A6EC77EFFA0FD12D83956CCBC94" box="[641,721,1651,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="586" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<date id="FF99FB9DC77EFFA0FD4FD8395772BC94" box="[732,879,1651,1677]" pageId="1" pageNumber="586" value="2003-01-19" valueMax="2003-01-27" valueMin="2003-01-19">
<collectingDate id="EFDD0275C77EFFA0FD4FD8395772BC94" box="[732,879,1651,1677]" pageId="1" pageNumber="586" value="2003-01-19" valueMax="2003-01-27" valueMin="2003-01-19">19-27.I.2003</collectingDate>
</date>
,
<collectorName id="26D2B88BC77EFFA0FCE9D83957F4BC94" box="[890,1001,1651,1677]" pageId="1" pageNumber="586">S. Murzin</collectorName>
leg. [
<specimenCount id="9D2116D4C77EFFA0FBBBD8395050BC94" box="[1064,1101,1651,1677]" pageId="1" pageNumber="586" type="male">1♂</specimenCount>
,
<collectionCode id="ED364598C77EFFA0FBCAD83850B2BC94" box="[1113,1199,1650,1677]" pageId="1" pageNumber="586">PMOC</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B4FD700C77EFFA0FB2CD83950F9BCA8" ID-GBIF-Occurrence="3403373310" collectionCode="BMNH" collectorName="R. V. de Salvaza" country="Laos" county="Laos" location="Aug" municipality="Paklay" pageId="1" pageNumber="586" specimenCount="1" specimenCount-male="1">
<emphasis id="B953014FC77EFFA0FB2CD83950E4BC94" bold="true" box="[1215,1273,1651,1677]" pageId="1" pageNumber="586">
<collectingCountry id="F3309DCDC77EFFA0FB2CD83950E4BC94" box="[1215,1273,1651,1677]" name="Laos" pageId="1" pageNumber="586">Laos</collectingCountry>
</emphasis>
:
<collectingCounty id="62F9A5D1C77EFFA0FA94D839515DBC94" box="[1287,1344,1651,1677]" pageId="1" pageNumber="586">Laos</collectingCounty>
,
<collectingMunicipality id="6BFC4727C77EFFA0FAD8D8395185BC94" box="[1355,1432,1651,1677]" pageId="1" pageNumber="586">Paklay</collectingMunicipality>
,
<location id="8EF88B86C77EFFA0FF04D8DC54D7BCA8" LSID="urn:lsid:plazi:treatment:038E6C4BC77EFFA2FF04DA1D55FCBEF0:8EF88B86C77EFFA0FF04D8DC54D7BCA8" box="[151,202,1686,1713]" country="Laos" county="Laos" municipality="Paklay" name="Aug" pageId="1" pageNumber="586">Aug</location>
[ust] 1917,
<collectorName id="26D2B88BC77EFFA0FEDFD8DD561FBCA8" box="[332,514,1687,1713]" pageId="1" pageNumber="586">R.V. de Salvaza</collectorName>
[
<specimenCount id="9D2116D4C77EFFA0FD80D8DD5625BCA8" box="[531,568,1687,1713]" pageId="1" pageNumber="586" type="male">1♂</specimenCount>
,
<collectionCode id="ED364598C77EFFA0FDD4D8DD5682BCA8" box="[583,671,1687,1713]" pageId="1" pageNumber="586">BMNH</collectionCode>
; specimen probably seen also by
<bibRefCitation id="EFB6A0ACC77EFFA0FBB0D8DD50C0BCA8" author="Janssens, A." box="[1059,1245,1686,1713]" pageId="1" pageNumber="586" pagination="1 - 118" refId="ref4838" refString="Janssens, A. (1953) Exploration du Parc National de l'Upemba. Mission G. F. De Witte (1946 - 1949). Oniticellini (Coleoptera Lamellicornia). Institut des parcs nationaux du Congo Belge, 11, 1 - 118." type="journal article" year="1953">Janssens (1953)</bibRefCitation>
]
</materialsCitation>
.
