543 lines
72 KiB
XML
543 lines
72 KiB
XML
<document id="8ED0972ED8BE2DB9422CFCF759F2A12C" ID-CLB-Dataset="31016" ID-DOI="10.5852/ejt.2018.394" ID-GBIF-Dataset="54a11801-d3d3-4b76-9f6c-7a9c27677825" ID-Zenodo-Dep="1146158" ID-ZooBank="94028C61-FAC5-4A21-BF2D-A75BBF3CCC4D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515747489213" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2018" docId="038D2864FFBFFFA1FDDAFBBA8BDBFA76" docLanguage="en" docName="ejt-394_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 394" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helicochetus mutaba Kraus 1960" docType="treatment" docVersion="17" lastPageNumber="17" masterDocId="FFB4501CFFB2FFB1FF94FFB0892EFF8E" masterDocTitle="A mountain of millipedes VI. New records, new species, a new genus and a general discussion of Odontopygidae from the Udzungwa Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="29" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547161722869" originalUpdateUser="yanell" pageNumber="14" updateTime="1698513679935" updateUser="ExternalLinkService">
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<mods:title id="B9ACDCCD1D8E93D531073B54F53FA291">A mountain of millipedes VI. New records, new species, a new genus and a general discussion of Odontopygidae from the Udzungwa Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="789B785ADA8A6B3363219A378DEE42FB">Enghoff, Henrik</mods:namePart>
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<mods:title id="09BA908A5EE20C3D431685CC3AED14DB">European Journal of Taxonomy</mods:title>
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<mods:part id="7985EEA2882F45DF5D04A309FC5EFCD8">
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<mods:date id="39A2CAE6DC1CD3EAC56331D7D37A0DB4">2018</mods:date>
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<mods:number id="F6A3FCAC6FC02772CF02D29B964E08E8">2018-01-11</mods:number>
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<mods:number id="25AF71E1652B25613737A3342E8ACDB1">394</mods:number>
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<treatment id="038D2864FFBFFFA1FDDAFBBA8BDBFA76" ID-DOI="http://doi.org/10.5281/zenodo.5613984" ID-GBIF-Taxon="140251437" ID-Zenodo-Dep="5613984" LSID="urn:lsid:plazi:treatment:038D2864FFBFFFA1FDDAFBBA8BDBFA76" httpUri="http://treatment.plazi.org/id/038D2864FFBFFFA1FDDAFBBA8BDBFA76" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
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<subSubSection id="C33ECAF9FFBFFFBCFDDAFBBA8AC8FBAA" box="[590,998,1034,1061]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8B9B9972FFBFFFBCFDDAFBBA8AC8FBAA" blockId="13.[590,998,1034,1095]" box="[590,998,1034,1061]" pageId="13" pageNumber="14">
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<heading id="D0D32E1EFFBFFFBCFDDAFBBA8AC8FBAA" box="[590,998,1034,1061]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
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<taxonomicName id="4C24E2F1FFBFFFBCFDDAFBBA8AC8FBAA" ID-CoL="3K82R" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[590,998,1034,1061]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mutaba">
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<emphasis id="B9504560FFBFFFBCFDDAFBBA8A60FBAB" bold="true" box="[590,846,1034,1061]" italics="true" pageId="13" pageNumber="14">Helicochetus mutaba</emphasis>
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<bibRefCitation id="EFB5E483FFBFFFBCFCC2FBBB8AC8FBAA" author="Kraus O." box="[854,998,1034,1061]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="13" pageNumber="14" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus, 1960</bibRefCitation>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C33ECAF9FFBFFFBCFD5BFB9D8A4BFBC9" box="[719,869,1069,1095]" pageId="13" pageNumber="14" type="description">
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<paragraph id="8B9B9972FFBFFFBCFD5BFB9D8A4BFBC9" blockId="13.[590,998,1034,1095]" box="[719,869,1069,1095]" pageId="13" pageNumber="14">
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<figureCitation id="131F85F7FFBFFFBCFD5BFB9D8A65FBC9" box="[719,843,1069,1095]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="13" pageNumber="14">Figs 4C–F</figureCitation>
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,
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<figureCitation id="131F85F7FFBFFFBCFCC2FB9D8A4BFBC9" box="[854,869,1069,1095]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="13" pageNumber="14">6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ECAF9FFBFFFBCFF29FBDF8AF6FA24" pageId="13" pageNumber="14" type="reference_group">
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<paragraph id="8B9B9972FFBFFFBCFF29FBDF8BAFFB07" blockId="13.[189,641,1135,1161]" box="[189,641,1135,1161]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB34BE5CFFBFFFBCFF29FBDF8BAFFB07" box="[189,641,1135,1161]" pageId="13" pageNumber="14">
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<taxonomicName id="4C24E2F1FFBFFFBCFF29FBDF8B53FB07" ID-CoL="3K82R" authority="Kraus, 1960: 40" authorityName="Kraus" authorityPageNumber="40" authorityYear="1960" box="[189,637,1135,1161]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mutaba">
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<emphasis id="B9504560FFBFFFBCFF29FBDF889BFB07" box="[189,437,1135,1161]" italics="true" pageId="13" pageNumber="14">Helicochetus mutaba</emphasis>
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<treatmentCitation id="0A85BF63FFBFFFBCFE29FBDF8B53FB07" author="Kraus" box="[445,637,1135,1161]" page="40" pageId="13" pageNumber="14" year="1960">
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<bibRefCitation id="EFB5E483FFBFFFBCFE29FBDF8B53FB07" author="Kraus O." box="[445,637,1135,1161]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" page="40" pageId="13" pageNumber="14" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus, 1960: 40</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9B9972FFBFFFBCFF29FB008818FB44" blockId="13.[189,310,1200,1226]" box="[189,310,1200,1226]" pageId="13" pageNumber="14">
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<heading id="D0D32E1EFFBFFFBCFF29FB008818FB44" bold="true" box="[189,310,1200,1226]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
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<emphasis id="B9504560FFBFFFBCFF29FB008818FB44" bold="true" box="[189,310,1200,1226]" pageId="13" pageNumber="14">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B9B9972FFBFFFBCFF29FB6F8AF6FA24" blockId="13.[189,1399,1247,1450]" pageId="13" pageNumber="14">
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Resembles
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<taxonomicName id="4C24E2F1FFBFFFBCFED2FB508BA1FB74" ID-CoL="6LJ74" authority="Kraus, 1957" authorityName="Kraus" authorityYear="1957" box="[326,655,1247,1274]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="digititarsus">
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<emphasis id="B9504560FFBFFFBCFED2FB5088DDFB77" box="[326,499,1247,1273]" italics="true" pageId="13" pageNumber="14">H. digititarsus</emphasis>
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Kraus, 1957
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</taxonomicName>
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,
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<taxonomicName id="4C24E2F1FFBFFFBCFD34FB508AC8FB74" ID-CoL="3K82K" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[672,998,1247,1274]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gregorii">
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<emphasis id="B9504560FFBFFFBCFD34FB508A08FB77" box="[672,806,1248,1273]" italics="true" pageId="13" pageNumber="14">H. gregorii</emphasis>
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||
(
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<bibRefCitation id="EFB5E483FFBFFFBCFCACFB6F8AF0FB74" author="Pocock R. I." box="[824,990,1247,1274]" firstAuthor="Pocock" journalOrPublisher="Annals and Magazine of Natural History, Series 6" pageId="13" pageNumber="14" pagination="425 - 444" part="17" publicationUrl="https://doi.org/10.1080/00222939608680394" refId="ref10362" refString="Pocock R. I. 1896. On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History, Series 6, 17: 425 - 444. https: // doi. org / 10.1080 / 00222939608680394" title="On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa" type="journal article" year="1896">Pocock, 1896</bibRefCitation>
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||
)
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||
</taxonomicName>
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and
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<taxonomicName id="4C24E2F1FFBFFFBCFBB1FB508C78FB77" ID-CoL="3K82Q" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[1061,1366,1247,1273]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="monodon">
