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<document id="FA11F0001A5170C5FDE699F818A4D113" ID-DOI="10.5281/zenodo.3677235" ID-GBIF-Dataset="1ee19d5d-27cf-4c93-989c-3fe3eb467203" ID-ISSN="1942-1354" ID-Zenodo-Dep="3677235" ID-ZooBank="BA35690A-FC73-4E5A-A805-FE9550275FEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582218729488" checkinUser="felipe" docAuthor="Cong, Qian, Zhang, Jing, Shen, Jinhui &amp; Grishin, Nick V." docDate="2019" docId="038E29227E7DFFEEFF3F75D4DCDBF8CA" docLanguage="en" docName="InsectaMundi.0731.1-56.pdf.imf" docOrigin="Insecta Mundi 731 (731)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Anaxas Cong &amp; Zhang &amp; Shen &amp; Grishin 2019, new genus" docType="treatment" docUuid="FE71A883-25C6-491B-B1BE-3EC4F82C62B1" docUuidSource="ZooBank" docVersion="11" lastPageNumber="15" masterDocId="FFB7515A7E72FFFEFFFF732ADF1EFFFB" masterDocTitle="Fifty new genera of Hesperiidae (Lepidoptera)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="14" updateTime="1698720609646" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="75125E9F7DD234CA8C180158BABAA023">Fifty new genera of Hesperiidae (Lepidoptera)</mods:title>
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<treatment id="038E29227E7DFFEEFF3F75D4DCDBF8CA" ID-DOI="http://doi.org/10.5281/zenodo.3681697" ID-GBIF-Taxon="162238027" ID-Zenodo-Dep="3681697" LSID="urn:lsid:zoobank.org:act:FE71A883-25C6-491B-B1BE-3EC4F82C62B1" httpUri="http://treatment.plazi.org/id/038E29227E7DFFEEFF3F75D4DCDBF8CA" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14">
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<paragraph id="8B9898347E7DFFF1FF3F75D4DD28F8ED" blockId="15.[192,566,1790,1814]" box="[192,566,1790,1814]" pageId="15" pageNumber="14">
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<emphasis id="B95344267E7DFFF1FF3F75D4DD28F8ED" bold="true" box="[192,566,1790,1814]" pageId="15" pageNumber="14">
<taxonomicName id="4C27E3B77E7DFFF1FF3F75D4DE89F8ED" authority="Grishin" authorityName="Cong &amp; Zhang &amp; Shen &amp; Grishin" authorityYear="2019" box="[192,407,1790,1814]" class="Insecta" family="Hesperiidae" genus="Anaxas" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95344267E7DFFF1FF3F75D4DE3BF8ED" bold="true" box="[192,293,1790,1814]" italics="true" pageId="15" pageNumber="14">Anaxas</emphasis>
Grishin
</taxonomicName>
,
<taxonomicNameLabel id="A260F95D7E7DFFF1FE5A75D4DD28F8ED" box="[421,566,1790,1814]" pageId="15" pageNumber="14" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C33DCBBF7E7DFFF1FF3F7401DCCDF8B8" box="[192,979,1835,1859]" pageId="15" pageNumber="14" type="description">
<paragraph id="8B9898347E7DFFF1FF3F7401DCCDF8B8" blockId="15.[192,979,1835,1859]" box="[192,979,1835,1859]" pageId="15" pageNumber="14">
http://zoobank.org/
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<paragraph id="8B9898347E7DFFF1FF3F745ADDEBF872" blockId="15.[192,757,1904,1929]" box="[192,757,1904,1929]" pageId="15" pageNumber="14">
<emphasis id="B95344267E7DFFF1FF3F745ADE69F873" bold="true" box="[192,375,1904,1928]" pageId="15" pageNumber="14">
<typeStatus id="549C26967E7DFFF1FF3F745ADE1AF873" box="[192,260,1904,1928]" pageId="15" pageNumber="14">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C27E3B77E7DFFF1FE81745ADDEFF872" ID-CoL="95WVF" authority="Plotz, 1884" authorityName="Plotz" authorityYear="1884" box="[382,753,1904,1929]" class="Insecta" family="Hesperiidae" genus="Antigonus" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis id="B95344267E7DFFF1FE81745ADD40F873" box="[382,606,1904,1928]" italics="true" pageId="15" pageNumber="14">Antigonus obliqua</emphasis>
Plötz, 1884
</taxonomicName>
.
