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<document id="7F10CED7D9CF93596FE2184F18E143E9" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629685057509" checkinUser="felipe" docAuthor="Scarbrough, A. G." docDate="2010" docId="038C87F8FF97FFE1FF7B6AE2E4B9FAD2" docLanguage="en" docName="zt02540p047.pdf" docOrigin="Zootaxa 2540" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Longibeccus Scarbrough 2010, new genus" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFB5FF80FF9CFFEAFFEC6A75E64FFF88" masterDocTitle="An overview of the Afrotropical Ommatiinae (Diptera: Asilidae) with a key to genera" masterLastPageNumber="47" masterPageNumber="1" pageNumber="12" updateTime="1699095850413" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F8D863A1EA7175760487E10332D1163E">An overview of the Afrotropical Ommatiinae (Diptera: Asilidae) with a key to genera</mods:title>
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<mods:namePart id="218737FF27AC445C6EF94ECFD005FF7C">Scarbrough, A. G.</mods:namePart>
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<treatment id="038C87F8FF97FFE1FF7B6AE2E4B9FAD2" ID-DOI="http://doi.org/10.5281/zenodo.5310833" ID-Zenodo-Dep="5310833" LSID="urn:lsid:plazi:treatment:038C87F8FF97FFE1FF7B6AE2E4B9FAD2" httpUri="http://treatment.plazi.org/id/038C87F8FF97FFE1FF7B6AE2E4B9FAD2" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="C33F6565FF97FFE1FF7B6AE2E7F0FF39" box="[151,447,151,177]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B9A36EEFF97FFE1FF7B6AE2E7F0FF39" blockId="11.[151,447,151,212]" box="[151,447,151,177]" pageId="11" pageNumber="12">
<heading id="D0D28182FF97FFE1FF7B6AE2E7F0FF39" bold="true" box="[151,447,151,177]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B951EAFCFF97FFE1FF7B6AE2E7F0FF39" bold="true" box="[151,447,151,177]" pageId="11" pageNumber="12">
<taxonomicName id="4C254D6DFF97FFE1FF7B6AE2E77FFF39" ID-CoL="89QLQ" ID-ENA="2173473" authority="Scarbrough, 2010" authorityName="Scarbrough" authorityYear="2010" box="[151,304,151,177]" class="Insecta" family="Asilidae" genus="Longibeccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B951EAFCFF97FFE1FF7B6AE2E77FFF39" bold="true" box="[151,304,151,177]" italics="true" pageId="11" pageNumber="12">Longibeccus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2625787FF97FFE1FED26AE2E7F0FF39" box="[318,447,151,177]" pageId="11" pageNumber="12" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FF97FFE1FF7B6ACFE7C0FF5C" box="[151,399,186,212]" pageId="11" pageNumber="12" type="description">
<paragraph id="8B9A36EEFF97FFE1FF7B6ACFE7C0FF5C" blockId="11.[151,447,151,212]" box="[151,399,186,212]" pageId="11" pageNumber="12">
<figureCitation id="131E2A6BFF97FFE1FF7B6ACFE6A7FF5C" box="[151,232,186,212]" captionStart="FIGURES 16" captionStartId="4.[150,270,1948,1972]" captionTargetBox="[396,1217,190,1930]" captionTargetId="figure-50@4.[384,1225,175,1944]" captionTargetPageId="4" captionText="FIGURES 16. Male antennae. 1. Afroestricus flavipes (Loew); 2. Cophinopoda pulchripes (Bigot); 3. Longibeccus fuscovittatus (Ricardo); 4. Michotamia coarctata (Macquart); 5. Ommatius neotenellus Bromley; 6. Thallosia congoicola (Oldroyd). Abbr.: flgm=flagellum; ped=pedicel; sc=scape." pageId="11" pageNumber="12">Figs. 2</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FF196ACFE749FF5C" box="[245,262,186,212]" captionStart="FIGURES 712" captionStartId="5.[151,276,1937,1961]" captionTargetBox="[225,1350,199,1918]" captionTargetId="figure-44@5.[224,1358,179,1918]" captionTargetPageId="5" captionText="FIGURES 712. Male heads, lateral. 7. Afroestricus flavipes (Loew); 8. Cophinopoda pulchripes (Bigot); 9. Longibeccus fuscovittatus (Ricardo); 10. Michotamia coarctata (Macquart); 11. Ommatius neotenellus Bromley; 12. Pygommatius litoreus (Scarbrough &amp; Marascia)." pageId="11" pageNumber="12">9</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FEFF6ACFE760FF5C" box="[275,303,186,212]" captionStart="FIGURES 3036" captionStartId="12.[150,271,1898,1922]" captionTargetBox="[154,1435,203,1857]" captionTargetId="figure-61@12.[150,1436,194,1877]" captionTargetPageId="12" captionText="FIGURES 3036. Longibeccus fuscovittatus (Ricardo), male wing and terminalia. 30. Right wing; 3133. Lateral, dorsal and ventral views; 34. Gonostylus; 3536. Aedeagus, dorsal and lateral views. Abbr.: r4, m3=cells; cerc=cercus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">30</figureCitation>
