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<document id="0730C1B131861D7CF2CC81A7B5BCD8F4" ID-DOI="10.24199/j.mmv.2003.60.28" ID-ISSN="1447-2554" ID-Zenodo-Dep="12209846" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718990989155" checkinUser="felipe" docAuthor="Bruce, N. L." docDate="2003" docId="038D87F1FFDBFFFDFF7AFA28439DF9B8" docLanguage="en" docName="MemMusVictoria.60.2.309-369.pdf" docOrigin="Memoirs of Museum Victoria 60 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-60-issue-2-2003/pages-309-369/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Cassidias Richardson 1906" docType="treatment" docVersion="2" lastPageNumber="320" masterDocId="FFB4FF89FFD1FFF6FFF1FFED414AFF94" masterDocTitle="New genera and species of sphaeromatid isopod crustaceans from Australian marine coastal waters" masterLastPageNumber="369" masterPageNumber="309" pageNumber="319" updateTime="1719000294287" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="77CB389E1EB938CEDF15E5BD79658288">New genera and species of sphaeromatid isopod crustaceans from Australian marine coastal waters</mods:title>
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<paragraph id="8B9B36E7FFDBFFFCFF7AFA284022FA4F" blockId="10.[139,360,1477,1499]" box="[139,360,1477,1499]" pageId="10" pageNumber="319">
<heading id="D0D3818BFFDBFFFCFF7AFA284022FA4F" box="[139,360,1477,1499]" fontSize="9" level="2" pageId="10" pageNumber="319" reason="5">
<taxonomicName id="4C244D64FFDBFFFCFF7AFA284022FA4F" ID-CoL="7NRZP" authority="Richardson" authorityName="Richardson" authorityYear="1906" box="[139,360,1477,1499]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDBFFFCFF7AFA2841A4FA4F" bold="true" box="[139,238,1477,1499]" italics="true" pageId="10" pageNumber="319">Cassidias</emphasis>
Richardson
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E656CFFDBFFFCFF57FA0243A0F98A" pageId="10" pageNumber="319" type="reference_group">
<paragraph id="8B9B36E7FFDBFFFCFF57FA0243A0F98A" blockId="10.[139,780,1519,1566]" pageId="10" pageNumber="319">
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<taxonomicName id="4C244D64FFDBFFFCFF57FA024085F997" ID-CoL="7NRZP" authority="Richardson, 1906: 20" authorityName="Richardson" authorityPageNumber="20" authorityYear="1906" box="[166,463,1519,1539]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDBFFFCFF57FA024024F997" box="[166,366,1519,1539]" italics="true" pageId="10" pageNumber="319">
Cassidias
<bibRefCitation id="EFB54B16FFDBFFFCFEF6FA024024F997" author="Richardson, H." box="[263,366,1519,1539]" pageId="10" pageNumber="319" pagination="1 - 22" refId="ref30150" refString="Richardson, H. 1906. Descriptions of a new isopod crustaceans of the family Sphaeromidae. Proceedings of the United States National Museum 31: 1 - 22." type="journal article" year="1906">Richardson</bibRefCitation>
</emphasis>
, 1906: 20
</taxonomicName>
.—
<treatmentCitation id="0A8510F6FFDBFFFCFE1BFA0243EBF997" author="Barnard, K. H." box="[490,673,1519,1539]" page="374" pageId="10" pageNumber="319" year="1920">
<bibRefCitation id="EFB54B16FFDBFFFCFE1BFA0243EBF997" author="Barnard, K. H." box="[490,673,1519,1539]" pageId="10" pageNumber="319" pagination="319 - 438" refId="ref27248" refString="Barnard, K. H. 1920. Contributions to the crustacean fauna of South Africa. 6. Further additions to the list of marine Isopoda. Annals of the South African Museum 17: 319 - 438." type="journal article" year="1920">Barnard, 1920: 374</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0A8510F6FFDBFFFCFD4DFA024191F98A" author="Kensley, B." page="87" pageId="10" pageNumber="319" year="1978">
