156 lines
18 KiB
XML
156 lines
18 KiB
XML
<document id="25D83A8DE04B6FD2F14481A55344E616" ID-CLB-Dataset="25163" ID-DOI="10.11646/zootaxa.4688.3.3" ID-GBIF-Dataset="a9af11dc-d705-48ed-96fb-3b323accc19e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3516791" ID-ZooBank="B8F6F528-F616-4EA1-8CA7-FA2FA89A34B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571832732088" checkinUser="plazi" docAuthor="Thandar, Ahmed S." docDate="2019" docId="038C87E5BF6EFF92FF50FE507DB66720" docLanguage="en" docName="zootaxa.4688.3.3.pdf" docOrigin="Zootaxa 4688 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyone Oken 1815" docType="treatment" docVersion="7" lastPageNumber="370" masterDocId="FFB5FF9DBF67FF9BFFC7FF8C7E106073" masterDocTitle="On some lesser known sea cucumbers in the Natural History Museum, UK (Echinodermata: Holothuroidea)" masterLastPageNumber="381" masterPageNumber="361" pageNumber="370" updateTime="1698760707621" updateUser="ExternalLinkService">
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<mods:title id="DD5543696F64861FCDCFCD62F32A94FD">On some lesser known sea cucumbers in the Natural History Museum, UK (Echinodermata: Holothuroidea)</mods:title>
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<mods:namePart id="E6B5E32F9AFCDF891491DAA1426DA861">Thandar, Ahmed S.</mods:namePart>
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<mods:typeOfResource id="F189C97365DE224F0560AF4F0BA9DDC7">text</mods:typeOfResource>
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<mods:title id="5605A159E40899AEE9453885289107EF">Zootaxa</mods:title>
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<mods:part id="89BC65695BCCEF9693F3895015D61F29">
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<mods:date id="9F1835715B8CBADD154D61CB0D90F298">2019</mods:date>
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<mods:number id="165C76369C9EC4793061411369CF9F7F">2019-10-23</mods:number>
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<mods:number id="37035C8B2C8E8710448D1E64BB9A719A">4688</mods:number>
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<treatment id="038C87E5BF6EFF92FF50FE507DB66720" ID-GBIF-Taxon="159812118" LSID="urn:lsid:plazi:treatment:038C87E5BF6EFF92FF50FE507DB66720" httpUri="http://treatment.plazi.org/id/038C87E5BF6EFF92FF50FE507DB66720" lastPageNumber="370" pageId="9" pageNumber="370">
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<subSubSection id="C33F6578BF6EFF92FF50FE507A236185" box="[151,1075,476,503]" pageId="9" pageNumber="370" type="nomenclature">
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<paragraph id="8B9A36F3BF6EFF92FF50FE507A236185" blockId="9.[151,1075,476,539]" box="[151,1075,476,503]" pageId="9" pageNumber="370">
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<heading id="D0D2819FBF6EFF92FF50FE507A236185" bold="true" box="[151,1075,476,503]" fontSize="11" level="1" pageId="9" pageNumber="370" reason="1">
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<emphasis id="B951EAE1BF6EFF92FF50FE507A236185" bold="true" box="[151,1075,476,503]" pageId="9" pageNumber="370">
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<taxonomicName id="4C254D70BF6EFF92FF50FE507EE26184" ID-CoL="7Q4FF" authorityName="Oken" authorityYear="1815" box="[151,242,476,503]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="genus">
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<emphasis id="B951EAE1BF6EFF92FF50FE507EE26184" bold="true" box="[151,242,476,503]" italics="true" pageId="9" pageNumber="370">Thyone</emphasis>
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</taxonomicName>
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?