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<emphasis id="B9504560FFBFFFBCFBB1FB508D97FB77" box="[1061,1209,1247,1273]" italics="true" pageId="13" pageNumber="14">H. monodon</emphasis>
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<bibRefCitation id="EFB5E483FFBFFFBCFB57FB6F8C78FB77" author="Kraus O." box="[1219,1366,1247,1273]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="13" pageNumber="14" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">
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Kraus,
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<quantity id="4CDC3497FFBFFFBCFA8FFB6F8C78FB77" box="[1307,1366,1247,1273]" metricMagnitude="1" metricUnit="m" metricValue="4.9784" pageId="13" pageNumber="14" unit="in" value="1960.0">1960</quantity>
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||
</bibRefCitation>
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||
</taxonomicName>
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in the combination of a broad, shovel-like process (“Tibialdorn”) at the level of the post-torsal narrowing; absence of ventral pads on male tibiae; presence of a basal lamella (“Grundblatt”) on the telomere; tip of telomere not drawn out into long process, with several short to moderately long spine-like outgrowths from the margin. Differs from these species in the shape of the basal lamella: slender, pointed and curved (with a few denticles on the concave margin in the new specimens).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33ECAF9FFBFFFBCFF29FA608A5AF9D1" pageId="13" pageNumber="14" type="materials_examined">
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<paragraph id="8B9B9972FFBFFFBCFF29FA608B03FA65" blockId="13.[189,1399,1488,1632]" box="[189,557,1488,1515]" pageId="13" pageNumber="14">
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<emphasis id="B9504560FFBFFFBCFF29FA6088A2FA64" bold="true" box="[189,396,1488,1514]" pageId="13" pageNumber="14">Material studied</emphasis>
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||
(total: 2 ♂♂)
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||
</paragraph>
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||
<paragraph id="8B9B9972FFBFFFBCFF29FA4F8A5AF9D1" blockId="13.[189,1399,1488,1632]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="3B4C932FFFBFFFBCFF29FA4F8B80F9B2" ID-GBIF-Occurrence="1802834929" collectionCode="VMNH" collectorName="W. A. Rodgers" country="Tanzania" county="Nyumbenito Mt" elevation="2500" location="KMH 2097" municipality="Udekwa" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1">
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<collectingCountry id="F333D9E2FFBFFFBCFF29FA4F8879F997" box="[189,343,1535,1561]" name="Tanzania" pageId="13" pageNumber="14">TANZANIA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D2252FBFFBFFFBCFECBFA4F88A5F997" box="[351,395,1535,1561]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
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||
,
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||
<location id="8EFBCFA9FFBFFFBCFE01FA4F8B32F997" LSID="urn:lsid:plazi:treatment:038D2864FFBFFFA1FDDAFBBA8BDBFA76:8EFBCFA9FFBFFFBCFE01FA4F8B32F997" box="[405,540,1535,1561]" country="Tanzania" county="Nyumbenito Mt" municipality="Udekwa" name="KMH 2097" pageId="13" pageNumber="14">KMH 2097</location>
|
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,
|
||
<collectingCounty id="62FAE1FEFFBFFFBCFDB1FA4F8BCCF997" box="[549,738,1535,1561]" pageId="13" pageNumber="14">Nyumbenito Mt</collectingCounty>
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,
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<location id="8EFBCFA9FFBFFFBCFD7FFA4F8AE1F997" LSID="urn:lsid:plazi:treatment:038D2864FFBFFFA1FDDAFBBA8BDBFA76:8EFBCFA9FFBFFFBCFD7FFA4F8AE1F997" box="[747,975,1535,1561]" country="Tanzania" county="Nyumbenito Mt" municipality="Udekwa" name="forest near Udekwa" pageId="13" pageNumber="14">forest near Udekwa</location>
|
||
,
|
||
<location id="8EFBCFA9FFBFFFBCFC4CFA4F8DE0F997" LSID="urn:lsid:plazi:treatment:038D2864FFBFFFA1FDDAFBBA8BDBFA76:8EFBCFA9FFBFFFBCFC4CFA4F8DE0F997" box="[984,1230,1535,1561]" country="Tanzania" county="Nyumbenito Mt" municipality="Udekwa" name="Uzungwa Mountains" pageId="13" pageNumber="14">Uzungwa Mountains</location>
|
||
,
|
||
<elevation id="00097E41FFBFFFBCFB43FA4F89DBF9B2" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="14" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">
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<quantity id="4CDC3497FFBFFFBCFB43FA4F8C59F997" box="[1239,1399,1535,1561]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="14" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">2000–3000 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, date?,
|
||
<collectorName id="26D1FCA4FFBFFFBCFEC0F99288D0F9B2" box="[340,510,1570,1596]" pageId="13" pageNumber="14">W.A. Rodgers</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED3501B7FFBFFFBCFDD2F9928B88F9B2" box="[582,678,1570,1596]" collectionName="USA, Virginia, Martinsville, Virginia Museum of Natural History" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="13" pageNumber="14">VMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4C932FFFBFFFBCFD29F9928A41F9D1" ID-GBIF-Occurrence="1802834931" collectingDate="1984-01" collectionCode="ZMUC" collectorName="Jan Kielland" country="Tanzania" elevation="1700" location="Mazombo" municipality="District" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region and District">
|
||
<specimenCount id="9D2252FBFFBFFFBCFD29F9928BC0F9B2" box="[701,750,1570,1596]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="49E05790FFBFFFBCFD6AF9928D13F9B2" box="[766,1085,1570,1596]" pageId="13" pageNumber="14">Iringa Region and District</collectingRegion>
|
||
,
|
||
<location id="8EFBCFA9FFBFFFBCFBD8F9928DE8F9B2" LSID="urn:lsid:plazi:treatment:038D2864FFBFFFA1FDDAFBBA8BDBFA76:8EFBCFA9FFBFFFBCFBD8F9928DE8F9B2" box="[1100,1222,1570,1596]" country="Tanzania" municipality="District" name="Mazombo" pageId="13" pageNumber="14" stateProvince="Iringa Region and District">Mazombo</location>
|
||
,
|
||
<locationDeviation id="9A298050FFBFFFBCFB41F9928828F9D1" pageId="13" pageNumber="14">
|
||
<quantity id="4CDC3497FFBFFFBCFB41F9928C0CF9B2" box="[1237,1314,1570,1596]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="km" value="20.0">20 km</quantity>
|
||
NE of Iringa
|
||
</locationDeviation>
|
||
,
|
||
<elevation id="00097E41FFBFFFBCFE86F9F58884F9D1" box="[274,426,1605,1632]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="m" value="1700.0">
|
||
<quantity id="4CDC3497FFBFFFBCFE86F9F58845F9EE" box="[274,363,1605,1632]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="m" value="1700.0">1700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FF9ABFB2FFBFFFBCFE23F9F68B04F9D1" box="[439,554,1605,1632]" pageId="13" pageNumber="14" value="1984-01">
|
||
<collectingDate id="EFDE465AFFBFFFBCFE23F9F68B04F9D1" box="[439,554,1605,1632]" pageId="13" pageNumber="14" value="1984-01">Jan. 1984</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26D1FCA4FFBFFFBCFDA2F9F68BE4F9D1" box="[566,714,1605,1632]" pageId="13" pageNumber="14">Jan Kielland</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED3501B7FFBFFFBCFC98F9F58A46F9D1" box="[780,872,1605,1631]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="13" pageNumber="14">ZMUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ECAF9FFBFFFBFFF29F9368817FAD4" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8B9B9972FFBFFFBCFF29F9368860F92E" blockId="13.[189,334,1670,1696]" box="[189,334,1670,1696]" pageId="13" pageNumber="14">
|
||
<heading id="D0D32E1EFFBFFFBCFF29F9368860F92E" bold="true" box="[189,334,1670,1696]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B9504560FFBFFFBCFF29F9368860F92E" bold="true" box="[189,334,1670,1696]" pageId="13" pageNumber="14">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBFFFBCFF7EF9058804F941" blockId="13.[189,1399,1717,1822]" box="[234,298,1717,1743]" pageId="13" pageNumber="14">
|
||
<emphasis id="B9504560FFBFFFBCFF7EF9058804F941" bold="true" box="[234,298,1717,1743]" pageId="13" pageNumber="14">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBFFFBCFF29F9508AB5F893" blockId="13.[189,1399,1717,1822]" pageId="13" pageNumber="14">
|
||
Based on studied specimens. Details from original description (
|
||
<bibRefCitation id="EFB5E483FFBFFFBCFC37F9508D00F975" author="Kraus O." box="[931,1070,1760,1787]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="13" pageNumber="14" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus 1960</bibRefCitation>
|
||
) in parentheses. Both of the new specimens are strongly fragmented and completely faded.