<bibRefCitation id="EFB54B16FFDBFFFCFD4DFA024191F98A" author="Kensley, B." pageId="10" pageNumber="319" refId="ref29121" refString="Kensley, B. 1978. Guide to the Marine Isopods of Southern Africa. South African Museum and The Rustica Press, Wynberg, Cape Town." type="book" year="1978">Kensley, 1978: 87</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0A8510F6FFDBFFFCFF07F9E740F2F98A" author="Harrison, K." box="[246,440,1546,1566]" page="373" pageId="10" pageNumber="319" year="1984">
<bibRefCitation id="EFB54B16FFDBFFFCFF07F9E740F2F98A" author="Harrison, K." box="[246,440,1546,1566]" pageId="10" pageNumber="319" pagination="363 - 407" refId="ref28513" refString="Harrison, K. 1984 a. The morphology of the sphaeromatid brood pouch (Crustacea: Isopoda: Sphaeromatidae). Zoological Journal of the Linnean Society 82: 363 - 407." type="journal article" year="1984">Harrison, 1984a: 373</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0A8510F6FFDBFFFCFE22F9E743ADF98A" author="Harrison, K. &amp; Ellis, J. P." box="[467,743,1546,1566]" page="934" pageId="10" pageNumber="319" year="1991">
<bibRefCitation id="EFB54B16FFDBFFFCFE22F9E743ADF98A" author="Harrison, K. &amp; Ellis, J. P." box="[467,743,1546,1566]" pageId="10" pageNumber="319" pagination="195 - 952" refId="ref28586" refString="Harrison, K. and Ellis, J. P. 1991. The genera of the Sphaeromatidae (Crustacea: Isopoda): a key and distribution list. Invertebrate Taxonomy 5: 195 - 952." type="journal article" year="1991">Harrison and Ellis, 1991: 934</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<paragraph id="8B9B36E7FFDBFFFCFF7AF9D94036F910" blockId="10.[139,780,1588,1668]" pageId="10" pageNumber="319">
<emphasis id="B950EAF5FFDBFFFCFF7AF9D94056F9DE" box="[139,284,1588,1610]" italics="true" pageId="10" pageNumber="319">
<typeStatus id="549F8845FFDBFFFCFF7AF9D941F1F9DE" box="[139,187,1588,1610]" pageId="10" pageNumber="319">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C244D64FFDBFFFCFEDDF9D94394F9DE" authority="Richardson, 1906" authorityName="Richardson" authorityYear="1906" box="[300,734,1588,1610]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="argentinea">
<emphasis id="B950EAF5FFDBFFFCFEDDF9D94347F9DE" box="[300,525,1588,1610]" italics="true" pageId="10" pageNumber="319">Cassidias argentinea</emphasis>
<bibRefCitation id="EFB54B16FFDBFFFCFDECF9D94394F9DE" author="Richardson, H." box="[541,734,1588,1610]" pageId="10" pageNumber="319" pagination="1 - 22" refId="ref30150" refString="Richardson, H. 1906. Descriptions of a new isopod crustaceans of the family Sphaeromidae. Proceedings of the United States National Museum 31: 1 - 22." type="journal article" year="1906">Richardson, 1906</bibRefCitation>
</taxonomicName>
, by monotypy.
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<typeStatus id="549F8845FFDBFFFCFEF7F9BC4008F9F3" box="[262,322,1617,1639]" pageId="10" pageNumber="319">Types</typeStatus>
held at
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,
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,
<collectingRegion id="49E0F805FFDBFFFCFF7AF983404CF910" box="[139,262,1646,1668]" country="United States of America" name="Washington" pageId="10" pageNumber="319">Washington</collectingRegion>
,
<collectionCode id="ED35AE22FFDBFFFCFEE0F983407DF910" box="[273,311,1646,1668]" pageId="10" pageNumber="319">DC</collectionCode>
,
<collectingCountry id="F3337677FFDBFFFCFEB3F9834032F910" box="[322,376,1646,1668]" name="United States of America" pageId="10" pageNumber="319">USA</collectingCountry>
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.