<materialsCitation id="3B4FD700C77EFFA0FB62D8DC5057BCCC" ID-GBIF-Occurrence="3403373308" collectingDate="2008-06-20" collectingDateMax="2019-07-15" collectingDateMin="2008-06-20" collectionCode="BMNH" country="Thailand" county="Ban Tao Phan" latitude="19.558107" location="Ban Tao Phan" longLatPrecision="1" longitude="99.07593" municipality="Chiang Dao" pageId="1" pageNumber="586" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai province">
<collectingCountry id="F3309DCDC77EFFA0FB62D8DC5143BCA9" box="[1265,1374,1686,1712]" name="Thailand" pageId="1" pageNumber="586">
<emphasis id="B953014FC77EFFA0FB62D8DC5143BCA9" bold="true" box="[1265,1374,1686,1712]" pageId="1" pageNumber="586">Thailand</emphasis>
</collectingCountry>
:
<collectingCounty id="62F9A5D1C77EFFA0FAFCD8DD551EBCCC" pageId="1" pageNumber="586">Ban Tao Phan</collectingCounty>
, 11.III.[19]18 [likely Ban Phan Tao in
<collectingRegion id="49E313BFC77EFFA0FD2AD8F157B1BCCC" box="[697,940,1723,1749]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="586">Chiang Mai province</collectingRegion>
;
<specimenCount id="9D2116D4C77EFFA0FC24D8F157C6BCCC" box="[951,987,1723,1749]" pageId="1" pageNumber="586" type="male">1♂</specimenCount>
,
<collectionCode id="ED364598C77EFFA0FC7AD8F0505EBCCC" box="[1001,1091,1722,1749]" pageId="1" pageNumber="586">BMNH</collectionCode>
]
</materialsCitation>
<materialsCitation id="3B4FD700C77EFFA0FBDAD8F05529BD04" ID-GBIF-Occurrence="3403373307" collectingDate="2019-07-02" collectingDateMax="2019-07-11" collectingDateMin="2019-07-02" collectionCode="GMOC" collectorName="Vishnyakov Alexey" country="Thailand" county="Ban Tao Phan" latitude="19.558107" location="Chiang Dao Hill Resort" longLatPrecision="1" longitude="99.07593" municipality="Chiang Dao" pageId="1" pageNumber="586" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai">
; N
<collectingCountry id="F3309DCDC77EFFA0FBE3D8F15119BCCC" box="[1136,1284,1723,1749]" pageId="1" pageNumber="586">THAILAND</collectingCountry>
,
<collectingRegion id="49E313BFC77EFFA0FA83D8F15185BCCC" box="[1296,1432,1723,1749]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="586">Chiang Mai</collectingRegion>
,
<collectingMunicipality id="6BFC4727C77EFFA0FF04D895553CBCE0" box="[151,289,1759,1785]" pageId="1" pageNumber="586">Chiang Dao</collectingMunicipality>
,
<location id="8EF88B86C77EFFA0FEBED8955625BCE0" LSID="urn:lsid:plazi:treatment:038E6C4BC77EFFA2FF04DA1D55FCBEF0:8EF88B86C77EFFA0FEBED8955625BCE0" box="[301,568,1759,1785]" country="Thailand" county="Ban Tao Phan" latitude="19.558107" longLatPrecision="1" longitude="99.07593" municipality="Chiang Dao" name="Chiang Dao Hill Resort" pageId="1" pageNumber="586" stateProvince="Chiang Mai">Chiang Dao Hill Resort</location>
env.,
<geoCoordinate id="EE13BB9AC77EFFA0FDEFD895573ABCE0" box="[636,807,1759,1785]" degrees="19" direction="north" minutes="33" orientation="latitude" pageId="1" pageNumber="586" precision="1" seconds="28.37" value="19.55788">19°3328.37”N</geoCoordinate>
<geoCoordinate id="EE13BB9AC77EFFA0FCBCD89557D0BCE0" box="[815,973,1759,1785]" degrees="99" direction="east" minutes="4" orientation="longitude" pageId="1" pageNumber="586" precision="1" seconds="34.20" value="99.076164">99°434.20”E</geoCoordinate>
,