51,108
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FF97FFE1FF7B6B71E580FE94" box="[151,975,260,284]" pageId="11" pageNumber="12" type="type_taxon">
<paragraph id="8B9A36EEFF97FFE1FF7B6B71E580FE94" blockId="11.[151,975,260,284]" box="[151,975,260,284]" pageId="11" pageNumber="12">
<emphasis id="B951EAFCFF97FFE1FF7B6B71E76CFE94" bold="true" box="[151,291,260,284]" pageId="11" pageNumber="12">
<typeStatus id="549E884CFF97FFE1FF7B6B71E680FE94" box="[151,207,260,284]" pageId="11" pageNumber="12">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C254D6DFF97FFE1FEDE6B70E483FE94" authority="Ricardo, 1900" authorityName="Ricardo" authorityYear="1900" box="[306,716,261,284]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fuscovittatus">
<emphasis id="B951EAFCFF97FFE1FEDE6B70E462FE94" box="[306,557,261,284]" italics="true" pageId="11" pageNumber="12">Ommatius fuscovittatus</emphasis>
<bibRefCitation id="EFB44B1FFF97FFE1FDD86B70E483FE94" author="Ricardo, G." box="[564,716,261,284]" pageId="11" pageNumber="12" pagination="161 - 178" refId="ref24375" refString="Ricardo, G. (1900) Notes on Diptera from South Africa. Tabanidae and Asilidae. Annals and Magazine of Natural History (7) 6, 161 - 178. London." type="journal article" year="1900">Ricardo, 1900</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FF97FFE1FF7B6B3FE698FCC7" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8B9A36EEFF97FFE1FF7B6B3FE698FCC7" blockId="11.[151,1436,330,1370]" pageId="11" pageNumber="12">
<emphasis id="B951EAFCFF97FFE1FF7B6B3FE758FEEC" bold="true" box="[151,279,330,356]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Male.
<emphasis id="B951EAFCFF97FFE1FE876B3FE7E5FEEB" box="[363,426,330,355]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
: Face moderately produced, gradually sloped or angled forward from antenna to oral margin (
<figureCitation id="131E2A6BFF97FFE1FF176B05E70BFE02" box="[251,324,368,394]" captionStart="FIGURES 712" captionStartId="5.[151,276,1937,1961]" captionTargetBox="[225,1350,199,1918]" captionTargetId="figure-44@5.[224,1358,179,1918]" captionTargetPageId="5" captionText="FIGURES 712. Male heads, lateral. 7. Afroestricus flavipes (Loew); 8. Cophinopoda pulchripes (Bigot); 9. Longibeccus fuscovittatus (Ricardo); 10. Michotamia coarctata (Macquart); 11. Ommatius neotenellus Bromley; 12. Pygommatius litoreus (Scarbrough &amp; Marascia)." pageId="11" pageNumber="12">Fig. 9</figureCitation>
); bristles limited to lower 1/2 of face, only short setae dorsally. Flagellum (
<figureCitation id="131E2A6BFF97FFE1FB296B05E343FE02" box="[1221,1292,368,394]" captionStart="FIGURES 16" captionStartId="4.[150,270,1948,1972]" captionTargetBox="[396,1217,190,1930]" captionTargetId="figure-50@4.[384,1225,175,1944]" captionTargetPageId="4" captionText="FIGURES 16. Male antennae. 1. Afroestricus flavipes (Loew); 2. Cophinopoda pulchripes (Bigot); 3. Longibeccus fuscovittatus (Ricardo); 4. Michotamia coarctata (Macquart); 5. Ommatius neotenellus Bromley; 6. Thallosia congoicola (Oldroyd). Abbr.: flgm=flagellum; ped=pedicel; sc=scape." pageId="11" pageNumber="12">Fig. 3</figureCitation>
) wider than and as long as scape+pedicel combined; surface sparsely punctuate. Proboscis equal to vertical height of compound eye, slightly compressed medially with vertical height greater than diameter; apex flattened dorsoventrally, awl-shape (
<figureCitation id="131E2A6BFF97FFE1FE396B95E451FE72" box="[469,542,480,506]" captionStart="FIGURES 712" captionStartId="5.[151,276,1937,1961]" captionTargetBox="[225,1350,199,1918]" captionTargetId="figure-44@5.[224,1358,179,1918]" captionTargetPageId="5" captionText="FIGURES 712. Male heads, lateral. 7. Afroestricus flavipes (Loew); 8. Cophinopoda pulchripes (Bigot); 9. Longibeccus fuscovittatus (Ricardo); 10. Michotamia coarctata (Macquart); 11. Ommatius neotenellus Bromley; 12. Pygommatius litoreus (Scarbrough &amp; Marascia)." pageId="11" pageNumber="12">Fig. 9</figureCitation>
). Anterior scutal setae much shorter than scape.