</paragraph>
<paragraph id="8B9B36E7FFDBFFFCFF7AF9764462FEA4" blockId="10.[139,780,1690,1918]" lastBlockId="10.[817,1457,224,304]" pageId="10" pageNumber="319">
<emphasis id="B950EAF5FFDBFFFCFF7AF97641B3F925" box="[139,249,1691,1713]" italics="true" pageId="10" pageNumber="319">Diagnosis.</emphasis>
Pleotelson with prominent median process; posterior margin with simple median notch and distinct ventral exit channel. Antennule peduncle article 1 more than twice as long as article 2, articles 1 and 2 robust; article 3 slender, all articles collinear. Maxilliped palp articles 24 with distomesial angle moderately produced, mesial margins with numerous setae. Pleopods 13 with both rami longitudinally oblique. Pleopod 2 with appendix masculina distally attached. Male uropods attached subdistally on pleon, endopod reduced to short stub, exopod round in section with apical spike, entire ramus extending beyond posterior margin of pleotelson.
</paragraph>
<paragraph id="8B9B36E7FFDBFFFCFCC0FEB84404FD3A" blockId="10.[817,1458,341,1918]" pageId="10" pageNumber="319">
<emphasis id="B950EAF5FFDBFFFCFCC0FEB8454EFEFF" box="[817,1028,341,363]" italics="true" pageId="10" pageNumber="319">Description of male.</emphasis>
Body stout, about twice as long as greatest width, strongly vaulted; dorsal surface finely granular, with scattered setae; unable to conglobate. Head weakly immersed in pereonite 1. Eyes large, lateral, facets distinct, prominent posterior lobe. Pereon segments without raised posterior margins. Coxae not distinctly demarcated, overlapping anterior over posterior, ventrally directed. Membrana cingula absent. Pleon of 4 segments, segment 1 largely concealed by pereonite 7, segments 24 indicated by 2 distinct suture lines running to lateral margins of pleon. Pleotelson with prominent median process; posterior margin with simple median notch and distinct ventral exit channel. Pleonal sternite present.
</paragraph>
<paragraph id="8B9B36E7FFDBFFFCFCA0FD5842E3FC0C" blockId="10.[817,1458,341,1918]" pageId="10" pageNumber="319">Antennule and antenna anteriorly positioned on head. Antennule peduncle articles 1 and 2 robust, article 1 more than twice as long as article 2; article 3 slender, all articles collinear; flagellum shorter than peduncle, extending to anterior of pereonite 1. Antenna peduncle article 1 and 2 short, subequal, shorter than 35, which become progressively longer; flagellum about as long as peduncle, extending to posterior of pereonite 1.</paragraph>
<paragraph id="8B9B36E7FFDBFFFCFCA0FC4D45DFFAFA" blockId="10.[817,1458,341,1918]" pageId="10" pageNumber="319">Epistome anteriorly acute, with weak mesial constriction; with indistinct ventral tubercle Labrum unornamented. Mandible incisor multicuspid; molar process prominent, crushing, provided with marginal scale teeth; left mandible with prominent lacinia mobilis both mandibles with spine row of 5 or 6 spines; palp article 1 longest, 3 shortest. Maxillule lateral lobe with about 13 RS on gnathal surface, most of which are serrate; mesial lobe with 5 long RS, 3 of which are prominently serrate. Maxilla with all articles well developed; lateral and middle lobes with flat strongly curved and finely serrate RS, mesial lobe with several and acute long RS, some of which are basally CP. Maxilliped endite distal margin numerous long acute CP setae and 3 long CP RS on distomesial margin; palp articles 24 each with distomesial angle produced, mesial margins with numerous setae, lateral margins of articles 2 and 3 without setae, article 4 with 1 seta.</paragraph>