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBFFFBCFF29F8F58B4DF828" blockId="13.[189,1399,1860,1958]" pageId="13" pageNumber="14">
|
||
SIZE. Diameter
|
||
<quantity id="4CDC3497FFBFFFBCFEE7F8F58B2CF8D1" box="[371,514,1861,1887]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.7" metricValueMin="4.1" pageId="13" pageNumber="14" unit="mm" value="4.4" valueMax="4.7" valueMin="4.1">4.1–4.7 mm</quantity>
|
||
(4.0 mm), specimen from Mazombo with ca 64 podous rings, no apodous rings in front of telson, specimen from Nyumbenito Mt probably incomplete (48 podous rings present, no apodous rings in front of telson).
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBFFFBCFF29F87D8A11F869" blockId="13.[189,831,1996,2023]" box="[189,831,1996,2023]" pageId="13" pageNumber="14">HEAD. Without peculiarities. Five supralabral setae (6).</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FEBB8881FEC7" blockId="14.[189,1399,267,329]" pageId="14" pageNumber="15">COLLUM. With two complete furrows on each side, none of them marginal (two distinct furrows, marginal furrow less distinct).</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FEDD888CFE7E" blockId="14.[189,1399,364,496]" pageId="14" pageNumber="15">
|
||
BODY RINGS. Almost perfect cylinders, not vaulted; suture straight; ozopores starting from ring 6, placed ca ⅓ of metazonite length between suture and limbus. Ca 14 metazonital striae, reaching almost up to ozopore (10–11 striae, not at all reaching ozopore level). Surface microsculpture (
|
||
<figureCitation id="131F85F7FFBCFFBFFB2CFE028C17FE43" box="[1208,1337,434,461]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="14" pageNumber="15">Fig. 4D, F</figureCitation>
|
||
), see Remarks for genus.
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FDA48A60FDDF" blockId="14.[189,1398,531,593]" pageId="14" pageNumber="15">ANAL VALVES. Dorsally drawn out into pointed triangular process, ventrally with small protruding knob; mesal margin raised, setiferous tubercles not detectable.</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FDC58BE7FD01" blockId="14.[189,713,628,655]" box="[189,713,628,655]" pageId="14" pageNumber="15">
|
||
LIMBUS (
|
||
<figureCitation id="131F85F7FFBCFFBFFEB5FDC488B0FD01" box="[289,414,628,655]" captionStart="Fig.4" captionStartId="11.[189,232,1865,1891]" captionTargetBox="[189,1399,522,1830]" captionTargetId="figure@11.[173,1414,507,1840]" captionTargetPageId="11" captionText="Fig.4.Helicochetusspp.,♂♂,fromUdzungwaMts.A–B.H.dimidiatus(Peters,1855).C–F.H.mutaba Kraus, 1960. A, C, E. Limbus; in E the limbus proper is damaged (remains are seen to the left), so the round-lobed “sublimbus” becomes visible.B, D, F. Surface microsculpture of limbus. Scalebars:A, C, E = 0.01 mm; B, D, F = 0.002 mm." httpUri="https://zenodo.org/record/1146164/files/figure.png" pageId="14" pageNumber="15">Fig. 4C, E</figureCitation>
|
||
). See Remarks for genus.
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FD028C66FD42" blockId="14.[189,1352,690,717]" box="[189,1352,690,717]" pageId="14" pageNumber="15">MALE LEGS. With postfemoral ventral pads from first post-gonopodal leg-pair 3–5 until ca midbody.</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FD408B66FCFD" blockId="14.[189,1399,751,884]" pageId="14" pageNumber="15">
|
||
GONOPOD COXA (
|
||
<figureCitation id="131F85F7FFBCFFBFFE14FD408B2AFC84" box="[384,516,751,778]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6A–B</figureCitation>
|
||
). Lateral margin convex in basal half. Proplica (
|
||
<emphasis id="B9504560FFBCFFBFFBDBFD408D42FC87" box="[1103,1132,752,777]" italics="true" pageId="14" pageNumber="15">pp</emphasis>
|
||
) simple, with straight mesal margin, ending in small proplical lobe (
|
||
<emphasis id="B9504560FFBCFFBFFC96FCA38A09FCA3" box="[770,807,787,813]" italics="true" pageId="14" pageNumber="15">ppl</emphasis>
|
||
). Metaplica (
|
||
<emphasis id="B9504560FFBCFFBFFC46FCA48AD8FCA3" box="[978,1014,788,813]" italics="true" pageId="14" pageNumber="15">mp</emphasis>
|
||
) higher than proplica, apically regularly rounded, subapically with low mesal flange (
|
||
<emphasis id="B9504560FFBCFFBFFCADFC868A42FCDE" box="[825,876,822,848]" italics="true" pageId="14" pageNumber="15">mmf</emphasis>
|
||
), mesal margin of flange shallowly concave; distal, basad process (
|
||
<emphasis id="B9504560FFBCFFBFFE2AFCE988F5FCFD" box="[446,475,857,883]" italics="true" pageId="14" pageNumber="15">bp</emphasis>
|
||
) slender.
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FC27883DFB60" blockId="14.[189,1399,919,1262]" pageId="14" pageNumber="15">
|
||
GONOPOD TELOPODITE (
|
||
<figureCitation id="131F85F7FFBCFFBFFE55FC278B6AFC3C" box="[449,580,919,946]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6C–G</figureCitation>
|
||
). Basomere including torsotope without spines, arculus 90°. A large, shovel-like flange (
|
||
<emphasis id="B9504560FFBCFFBFFE08FC0A8890FC5A" box="[412,446,954,980]" italics="true" pageId="14" pageNumber="15">shl</emphasis>
|
||
) (“Tibialdorn”) present at level of post-torsal narrowing (
|
||
<figureCitation id="131F85F7FFBCFFBFFBC3FC0A8D81FC5B" box="[1111,1199,954,981]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6F</figureCitation>
|
||
). A long, slender, curved spine (
|
||
<emphasis id="B9504560FFBCFFBFFEFEFC6E88AAFC79" box="[362,388,990,1015]" italics="true" pageId="14" pageNumber="15">ps</emphasis>
|
||
) at base of solenomere (
|
||
<figureCitation id="131F85F7FFBCFFBFFD21FC6D8A1AFC76" box="[693,820,989,1016]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6E–F</figureCitation>
|
||
). Solenomere (
|
||
<emphasis id="B9504560FFBCFFBFFC78FC6D8D3BFC79" box="[1004,1045,989,1015]" italics="true" pageId="14" pageNumber="15">slm</emphasis>
|
||
) slender, round in transverse section, as long as telomere (
|
||
<emphasis id="B9504560FFBCFFBFFD9BFBB18B02FB94" box="[527,556,1025,1050]" italics="true" pageId="14" pageNumber="15">tm</emphasis>
|
||
), apically wound up in tight, corkscrew-like spiral; spiraled part basally delimited by tight, knob-like turn (
|
||
<emphasis id="B9504560FFBCFFBFFDF5FB948B53FBB0" box="[609,637,1060,1086]" italics="true" pageId="14" pageNumber="15">kn</emphasis>
|
||
) (“knotige Drehung”) of solenomere shaft (
|
||
<figureCitation id="131F85F7FFBCFFBFFB04FB948DDFFBB1" box="[1168,1265,1060,1087]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6G</figureCitation>
|
||
). Telomere basically a moderately narrow band curved in almost complete circle; with a curved, pointed basal lamella (
|
||
<emphasis id="B9504560FFBCFFBFFEBDFBDA8866FB0A" box="[297,328,1130,1156]" italics="true" pageId="14" pageNumber="15">btl</emphasis>
|
||
) (“Grundblatt”) pointing into main telomere curvature and with 1–2 small denticles on concave side (
|
||
<figureCitation id="131F85F7FFBCFFBFFEF7FB3D88F3FB26" box="[355,477,1165,1192]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6C, E</figureCitation>
|
||
) (no denticles); tip of telomere on specimen from Mazombo with several long, thin, spinelike processes from margin (
|
||
<figureCitation id="131F85F7FFBCFFBFFD0CFB008BD4FB45" box="[664,762,1200,1227]" captionStart="Fig. 6" captionStartId="15.[190,233,1779,1805]" captionTargetBox="[227,1365,267,1744]" captionTargetId="figure@15.[216,1373,262,1747]" captionTargetPageId="15" captionText="Fig. 6. Helicochetus mutaba Kraus, 1960, ♂, from Mazombo. Left gonopod. A–B. Coxa. A. Anterior view. B. Mesal view. C–G. Telopodite. C. Anterior-lateral view. D. Detail of C (tip of telomere). E. Mesal-posterior view. F. Anterior-ventral view. G. Detail of F (tip of solenomere). Abbreviations: bp = apical basad process of coxa; btl = basal lamella of telomere; kn = knob-like turn of solenomere; mmf = mesal metaplical flange; mp = metaplica; pp = proplica; ppl = proplical lobe; ps = proximal solenomeral spine; shl = shovel-like flange; slm = solenomere; tm = telomere. Scale bars:A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm." httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="14" pageNumber="15">Fig. 6D</figureCitation>
|
||
), in specimen from Nyumbenito Mt more shallowly serrate.