<emphasis id="B951EAFCFF97FFE1FBB16B95E2D6FE71" box="[1117,1177,480,505]" italics="true" pageId="11" pageNumber="12">Wing</emphasis>
(
<figureCitation id="131E2A6BFF97FFE1FB466B95E342FE72" box="[1194,1293,480,506]" captionStart="FIGURES 3036" captionStartId="12.[150,271,1898,1922]" captionTargetBox="[154,1435,203,1857]" captionTargetId="figure-61@12.[150,1436,194,1877]" captionTargetPageId="12" captionText="FIGURES 3036. Longibeccus fuscovittatus (Ricardo), male wing and terminalia. 30. Right wing; 3133. Lateral, dorsal and ventral views; 34. Gonostylus; 3536. Aedeagus, dorsal and lateral views. Abbr.: r4, m3=cells; cerc=cercus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">Figs. 30</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FAF66B95E374FE72" box="[1306,1339,480,506]" captionStart="FIGURES 4147" captionStartId="18.[150,268,1749,1773]" captionTargetBox="[151,1431,198,1719]" captionTargetId="figure-196@18.[151,1436,197,1729]" captionTargetPageId="18" captionText="FIGURES 4147. Longibeccus imperator (Oldroyd), male, wing and terminalia. 41. Left wing. Figures 4244. Lateral, dorsal, and ventral views; 45. Gonostylus; 4647. Aedeagus, dorsal and lateral views. Abbr.: cerc=cercus; distph=disti phallus; epand=epandrium; ep iph=epiphallus; goncx=gon ocoxit e; hypd=hyp andrium; pvapod=posteroventral apodeme." pageId="11" pageNumber="12">41</figureCitation>
): Cell r
<subScript id="17A134ABFF97FFE1FA786B85E3D3FE76" attach="left" box="[1428,1436,496,510]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
narrowly pointed; base of cell r
<subScript id="17A134ABFF97FFE1FDE1686DE45AFDAE" attach="left" box="[525,533,536,550]" fontSize="6" pageId="11" pageNumber="12">4</subScript>
at or just before apex of cell d; R
<subScript id="17A134ABFF97FFE1FC45686DE5FEFDAE" attach="left" box="[937,945,536,550]" fontSize="6" pageId="11" pageNumber="12">4</subScript>
slightly concave apically, joining C at or just below wing apex.
<emphasis id="B951EAFCFF97FFE1FE406844E796FDC2" box="[428,473,561,586]" italics="true" pageId="11" pageNumber="12">Leg</emphasis>
: Metathoracic femur with short, brown, spur-like, anteroventral bristles, about twice as long as ventral setae.
<emphasis id="B951EAFCFF97FFE1FDE36823E4DCFDE7" box="[527,659,598,623]" italics="true" pageId="11" pageNumber="12">Terminalia</emphasis>
(
<figureCitation id="131E2A6BFF97FFE1FD496820E575FDE7" box="[677,826,597,623]" captionStart="FIGURES 3036" captionStartId="12.[150,271,1898,1922]" captionTargetBox="[154,1435,203,1857]" captionTargetId="figure-61@12.[150,1436,194,1877]" captionTargetPageId="12" captionText="FIGURES 3036. Longibeccus fuscovittatus (Ricardo), male wing and terminalia. 30. Right wing; 3133. Lateral, dorsal and ventral views; 34. Gonostylus; 3536. Aedeagus, dorsal and lateral views. Abbr.: r4, m3=cells; cerc=cercus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">Figs. 3136</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FCA46820E5D5FDE7" box="[840,922,597,623]" captionStart="FIGURES 4147" captionStartId="18.[150,268,1749,1773]" captionTargetBox="[151,1431,198,1719]" captionTargetId="figure-196@18.[151,1436,197,1729]" captionTargetPageId="18" captionText="FIGURES 4147. Longibeccus imperator (Oldroyd), male, wing and terminalia. 41. Left wing. Figures 4244. Lateral, dorsal, and ventral views; 45. Gonostylus; 4647. Aedeagus, dorsal and lateral views. Abbr.: cerc=cercus; distph=disti phallus; epand=epandrium; ep iph=epiphallus; goncx=gon ocoxit e; hypd=hyp andrium; pvapod=posteroventral apodeme." pageId="11" pageNumber="12">4247</figureCitation>