<paragraph id="8B9B36E7FFDBFFFCFCA0FA9844EEF9AF" blockId="10.[817,1458,341,1918]" pageId="10" pageNumber="319">Pereopods all ambulatory, pereopods 13 subsimilar, more robust than 47; inferior margins of merus, carpus and propodus with serrate and CP RS; setulose fringe weakly developed; dactylus with prominent simple secondary unguis and 1 flattened seta arising at lateral margin, 2 flat setae at distolateral margin. Pereopods 6 and 7 inferior and distal margins of merus, carpus and propodus with numerous serrate and biserrate RS.</paragraph>
<paragraph id="8B9B36E7FFDBFFFCFCA0F9AF445CF9E2" blockId="10.[817,1458,341,1918]" pageId="10" pageNumber="319">Penes paired, separated by about basal width of penial process; short, not reaching pleopod peduncles.</paragraph>
<paragraph id="8B9B36E7FFDBFFFDFCA0F990435AFEDA" blockId="10.[817,1458,341,1918]" lastBlockId="11.[131,772,224,657]" lastPageId="11" lastPageNumber="320" pageId="10" pageNumber="319">
Pleopods 13 both rami with
<collectionCode id="ED35AE22FFDBFFFCFB8AF99045FAF907" box="[1147,1200,1661,1683]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="10" pageNumber="319">PMS</collectionCode>
, longitudinal axis of both rami oblique Pleopod 1 exopod extending beyond endopod distal margin subtruncate, proximolateral angle with single short acute RS; endopod distinctly triangular in shape. Pleopod 2 with appendix masculina distally attached on mesial margin. Pleopod 3 exopod with complete suture, suture not distinct on pleopod 4 and incomplete on pleopod 5. Pleopods 4 and 5 exopod and endopods with well-developed transverse thickened ridges; pleopod 5 endopod with 3 lobate scale patches and transverse suture. Uropod exopod 3 times as long as wide, subdistally attached on pleon, endopod reduced to short stub, exopod round in section with apical spike, entire ramus extending beyond posterior margin of pleotelson.
</paragraph>
<paragraph id="8B9B36E7FFDAFFFDFF52FEB84039FD8F" blockId="11.[131,772,224,657]" pageId="11" pageNumber="320">
<emphasis id="B950EAF5FFDAFFFDFF52FEB841BEFEFF" box="[163,244,341,363]" italics="true" pageId="11" pageNumber="320">Female.</emphasis>
Pleotelsonic process considerably smaller than in the male; uropods with both rami lamellar, positioned midlaterally on pleon, endopod extending only slightly beyond posteri- or margin of pleotelson; both rami with numerous CP setae. Mouthparts metamorphosed. Brood pouch of the
<typeStatus id="549F8845FFDAFFFDFD77FE2743F8FE74" box="[646,690,458,480]" pageId="11" pageNumber="320">type</typeStatus>
species made up of 4 pairs of oostegites on pereonites 14 (Harrison 1984, but see remarks).