</paragraph>
<caption id="DF58C8BC7E62FFEEFF3F77B0DCA4FB4B" ID-DOI="http://doi.org/10.5281/zenodo.3677247" ID-Zenodo-Dep="3677247" box="[192,954,1178,1200]" httpUri="https://zenodo.org/record/3677247/files/figure.png" pageId="16" pageNumber="15" startId="16.[192,275,1178,1200]" targetBox="[192,1438,214,1144]" targetPageId="16">
<paragraph id="8B9898347E62FFEEFF3F77B0DCA4FB4B" blockId="16.[192,954,1178,1200]" box="[192,954,1178,1200]" pageId="16" pageNumber="15">
<emphasis id="B95344267E62FFEEFF3F77B0DE31FB54" bold="true" box="[192,303,1178,1200]" pageId="16" pageNumber="15">Figure 6.</emphasis>
Genomic trees of
<taxonomicName id="4C27E3B77E62FFEEFE0B77B0DD45FB4B" box="[500,603,1178,1200]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="tribe" tribe="Erynnini">Erynnini</taxonomicName>
. See Fig. 2 legend for notations.
</paragraph>
</caption>
<paragraph id="8B9898347E62FFEEFF3F77C2DC60F99A" blockId="16.[192,1440,1256,1633]" pageId="16" pageNumber="15">
<emphasis id="B95344267E62FFEEFF3F77C2DE4DFAFB" bold="true" box="[192,339,1256,1280]" pageId="16" pageNumber="15">Definition.</emphasis>
A likely sister genus to
<taxonomicName id="4C27E3B77E62FFEEFD6B77C3DCE7FAFB" ID-CoL="92KHC" authority="Lindsey, 1925" authorityName="Lindsey" authorityYear="1925" box="[660,1017,1256,1280]" class="Insecta" family="Hesperiidae" genus="Potamanaxas" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E62FFEEFD6B77C3DC25FAFB" box="[660,827,1257,1280]" italics="true" pageId="16" pageNumber="15">Potamanaxas</emphasis>
Lindsey, 1925
</taxonomicName>
, and thus not monophyletic with
<taxonomicName id="4C27E3B77E62FFEEFF3F7622DEEAFADA" ID-CoL="8ZZV2" authority="Hubner, [1824]" authorityName="Hubner" authorityYear="1824" box="[192,500,1288,1313]" class="Insecta" family="Hesperiidae" genus="Anastrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E62FFEEFF3F7622DE31FAE4" box="[192,303,1288,1311]" italics="true" pageId="16" pageNumber="15">Anastrus</emphasis>
Hübner, [1824]
</taxonomicName>
(
<typeStatus id="549C26967E62FFEEFDFA7623DD25FADA" box="[517,571,1289,1313]" pageId="16" pageNumber="15">type</typeStatus>
species
<taxonomicName id="4C27E3B77E62FFEEFD5B7622DB51FADA" ID-CoL="DHMY" authority="Hubner, [1824]" authorityName="Hubner" authorityYear="1824" box="[676,1103,1288,1313]" class="Insecta" family="Hesperiidae" genus="Anastrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B95344267E62FFEEFD5B7622DC99FADB" box="[676,903,1288,1312]" italics="true" pageId="16" pageNumber="15">Anastrus obscurus</emphasis>
Hübner, [1824]
</taxonomicName>
), where these species were formerly placed (
<figureCitation id="131C84B17E62FFEEFE707602DECBFABB" box="[399,469,1320,1344]" captionStart="Figure 6" captionStartId="16.[192,275,1178,1200]" captionTargetBox="[192,1438,214,1144]" captionTargetId="figure@16.[192,1440,213,1144]" captionTargetPageId="16" captionText="Figure 6. Genomic trees of Erynnini. See Fig. 2 legend for notations." figureDoi="http://doi.org/10.5281/zenodo.3677247" httpUri="https://zenodo.org/record/3677247/files/figure.png" pageId="16" pageNumber="15">Fig. 6</figureCitation>
). The need for this genus has been suggested previously (
<bibRefCitation id="EFB6E5C57E62FFEEFB687602DA2FFABA" author="Austin" box="[1175,1329,1320,1345]" firstAuthor="Austin" pageId="16" pageNumber="15" pagination="19 - 25" refId="ref26322" refString="Austin, G. T. 1998. Hesperiidae of Rondonia, Brazil: Anastrus and Tosta, with descriptions of two new species (Lepidoptera: Hesperiidae: Pyrginae). Tropical Lepidoptera 9 (suppl. 2): 19 - 25." type="journal article" year="1998">Austin 1998</bibRefCitation>