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<taxonomicName id="4C254D70BF6EFF92FECFFE507C0D6185" authority="Norman, 1869" authorityName="Norman" authorityYear="1869" box="[264,541,476,503]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="elegans">
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<emphasis id="B951EAE1BF6EFF92FECFFE507F736184" bold="true" box="[264,355,476,503]" italics="true" pageId="9" pageNumber="370">elegans</emphasis>
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<bibRefCitation id="EFB44B02BF6EFF92FEACFE517C0D6185" author="Norman, A. M." box="[363,541,476,503]" pageId="9" pageNumber="370" pagination="47 - 336" refId="ref12062" refString="Norman, A. M. (1869) Shetland final dredging report ii. On the Crustacea, Tunicata, Polyzoa, Echinodermata, Actinozoa, Hydrozoa and Porifera. Report of the British Association of the Advancement of Science, 1869, 47 - 336." type="journal article" year="1869">Norman, 1869</bibRefCitation>
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</taxonomicName>
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(? synonym of
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<taxonomicName id="4C254D70BF6EFF92FD1CFE517A236185" authority="Herouard, 1889" authorityName="Herouard" authorityYear="1889" box="[731,1075,476,503]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="roscovita">
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<emphasis id="B951EAE1BF6EFF92FD1CFE517D766184" bold="true" box="[731,870,477,503]" italics="true" pageId="9" pageNumber="370">T. roscovita</emphasis>
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<bibRefCitation id="EFB44B02BF6EFF92FCA9FE507A236185" author="Herouard, R." box="[878,1075,476,503]" pageId="9" pageNumber="370" pagination="535 - 704" refId="ref11494" refString="Herouard, R. (1889) Recherches sur les holothuries des cotes de France. Archives de Zoologie Experimentale et Generale, 2, 535 - 704." type="journal article" year="1889">Hérouard, 1889</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6578BF6EFF92FF50FD8C7EE56268" box="[151,245,512,539]" pageId="9" pageNumber="370" type="description">
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<paragraph id="8B9A36F3BF6EFF92FF50FD8C7EE56268" blockId="9.[151,1075,476,539]" box="[151,245,512,539]" pageId="9" pageNumber="370">
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<figureCitation id="131E2A76BF6EFF92FF50FD8C7EE56268" box="[151,245,512,539]" captionStart="FIGURE 5" captionStartId="9.[151,250,1453,1478]" captionTargetBox="[212,1335,653,1434]" captionTargetId="figure@9.[212,1335,653,1434]" captionTargetPageId="9" captionText="FIGURE 5. Thyone?elegans Norman, 1869 /?T. roscovita Hérouard, 1889." figureDoi="http://doi.org/10.5281/zenodo.3516801" httpUri="https://zenodo.org/record/3516801/files/figure.png" pageId="9" pageNumber="370">Figure 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6578BF6EFF92FF50FDC47F8362F4" pageId="9" pageNumber="370" type="materials_examined">
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<paragraph id="8B9A36F3BF6EFF92FF50FDC47F8362F4" blockId="9.[151,1436,584,647]" pageId="9" pageNumber="370">
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<emphasis id="B951EAE1BF6EFF92FF50FDC47F906210" bold="true" box="[151,384,584,611]" pageId="9" pageNumber="370">Material examined.</emphasis>
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<materialsCitation id="3B4D3CAEBF6EFF92FE40FDC57F8362F4" ID-GBIF-Occurrence="2429072687" collectionCode="NHMUK" collectorName="Siteb, E. & Seagull Bank & Rhum & Dr B. Earll & Marine Conservation Society" elevation="285" pageId="9" pageNumber="370" specimenCode="NHMUK 1985.10" specimenCount="1">
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<specimenCode id="DB839E88BF6EFF92FE40FDC57C466210" box="[391,598,584,611]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="370" type="Museum">NHMUK 1985.10</specimenCode>
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.4.14,
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<collectorName id="26D05325BF6EFF92FD5FFDC47CE76210" box="[664,759,584,611]" pageId="9" pageNumber="370">Siteb, E.</collectorName>
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of
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<collectorName id="26D05325BF6EFF92FCDCFDC47DA46211" box="[795,948,584,610]" pageId="9" pageNumber="370">Seagull Bank</collectorName>
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,
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<collectorName id="26D05325BF6EFF92FC79FDC47A156211" box="[958,1029,584,610]" pageId="9" pageNumber="370">Rhum</collectorName>
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,
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<quantity id="4CDD9B16BF6EFF92FBD7FDC47A496210" box="[1040,1113,584,611]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="9" pageNumber="370" unit="m" value="285.0">