): Epandrium deeply excised apically with contrasting shorter dorsal and longer ventral arms, former with several minute denticles along ventral margin. Aedeagal sheath inflated and produced dorsally, forming an epiphallus which extends above and beyond distiphallus in dorsal view. Female (
<figureCitation id="131E2A6BFF97FFE1FDAC68B0E481FD57" box="[576,718,709,735]" captionStart="FIGURES 3740" captionStartId="13.[151,274,1477,1501]" captionTargetBox="[162,1433,207,1417]" captionTargetId="figure-234@13.[151,1436,194,1456]" captionTargetPageId="13" captionText="FIGURES 3740. Longibeccus fuscovittatus (Ricardo), female terminalia. 3738. Dorsal and ventral views; 39. Spermathecae; 40. Genital fork. Abbr.: cerc=cercus; tg8, tg9=tergite 8, tergite 9; st8=sternite 8." pageId="11" pageNumber="12">Figs. 3740</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FD3668B0E567FD57" box="[730,808,709,735]" captionStart="FIGURES 4851" captionStartId="19.[151,276,1926,1950]" captionTargetBox="[159,1428,693,1895]" captionTargetId="figure-293@19.[151,1436,665,1905]" captionTargetPageId="19" captionText="FIGURES 4851. Longibeccus imperator (Oldroyd), female terminalia. 4849. Dorsal and ventral views; 50. Spermathecae; 51. Genital fork. Abbr.: cerc=cercus; tg8= tergite 8, tg9=tergite 9; st7=sternite 7, st8=sternite 8." pageId="11" pageNumber="12">4851</figureCitation>
). Sparse, unusually stout, bristles on apical margin of abdominal segments 7. Cerci triangular, broad basally, fused medially, without membranous line between cerci; base of cercus extends ventrally, fusing with ventral lamella. Spermathecae oval, slightly longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FF97FFE1FF2A692FE400FCFD" box="[198,591,858,885]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8B9A36EEFF97FFE1FF2A692FE400FCFD" blockId="11.[151,1436,330,1370]" box="[198,591,858,885]" pageId="11" pageNumber="12">
<emphasis id="B951EAFCFF97FFE1FF2A692FE710FCFC" bold="true" box="[198,351,858,884]" pageId="11" pageNumber="12">Distribution</emphasis>
. Subsaharan Africa.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FF97FFE1FF2A69F5E340FC12" box="[198,1295,896,922]" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8B9A36EEFF97FFE1FF2A69F5E340FC12" blockId="11.[151,1436,330,1370]" box="[198,1295,896,922]" pageId="11" pageNumber="12">
<emphasis id="B951EAFCFF97FFE1FF2A69F5E704FC12" bold="true" box="[198,331,896,922]" pageId="11" pageNumber="12">Etymology</emphasis>
. Latin combination
<taxonomicName id="4C254D6DFF97FFE1FDD569F4E482FC12" authorityName="Scarbrough" authorityYear="2010" box="[569,717,897,922]" class="Insecta" family="Asilidae" genus="Longibeccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FDD569F4E482FC12" box="[569,717,897,922]" italics="true" pageId="11" pageNumber="12">Longibeccus</emphasis>
</taxonomicName>
. Refers to the long proboscis. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FF97FFE1FF2A69D0E4B9FAD2" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8B9A36EEFF97FFE1FF2A69D0E593FA98" blockId="11.[151,1436,330,1370]" pageId="11" pageNumber="12">
<emphasis id="B951EAFCFF97FFE1FF2A69D0E777FC37" bold="true" box="[198,312,933,959]" pageId="11" pageNumber="12">Remarks</emphasis>
.