<date id="FF99FB9DC77EFFA0FC4AD8955066BCE0" box="[985,1147,1759,1785]" pageId="1" pageNumber="586" value="2019-07-02" valueMax="2019-07-11" valueMin="2019-07-02">
<collectingDate id="EFDD0275C77EFFA0FC4AD8955066BCE0" box="[985,1147,1759,1785]" pageId="1" pageNumber="586" value="2019-07-02" valueMax="2019-07-11" valueMin="2019-07-02">2-11.VII.2019</collectingDate>
</date>
,
<collectorName id="26D2B88BC77EFFA0FB15D895517EBCE0" box="[1158,1379,1759,1785]" pageId="1" pageNumber="586">Vishnyakov Alexey</collectorName>
legit [
<specimenCount id="9D2116D4C77EFFA0FF33D94954D9BD04" box="[160,196,1795,1821]" pageId="1" pageNumber="586" type="male">1♂</specimenCount>
,
<collectionCode id="ED364598C77EFFA0FF40D9485530BD04" box="[211,301,1794,1821]" pageId="1" pageNumber="586">GMOC</collectionCode>
]
</materialsCitation>
<materialsCitation id="3B4FD700C77EFFA0FEA7D94854E5BD58" ID-GBIF-Occurrence="3403373303" collectingDate="2017-06-14" collectingDateMax="2019-07-15" collectingDateMin="2008-06-20" collectionCode="GMOC" collectorName="Prosvirov" country="Thailand" county="Ban Tao Phan" latitude="19.558107" location="Chiang Mai" longLatPrecision="1" longitude="99.07593" municipality="Chiang Dao" pageId="1" pageNumber="586" specimenCount="1" specimenCount-female="1" specimenCount-male="5" stateProvince="Chiang Mai">
;
<collectingCountry id="F3309DCDC77EFFA0FED2D94955C8BD04" box="[321,469,1795,1821]" pageId="1" pageNumber="586">THAILAND</collectingCountry>
,
<collectingRegion id="49E313BFC77EFFA0FE72D9495676BD04" box="[481,619,1795,1821]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="586">Chiang Mai</collectingRegion>
, Chiang Dao,
<geoCoordinate id="EE13BB9AC77EFFA0FC83D9495784BD04" box="[784,921,1795,1821]" degrees="19" direction="north" minutes="33" orientation="latitude" pageId="1" pageNumber="586" precision="15" seconds="28" value="19.557777">19°3328”N</geoCoordinate>
<geoCoordinate id="EE13BB9AC77EFFA0FC31D9495001BD04" box="[930,1052,1795,1821]" degrees="99" direction="east" minutes="4" orientation="longitude" pageId="1" pageNumber="586" precision="15" seconds="34" value="99.07611">99°434”E</geoCoordinate>
,
<date id="FF99FB9DC77EFFA0FBBAD94950B6BD04" box="[1065,1195,1795,1821]" pageId="1" pageNumber="586" value="2017-06-14">
<collectingDate id="EFDD0275C77EFFA0FBBAD94950B6BD04" box="[1065,1195,1795,1821]" pageId="1" pageNumber="586" value="2017-06-14">14.VI.2017</collectingDate>
</date>
,
<collectorName id="26D2B88BC77EFFA0FB2AD949513BBD04" box="[1209,1318,1795,1821]" pageId="1" pageNumber="586">Prosvirov</collectorName>
legit [
<specimenCount id="9D2116D4C77EFFA0FAE0D949518BBD04" box="[1395,1430,1795,1821]" pageId="1" pageNumber="586" type="female">1♀</specimenCount>
,
<collectionCode id="ED364598C77EFFA0FF04D96C54ECBD58" box="[151,241,1830,1857]" pageId="1" pageNumber="586">GMOC</collectionCode>
]
</materialsCitation>
<materialsCitation id="3B4FD700C77EFFA0FF6BD96C55F8BD7C" ID-GBIF-Occurrence="3403373302" collectingDate="2019-07-01" collectingDateMax="2019-07-15" collectingDateMin="2019-07-01" collectionCode="GMOC" collectorName="Ustinov V." country="Thailand" county="Ban Tao Phan" latitude="19.558107" location="Chiangmai" longLatPrecision="1" longitude="99.07593" municipality="Chiang Dao" pageId="1" pageNumber="586" specimenCount="2" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiangmai">