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF7EFAA5886BFAA1" blockId="14.[189,325,1301,1370]" box="[234,325,1301,1327]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9504560FFBCFFBFFF7EFAA5886BFAA1" bold="true" box="[234,325,1301,1327]" pageId="14" pageNumber="15">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FAF08817FAD4" blockId="14.[189,325,1301,1370]" box="[189,313,1344,1370]" pageId="14" pageNumber="15">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ECAF9FFBCFFBFFF29FA318854F9BD" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FA318878FA15" blockId="14.[189,342,1409,1435]" box="[189,342,1409,1435]" pageId="14" pageNumber="15">
|
||
<heading id="D0D32E1EFFBCFFBFFF29FA318878FA15" bold="true" box="[189,342,1409,1435]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B9504560FFBCFFBFFF29FA318878FA15" bold="true" box="[189,342,1409,1435]" pageId="14" pageNumber="15">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29FA008854F9BD" blockId="14.[189,1399,1456,1587]" pageId="14" pageNumber="15">
|
||
Hitherto known only from the
|
||
<typeStatus id="549F27D0FFBCFFBFFDBCFA008B75FA44" box="[552,603,1456,1482]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality,
|
||
<collectingCounty id="62FAE1FEFFBCFFBFFD5CFA008A0CFA44" box="[712,802,1456,1482]" pageId="14" pageNumber="15">Mutaba</collectingCounty>
|
||
in the vicinity of
|
||
<collectingMunicipality id="6BFF0308FFBCFFBFFC63FA008D77FA44" box="[1015,1113,1456,1482]" pageId="14" pageNumber="15">Kirungu</collectingMunicipality>
|
||
(=
|
||
<location id="8EFBCFA9FFBCFFBFFB17FA008C31FA45" LSID="urn:lsid:plazi:treatment:038D2864FFBFFFA1FDDAFBBA8BDBFA76:8EFBCFA9FFBCFFBFFB17FA008C31FA45" box="[1155,1311,1456,1483]" country="Democratic Republic of the Congo" county="Mutaba" municipality="Kirungu" name="Baudoinville" pageId="14" pageNumber="15">Baudoinville</location>
|
||
), in
|
||
<collectingCountry id="F333D9E2FFBCFFBFFAC7FA008B73FA63" name="Democratic Republic of the Congo" pageId="14" pageNumber="15">the Democratic Republic of the Congo</collectingCountry>
|
||
. The two records from
|
||
<collectingCountry id="F333D9E2FFBCFFBFFCE4FA638AF3FA63" box="[880,989,1491,1517]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
|
||
, Nyumbenito Mt in the Udzungwa Range and
|
||
<collectorName id="26D1FCA4FFBCFFBFFEABFA4688FFF99F" box="[319,465,1526,1553]" pageId="14" pageNumber="15">Mazombo N</collectorName>
|
||
of the Udzungwas, thus represent a considerable range extension. See, however, Remarks below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ECAF9FFBCFFA1FF29F9EB88B4FA8E" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="8B9B9972FFBCFFBFFF29F9EB8801F9FB" blockId="14.[189,303,1627,1653]" box="[189,303,1627,1653]" pageId="14" pageNumber="15">
|
||
<heading id="D0D32E1EFFBCFFBFFF29F9EB8801F9FB" bold="true" box="[189,303,1627,1653]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B9504560FFBCFFBFFF29F9EB8801F9FB" bold="true" box="[189,303,1627,1653]" pageId="14" pageNumber="15">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFBCFFBFFF29F939888CF86E" blockId="14.[189,1399,1673,2017]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="4C24E2F1FFBCFFBFFF29F939889BF92D" authorityName="Kraus" authorityYear="1960" box="[189,437,1673,1699]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mutaba">
|
||
<emphasis id="B9504560FFBCFFBFFF29F939889BF92D" box="[189,437,1673,1699]" italics="true" pageId="14" pageNumber="15">Helicochetus mutaba</emphasis>
|
||
</taxonomicName>
|
||
belongs to a group of very similar nominal species characterized by the following combination of key characters: a shovel-like lamella (“Tibialdorn”) at the level of the post-torsal narrowing; no ventral pads on male tibiae; telomere with a basal lamella (“Grundblatt”); tip of telomere not drawn out into a long process, with several short to moderately long, spine-like outgrowths from the margin. This group consists of
|
||
<taxonomicName id="4C24E2F1FFBCFFBFFDF0F8A78A83F8BF" authority="Kraus, 1957" authorityName="Kraus" authorityYear="1957" box="[612,941,1814,1841]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="digititarsus">
|
||
<emphasis id="B9504560FFBCFFBFFDF0F8A78A3FF8BE" box="[612,785,1814,1840]" italics="true" pageId="14" pageNumber="15">H. digititarsus</emphasis>
|
||
Kraus, 1957
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F333D9E2FFBCFFBFFC2AF8A68C44F8BE" box="[958,1386,1814,1840]" name="Democratic Republic of the Congo" pageId="14" pageNumber="15">
|
||
Democratic Republic of the
|
||
<collectingRegion id="49E05790FFBCFFBFFA83F8A68C44F8BE" box="[1303,1386,1814,1840]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="14" pageNumber="15">Congo</collectingRegion>
|
||
</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C24E2F1FFBCFFBFFF29F88A8B49F8DA" authority="(Pocock, 1896) (Kenya)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[189,615,1849,1876]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gregorii">
|
||
<emphasis id="B9504560FFBCFFBFFF29F88A886EF8DD" box="[189,320,1850,1875]" italics="true" pageId="14" pageNumber="15">H. gregorii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB5E483FFBCFFBFFEC4F88988DAF8DA" author="Pocock R. I." box="[336,500,1849,1876]" firstAuthor="Pocock" journalOrPublisher="Annals and Magazine of Natural History, Series 6" pageId="14" pageNumber="15" pagination="425 - 444" part="17" publicationUrl="https://doi.org/10.1080/00222939608680394" refId="ref10362" refString="Pocock R. I. 1896. On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History, Series 6, 17: 425 - 444. https: // doi. org / 10.1080 / 00222939608680394" title="On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa" type="journal article" year="1896">Pocock, 1896</bibRefCitation>
|
||
) (
|
||
<collectingCountry id="F333D9E2FFBCFFBFFD98F88A8B70F8DA" box="[524,606,1850,1876]" name="Kenya" pageId="14" pageNumber="15">Kenya</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C24E2F1FFBCFFBFFDE7F88A8D0BF8DA" authority="(Kraus, 1960) (Zambia)" baseAuthorityName="Kraus" baseAuthorityYear="1960" box="[627,1061,1849,1876]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="monodon">
|
||
<emphasis id="B9504560FFBCFFBFFDE7F88A8A2BF8DD" box="[627,773,1849,1875]" italics="true" pageId="14" pageNumber="15">H. monodon</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB5E483FFBCFFBFFC81F88A8A86F8DA" author="Kraus O." box="[789,936,1849,1876]" firstAuthor="Kraus" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="14" pageNumber="15" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus, 1960</bibRefCitation>
|
||
) (
|
||
<collectingCountry id="F333D9E2FFBCFFBFFC54F8898D33F8DA" box="[960,1053,1849,1876]" name="Zambia" pageId="14" pageNumber="15">Zambia</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C24E2F1FFBCFFBFFBF4F88A8DF2F8DD" authorityName="Kraus" authorityYear="1960" box="[1120,1244,1849,1875]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mutaba">
|
||
<emphasis id="B9504560FFBCFFBFFBF4F88A8DF2F8DD" box="[1120,1244,1849,1875]" italics="true" pageId="14" pageNumber="15">H. mutaba</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F333D9E2FFBCFFBFFB78F88988F7F8F9" name="Democratic Republic of the Congo" pageId="14" pageNumber="15">
|
||
Democratic Republic of the
|
||
<collectingRegion id="49E05790FFBCFFBFFE1CF8ED88F7F8F9" box="[392,473,1885,1911]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="14" pageNumber="15">Congo</collectingRegion>
|
||
</collectingCountry>
|
||
,
|
||
<collectingCountry id="F333D9E2FFBCFFBFFE7EF8ED8B77F8F6" box="[490,601,1885,1912]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
|
||
).