<taxonomicName id="4C254D6DFF97FFE1FEA069D3E7AFFC37" authorityName="Scarbrough" authorityYear="2010" box="[332,480,934,959]" class="Insecta" family="Asilidae" genus="Longibeccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FEA069D3E7AFFC37" box="[332,480,934,959]" italics="true" pageId="11" pageNumber="12">Longibeccus</emphasis>
</taxonomicName>
is proposed to accommodate
<taxonomicName id="4C254D6DFF97FFE1FCB669D3E23BFC37" authorityName="Ricardo" authorityYear="1900" box="[858,1140,934,959]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fuscovittatus">
<emphasis id="B951EAFCFF97FFE1FCB669D3E23BFC37" box="[858,1140,934,959]" italics="true" pageId="11" pageNumber="12">Ommatius fuscovittatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D6DFF97FFE1FB5469D3E31AFC37" authorityName="Oldroyd" authorityYear="1936" box="[1208,1365,934,959]" class="Insecta" family="Asilidae" genus="Longibeccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imperator">
<emphasis id="B951EAFCFF97FFE1FB5469D3E31AFC37" box="[1208,1365,934,959]" italics="true" pageId="11" pageNumber="12">O. imperator</emphasis>
</taxonomicName>
. It is distinguished by the characters in the key and diagnosis, especially the gently sloped face, unusual shape of the flagellum, long proboscis, wing venation (
<figureCitation id="131E2A6BFF97FFE1FD426985E4B0FB82" box="[686,767,1008,1034]" captionStart="FIGURES 16" captionStartId="4.[150,270,1948,1972]" captionTargetBox="[396,1217,190,1930]" captionTargetId="figure-50@4.[384,1225,175,1944]" captionTargetPageId="4" captionText="FIGURES 16. Male antennae. 1. Afroestricus flavipes (Loew); 2. Cophinopoda pulchripes (Bigot); 3. Longibeccus fuscovittatus (Ricardo); 4. Michotamia coarctata (Macquart); 5. Ommatius neotenellus Bromley; 6. Thallosia congoicola (Oldroyd). Abbr.: flgm=flagellum; ped=pedicel; sc=scape." pageId="11" pageNumber="12">Figs. 3</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FCE06985E552FB82" box="[780,797,1008,1034]" captionStart="FIGURES 712" captionStartId="5.[151,276,1937,1961]" captionTargetBox="[225,1350,199,1918]" captionTargetId="figure-44@5.[224,1358,179,1918]" captionTargetPageId="5" captionText="FIGURES 712. Male heads, lateral. 7. Afroestricus flavipes (Loew); 8. Cophinopoda pulchripes (Bigot); 9. Longibeccus fuscovittatus (Ricardo); 10. Michotamia coarctata (Macquart); 11. Ommatius neotenellus Bromley; 12. Pygommatius litoreus (Scarbrough &amp; Marascia)." pageId="11" pageNumber="12">9</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FCC56985E506FB82" box="[809,841,1008,1034]" captionStart="FIGURES 3036" captionStartId="12.[150,271,1898,1922]" captionTargetBox="[154,1435,203,1857]" captionTargetId="figure-61@12.[150,1436,194,1877]" captionTargetPageId="12" captionText="FIGURES 3036. Longibeccus fuscovittatus (Ricardo), male wing and terminalia. 30. Right wing; 3133. Lateral, dorsal and ventral views; 34. Gonostylus; 3536. Aedeagus, dorsal and lateral views. Abbr.: r4, m3=cells; cerc=cercus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">30</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FCB86985E53BFB82" box="[852,884,1008,1034]" captionStart="FIGURES 4147" captionStartId="18.[150,268,1749,1773]" captionTargetBox="[151,1431,198,1719]" captionTargetId="figure-196@18.[151,1436,197,1729]" captionTargetPageId="18" captionText="FIGURES 4147. Longibeccus imperator (Oldroyd), male, wing and terminalia. 41. Left wing. Figures 4244. Lateral, dorsal, and ventral views; 45. Gonostylus; 4647. Aedeagus, dorsal and lateral views. Abbr.: cerc=cercus; distph=disti phallus; epand=epandrium; ep iph=epiphallus; goncx=gon ocoxit e; hypd=hyp andrium; pvapod=posteroventral apodeme." pageId="11" pageNumber="12">41</figureCitation>
), and the combined characters of the terminalia (