;
<collectingCountry id="F3309DCDC77EFFA0FE96D96D5584BD58" box="[261,409,1831,1857]" pageId="1" pageNumber="586">THAILAND</collectingCountry>
,
<collectingRegion id="49E313BFC77EFFA0FE35D96D5639BD58" box="[422,548,1831,1857]" pageId="1" pageNumber="586">Chiangmai</collectingRegion>
, Chi[a]ngdao Hill Resort,
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,
<collectorName id="26D2B88BC77EFFA0FAE7D96D54ECBD7C" pageId="1" pageNumber="586">Ustinov V.</collectorName>
legit [
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; N
<collectingCountry id="F3309DCDC77EFFA0FD9CD90156BEBD7C" box="[527,675,1867,1893]" pageId="1" pageNumber="586">THAILAND</collectingCountry>
,
<quantity id="4CDF70B8C77EFFA0FD3CD90156E3BD7C" box="[687,766,1867,1893]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="586" unit="km" value="100.0">
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N
<collectingRegion id="49E313BFC77EFFA0FCB0D90157B1BD7C" box="[803,940,1867,1893]" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="586">Chiang Mai</collectingRegion>
,
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,
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<collectingDate id="EFDD0275C77EFFA0FB42D9015164BD7C" box="[1233,1401,1867,1893]" pageId="1" pageNumber="586" value="2008-06-20" valueMax="2008-06-30" valueMin="2008-06-20">20-30.VI.2008</collectingDate>
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,
<collectorName id="26D2B88BC77EFFA0FA15D90154F4BD90" pageId="1" pageNumber="586">S. Murzin</collectorName>
[
<specimenCount id="9D2116D4C77EFFA0FF6AD9255503BD90" box="[249,286,1903,1929]" pageId="1" pageNumber="586" type="male">1♂</specimenCount>
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.
</paragraph>
</subSubSection>
<caption id="DF588DD5C77DFFA3FF04DABA5177BF35" ID-DOI="http://doi.org/10.5281/zenodo.5683453" ID-Zenodo-Dep="5683453" httpUri="https://zenodo.org/record/5683453/files/figure.png" pageId="2" pageNumber="587" startId="2.[151,264,1264,1288]" targetBox="[191,1394,188,1229]" targetPageId="2">
<paragraph id="8B98DD5DC77DFFA3FF04DABA5177BF35" blockId="2.[151,1436,1264,1324]" pageId="2" pageNumber="587">
<emphasis id="B953014FC77DFFA3FF04DABA5521BF11" bold="true" box="[151,316,1264,1288]" pageId="2" pageNumber="587">FIGURES 1, 2.</emphasis>
<taxonomicName id="4C27A6DEC77DFFA3FED0DABA5609BF11" box="[323,532,1264,1288]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B953014FC77DFFA3FED0DABA55D0BF11" box="[323,461,1264,1288]" italics="true" pageId="2" pageNumber="587">Sinodrepanus</emphasis>
spp. 1
</taxonomicName>
: habitus of a male specimen of
<taxonomicName id="4C27A6DEC77DFFA3FCC8DABA505EBF11" authority="(Sharp, 1875)" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[859,1091,1264,1288]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="B953014FC77DFFA3FCC8DABA57ADBF11" box="[859,944,1264,1288]" italics="true" pageId="2" pageNumber="587">S. falsus</emphasis>
(
<bibRefCitation id="EFB6A0ACC77DFFA3FC2DDABA5021BF11" author="Sharp, D." box="[958,1084,1264,1288]" pageId="2" pageNumber="587" pagination="33 - 54" refId="ref5611" refString="Sharp, D. (1875) Description of some new genera and species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 13, 33 - 54." type="journal article" year="1875">Sharp, 1875</bibRefCitation>