</paragraph>
<paragraph id="8B9B36E7FFDAFFFDFF52FDCE4384FD04" blockId="11.[131,772,224,657]" pageId="11" pageNumber="320">
<emphasis id="B950EAF5FFDAFFFDFF52FDCE4066FDAD" box="[163,300,547,569]" italics="true" pageId="11" pageNumber="320">Composition.</emphasis>
<taxonomicName id="4C244D64FFDAFFFDFEC9FDCE4391FDAC" authority="Richardson, 1906" authorityName="Richardson" authorityYear="1906" box="[312,731,546,569]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="argentinea">
<emphasis id="B950EAF5FFDAFFFDFEC9FDCE435EFDAD" box="[312,532,547,569]" italics="true" pageId="11" pageNumber="320">Cassidias argentinea</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFDD1FDCF4391FDAC" author="Richardson, H." box="[544,731,546,568]" pageId="11" pageNumber="320" pagination="1 - 22" refId="ref30150" refString="Richardson, H. 1906. Descriptions of a new isopod crustaceans of the family Sphaeromidae. Proceedings of the United States National Museum 31: 1 - 22." type="journal article" year="1906">Richardson, 1906</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C244D64FFDAFFFDFD1DFDCE404EFDC2" authorityName="Bruce" authorityYear="2003" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="australiensis" status="sp. nov.">
<emphasis id="B950EAF5FFDAFFFDFD1DFDCE404EFDC2" italics="true" pageId="11" pageNumber="320">C. australiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263578EFFDAFFFDFEE1FDAD402CFDC2" box="[272,358,576,598]" pageId="11" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C244D64FFDAFFFDFE84FDAD43D8FDC2" authority="Barnard, 1920" authorityName="Barnard" authorityYear="1920" box="[373,658,576,598]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B950EAF5FFDAFFFDFE84FDAD40A7FDC2" box="[373,493,576,598]" italics="true" pageId="11" pageNumber="320">C. africana</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFE08FDAD43D8FDC2" author="Barnard, K. H." box="[505,658,576,598]" pageId="11" pageNumber="320" pagination="319 - 438" refId="ref27248" refString="Barnard, K. H. 1920. Contributions to the crustacean fauna of South Africa. 6. Further additions to the list of marine Isopoda. Annals of the South African Museum 17: 319 - 438." type="journal article" year="1920">Barnard, 1920</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C244D64FFDAFFFDFD51FDAD409CFDE7" authority="Thielemann, 1910" authorityName="Thielemann" authorityYear="1910" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="trituberculata">
<emphasis id="B950EAF5FFDAFFFDFD51FDAD4045FDE7" italics="true" pageId="11" pageNumber="320">Cassidias trituberculata</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFEE8FDB0409CFDE7" author="Thielemann, M." box="[281,470,605,627]" pageId="11" pageNumber="320" pagination="1 - 109" refId="ref30653" refString="Thielemann, M. 1910. Beitrage zur Kenntnis der Naturgechichte Ostasiens. Herausgegeben von F. Doflein. Band II. No. 9. Beitrage zur Kenntnis der Isopodenfauna Ostasiens. Abhandlungen der Matemathetisch-Naturwissenschaftlichen Klasse der K. Bayer. Akademia der Wissenschaften (suppl) 2: 1 - 109." type="journal article" year="1910">Thielemann, 1910</bibRefCitation>
</taxonomicName>
has been synonymised with
<taxonomicName id="4C244D64FFDAFFFDFF72FD964378FD04" authority="Richardson, 1909" authorityName="Richardson" authorityYear="1909" box="[131,562,634,657]" class="Malacostraca" family="Sphaeromatidae" genus="Holotelson" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="B950EAF5FFDAFFFDFF72FD96403DFD05" box="[131,375,635,657]" italics="true" pageId="11" pageNumber="320">Holotelson tuberculatus</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFE8FFD974378FD04" author="Richardson, H." box="[382,562,634,656]" pageId="11" pageNumber="320" pagination="75 - 129" refId="ref30212" refString="Richardson, H. 1909. Isopods collected in the Northwest Pacific by the U. S. Bureau of Fisheries Steamer &quot; Albatross &quot; in 1906. Proceedings of the U. S. National Museum 37: 75 - 129." type="journal article" year="1909">Richardson, 1909</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFB54B16FFDAFFFDFDCEFD974389FD04" author="Kwon, D. H." box="[575,707,634,656]" pageId="11" pageNumber="320" pagination="151 - 192" refId="ref29320" refString="Kwon, D. H. 1990. A systematic study on the Korean marine isopod crustaceans. Inje Journal 6: 151 - 192." type="journal article" year="1990">Kwon, 1990</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B36E7FFDAFFFDFF72FD4A4304FBAE" blockId="11.[131,772,678,1581]" pageId="11" pageNumber="320">