<figureCitation id="131E2A6BFF97FFE1FF736E60E760FBA7" box="[159,303,1045,1071]" captionStart-0="FIGURES 3036" captionStart-1="FIGURES 3740" captionStartId-0="12.[150,271,1898,1922]" captionStartId-1="13.[151,274,1477,1501]" captionTargetBox-0="[154,1435,203,1857]" captionTargetBox-1="[162,1433,207,1417]" captionTargetId-0="figure-61@12.[150,1436,194,1877]" captionTargetId-1="figure-234@13.[151,1436,194,1456]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 3036. Longibeccus fuscovittatus (Ricardo), male wing and terminalia. 30. Right wing; 3133. Lateral, dorsal and ventral views; 34. Gonostylus; 3536. Aedeagus, dorsal and lateral views. Abbr.: r4, m3=cells; cerc=cercus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." captionText-1="FIGURES 3740. Longibeccus fuscovittatus (Ricardo), female terminalia. 3738. Dorsal and ventral views; 39. Spermathecae; 40. Genital fork. Abbr.: cerc=cercus; tg8, tg9=tergite 8, tergite 9; st8=sternite 8." pageId="11" pageNumber="12">Figs. 3140</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FED06E60E7C5FBA7" box="[316,394,1045,1071]" captionStart-0="FIGURES 4147" captionStart-1="FIGURES 4851" captionStartId-0="18.[150,268,1749,1773]" captionStartId-1="19.[151,276,1926,1950]" captionTargetBox-0="[151,1431,198,1719]" captionTargetBox-1="[159,1428,693,1895]" captionTargetId-0="figure-196@18.[151,1436,197,1729]" captionTargetId-1="figure-293@19.[151,1436,665,1905]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 4147. Longibeccus imperator (Oldroyd), male, wing and terminalia. 41. Left wing. Figures 4244. Lateral, dorsal, and ventral views; 45. Gonostylus; 4647. Aedeagus, dorsal and lateral views. Abbr.: cerc=cercus; distph=disti phallus; epand=epandrium; ep iph=epiphallus; goncx=gon ocoxit e; hypd=hyp andrium; pvapod=posteroventral apodeme." captionText-1="FIGURES 4851. Longibeccus imperator (Oldroyd), female terminalia. 4849. Dorsal and ventral views; 50. Spermathecae; 51. Genital fork. Abbr.: cerc=cercus; tg8= tergite 8, tg9=tergite 9; st7=sternite 7, st8=sternite 8." pageId="11" pageNumber="12">4251</figureCitation>
). The large plate-like epiphallus concealing the distiphallus dorsally is diagnostic of the genus (35, 36, 46, 47). The epiphallus of
<taxonomicName id="4C254D6DFF97FFE1FD686E4EE55CFBDC" authorityName="Scarbrough" authorityYear="2005" box="[644,787,1083,1108]" class="Insecta" family="Asilidae" genus="Afroestricus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FD686E4EE55CFBDC" box="[644,787,1083,1108]" italics="true" pageId="11" pageNumber="12">Afroestricus</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A6BFF97FFE1FCC86E4EE534FBDD" box="[804,891,1083,1109]" captionStart="FIGURES 1318" captionStartId="7.[151,270,1918,1942]" captionTargetBox="[163,1423,297,1889]" captionTargetId="figure-89@7.[151,1436,269,1897]" captionTargetPageId="7" captionText="FIGURES 1318. Afroestricus flavipes (Loew), male terminalia (reillustrated after Scarbrough 2005). 1315. Lateral, dorsal, and ventral views; 16. Aedeagus, lateral view; 17ac. Spermathecae, A. flavipes; 18. Type labels, A. flavipes. Abbr.: cerc=cercus; distph=distiphallus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">Fig. 16</figureCitation>
),
<taxonomicName id="4C254D6DFF97FFE1FC7F6E4EE27CFBDC" authorityName="Hull" authorityYear="1958" box="[915,1075,1083,1108]" class="Insecta" family="Asilidae" genus="Cophinopoda" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FC7F6E4EE27CFBDC" box="[915,1075,1083,1108]" italics="true" pageId="11" pageNumber="12">Cophinopoda</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A6BFF97FFE1FBA86E4EE2D3FBDD" box="[1092,1180,1083,1109]" captionStart="FIGURES 1923" captionStartId="9.[151,269,1653,1677]" captionTargetBox="[164,1424,205,1624]" captionTargetId="figure-139@9.[161,1426,194,1632]" captionTargetPageId="9" captionText="FIGURES 1923. Cophinopoda pulchripes Bigot, male terminalia. 19. Epandrium; 20. Gonocoxite and hypandrium; 21. Aedeagus; 22. Female, sternite 8; 23. Spermathecae. Abbr.: distph=distiphallus; epiph=epiphallus; goncx=gonocoxite; pvapod=posteroventral aedeagal apodeme; spmth caps= spermathecal capsule." pageId="11" pageNumber="12">Fig. 21</figureCitation>
),
<taxonomicName id="4C254D6DFF97FFE1FB5F6E4EE314FBDC" authorityName="Schiner" authorityYear="1866" box="[1203,1371,1083,1108]" class="Insecta" family="Asilidae" genus="Emphysomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FB5F6E4EE314FBDC" box="[1203,1371,1083,1108]" italics="true" pageId="11" pageNumber="12">Emphysomera</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A6BFF97FFE1FA806E4EE6F9FBF2" captionStart="FIGURES 2429" captionStartId="10.[151,271,1811,1835]" captionTargetBox="[164,1423,389,1783]" captionTargetId="figure-129@10.[151,1436,381,1790]" captionTargetPageId="10" captionText="FIGURES 2429. Emphysomera atrogaster Bigot (reillustrated after Scarbrough &amp; Marascia 1996). 24. Right wing. 2526. Lateral and dorsal views, terminalia; 27. Aedeagus, lateral view; 28.Male abdomen; 29. Spermathecae. Abbr.; epand=epandrium, epiph=epiphallus, distph=distiphallus, hypd=hypandrium, goncxp=gonocoxital process, r =4th radial 4 cell; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">Fig. 27</figureCitation>