)
</taxonomicName>
from Thailand (picture by Marco Uliana); 2: habitus of the male holotype of
<taxonomicName id="4C27A6DEC77DFFA3FDDEDB5E5733BF35" authority="(Sharp, 1875)" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[589,814,1300,1324]" class="Insecta" family="Scarabaeidae" genus="Sinodrepanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="exsul">
<emphasis id="B953014FC77DFFA3FDDEDB5E5687BF35" box="[589,666,1300,1324]" italics="true" pageId="2" pageNumber="587">S. exsul</emphasis>
(
<bibRefCitation id="EFB6A0ACC77DFFA3FD3BDB5E573ABF35" author="Sharp, D." box="[680,807,1300,1324]" pageId="2" pageNumber="587" pagination="33 - 54" refId="ref5611" refString="Sharp, D. (1875) Description of some new genera and species of Scarabaeidae from tropical Asia and Malaysia. Coleopterologische Hefte, 13, 33 - 54." type="journal article" year="1875">Sharp, 1875</bibRefCitation>
)
</taxonomicName>
, from Thailand (picture by Antoine Mantilleri, MNHN).
</paragraph>
</caption>
<subSubSection id="C33D8ED6C77DFFA2FF54DB10514ABE08" lastPageId="3" lastPageNumber="588" pageId="2" pageNumber="587" type="description">
<paragraph id="8B98DD5DC77DFFA2FF54DB10514ABE08" blockId="2.[151,1437,1370,2009]" lastBlockId="3.[151,1437,151,1257]" lastPageId="3" lastPageNumber="588" pageId="2" pageNumber="587">
<emphasis id="B953014FC77DFFA3FF54DB1055C0BF6D" bold="true" box="[199,477,1370,1396]" pageId="2" pageNumber="587">Description of the male</emphasis>
(
<figureCitation id="131CC1D8C77DFFA3FE7FDB105627BF6D" box="[492,570,1370,1396]" captionStart="FIGURES 1, 2" captionStartId="2.[151,264,1264,1288]" captionTargetBox="[191,1394,188,1229]" captionTargetId="figure-19@2.[177,1411,179,1238]" captionTargetPageId="2" captionText="FIGURES 1, 2. Sinodrepanus spp. 1: habitus of a male specimen of S. falsus (Sharp, 1875) from Thailand (picture by Marco Uliana); 2: habitus of the male holotype of S. exsul (Sharp, 1875), from Thailand (picture by Antoine Mantilleri, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5683453" httpUri="https://zenodo.org/record/5683453/files/figure.png" pageId="2" pageNumber="587">Figs. 1</figureCitation>
,
<figureCitation id="131CC1D8C77DFFA3FDDBDB10566BBF6D" box="[584,630,1370,1396]" captionStart="FIGURES 39" captionStartId="4.[151,264,1228,1252]" captionTargetBox="[157,1414,199,1205]" captionTargetId="figure-19@4.[151,1435,166,1205]" captionTargetPageId="4" captionText="FIGURES 39. Sinodrepanus spp.—S. falsus (Sharp, 1875), specimens from Thailand (37) (pictures by Marco Uliana). 3: female forebody; 4: male metasternum; 5: lamella copulatrix; 6: parameres in dorsal view; 7: aedeagus in lateral view.—S. exsul (Sharp, 1875), male holotype from Thailand (8, 9) (pictures by Christophe Rivier, MNHN). 8: parameres in dorsal view; 9: aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5683455" httpUri="https://zenodo.org/record/5683455/files/figure.png" pageId="2" pageNumber="587">47</figureCitation>
). Flat, elongated, integument microreticulated, dark brown and partially with metallic lustre; body surface almost wholly covered by simple or ocellate punctures which carry light brown scale-like bristles.