|
||
<tableCitation id="C6A6ACC9FFBCFFBFFDE4F8ED8BE2F8F9" box="[624,716,1885,1911]" captionStart="Table 2" captionStartId="16.[189,258,268,294]" captionTargetPageId="16" captionText="Table 2. Comparison of four species of HelicochetusAttems, 1909." httpUri="http://table.plazi.org/id/DF5BC9FAFFA2FFA1FF29FEBC8AF8FEAB" pageId="14" pageNumber="15" tableUuid="DF5BC9FAFFA2FFA1FF29FEBC8AF8FEAB">Table 2</tableCitation>
|
||
shows the differences between these four species. The outline of the basal lamella of the telomere differs between the species, but it is not unlikely that future collections will bridge the small morphological gaps and that some or all of the four species will have to be synonymized.
|
||
</paragraph>
|
||
<caption id="DF5BC9FAFFBDFFBEFF2AF9438B47F86F" httpUri="https://zenodo.org/record/1146168/files/figure.png" pageId="15" pageNumber="16" targetBox="[227,1365,267,1744]" targetPageId="15">
|
||
<paragraph id="8B9B9972FFBDFFBEFF2AF9438B47F86F" blockId="15.[190,1400,1779,2017]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9504560FFBDFFBEFF2AF9438823F883" bold="true" box="[190,269,1779,1805]" pageId="15" pageNumber="16">Fig. 6.</emphasis>
|
||
<taxonomicName id="4C24E2F1FFBDFFBEFE82F9438B82F883" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[278,684,1779,1806]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mutaba">
|
||
<emphasis id="B9504560FFBDFFBEFE82F9438B3EF883" box="[278,528,1779,1805]" italics="true" pageId="15" pageNumber="16">Helicochetus mutaba</emphasis>
|
||
<bibRefCitation id="EFB5E483FFBDFFBEFD8DF9448B82F883" author="Kraus O." box="[537,684,1779,1806]" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="15" pageNumber="16" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
, ♂, from Mazombo. Left gonopod.
|
||
<emphasis id="B9504560FFBDFFBEFBCEF9438DBCF883" bold="true" box="[1114,1170,1779,1805]" pageId="15" pageNumber="16">A–B</emphasis>
|
||
. Coxa.
|
||
<emphasis id="B9504560FFBDFFBEFB64F9438C2BF883" bold="true" box="[1264,1285,1779,1805]" pageId="15" pageNumber="16">A</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="B9504560FFBDFFBEFE9DF8A78833F8BF" bold="true" box="[265,285,1815,1841]" pageId="15" pageNumber="16">B</emphasis>
|
||
. Mesal view.
|
||
<emphasis id="B9504560FFBDFFBEFE5BF8A68B24F8BE" bold="true" box="[463,522,1814,1840]" pageId="15" pageNumber="16">C–G</emphasis>
|
||
. Telopodite.
|
||
<emphasis id="B9504560FFBDFFBEFD3AF8A68BEDF8BE" bold="true" box="[686,707,1814,1840]" pageId="15" pageNumber="16">C</emphasis>
|
||
. Anterior-lateral view.
|
||
<emphasis id="B9504560FFBDFFBEFC77F8A78AD6F8BF" bold="true" box="[995,1016,1815,1841]" pageId="15" pageNumber="16">D</emphasis>
|
||
. Detail of C (tip of telomere).
|
||
<emphasis id="B9504560FFBDFFBEFF2AF88A89FCF8DA" bold="true" box="[190,210,1850,1876]" pageId="15" pageNumber="16">E</emphasis>
|
||
. Mesal-posterior view.
|
||
<emphasis id="B9504560FFBDFFBEFE7AF88A8B2EF8DA" bold="true" box="[494,512,1850,1876]" pageId="15" pageNumber="16">F</emphasis>
|
||
. Anterior-ventral view.
|
||
<emphasis id="B9504560FFBDFFBEFC8BF8898A18F8DD" bold="true" box="[799,822,1849,1875]" pageId="15" pageNumber="16">G</emphasis>
|
||
. Detail of F (tip of solenomere). Abbreviations:
|
||
<emphasis id="B9504560FFBDFFBEFF2AF8ED89F5F8F9" box="[190,219,1885,1911]" italics="true" pageId="15" pageNumber="16">bp</emphasis>
|
||
= apical basad process of coxa;
|
||
<emphasis id="B9504560FFBDFFBEFDCAF8ED8B53F8F9" box="[606,637,1885,1911]" italics="true" pageId="15" pageNumber="16">btl</emphasis>
|
||
= basal lamella of telomere;
|
||
<emphasis id="B9504560FFBDFFBEFC4EF8ED8AD8F8F9" box="[986,1014,1885,1911]" italics="true" pageId="15" pageNumber="16">kn</emphasis>
|
||
= knob-like turn of solenomere;
|
||
<emphasis id="B9504560FFBDFFBEFF2AF83089DFF814" box="[190,241,1920,1946]" italics="true" pageId="15" pageNumber="16">mmf</emphasis>
|
||
= mesal metaplical flange;
|
||
<emphasis id="B9504560FFBDFFBEFDD1F8318B47F814" box="[581,617,1921,1946]" italics="true" pageId="15" pageNumber="16">mp</emphasis>
|
||
= metaplica;
|
||
<emphasis id="B9504560FFBDFFBEFC87F8318A1EF814" box="[787,816,1921,1946]" italics="true" pageId="15" pageNumber="16">pp</emphasis>
|
||
= proplica;
|
||
<emphasis id="B9504560FFBDFFBEFC5DF8308AC0F814" box="[969,1006,1920,1946]" italics="true" pageId="15" pageNumber="16">ppl</emphasis>
|
||
= proplical lobe;
|
||
<emphasis id="B9504560FFBDFFBEFB59F8318DC9F814" box="[1229,1255,1921,1946]" italics="true" pageId="15" pageNumber="16">ps</emphasis>
|
||
= proximal solenomeral spine;
|
||
<emphasis id="B9504560FFBDFFBEFE37F81388EBF833" box="[419,453,1955,1981]" italics="true" pageId="15" pageNumber="16">shl</emphasis>
|
||
= shovel-like flange;
|
||
<emphasis id="B9504560FFBDFFBEFD5DF8138BDCF833" box="[713,754,1955,1981]" italics="true" pageId="15" pageNumber="16">slm</emphasis>
|
||
= solenomere;
|
||
<emphasis id="B9504560FFBDFFBEFC3FF8148AE6F833" box="[939,968,1956,1981]" italics="true" pageId="15" pageNumber="16">tm</emphasis>
|
||
= telomere. Scale bars: A–C, E–F = 0.2 mm; D = 0.05 mm; G = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5BC9FAFFA2FFA1FF29FEBC8AF8FEAB" ID-Table-UUID="DF5BC9FAFFA2FFA1FF29FEBC8AF8FEAB" box="[189,982,267,294]" httpUri="http://table.plazi.org/id/DF5BC9FAFFA2FFA1FF29FEBC8AF8FEAB" pageId="16" pageNumber="17" targetBox="[203,1389,335,1065]" targetIsTable="true" targetPageId="16">
|
||
<paragraph id="8B9B9972FFA2FFA1FF29FEBC8AF8FEAB" blockId="16.[189,982,267,294]" box="[189,982,267,294]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9504560FFA2FFA1FF29FEBC8831FEAB" bold="true" box="[189,287,267,294]" pageId="16" pageNumber="17">Table 2.</emphasis>
|
||
Comparison of four species of
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FD05FEBB8AFFFEAB" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" box="[657,977,267,294]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9504560FFA2FFA1FD05FEBB8A04FEAB" box="[657,810,267,293]" italics="true" pageId="16" pageNumber="17">Helicochetus</emphasis>
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FCA4FEBC8AFFFEAB" author="Attems C." box="[816,977,267,294]" journalOrPublisher="Zoologischer Anzeiger" pageId="16" pageNumber="17" pagination="156 - 159" part="34" publicationUrl="https://biodiversitylibrary.org/page/30144159" refId="ref8995" refString="Attems C. 1909 a. Zur Systematik der Spirostreptoidea. Zoologischer Anzeiger 34 (5): 156 - 159. Available from https: // biodiversitylibrary. org / page / 30144159 [accessed 27 Nov. 2017]." title="Zur Systematik der Spirostreptoidea" type="journal article" year="1909">Attems, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B9972FFA2FFA1FE21FEE08C0DFC7A" pageId="16" pageNumber="17">