). Keys to F.6.3. in
<bibRefCitation id="EFB6E5C57E62FFEEFED77663DED4FA9B" author="Evans" box="[296,458,1352,1376]" firstAuthor="Evans" pageId="16" pageNumber="15" pagination="26 - 53" refId="ref27534" refString="Evans, W. H. 1953. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III. Pyrginae. Section 2. British Museum (Natural History); London. v + 246 p., pl. 26 - 53" type="book chapter" year="1953">Evans (1953)</bibRefCitation>
. Genitalia illustrated by Grishin in his fig. 3039 (
<bibRefCitation id="EFB6E5C57E62FFEEFBDE7662DBDDFA9A" author="Grishin" box="[1057,1219,1352,1377]" firstAuthor="Grishin" pageId="16" pageNumber="15" pagination="1 - 7" refId="ref27647" refString="Grishin, N. V. 2012. A new Central American Anastrus with unexpectedly distinct genitalia (Lepidoptera, Hesperiidae, Pyrginae). Tropical Lepidoptera Research 22 (1): 1 - 7." type="journal article" year="2012">Grishin 2012</bibRefCitation>
). Morphologically, distinguished from
<taxonomicName id="4C27E3B77E62FFEEFE507642DD00FA84" ID-CoL="8ZZV2" authorityName="Hubner" authorityYear="1824" box="[431,542,1384,1407]" class="Insecta" family="Hesperiidae" genus="Anastrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E62FFEEFE507642DD00FA84" box="[431,542,1384,1407]" italics="true" pageId="16" pageNumber="15">Anastrus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C27E3B77E62FFEEFDA47642DDCDFA7B" box="[603,723,1384,1408]" class="Insecta" family="Hesperiidae" genus="Echelatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E62FFEEFDA47642DDCDFA7B" box="[603,723,1384,1408]" italics="true" pageId="16" pageNumber="15">Echelatus</emphasis>
</taxonomicName>
by having processes on sacculus of the valva and a second pair of uncus arms, from the ventral side of uncus, similar to some species of its sister
<taxonomicName id="4C27E3B77E62FFEEFB0C76A3DA87FA5B" ID-CoL="92KHC" authorityName="Lindsey" authorityYear="1925" box="[1267,1433,1417,1440]" class="Insecta" family="Hesperiidae" genus="Potamanaxas" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E62FFEEFB0C76A3DA87FA5B" box="[1267,1433,1417,1440]" italics="true" pageId="16" pageNumber="15">Potamanaxas</emphasis>
</taxonomicName>
, but uncus itself is essentially undivided with arms reduced to small knobs. Primary uncus arms are well-developed in
<taxonomicName id="4C27E3B77E62FFEEFE6776E3DD20FA1B" ID-CoL="92KHC" authorityName="Lindsey" authorityYear="1925" box="[408,574,1481,1504]" class="Insecta" family="Hesperiidae" genus="Potamanaxas" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E62FFEEFE6776E3DD20FA1B" box="[408,574,1481,1504]" italics="true" pageId="16" pageNumber="15">Potamanaxas</emphasis>
</taxonomicName>
. In facies, characterized by brown wings above with several dark-brown stripes and often with some bluish scaling between the stripes. Below, wings plain brown and some species with slate overscaling on the posterior third of hindwing. In DNA, a combination of the following base pairs is diagnostic: aly3570.7.4:A85T, aly3570.7.4:G86C, aly9673.2.5:T298A, aly
<date id="FF99BEF47E62FFEEFB217502DA56F9BB" box="[1246,1352,1576,1600]" pageId="16" pageNumber="15" value="1735-08-01">1735.8.1</date>
:C820C (not T), aly16.28.2:C88C (not A), aly16.28.2:A89A (not T).