<emphasis id="B953014FC77DFFA3FEFDDBE855D9BFA5" bold="true" box="[366,452,1442,1468]" pageId="2" pageNumber="587">Length</emphasis>
: 8.29.8 mm (N=7).
<emphasis id="B953014FC77DFFA3FD2EDBE856E0BFA5" bold="true" box="[701,765,1442,1468]" pageId="2" pageNumber="587">Head</emphasis>
. Subhexagonal, clypeus subtrapezoidal, covered by small, ocellate, setigerous punctures. Frontovertex with two short longitudinal median keels carrying long testaceous bristles; keels dividing the interocular space into three almost equal segments. A third short keel runs along the frontoclypeal suture, bulging especially in the middle and towards the anterior part of clypeus, connecting anteriorly the two longitudinal keels and delimiting with them a square surface. Maximum head width at about anterior 1/3 of genae. External edge of genae slightly obtusely angular. Anterior edge of clypeus reflexed and slightly depressed transversely behind, bearing two upturned triangular denticles separated by a largely obtuse angle, this angle narrower and more angular than in females.
<emphasis id="B953014FC77DFFA3FDE9D8D456EEBCA1" bold="true" box="[634,755,1694,1720]" pageId="2" pageNumber="587">Pronotum</emphasis>
. Subquadrangular, widening anteriorly, maximum width at about anterior 1/3, where lateral edges are slightly and broadly angular. Lateral edges slightly and broadly sinuous on their basal third. Anterior angles somehow truncated anteriorly; posterior angles broadly rounded. Covered with ocellate setigerous punctures of uneven size, largest on the median basal part of disc. Disc with eight longitudinal keels carrying long testaceous bristles (in order, from medial to lateral): two keels on either side of the midline, reaching the anterior pronotal border but expiring before posterior edge, converging at about the anterior 2/5 of their length, where they are connected by a very short transverse keel. External to these, two shorter keels almost reaching the posterior pronotal border but ending clearly before the anterior one, connected anteriorly to the medial keels by two weak oblique keels, and almost devoid of long bristles in their middle. Other two thick keels originate from the anterolateral corners of pronotum, obliquely reaching the anterior end of the previous keels. Finally, two short keels are present on the basal 1/3 of lateral edge, they are placed close to the pronotal edge and do not reach the posterior border.
<emphasis id="B953014FC77CFFA2FE14DE9555CEBAE0" bold="true" box="[391,467,223,249]" pageId="3" pageNumber="588">Elytra</emphasis>
. Flat on the whole, subquadrangular. Striae broad, sinuous except the 1st one which is straight, their borders clearly ridged, punctures marked, each one carrying a scale-like seta. Interstriae covered by small and dense setigerous punctures; the 1st costate, the 5th strongly but gradually raised, the 7th ridge-like and concealing the 8th in dorsal view. The 1st, 3rd, 5th and 7th interstriae clothed in progressively longer, more abundant and thicker setae, with a tuft of long setae apically. Interstriae 5, 6 and 7 obliquely elevated near humeral callus, the 7th depressed a little behind the callus. The 3rd interstria becomes progressively narrow towards the apex, then widens again near the apex; the 6th interstria widens in basal third, then remains more or less parallel on the remaining two thirds. Scale-like bristles on epipleura gradually disappearing posteriorly. Scutellum lingulate, smooth.
<emphasis id="B953014FC77CFFA2FF69DFB55524B800" bold="true" box="[250,313,511,537]" pageId="3" pageNumber="588">Legs.</emphasis>
Ventral surfaces of femurs covered by dense small setigerous punctures and by scattered ocellate punctures carrying thick bristles. Dorsal surface of fore legs glabrous and quite shiny. Protibiae obliquely truncated at apex, apical edge linear; with two teeth on outer edge, the distal one acute-angled, the proximal one much broader. Apical spur with distal half bent downwards.