|
||
<table id="F9246BD2FFA2004EFF5FFEFF8C43FBA7" box="[203,1389,335,1065]" gridcols="6" gridrows="9" pageId="16" pageNumber="17">
|
||
<tr id="35149B30FFA2004EFF5FFEFF8C43FE14" box="[203,1389,335,410]" gridrow="0" pageId="16" pageNumber="17" rowspan-0="1">
|
||
<th id="76C5F24CFFA2004EFE0EFEFF8B14FE14" box="[410,570,335,410]" gridcol="1" gridrow="0" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FE21FEE08B31FEEB" authorityName="Kraus" authorityYear="1960" box="[437,543,335,357]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mutaba">
|
||
<emphasis id="B9504560FFA2FFA1FE21FEE08B31FEEB" bold="true" box="[437,543,335,357]" italics="true" pageId="16" pageNumber="17">H. mutaba</emphasis>
|
||
</taxonomicName>
|
||
Tanzania
|
||
</th>
|
||
<th id="76C5F24CFFA2004EFDC5FEFF8A2BFE14" box="[593,773,335,410]" gridcol="2" gridrow="0" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FDE2FEE08BCEFEEB" authorityName="Kraus" authorityYear="1960" box="[630,736,335,357]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mutaba">
|
||
<emphasis id="B9504560FFA2FFA1FDE2FEE08BCEFEEB" bold="true" box="[630,736,335,357]" italics="true" pageId="16" pageNumber="17">H. mutaba</emphasis>
|
||
</taxonomicName>
|
||
D.R. of the Congo (from
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FD19FE358BD0FE14" author="Kraus O." box="[653,766,388,410]" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="16" pageNumber="17" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus 1960</bibRefCitation>
|
||
)
|
||
</th>
|
||
<th id="76C5F24CFFA2004EFC8EFEFF8AE0FE14" box="[794,974,335,410]" gridcol="3" gridrow="0" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FCB9FEE08A95FEEB" authorityName="Kraus" authorityYear="1957" box="[813,955,335,357]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="digititarsus">
|
||
<emphasis id="B9504560FFA2FFA1FCB9FEE08A95FEEB" bold="true" box="[813,955,335,357]" italics="true" pageId="16" pageNumber="17">H. digititarsus</emphasis>
|
||
</taxonomicName>
|
||
(from
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FCC3FEDA8AE9FE0E" author="Kraus O." box="[855,967,362,384]" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="16" pageNumber="17" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus 1960</bibRefCitation>
|
||
)
|
||
</th>
|
||
<th id="76C5F24CFFA2004EFC72FEFF8D8FFE14" box="[998,1185,335,410]" gridcol="4" gridrow="0" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FB98FEE08D54FEEB" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[1036,1146,336,357]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gregorii">
|
||
<emphasis id="B9504560FFA2FFA1FB98FEE08D54FEEB" bold="true" box="[1036,1146,336,357]" italics="true" pageId="16" pageNumber="17">H. gregorii</emphasis>
|
||
</taxonomicName>
|
||
(from
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FBB7FEDA8D8EFE0E" author="Pocock R. I." box="[1059,1184,362,384]" journalOrPublisher="Annals and Magazine of Natural History, Series 6" pageId="16" pageNumber="17" pagination="425 - 444" part="17" publicationUrl="https://doi.org/10.1080/00222939608680394" refId="ref10362" refString="Pocock R. I. 1896. On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History, Series 6, 17: 425 - 444. https: // doi. org / 10.1080 / 00222939608680394" title="On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa" type="journal article" year="1896">Pocock 1896</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FB88FE358DA2FE14" author="Kraus O." box="[1052,1164,388,410]" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="16" pageNumber="17" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus 1960</bibRefCitation>
|
||
)
|
||
</th>
|
||
<th id="76C5F24CFFA2004EFB2DFEFF8C43FE14" box="[1209,1389,335,410]" gridcol="5" gridrow="0" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FB41FEE08C7FFEEB" baseAuthorityName="Kraus" baseAuthorityYear="1960" box="[1237,1361,335,357]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="monodon">
|
||
<emphasis id="B9504560FFA2FFA1FB41FEE08C7FFEEB" bold="true" box="[1237,1361,335,357]" italics="true" pageId="16" pageNumber="17">H. monodon</emphasis>
|
||
</taxonomicName>
|
||
(from
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FB62FEDA8C48FE0E" author="Kraus O." box="[1270,1382,362,384]" journalOrPublisher="Annalen van het Koninklijk Museum van Belgisch-Congo" pageId="16" pageNumber="17" pagination="1 - 207" part="82" refId="ref10124" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." title="Athiopische Diplopoden I. Monographie der Odontopygidae - Odontopyginae (Diplopoda, Spirostreptoidea)" type="journal article" year="1960">Kraus 1960</bibRefCitation>
|
||
)
|
||
</th>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFE028C43FE46" box="[203,1389,434,456]" gridrow="1" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFE0288ACFE46" box="[203,386,434,456]" gridcol="0" gridrow="1" pageId="16" pageNumber="17">No. of body rings</th>
|
||
<td id="76C5F24CFFA2004EFE0EFE028B14FE46" box="[410,570,434,456]" gridcol="1" gridrow="1" pageId="16" pageNumber="17">ca 64</td>
|
||
<td id="76C5F24CFFA2004EFDC5FE028A2BFE46" box="[593,773,434,456]" gridcol="2" gridrow="1" pageId="16" pageNumber="17">?</td>
|
||
<td id="76C5F24CFFA2004EFC8EFE028AE0FE46" box="[794,974,434,456]" gridcol="3" gridrow="1" pageId="16" pageNumber="17">60</td>
|
||
<td id="76C5F24CFFA2004EFC72FE028D8FFE46" box="[998,1185,434,456]" gridcol="4" gridrow="1" pageId="16" pageNumber="17">71–72</td>
|
||
<td id="76C5F24CFFA2004EFB2DFE028C43FE46" box="[1209,1389,434,456]" gridcol="5" gridrow="1" pageId="16" pageNumber="17">64</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFE688C43FE60" box="[203,1389,472,494]" gridrow="2" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFE6888ACFE60" box="[203,386,472,494]" gridcol="0" gridrow="2" pageId="16" pageNumber="17">Body diameter (♂)</th>
|
||
<td id="76C5F24CFFA2004EFE0EFE688B14FE60" box="[410,570,472,494]" gridcol="1" gridrow="2" pageId="16" pageNumber="17">4.1–4.7 mm</td>
|
||
<td id="76C5F24CFFA2004EFDC5FE688A2BFE60" box="[593,773,472,494]" gridcol="2" gridrow="2" pageId="16" pageNumber="17">4.0</td>
|
||
<td id="76C5F24CFFA2004EFC8EFE688AE0FE60" box="[794,974,472,494]" gridcol="3" gridrow="2" pageId="16" pageNumber="17">3.7</td>
|
||
<td id="76C5F24CFFA2004EFC72FE688D8FFE60" box="[998,1185,472,494]" gridcol="4" gridrow="2" pageId="16" pageNumber="17">?</td>
|
||
<td id="76C5F24CFFA2004EFB2DFE688C43FE60" box="[1209,1389,472,494]" gridcol="5" gridrow="2" pageId="16" pageNumber="17">3.5</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFE4E8C43FDC7" box="[203,1389,510,585]" gridrow="3" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFE4E88ACFDC7" box="[203,386,510,585]" gridcol="0" gridrow="3" pageId="16" pageNumber="17">