<emphasis id="B950EAF5FFDAFFFDFF72FD4A41A9FD29" box="[131,227,679,701]" italics="true" pageId="11" pageNumber="320">Remarks.</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFF01FD4B40EDFD28" author="Harrison, K." box="[240,423,678,700]" pageId="11" pageNumber="320" pagination="363 - 407" refId="ref28513" refString="Harrison, K. 1984 a. The morphology of the sphaeromatid brood pouch (Crustacea: Isopoda: Sphaeromatidae). Zoological Journal of the Linnean Society 82: 363 - 407." type="journal article" year="1984">Harrison (1984a)</bibRefCitation>
, on the basis of differences in brood-pouch morphology, excluded
<taxonomicName id="4C244D64FFDAFFFDFDDFFD29405FFD63" authority="Barnard, 1920" authorityName="Barnard" authorityYear="1920" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B950EAF5FFDAFFFDFDDFFD294249FD4E" box="[558,771,708,730]" italics="true" pageId="11" pageNumber="320">Cassidias africana</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFF72FD0C405FFD63" author="Barnard, K. H." box="[131,277,737,759]" pageId="11" pageNumber="320" pagination="319 - 438" refId="ref27248" refString="Barnard, K. H. 1920. Contributions to the crustacean fauna of South Africa. 6. Further additions to the list of marine Isopoda. Annals of the South African Museum 17: 319 - 438." type="journal article" year="1920">Barnard, 1920</bibRefCitation>
</taxonomicName>
from the genus. The
<typeStatus id="549F8845FFDAFFFDFE01FD0C4356FD63" box="[496,540,737,759]" pageId="11" pageNumber="320">type</typeStatus>
species is known only from females (
<bibRefCitation id="EFB54B16FFDAFFFDFEE9FD13409AFC80" author="Richardson, H." box="[280,464,766,788]" pageId="11" pageNumber="320" pagination="1 - 22" refId="ref30150" refString="Richardson, H. 1906. Descriptions of a new isopod crustaceans of the family Sphaeromidae. Proceedings of the United States National Museum 31: 1 - 22." type="journal article" year="1906">Richardson, 1906</bibRefCitation>
), and until such time as males are known it is impossible to assess the generic characters for this genus. Although brood-pouch morphology is generally consistent within sphaeromatid genera, there is insufficient evidence to consider such differences to be axiomatically of generic merit, and this character is known to vary in
<taxonomicName id="4C244D64FFDAFFFDFD60FC7C407DFC50" authority="(Harrison, 1984 a)" baseAuthorityName="Harrison" baseAuthorityYear="1984" class="Malacostraca" family="Sphaeromatidae" genus="Sphaeroma" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDAFFFDFD60FC7C4249FC33" box="[657,771,913,935]" italics="true" pageId="11" pageNumber="320">Sphaeroma</emphasis>
(
<bibRefCitation id="EFB54B16FFDAFFFDFF7BFC43407AFC50" author="Harrison, K." box="[138,304,942,964]" pageId="11" pageNumber="320" pagination="363 - 407" refId="ref28513" refString="Harrison, K. 1984 a. The morphology of the sphaeromatid brood pouch (Crustacea: Isopoda: Sphaeromatidae). Zoological Journal of the Linnean Society 82: 363 - 407." type="journal article" year="1984">Harrison, 1984a</bibRefCitation>
)
</taxonomicName>
and now also in
<taxonomicName id="4C244D64FFDAFFFDFE15FC424315FC51" authorityName="Bruce" authorityYear="1993" box="[484,607,943,965]" class="Malacostraca" family="Sphaeromatidae" genus="Margueritta" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDAFFFDFE15FC424315FC51" box="[484,607,943,965]" italics="true" pageId="11" pageNumber="320">Margueritta</emphasis>
</taxonomicName>
(present study). At present it seems appropriate to retain the genus, diagnosed on the basis of the known males, with recognition that its status is uncertain. A new genus cannot adequately be differentiated from
<taxonomicName id="4C244D64FFDAFFFDFF4AFBC94054FBAE" authorityName="Richardson" authorityYear="1906" box="[187,286,1060,1082]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDAFFFDFF4AFBC94054FBAE" box="[187,286,1060,1082]" italics="true" pageId="11" pageNumber="320">Cassidias</emphasis>
</taxonomicName>
on the basis of
<typeStatus id="549F8845FFDAFFFDFE30FBC940A7FBAE" box="[449,493,1060,1082]" pageId="11" pageNumber="320">type</typeStatus>
material.