<emphasis id="B953014FC77CFFA2FD32DC205701B89D" bold="true" box="[673,796,618,644]" pageId="3" pageNumber="588">Pygidium.</emphasis>
Densely setigerous, longer in males than females, more broadly curved at apex, with the same two tubercules and a longitudinal basal keel, all carrying long thick bristles. Strongly depressed along basal border.
<emphasis id="B953014FC77CFFA2FDDEDCF8574FB8D5" bold="true" box="[589,850,690,717]" pageId="3" pageNumber="588">Inferior body surface.</emphasis>
Abdominal sternites evenly covered by dense, small setigerous punctures, the posterior sternites with a tuft of long bristles on their distal edge. Metasternum (
<figureCitation id="131CC1D8C77CFFA2FADCDC9D5189B8E8" box="[1359,1428,727,753]" captionStart="FIGURES 39" captionStartId="4.[151,264,1228,1252]" captionTargetBox="[157,1414,199,1205]" captionTargetId="figure-19@4.[151,1435,166,1205]" captionTargetPageId="4" captionText="FIGURES 39. Sinodrepanus spp.—S. falsus (Sharp, 1875), specimens from Thailand (37) (pictures by Marco Uliana). 3: female forebody; 4: male metasternum; 5: lamella copulatrix; 6: parameres in dorsal view; 7: aedeagus in lateral view.—S. exsul (Sharp, 1875), male holotype from Thailand (8, 9) (pictures by Christophe Rivier, MNHN). 8: parameres in dorsal view; 9: aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5683455" httpUri="https://zenodo.org/record/5683455/files/figure.png" pageId="3" pageNumber="588">Fig. 4</figureCitation>
) covered by ocellate punctures of uneven size; its basal part with a deep pentagonal hollow in which punctures are sparser, delimited anteriorly by dense, coricate, thick and sharp bristles.
<emphasis id="B953014FC77CFFA2FC52DD54502AB921" bold="true" box="[961,1079,798,824]" pageId="3" pageNumber="588">Genitalia.</emphasis>
Basolateral plates of parameres greatly developed, almost touching parameral apices; apices of parameres relatively short, truncated, divergent and, in dorsal view, directed obliquely downwards; parameral apices not reaching the inferior edge of basolateral plates in lateral view (
<figureCitation id="131CC1D8C77CFFA2FED5DDC155ACB9BC" box="[326,433,907,933]" captionStart="FIGURES 39" captionStartId="4.[151,264,1228,1252]" captionTargetBox="[157,1414,199,1205]" captionTargetId="figure-19@4.[151,1435,166,1205]" captionTargetPageId="4" captionText="FIGURES 39. Sinodrepanus spp.—S. falsus (Sharp, 1875), specimens from Thailand (37) (pictures by Marco Uliana). 3: female forebody; 4: male metasternum; 5: lamella copulatrix; 6: parameres in dorsal view; 7: aedeagus in lateral view.—S. exsul (Sharp, 1875), male holotype from Thailand (8, 9) (pictures by Christophe Rivier, MNHN). 8: parameres in dorsal view; 9: aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5683455" httpUri="https://zenodo.org/record/5683455/files/figure.png" pageId="3" pageNumber="588">Figs. 6, 7</figureCitation>
). Lamella copulatrix (LC) (
<figureCitation id="131CC1D8C77CFFA2FD75DDC15734B9BC" box="[742,809,906,933]" captionStart="FIGURES 39" captionStartId="4.[151,264,1228,1252]" captionTargetBox="[157,1414,199,1205]" captionTargetId="figure-19@4.[151,1435,166,1205]" captionTargetPageId="4" captionText="FIGURES 39. Sinodrepanus spp.—S. falsus (Sharp, 1875), specimens from Thailand (37) (pictures by Marco Uliana). 3: female forebody; 4: male metasternum; 5: lamella copulatrix; 6: parameres in dorsal view; 7: aedeagus in lateral view.—S. exsul (Sharp, 1875), male holotype from Thailand (8, 9) (pictures by Christophe Rivier, MNHN). 8: parameres in dorsal view; 9: aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5683455" httpUri="https://zenodo.org/record/5683455/files/figure.png" pageId="3" pageNumber="588">Fig. 5</figureCitation>
) four-lobed; superior left lobe (SL) with a dorsally-bent expansion; superior right lobe (SR) simple, subtriangular, with a small and a little hooked lobule internally, in correspondence of which SL and SR are fused; inferior left lobe (IL) separated from the rest of the LC, elongated, with an incospicuous additional sclerite on its left; inferior right lobe (IR) very slender and strongly bent inwards.