|
||
Distal, basad process (
|
||
<emphasis id="B9504560FFA2FFA1FEA6FDA98864FDA0" box="[306,330,537,558]" italics="true" pageId="16" pageNumber="17">bp</emphasis>
|
||
) on coxa
|
||
</th>
|
||
<td id="76C5F24CFFA2004EFE0EFE4E8B14FDC7" box="[410,570,510,585]" gridcol="1" gridrow="3" pageId="16" pageNumber="17">long</td>
|
||
<td id="76C5F24CFFA2004EFDC5FE4E8A2BFDC7" box="[593,773,510,585]" gridcol="2" gridrow="3" pageId="16" pageNumber="17">long</td>
|
||
<td id="76C5F24CFFA2004EFC8EFE4E8AE0FDC7" box="[794,974,510,585]" gridcol="3" gridrow="3" pageId="16" pageNumber="17">long</td>
|
||
<td id="76C5F24CFFA2004EFC72FE4E8D8FFDC7" box="[998,1185,510,585]" gridcol="4" gridrow="3" pageId="16" pageNumber="17">short</td>
|
||
<td id="76C5F24CFFA2004EFB2DFE4E8C43FDC7" box="[1209,1389,510,585]" gridcol="5" gridrow="3" pageId="16" pageNumber="17">long</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFDE98C43FD07" box="[203,1389,601,649]" gridrow="4" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFDE988ACFD07" box="[203,386,601,649]" gridcol="0" gridrow="4" pageId="16" pageNumber="17">Mesal margin of distal part of coxa</th>
|
||
<td id="76C5F24CFFA2004EFE0EFDE98B14FD07" box="[410,570,601,649]" gridcol="1" gridrow="4" pageId="16" pageNumber="17">very shallowly concave</td>
|
||
<td id="76C5F24CFFA2004EFDC5FDE98A2BFD07" box="[593,773,601,649]" gridcol="2" gridrow="4" pageId="16" pageNumber="17">shallowly concave</td>
|
||
<td id="76C5F24CFFA2004EFC8EFDE98AE0FD07" box="[794,974,601,649]" gridcol="3" gridrow="4" pageId="16" pageNumber="17">with a deep sinus</td>
|
||
<td id="76C5F24CFFA2004EFC72FDE98D8FFD07" box="[998,1185,601,649]" gridcol="4" gridrow="4" pageId="16" pageNumber="17">shallowly concave</td>
|
||
<td id="76C5F24CFFA2004EFB2DFDE98C43FD07" box="[1209,1389,601,649]" gridcol="5" gridrow="4" pageId="16" pageNumber="17">shallowly concave</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFD298C43FD71" box="[203,1389,665,767]" gridrow="5" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFD2988ACFD71" box="[203,386,665,767]" gridcol="0" gridrow="5" pageId="16" pageNumber="17">
|
||
Basal lamella of telomere (
|
||
<emphasis id="B9504560FFA2FFA1FED3FD048849FD47" box="[327,359,692,713]" italics="true" pageId="16" pageNumber="17">btl)</emphasis>
|
||
</th>
|
||
<td id="76C5F24CFFA2004EFE0EFD298B14FD71" box="[410,570,665,767]" gridcol="1" gridrow="5" pageId="16" pageNumber="17">slender, pointed, curved, with a few denticles on concave margin</td>
|
||
<td id="76C5F24CFFA2004EFDC5FD298A2BFD71" box="[593,773,665,767]" gridcol="2" gridrow="5" pageId="16" pageNumber="17">slender, pointed, curved, smooth</td>
|
||
<td id="76C5F24CFFA2004EFC8EFD298AE0FD71" box="[794,974,665,767]" gridcol="3" gridrow="5" pageId="16" pageNumber="17">very slender, pointed, smooth</td>
|
||
<td id="76C5F24CFFA2004EFC72FD298D8FFD71" box="[998,1185,665,767]" gridcol="4" gridrow="5" pageId="16" pageNumber="17">broad, subrectangular, smooth</td>
|
||
<td id="76C5F24CFFA2004EFB2DFD298C43FD71" box="[1209,1389,665,767]" gridcol="5" gridrow="5" pageId="16" pageNumber="17">slender, margin in part serrate</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFCBF8C43FCFA" box="[203,1389,783,884]" gridrow="6" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFCBF88ACFCFA" box="[203,386,783,884]" gridcol="0" gridrow="6" pageId="16" pageNumber="17">Tip of telomere</th>
|
||
<td id="76C5F24CFFA2004EFE0EFCBF8B14FCFA" box="[410,570,783,884]" gridcol="1" gridrow="6" pageId="16" pageNumber="17">margin with short or long, slender denticles</td>
|
||
<td id="76C5F24CFFA2004EFDC5FCBF8A2BFCFA" box="[593,773,783,884]" gridcol="2" gridrow="6" pageId="16" pageNumber="17">margin with short denticles</td>
|
||
<td id="76C5F24CFFA2004EFC8EFCBF8AE0FCFA" box="[794,974,783,884]" gridcol="3" gridrow="6" pageId="16" pageNumber="17">margin with short denticles</td>
|
||
<td id="76C5F24CFFA2004EFC72FCBF8D8FFCFA" box="[998,1185,783,884]" gridcol="4" gridrow="6" pageId="16" pageNumber="17">with many short and long denticles, apparently not only on margin</td>
|
||
<td id="76C5F24CFFA2004EFB2DFCBF8C43FCFA" box="[1209,1389,783,884]" gridcol="5" gridrow="6" pageId="16" pageNumber="17">margin with short denticles</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFC348C43FC41" box="[203,1389,900,975]" gridrow="7" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFC3488ACFC41" box="[203,386,900,975]" gridcol="0" gridrow="7" pageId="16" pageNumber="17">
|
||
Solenomere with a ‘knot’ (
|
||
<emphasis id="B9504560FFA2FFA1FE87FC2E8804FC3D" box="[275,298,926,947]" italics="true" pageId="16" pageNumber="17">kn</emphasis>
|
||
) at base of terminal spiral
|
||
</th>
|
||
<td id="76C5F24CFFA2004EFE0EFC348B14FC41" box="[410,570,900,975]" gridcol="1" gridrow="7" pageId="16" pageNumber="17">yes</td>
|
||
<td id="76C5F24CFFA2004EFDC5FC348A2BFC41" box="[593,773,900,975]" gridcol="2" gridrow="7" pageId="16" pageNumber="17">yes</td>
|
||
<td id="76C5F24CFFA2004EFC8EFC348AE0FC41" box="[794,974,900,975]" gridcol="3" gridrow="7" pageId="16" pageNumber="17">yes</td>
|
||
<td id="76C5F24CFFA2004EFC72FC348D8FFC41" box="[998,1185,900,975]" gridcol="4" gridrow="7" pageId="16" pageNumber="17">no</td>
|
||
<td id="76C5F24CFFA2004EFB2DFC348C43FC41" box="[1209,1389,900,975]" gridcol="5" gridrow="7" pageId="16" pageNumber="17">yes</td>
|
||
</tr>
|
||
<tr id="35149B30FFA2004EFF5FFC6E8C43FBA7" box="[203,1389,990,1065]" gridrow="8" pageId="16" pageNumber="17">
|
||
<th id="76C5F24CFFA2004EFF5FFC6E88ACFBA7" box="[203,386,990,1065]" gridcol="0" gridrow="8" pageId="16" pageNumber="17">Terminal spiral of solenomere with an accessory spine</th>
|
||
<td id="76C5F24CFFA2004EFE0EFC6E8B14FBA7" box="[410,570,990,1065]" gridcol="1" gridrow="8" pageId="16" pageNumber="17">yes</td>
|
||
<td id="76C5F24CFFA2004EFDC5FC6E8A2BFBA7" box="[593,773,990,1065]" gridcol="2" gridrow="8" pageId="16" pageNumber="17">yes</td>
|
||
<td id="76C5F24CFFA2004EFC8EFC6E8AE0FBA7" box="[794,974,990,1065]" gridcol="3" gridrow="8" pageId="16" pageNumber="17">yes</td>
|
||
<td id="76C5F24CFFA2004EFC72FC6E8D8FFBA7" box="[998,1185,990,1065]" gridcol="4" gridrow="8" pageId="16" pageNumber="17">no</td>
|
||
<td id="76C5F24CFFA2004EFB2DFC6E8C43FBA7" box="[1209,1389,990,1065]" gridcol="5" gridrow="8" pageId="16" pageNumber="17">yes</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B9B9972FFA2FFA1FF29FBCC88B4FA8E" blockId="16.[189,1399,1148,1280]" pageId="16" pageNumber="17">