) and
<taxonomicName id="4C254D6DFF97FFE1FF166E14E725FBF2" authorityName="Oldroyd" authorityYear="1970" box="[250,362,1121,1146]" class="Insecta" family="Asilidae" genus="Thallosia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FF166E14E725FBF2" box="[250,362,1121,1146]" italics="true" pageId="11" pageNumber="12">Thallosia</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A6BFF97FFE1FE976E15E7ADFBF2" box="[379,482,1120,1146]" captionStart="FIGURES 102105" captionStartId="39.[151,270,1878,1902]" captionTargetBox="[155,1430,195,1827]" captionTargetId="figure-52@39.[150,1436,194,1857]" captionTargetPageId="39" captionText="FIGURES 102105. Thallosia congiola Oldroyd (reillustrated from Tsacas &amp; Artigas 1994). 102. Epandrium, lateral; 103. Gonocoxite and hypandrium, lateral; 104. Aedeagus, lateral; 105. Female terminalia, ventral view. Abbr.: distph=distiphallus; epiph=epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme; tg8=tergite 8." pageId="11" pageNumber="12">Fig. 104</figureCitation>
) is also dorsal prominent but otherwise structurally different. The cercus which extends onto the ventral surface of the ventral lamella and the oval spermathecal bulbs are distinctive of females (
<figureCitation id="131E2A6BFF97FFE1FEEA6EDEE724FB4D" box="[262,363,1195,1221]" captionStart="FIGURES 3740" captionStartId="13.[151,274,1477,1501]" captionTargetBox="[162,1433,207,1417]" captionTargetId="figure-234@13.[151,1436,194,1456]" captionTargetPageId="13" captionText="FIGURES 3740. Longibeccus fuscovittatus (Ricardo), female terminalia. 3738. Dorsal and ventral views; 39. Spermathecae; 40. Genital fork. Abbr.: cerc=cercus; tg8, tg9=tergite 8, tergite 9; st8=sternite 8." pageId="11" pageNumber="12">Figs. 39</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FE956EDEE7D4FB4D" box="[377,411,1195,1221]" captionStart="FIGURES 4851" captionStartId="19.[151,276,1926,1950]" captionTargetBox="[159,1428,693,1895]" captionTargetId="figure-293@19.[151,1436,665,1905]" captionTargetPageId="19" captionText="FIGURES 4851. Longibeccus imperator (Oldroyd), female terminalia. 4849. Dorsal and ventral views; 50. Spermathecae; 51. Genital fork. Abbr.: cerc=cercus; tg8= tergite 8, tg9=tergite 9; st7=sternite 7, st8=sternite 8." pageId="11" pageNumber="12">50</figureCitation>
). In other genera, the cercus covers only the dorsal surface of the pygidium, and the spermathecal bulbs are usually 3 or more times longer than wide and recurved (
<figureCitation id="131E2A6BFF97FFE1FBBE6EA5E2F9FB62" box="[1106,1206,1232,1258]" captionStart="FIGURES 1318" captionStartId="7.[151,270,1918,1942]" captionTargetBox="[163,1423,297,1889]" captionTargetId="figure-89@7.[151,1436,269,1897]" captionTargetPageId="7" captionText="FIGURES 1318. Afroestricus flavipes (Loew), male terminalia (reillustrated after Scarbrough 2005). 1315. Lateral, dorsal, and ventral views; 16. Aedeagus, lateral view; 17ac. Spermathecae, A. flavipes; 18. Type labels, A. flavipes. Abbr.: cerc=cercus; distph=distiphallus; epand=epandrium; epiph= epiphallus; goncx=gonocoxite; hypd=hypandrium; pvapod=posteroventral aedeagal apodeme." pageId="11" pageNumber="12">Figs. 17</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FB286EA5E2ABFB62" box="[1220,1252,1232,1258]" captionStart="FIGURES 1923" captionStartId="9.[151,269,1653,1677]" captionTargetBox="[164,1424,205,1624]" captionTargetId="figure-139@9.[161,1426,194,1632]" captionTargetPageId="9" captionText="FIGURES 1923. Cophinopoda pulchripes Bigot, male terminalia. 19. Epandrium; 20. Gonocoxite and hypandrium; 21. Aedeagus; 22. Female, sternite 8; 23. Spermathecae. Abbr.: distph=distiphallus; epiph=epiphallus; goncx=gonocoxite; pvapod=posteroventral aedeagal apodeme; spmth caps= spermathecal capsule." pageId="11" pageNumber="12">23</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FB1D6EA5E35EFB62" box="[1265,1297,1232,1258]" captionStart="FIGURES 6466" captionStartId="28.[150,269,1612,1636]" captionTargetBox="[190,1406,180,1598]" captionTargetId="figure-168@28.