</paragraph>
<paragraph id="8B9B36E7FFDAFFFDFF52FBAC40B2FAD6" blockId="11.[131,772,678,1581]" pageId="11" pageNumber="320">Males of this genus, as here defined can be readily recognised by subterminal uropods which have the endopod reduced in size, by the presence of a broad, blunt, posteriorly directed process on the pleotelson and by the terminal appendix masculina. In addition, the posterior margin of the pleotelson has a distinct simple median notch. Females have the uropods in a more midlateral position and both rami are flat. The process on the pleotelson is reduced compared with that of the male, but does allow identification of females.</paragraph>
<paragraph id="8B9B36E7FFDAFFFDFF52FAA4439DF9B8" blockId="11.[131,772,678,1581]" pageId="11" pageNumber="320">
The most similar genus is
<taxonomicName id="4C244D64FFDAFFFDFE45FAA44189FAE8" authority="Harrison and Holdich, 1982 b" authorityName="Harrison and Holdich" authorityYear="1982" class="Malacostraca" family="Sphaeromatidae" genus="Neonaesa" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDAFFFDFE45FAA44352FACB" box="[436,536,1353,1375]" italics="true" pageId="11" pageNumber="320">Neonaesa</emphasis>
<bibRefCitation id="EFB54B16FFDAFFFDFDEEFAA44189FAE8" author="Harrison, K. &amp; Holdich, D. M." pageId="11" pageNumber="320" pagination="421 - 446" refId="ref28681" refString="Harrison, K. and Holdich, D. M. 1982 b. New eubrachiate sphaeromatid isopods from Queensland waters. Memoirs of the Queensland Museum 20: 421 - 446." type="journal article" year="1982">Harrison and Holdich, 1982b</bibRefCitation>
</taxonomicName>
which is readily differentiated by the males having the appendix masculina mesially attached, the pleotelson posterior margin with three small notches and the pleotelson dorsal surface with a small median boss flanked by two low submedian bosses. Females of
<taxonomicName id="4C244D64FFDAFFFDFEF5FA314022FA66" authorityName="Harrison and Holdich" authorityYear="1982" box="[260,360,1500,1522]" class="Malacostraca" family="Sphaeromatidae" genus="Neonaesa" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDAFFFDFEF5FA314022FA66" box="[260,360,1500,1522]" italics="true" pageId="11" pageNumber="320">Neonaesa</emphasis>
</taxonomicName>
are separated by having a cylindrical uropodal exopod and a reduced endopod in contrast to female
<taxonomicName id="4C244D64FFDAFFFDFF72F9FA41ACF9B9" authorityName="Richardson" authorityYear="1906" box="[131,230,1559,1581]" class="Malacostraca" family="Sphaeromatidae" genus="Cassidias" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAF5FFDAFFFDFF72F9FA41ACF9B9" box="[131,230,1559,1581]" italics="true" pageId="11" pageNumber="320">Cassidias</emphasis>
</taxonomicName>
which have uropodal rami both present and flat.
</paragraph>
</subSubSection>
</treatment>
</document>