</paragraph>
</subSubSection>
<subSubSection id="C33DCBBF7E62FFEEFF3F7552DCDBF8CA" pageId="16" pageNumber="15" type="etymology">
<paragraph id="8B9898347E62FFEEFF3F7552DB8FF94B" blockId="16.[192,1440,1656,1712]" pageId="16" pageNumber="15">
<emphasis id="B95344267E62FFEEFF3F7552DE45F96B" bold="true" box="[192,347,1656,1680]" pageId="16" pageNumber="15">Etymology.</emphasis>
The name is a masculine noun in the nominative singular, a group of species that were formerly placed in
<emphasis id="B95344267E62FFEEFE5975B2DEC6F954" box="[422,472,1688,1711]" italics="true" pageId="16" pageNumber="15">Ana</emphasis>
[strus], but are phylogenetically closer to [Potamana]
<emphasis id="B95344267E62FFEEFB9D75B3DB94F94B" box="[1122,1162,1689,1712]" italics="true" pageId="16" pageNumber="15">xas</emphasis>
.
</paragraph>
<paragraph id="8B9898347E62FFEEFF3F75E2DE1CF8FB" blockId="16.[192,1440,1736,1793]" pageId="16" pageNumber="15">
<emphasis id="B95344267E62FFEEFF3F75E2DEB4F91B" bold="true" box="[192,426,1736,1760]" pageId="16" pageNumber="15">Species included.</emphasis>
The
<typeStatus id="549C26967E62FFEEFE1D75E3DD08F91A" box="[482,534,1737,1761]" pageId="16" pageNumber="15">type</typeStatus>
species, its sister
<taxonomicName id="4C27E3B77E62FFEEFD1775E2DB59F91A" authority="Grishin, 2012" authorityName="Grishin" authorityYear="2012" box="[744,1095,1736,1761]" class="Insecta" family="Hesperiidae" genus="Anastrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="isidro">
<emphasis id="B95344267E62FFEEFD1775E2DC81F91B" box="[744,927,1736,1760]" italics="true" pageId="16" pageNumber="15">Anastrus isidro</emphasis>
<bibRefCitation id="EFB6E5C57E62FFEEFC5B75E2DB59F91A" author="Grishin" box="[932,1095,1736,1761]" firstAuthor="Grishin" pageId="16" pageNumber="15" pagination="1 - 7" refId="ref27647" refString="Grishin, N. V. 2012. A new Central American Anastrus with unexpectedly distinct genitalia (Lepidoptera, Hesperiidae, Pyrginae). Tropical Lepidoptera Research 22 (1): 1 - 7." type="journal article" year="2012">Grishin, 2012</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C27E3B77E62FFEEFB8075E2DFE5F8FA" authority="Moschler, 1877" authorityName="Moschler" authorityYear="1877" class="Insecta" family="Hesperiidae" genus="Pellicia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="petius">
<emphasis id="B95344267E62FFEEFB8075E2DA38F91B" box="[1151,1318,1736,1760]" italics="true" pageId="16" pageNumber="15">Pellicia petius</emphasis>
Möschler, 1877
</taxonomicName>
.
</paragraph>
<paragraph id="8B9898347E62FFEEFF3F7432DCDBF8CA" blockId="16.[192,965,1816,1841]" box="[192,965,1816,1841]" pageId="16" pageNumber="15">
<emphasis id="B95344267E62FFEEFF3F7432DE62F8CB" bold="true" box="[192,380,1816,1840]" pageId="16" pageNumber="15">Parent taxon.</emphasis>
Subtribe Erynnina Brues and Carpenter, 1932.
</paragraph>
</subSubSection>
</treatment>
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