[183,1412,170,1608]" captionTargetPageId="28" captionText="FIGURES 6466. Ommatius coperitus n. sp., female terminalia. 64. Ventral view; 65. Genital fork; 66. Spermathecae, posterior view. Abbr.: cerc=cercus; st8=sternite 8; vl=ventral lamella; spmth=spermatheca; spmth duct=spermatheca with coiled duct; spmth vlv=spermathecal valve." pageId="11" pageNumber="12">66</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FAF36EA5E370FB62" box="[1311,1343,1232,1258]" captionStart="FIGURES 7982" captionStartId="31.[151,277,1974,1998]" captionTargetBox="[204,1392,446,1943]" captionTargetId="figure-132@31.[191,1402,433,1972]" captionTargetPageId="31" captionText="FIGURES 7982. Ommatius neotenellus Bromley, female terminalia. 7980. Dorsal and ventral views; 81. Spermathecae; 82. Genital fork." pageId="11" pageNumber="12">81</figureCitation>
,
<figureCitation id="131E2A6BFF97FFE1FAA06EA5E321FB62" box="[1356,1390,1232,1258]" captionStart="FIGURES 9598" captionStartId="36.[151,269,1976,2000]" captionTargetBox="[210,1392,448,1946]" captionTargetId="figure-142@36.[192,1407,430,1990]" captionTargetPageId="36" captionText="FIGURES 9598. Ommatius ultimus n. sp., female terminalia. 9596. Dorsal and ventral views; 97. Spermathecae; 98. Genital fork." pageId="11" pageNumber="12">97</figureCitation>
). In
<taxonomicName id="4C254D6DFF97FFE1FF7B6E83E76AFA87" authorityName="Macquart" authorityYear="1839" box="[151,293,1270,1295]" class="Insecta" family="Asilidae" genus="Michotamia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FF7B6E83E76AFA87" box="[151,293,1270,1295]" italics="true" pageId="11" pageNumber="12">Michotamia</emphasis>
</taxonomicName>
, the spermathecal duct is looped or tightly coiled (
<figureCitation id="131E2A6BFF97FFE1FC9B6E83E581FA98" box="[887,974,1270,1296]" captionStart="FIGURES 5257" captionStartId="22.[151,274,1863,1887]" captionTargetBox="[190,1395,266,1833]" captionTargetId="figure-94@22.[186,1401,239,1871]" captionTargetPageId="22" captionText="FIGURES 5257. Michotamia coarctata Macquart, female. 52. Right wing; 53. Sternite 8 with genital apodeme (sternite 10?); 54. Genital furca; 55. Spermatheca with coiled duct; 56. Michotamia aurata (Fabricius 1794) (Oriental species; Scarrough &amp; Hill 2000), male terminalia, lateral view; 57. Aedeagus, lateral view. Abbr.: genapod=genital apodeme; flgm=flagellum; spmth=spermatheca; spmth duct=spermatheca with coiled duct; spmth vlv=spermathecal valve; st8=sternite 8; tg8=tergite 8." pageId="11" pageNumber="12">Fig. 55</figureCitation>
).
</paragraph>
<paragraph id="8B9A36EEFF97FFE1FF296F6EE4B9FAD2" blockId="11.[151,1436,330,1370]" pageId="11" pageNumber="12">
<taxonomicName id="4C254D6DFF97FFE1FF296F6EE715FABC" authorityName="Scarbrough" authorityYear="2010" box="[197,346,1307,1332]" class="Insecta" family="Asilidae" genus="Longibeccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAFCFF97FFE1FF296F6EE715FABC" box="[197,346,1307,1332]" italics="true" pageId="11" pageNumber="12">Longibeccus</emphasis>
</taxonomicName>
is confined to the African Continent with
<taxonomicName id="4C254D6DFF97FFE1FCBF6F6EE247FABC" authorityName="Ricardo" authorityYear="1900" box="[851,1032,1307,1332]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fuscovittatus">
<emphasis id="B951EAFCFF97FFE1FCBF6F6EE247FABC" box="[851,1032,1307,1332]" italics="true" pageId="11" pageNumber="12">L. fuscovittatus</emphasis>
</taxonomicName>
widespread whereas
<taxonomicName id="4C254D6DFF97FFE1FAEA6F6EE3D3FABC" authorityName="Oldroyd" authorityYear="1936" box="[1286,1436,1307,1332]" class="Insecta" family="Asilidae" genus="Longibeccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imperator">
<emphasis id="B951EAFCFF97FFE1FAEA6F6EE3D3FABC" box="[1286,1436,1307,1332]" italics="true" pageId="11" pageNumber="12">L. imperator</emphasis>
</taxonomicName>
are only recorded from equatorial Africa (
<figureCitation id="131E2A6BFF97FFE1FD6E6F35E4A6FAD2" box="[642,745,1344,1370]" captionStart="FIGURE 108" captionStartId="42.[151,255,1771,1795]" captionTargetBox="[157,1428,461,1746]" captionTargetId="figure-217@42.[150,1436,455,1750]" captionTargetPageId="42" captionText="FIGURE 108. Distribution of Longibeccus, new genus and Metommatus, new status in the Afrotropical Region." pageId="11" pageNumber="12">Fig. 108</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>