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FF29FBCC889CFB19" author="Enghoff" box="[189,434,1148,1175]" etAl="et al." firstAuthor="Enghoff" journalOrPublisher="European Journal of Taxonomy" pageId="16" pageNumber="17" pagination="1 - 19" part="177" publicationUrl="https://doi.org/10.5852/ejt.2016.177" refId="ref9343" refString="Enghoff H. 2016 a. A mountain of millipedes III: A new genus for three new species from the Udzungwa Mountains and surroundings, Tanzania, as well as several ' orphaned' species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 177: 1 - 19. https: // doi. org / 10.5852 / ejt. 2016.177" title="A mountain of millipedes III: A new genus for three new species from the Udzungwa Mountains and surroundings, Tanzania, as well as several ' orphaned' species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae)" type="journal article" year="2016">
|
||
Enghoff
|
||
<emphasis id="B9504560FFA2FFA1FEB7FBCD8870FB18" box="[291,350,1148,1174]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FDB3FBCD8BFEFB18" authorityName="Kraus" authorityYear="1957" box="[551,720,1148,1174]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="digititarsus">
|
||
<emphasis id="B9504560FFA2FFA1FDB3FBCD8BFEFB18" box="[551,720,1148,1174]" italics="true" pageId="16" pageNumber="17">H. digititarsus</emphasis>
|
||
</taxonomicName>
|
||
from two Tanzanian localities: 1)
|
||
<collectingRegion id="49E05790FFA2FFA1FBF4FBCD8DD3FB18" box="[1120,1277,1148,1175]" country="Tanzania" name="Mara" pageId="16" pageNumber="17">Mara Region</collectingRegion>
|
||
, Serengeti District, Seronera, Serengeti National Park; 2)
|
||
<collectingRegion id="49E05790FFA2FFA1FD78FB2F8AB1FB37" box="[748,927,1183,1209]" country="Tanzania" name="Tabora" pageId="16" pageNumber="17">Tabora Region</collectingRegion>
|
||
, Nzega District, Nzega City. In light of the above discussion, it cannot be excluded that these specimens are conspecific with the one recorded here as
|
||
<emphasis id="B9504560FFA2FFA1FE83FB5688B4FA8E" box="[279,410,1254,1280]" italics="true" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FE83FB5688B8FA8E" authorityName="Kraus" authorityYear="1960" box="[279,406,1254,1280]" class="Diplopoda" family="Odontopygidae" genus="Helicochetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mutaba">H. mutaba</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ECAF9FFA2FFA1FF29FA9E8BDBFA76" pageId="16" pageNumber="17" type="diagnosis">
|
||
<paragraph id="8B9B9972FFA2FFA1FF29FA9E8BDBFA76" blockId="16.[189,1399,1326,1529]" pageId="16" pageNumber="17">
|
||
The vial containing the male from Mazombo also contained a microvial with a mite which was presumably collected in association with the millipede. The mite was infested with numerous thalli of the fungus
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FED5FAC588A2FA01" authorityName="Cavara" authorityYear="1899" box="[321,396,1397,1423]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" higherTaxonomySource="GBIF" kingdom="Fungi" order="Laboulbeniales" pageId="16" pageNumber="17" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B9504560FFA2FFA1FED5FAC588A2FA01" box="[321,396,1397,1423]" italics="true" pageId="16" pageNumber="17">Rickia</emphasis>
|
||
</taxonomicName>
|
||
sp. (Ascomycota,
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FDFEFAC58A0DFA1E" authorityName="" authorityYear="1898" box="[618,803,1397,1424]" class="Laboulbeniomycetes" higherTaxonomySource="GBIF" kingdom="Fungi" order="Laboulbeniales" pageId="16" pageNumber="17" phylum="Ascomycota" rank="order">Laboulbeniales</taxonomicName>
|
||
). Several species of mites are known to be closely associated with millipedes (
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FD90FA288A03FA3C" author="Farfan M. A. & Klompen H." box="[516,813,1432,1458]" firstAuthor="Farfan" journalOrPublisher="International Journal of Myriapodology" pageId="16" pageNumber="17" pagination="62 - 91" part="7" publicationUrl="https://doi.org/10.3897/ijm.7.3064" refId="ref9899" refString="Farfan M. A. & Klompen H. 2012. Phoretic mite associates of millipedes (Diplopoda, Julidae) in the northern Atlantic region (North America, Europe). International Journal of Myriapodology 7: 62 - 91. https: // doi. org / 10.3897 / ijm. 7.3064" title="Phoretic mite associates of millipedes (Diplopoda, Julidae) in the northern Atlantic region (North America, Europe)" type="journal article" year="2012">Farfan & Klompen 2012</bibRefCitation>
|
||
), and several species of
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FBD9FA288DDDFA3C" authority="Cavara" authorityName="Cavara" authorityYear="1899" box="[1101,1267,1432,1458]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" higherTaxonomySource="GBIF" kingdom="Fungi" order="Laboulbeniales" pageId="16" pageNumber="17" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B9504560FFA2FFA1FBD9FA288DB6FA3C" box="[1101,1176,1432,1458]" italics="true" pageId="16" pageNumber="17">Rickia</emphasis>
|
||
Cavara
|
||
</taxonomicName>
|
||
are known to use mites as their host (
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FE7FFA0B8BA3FA58" author="Tavares I. I." box="[491,653,1467,1494]" firstAuthor="Tavares" journalOrPublisher="Mycologia Memoir" pageId="16" pageNumber="17" pagination="9" refId="ref10459" refString="Tavares I. I. 1985. Laboulbeniales (Fungi, Ascomycetes). Mycologia Memoir 9. J. Cramer, Braunschweig, Germany." title="Laboulbeniales (Fungi, Ascomycetes)." type="book chapter" year="1985">Tavares 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB5E483FFA2FFA1FD0DFA0B8AE9FA5B" author="Seeman O. D. & Nahrung H. F." box="[665,967,1467,1493]" firstAuthor="Seeman" journalOrPublisher="Australasian Mycologist" pageId="16" pageNumber="17" pagination="3 - 9" part="19" refId="ref10418" refString="Seeman O. D. & Nahrung H. F. 2000. Mites as fungal vectors? The ectoparasitic fungi of mites and their arthropod associates in Queensland. Australasian Mycologist 19 (1): 3 - 9." title="Mites as fungal vectors? The ectoparasitic fungi of mites and their arthropod associates in Queensland" type="journal article" year="2000">Seeman & Nahrung 2000</bibRefCitation>
|
||
), but this seems to be the first known instance of a
|
||
<taxonomicName id="4C24E2F1FFA2FFA1FECDFA6F888AFA77" authorityName="Cavara" authorityYear="1899" box="[345,420,1503,1529]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" higherTaxonomySource="GBIF" kingdom="Fungi" order="Laboulbeniales" pageId="16" pageNumber="17" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B9504560FFA2FFA1FECDFA6F888AFA77" box="[345,420,1503,1529]" italics="true" pageId="16" pageNumber="17">Rickia</emphasis>
|
||
</taxonomicName>
|
||
–mite–millipede association.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |