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<document id="F4B01DE28DD5031FE734FE2AC379BA89" ID-DOI="10.59893/bjc.23(2).009" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203106" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657248871" checkinUser="felipe" docAuthor="Sarker, Subir &amp; Talukder, Saranika" docDate="2023" docId="038C9D1E0462FFDCFD10FDD382BD7770" docLanguage="en" docName="BalticJColeopterol.23.2.233-239.pdf" docOrigin="Baltic Journal of Coleopterology 23 (2)" docSource="http://dx.doi.org/10.1128/mra.00329-24" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Malthininae Kiesenwetter 1852" docType="treatment" docVersion="2" lastPageNumber="239" masterDocId="FFB5E5660460FFDFFFABFFD081247205" masterDocTitle="Two new taxa of fossil Cantharidae from Baltic amber" masterLastPageNumber="239" masterPageNumber="233" pageNumber="233" updateTime="1722696919189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="B2C7D9919FA1E65E97CBF01B400740B0">Two new taxa of fossil Cantharidae from Baltic amber</mods:title>
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<mods:namePart id="4083D5EDFDB58D05FEACB975A9D173A7">Sarker, Subir</mods:namePart>
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<mods:namePart id="11018BF129BAF7DF3F202B237489B732">Talukder, Saranika</mods:namePart>
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<mods:title id="D3A8DC29CA9982C014FD2B696AB8E7D9">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="8DC4690AE1752C494C9548FE05942F09">2023</mods:date>
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<mods:number id="13B8EC1FAD6848FE0E00AA40E295794E">23</mods:number>
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<mods:identifier id="FB9DDDA2C72832CC09DD99EF76415D93" type="DOI">10.59893/bjc.23(2).009</mods:identifier>
<mods:identifier id="62A53443D112CE4BB659EE39DF99E0CB" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="E4BE64DC5193D315C8EB5F1D3D181165" type="Zenodo-Dep">13203106</mods:identifier>
</mods:mods>
<treatment id="038C9D1E0462FFDCFD10FDD382BD7770" LSID="urn:lsid:plazi:treatment:038C9D1E0462FFDCFD10FDD382BD7770" httpUri="http://treatment.plazi.org/id/038C9D1E0462FFDCFD10FDD382BD7770" lastPageId="3" pageId="2">
<subSubSection id="C33F7F830462FFDDFD10FDD383D3703C" pageId="2" type="nomenclature">
<paragraph id="8B9A2C080462FFDDFD10FDD385BE701E" blockId="2.[699,1178,515,631]" box="[699,1178,515,539]" pageId="2">
<heading id="D0D29B640462FFDDFD10FDD385BE701E" bold="true" box="[699,1178,515,539]" centered="true" fontSize="10" level="2" pageId="2" reason="3">
<emphasis id="B951F01A0462FFDDFD10FDD385BE701E" bold="true" box="[699,1178,515,539]" pageId="2">
Subfamily
<taxonomicName id="4C25578B0462FFDDFCE4FDD385B1701E" authority="Kiesenwetter" authorityName="Kiesenwetter" authorityYear="1852" box="[847,1173,515,539]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="subFamily" subFamily="Malthininae">Malthininae Kiesenwetter</taxonomicName>
,
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FDF183D3703C" blockId="2.[699,1178,515,631]" box="[699,759,545,569]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FDF183D3703C" bold="true" box="[699,759,545,569]" pageId="2">1852</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFD10FD90822F70D6" pageId="2" type="description">
<paragraph id="8B9A2C080462FFDDFD10FD908572705D" blockId="2.[699,1178,515,631]" box="[699,1110,576,600]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FD908572705D" bold="true" box="[699,1110,576,600]" pageId="2">
Tribe
<taxonomicName id="4C25578B0462FFDDFD54FD908572705D" authority="Kiesenwetter, 1852" authorityName="Kiesenwetter" authorityYear="1852" box="[767,1110,576,600]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="tribe" tribe="Malthinini">Malthinini Kiesenwetter, 1852</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FD8F850C7072" blockId="2.[699,1178,515,631]" box="[699,1064,607,631]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FD8F850C7072" bold="true" box="[699,1064,607,631]" pageId="2">
Genus
<taxonomicName id="4C25578B0462FFDDFCA1FD8F850C7072" authority="Latreille, 1806" authorityName="Latreille" authorityYear="1806" box="[778,1064,607,631]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B951F01A0462FFDDFCA1FD8F82A77072" bold="true" box="[778,899,607,631]" italics="true" pageId="2">Malthinus</emphasis>
Latreille, 1806
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FD4C85B770B1" blockId="2.[699,1171,668,723]" box="[699,1171,668,692]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FD4C83EC70B1" bold="true" box="[699,712,668,692]" pageId="2"></emphasis>
<taxonomicName id="4C25578B0462FFDDFD62FD4C851570B1" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[713,1073,668,692]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="amicitiae" status="sp. nov." subGenus="Malthinus">
<emphasis id="B951F01A0462FFDDFD62FD4C829870B1" bold="true" box="[713,956,668,692]" pageId="2">
<emphasis id="B951F01A0462FFDDFD62FD4C826570B1" bold="true" box="[713,833,668,692]" italics="true" pageId="2">Malthinus</emphasis>
(
<emphasis id="B951F01A0462FFDDFCE1FD4C829870B1" bold="true" box="[842,956,668,692]" italics="true" pageId="2">Malthinus</emphasis>
</emphasis>
)amicitiae
</taxonomicName>
<emphasis id="B951F01A0462FFDDFB9CFD4C85B770B1" bold="true" box="[1079,1171,668,692]" pageId="2">
<taxonomicNameLabel id="A2624D610462FFDDFB9CFD4C85B770B1" box="[1079,1171,668,692]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FD6B822F70D6" blockId="2.[699,1171,668,723]" box="[699,779,699,723]" pageId="2">
(
<figureCitation id="131E308D0462FFDDFD68FD6B822770D6" box="[707,771,699,723]" captionStart="Fig" captionStartId="2.[699,738,1082,1106]" captionTargetBox="[699,1184,754,1045]" captionTargetId="figure-329@2.[699,1184,754,1045]" captionTargetPageId="2" captionText="Fig. 2.Malthinus(Malthinus)amicitiaesp. nov. in Baltic amber (code BaA09RU). Holotype, dorso-lateral view, scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203112" httpUri="https://zenodo.org/record/13203112/files/figure.png" pageId="2">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFD10FB038253741B" pageId="2" type="materials_examined">
<paragraph id="8B9A2C080462FFDDFD10FB0382367742" blockId="2.[699,1178,1235,1351]" pageId="2">
<materialsCitation id="3B4D26550462FFDDFD10FB0382367742" country="Russia" county="Baltic" location="Baltic" municipality="Fanti" pageId="2" specimenCount="1" typeStatus="holotype">
<emphasis id="B951F01A0462FFDDFD10FB03821476EE" bold="true" box="[699,816,1235,1259]" pageId="2">
<typeStatus id="549E92AA0462FFDDFD10FB03820F76EE" box="[699,811,1235,1259]" pageId="2" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EFA7AD30462FFDDFC92FB03825C76EE" LSID="urn:lsid:plazi:treatment:038C9D1E0462FFDCFD10FDD382BD7770:8EFA7AD30462FFDDFC92FB03825C76EE" box="[825,888,1235,1259]" country="Russia" county="Baltic" municipality="Fanti" name="Baltic" pageId="2">Baltic</location>
amber, collection
<collectingMunicipality id="6BFEB6720462FFDDFBFDFB0385B176EE" box="[1110,1173,1235,1259]" pageId="2">Fanti</collectingMunicipality>
, access code BaA09RU (BaA =
<collectingCounty id="62FB54840462FFDDFBFFFB2285BE770F" box="[1108,1178,1266,1290]" pageId="2">Baltic</collectingCounty>
amber; 09 = sequential number; RU =
<collectingCountry id="F3326C980462FFDDFD10FAFF82237742" box="[699,775,1327,1351]" name="Russia" pageId="2">Russia</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FABC827F77A6" blockId="2.[699,1178,1388,1443]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FABC82797781" bold="true" box="[699,861,1388,1412]" pageId="2">Type locality.</emphasis>
Yantarny mine,
<collectingRegion id="49E1E2EA0462FFDDFBBFFABC822677A6" country="Russia" name="Kaliningrad" pageId="2">Kaliningrad region</collectingRegion>
,
<collectingCountry id="F3326C980462FFDDFCA6FA5B827377A6" box="[781,855,1419,1443]" name="Russia" pageId="2">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FA188253741B" blockId="2.[699,1178,1480,1566]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FA18824777E5" bold="true" box="[699,867,1480,1504]" pageId="2">Type horizon.</emphasis>
Middle Eocene: Lutetian (47.841.2 Mya) to late Eocene: Priabonian (37.833.9 Mya).
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDCFD10F99381D173D8" lastPageId="3" pageId="2" type="diagnosis">
<paragraph id="8B9A2C080462FFDDFD10F99385BE747F" blockId="2.[699,1178,1603,1658]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10F993829C745E" bold="true" box="[699,952,1603,1627]" pageId="2">Differential diagnosis.</emphasis>
The new species is easily recognizable by its neck gradually
</paragraph>
<paragraph id="8B9A2C080462FFDCFBC7F97681D173D8" blockId="2.[1132,1172,1702,1726]" lastBlockId="3.[114,593,178,477]" lastPageId="3" pageId="2">
235 constricted posteriorly. It is similar only to the extinct
<taxonomicName id="4C25578B0463FFDCFF55FF01818E7302" authority="Kuska &amp; Kania, 2010" authorityName="Kuska &amp; Kania" authorityYear="2010" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B951F01A0463FFDCFF55FF01805272ED" box="[254,374,208,232]" italics="true" pageId="3">M. danieli</emphasis>
<bibRefCitation id="EFB451F90463FFDCFE23FF00818E7302" author="Kuska A. &amp; Kania I." pageId="3" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania, 2010</bibRefCitation>
</taxonomicName>
, from which it differs in the pronotum not narrower anteriorly, the elytra without impressed punctuation and the different colour (
<bibRefCitation id="EFB451F90463FFDCFF61FE9B80E27366" author="Kuska A. &amp; Kania I." box="[202,454,331,355]" pageId="3" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania 2010</bibRefCitation>
). The other fossil
<taxonomicName id="4C25578B0463FFDCFF19FEBA80567387" box="[178,370,362,386]" pageId="3">
<emphasis id="B951F01A0463FFDCFF19FEBA803A7387" box="[178,286,362,386]" italics="true" pageId="3">Malthinus</emphasis>
species
</taxonomicName>
show head strongly restricted behind eyes and dotted elytra (
<bibRefCitation id="EFB451F90463FFDCFFD2FE7780B973BA" author="Fanti F. &amp; Damgaard A. L." box="[121,413,423,447]" pageId="3" pagination="1 - 32" refId="ref2486" refString="Fanti F., Damgaard A. L. 2018. Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology 18 (1): 1 - 32." type="journal article" year="2018">Fanti &amp; Damgaard 2018</bibRefCitation>
;
<bibRefCitation id="EFB451F90463FFDCFE1AFE7781CD73D8" author="Pankowski M. G. &amp; Fanti F." pageId="3" pagination="548 - 556" refId="ref3008" refString="Pankowski M. G., Fanti F. 2022. Two new fossil species of soldier beetles (Coleoptera, Cantharidae, Malthi-ninae) from Baltic amber. Zootaxa 5165 (4): 548 - 556." type="journal article" year="2022">Pankowski &amp; Fanti 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830463FFDCFFD9FDD382CA71CD" pageId="3" type="description">
<paragraph id="8B9A2C080463FFDCFFD9FDD381F6705D" blockId="3.[114,593,515,1643]" pageId="3">
<emphasis id="B951F01A0463FFDCFFD9FDD38027701E" bold="true" box="[114,259,515,539]" pageId="3">Description.</emphasis>
Female, adult, robust, winged. Body length: about 6.0-
<quantity id="4CDD81ED0463FFDCFE2EFDF280C7703F" box="[389,483,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="3" unit="mm" value="6.2">6.2 mm</quantity>
. Entirely blackish.
</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9FD8F8000718E" blockId="3.[114,593,515,1643]" pageId="3">Head almost completely exposed, neck gradually constricted posteriorly, equipped with several long setae and impressed small punctation. Eyes large, convex, rounded, inserted in the lateral-upper part of head. Mandibles falciform. Maxillary palps 4- segmented; last palpomere globularelongated and apically pointed. Labial palps 3-segmented; last palpomere globular and apically pointed.</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9FC42800076FE" blockId="3.[114,593,515,1643]" pageId="3">Antennae 11-segmented, filiform, relatively short, surpassing the elytral half, covered with several setae; scape elongated, club-shaped; antennomere II short; antennomeres III-IV sub-equal, elongated, rather robust; antennomeres V-VII sub-equal, slightly longer than previous ones; antennomere VIII longer and thinner than previous ones; antennomeres IX-X sub-equal, shorter and very slightly sturdier than previous one; antennomere XI elongated, rounded at apex.</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9FAD18064746E" blockId="3.[114,593,515,1643]" pageId="3">Pronotum rectangular, slightly longer than wide, slightly narrower than head; apical margin very slightly rounded and not bordered; anterior corners rounded; sides straight and strongly bordered; basal margin straight and strongly bordered; posterior corners almost acute; surface irregular, gibbous on the basal part and concave near the apical margin, equipped with some long setae, without punctation. Scutellum triangular, wide at base, pointed apically, with rounded apex.</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9F97683F57387" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,178,968]" pageId="3">236 Elytra not particularly short, revealing the last abdominal segment uncovered, slightly wider than pronotum, parallel-sided, not dehiscent, rounded at apex; surface rather wrinkled, covered with several long setae and without punctation. Hind wings as long as elytra.</paragraph>
<paragraph id="8B9A2C080463FFDCFDDBFE5882857115" blockId="3.[624,1103,178,968]" pageId="3">Metasternum with slightly rounded posterior margin, covered with dispersed short setae and very shallow punctation. Sternites transverse, wide, pubescent. Last tergite and last sternite wide and rounded. Legs robust, pubescent; coxae elongated, stout, enlarged at base; trochanters elongated, rounded at apex; femora enlarged, curved in the middle; tibiae cylindrical and thin, equipped with an apical spur, protibiae shorter than profemora, meso- and metatibiae longer than meso- and metafemora.</paragraph>
<paragraph id="8B9A2C080463FFDCFDDBFCC782CA71CD" blockId="3.[624,1103,178,968]" pageId="3">Tarsi 5-segmented, pubescent; tarsomere I elongated; tarsomere II about 1.4 times shorter than tarsomere I; tarsomere III shorter than previous one; tarsomere IV bilobed; tarsomere V short, thin, curved; claws simple, robust, without tooth.</paragraph>
</subSubSection>
<subSubSection id="C33F7F830463FFDCFDDBFC3D827676B8" pageId="3" type="etymology">
<paragraph id="8B9A2C080463FFDCFDDBFC3D82897664" blockId="3.[624,1103,1005,1121]" pageId="3">
<emphasis id="B951F01A0463FFDCFDDBFC3D83D47600" bold="true" box="[624,752,1005,1029]" pageId="3">Etymology.</emphasis>
The specific epithet derives from the Latin noun “
<emphasis id="B951F01A0463FFDCFC1CFBDC852A7621" box="[951,1038,1036,1060]" italics="true" pageId="3">ămīcĭtĭa</emphasis>
” = friendship, in reference to the friendship that binds the two co-authors.
</paragraph>
<paragraph id="8B9A2C080463FFDCFDDBFB57827676B8" blockId="3.[624,1102,1159,1213]" pageId="3">
<emphasis id="B951F01A0463FFDCFDDBFB578236769A" bold="true" box="[624,786,1159,1183]" pageId="3">Syninclusions.</emphasis>
Botanical fragments, air bubbles, stellate hair.
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830463FFDCFDDBFB3382BD7770" pageId="3" type="discussion">
<paragraph id="8B9A2C080463FFDCFDDBFB3382BD7770" blockId="3.[624,1103,1251,1397]" pageId="3">
<emphasis id="B951F01A0463FFDCFDDBFB3383FB76FE" bold="true" box="[624,735,1251,1275]" pageId="3">Remarks.</emphasis>
The oblong yellow amber piece measures approximately 30x20x
<quantity id="4CDD81ED0463FFDCFC7BFAD18537771C" box="[976,1043,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" unit="mm" value="9.0">9 mm</quantity>
. The inclusion is complete and clearly visible on one side, while it has shows some oxidations on the other one.
</paragraph>
</subSubSection>
</treatment>
</document>