</paragraph>
</subSubSection>
<subSubSection id="C33D8ED6C77CFFA2FF54DA505640BEDC" pageId="3" pageNumber="588" type="discussion">
<paragraph id="8B98DD5DC77CFFA2FF54DA505640BEDC" blockId="3.[151,1437,151,1257]" pageId="3" pageNumber="588">
<emphasis id="B953014FC77CFFA2FF54DA505555BE2D" bold="true" box="[199,328,1050,1076]" pageId="3" pageNumber="588">Comments</emphasis>
. Externally, males are almost identical to females (
<figureCitation id="131CC1D8C77CFFA2FC15DA5157CEBE2C" box="[902,979,1051,1077]" captionStart="FIGURES 1, 2" captionStartId="2.[151,264,1264,1288]" captionTargetBox="[191,1394,188,1229]" captionTargetId="figure-19@2.[177,1411,179,1238]" captionTargetPageId="2" captionText="FIGURES 1, 2. Sinodrepanus spp. 1: habitus of a male specimen of S. falsus (Sharp, 1875) from Thailand (picture by Marco Uliana); 2: habitus of the male holotype of S. exsul (Sharp, 1875), from Thailand (picture by Antoine Mantilleri, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5683453" httpUri="https://zenodo.org/record/5683453/files/figure.png" pageId="3" pageNumber="588">Figs. 1</figureCitation>
and
<figureCitation id="131CC1D8C77CFFA2FB99DA515004BE2C" box="[1034,1049,1051,1077]" captionStart="FIGURES 39" captionStartId="4.[151,264,1228,1252]" captionTargetBox="[157,1414,199,1205]" captionTargetId="figure-19@4.[151,1435,166,1205]" captionTargetPageId="4" captionText="FIGURES 39. Sinodrepanus spp.—S. falsus (Sharp, 1875), specimens from Thailand (37) (pictures by Marco Uliana). 3: female forebody; 4: male metasternum; 5: lamella copulatrix; 6: parameres in dorsal view; 7: aedeagus in lateral view.—S. exsul (Sharp, 1875), male holotype from Thailand (8, 9) (pictures by Christophe Rivier, MNHN). 8: parameres in dorsal view; 9: aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5683455" httpUri="https://zenodo.org/record/5683455/files/figure.png" pageId="3" pageNumber="588">3</figureCitation>
), except for the following secondary sexual characters: females have three protibial teeth instead of two (but see also the discussion below); their cylpeal teeth are sharper and the space between them is broader and more evenly rounded than in males; their metasternum is only shallowly hollowed posteriorly, the hollow being less defined than in males; their pygidium is shorter. For species diagnosis see below.
</paragraph>
</subSubSection>
<subSubSection id="C33D8ED6C77CFFA2FF54DA8455FCBEF0" box="[199,481,1230,1257]" pageId="3" pageNumber="588" type="distribution">
<paragraph id="8B98DD5DC77CFFA2FF54DA8455FCBEF0" blockId="3.[151,1437,151,1257]" box="[199,481,1230,1257]" pageId="3" pageNumber="588">
<emphasis id="B953014FC77CFFA2FF54DA845544BEF1" bold="true" box="[199,345,1230,1256]" pageId="3" pageNumber="588">Distribution</emphasis>
. See below.
</paragraph>
</subSubSection>
</treatment>
</document>