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<elevation id="0008D1C0BF6EFF92FBD7FDC47A496210" box="[1040,1113,584,611]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="9" pageNumber="370" unit="m" value="285.0">285 m</elevation>
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</quantity>
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,
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<collectorName id="26D05325BF6EFF92FBA3FDC57AF26211" box="[1124,1250,584,611]" pageId="9" pageNumber="370">Dr B. Earll</collectorName>
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,
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<collectorName id="26D05325BF6EFF92FB2AFDC47F2962F5" pageId="9" pageNumber="370">Marine Conservation Society</collectorName>
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,
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<specimenCount id="9D23FD7ABF6EFF92FE83FDE17F8362F4" box="[324,403,621,647]" pageId="9" pageNumber="370" type="generic">1 spec.</specimenCount>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<caption id="DF5A667BBF6EFF92FF50FA217D8F65B5" ID-DOI="http://doi.org/10.5281/zenodo.3516801" ID-Zenodo-Dep="3516801" box="[151,927,1453,1478]" httpUri="https://zenodo.org/record/3516801/files/figure.png" pageId="9" pageNumber="370" startId="9.[151,250,1453,1478]" targetBox="[212,1335,653,1434]" targetPageId="9">
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<paragraph id="8B9A36F3BF6EFF92FF50FA217D8F65B5" blockId="9.[151,927,1453,1478]" box="[151,927,1453,1478]" pageId="9" pageNumber="370">
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<emphasis id="B951EAE1BF6EFF92FF50FA217F0365B5" bold="true" box="[151,275,1453,1478]" pageId="9" pageNumber="370">FIGURE 5.</emphasis>
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<emphasis id="B951EAE1BF6EFF92FEDEFA227F6465B5" box="[281,372,1454,1478]" italics="true" pageId="9" pageNumber="370">
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<taxonomicName id="4C254D70BF6EFF92FEDEFA227F7465B5" authorityName="Oken" authorityYear="1815" box="[281,356,1454,1478]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="genus">Thyone</taxonomicName>
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?
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</emphasis>
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<taxonomicName id="4C254D70BF6EFF92FEB3FA227C4D65B5" authority="Norman, 1869" authorityName="Norman" authorityYear="1869" box="[372,605,1453,1478]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="elegans">
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<emphasis id="B951EAE1BF6EFF92FEB3FA227FD465B5" box="[372,452,1454,1478]" italics="true" pageId="9" pageNumber="370">elegans</emphasis>
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<bibRefCitation id="EFB44B02BF6EFF92FE0CFA227C4D65B5" author="Norman, A. M." box="[459,605,1453,1478]" pageId="9" pageNumber="370" pagination="47 - 336" refId="ref12062" refString="Norman, A. M. (1869) Shetland final dredging report ii. On the Crustacea, Tunicata, Polyzoa, Echinodermata, Actinozoa, Hydrozoa and Porifera. Report of the British Association of the Advancement of Science, 1869, 47 - 336." type="journal article" year="1869">Norman, 1869</bibRefCitation>
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</taxonomicName>
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/?
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<taxonomicName id="4C254D70BF6EFF92FDBBFA237D8B65B5" authority="Herouard, 1889" authorityName="Herouard" authorityYear="1889" box="[636,923,1453,1478]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="roscovita">
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<emphasis id="B951EAE1BF6EFF92FDBBFA237CE265B5" box="[636,754,1455,1478]" italics="true" pageId="9" pageNumber="370">T. roscovita</emphasis>
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<bibRefCitation id="EFB44B02BF6EFF92FD3EFA217D8B65B5" author="Herouard, R." box="[761,923,1453,1478]" pageId="9" pageNumber="370" pagination="535 - 704" refId="ref11494" refString="Herouard, R. (1889) Recherches sur les holothuries des cotes de France. Archives de Zoologie Experimentale et Generale, 2, 535 - 704." type="journal article" year="1889">Hérouard, 1889</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<subSubSection id="C33F6578BF6EFF92FF00FA7F7D1566ED" pageId="9" pageNumber="370" type="description">
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<paragraph id="8B9A36F3BF6EFF92FF00FA7F7D1566ED" blockId="9.[151,1437,1523,1875]" pageId="9" pageNumber="370">
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<emphasis id="B951EAE1BF6EFF92FF00FA7F7F49667D" bold="true" box="[199,345,1523,1550]" pageId="9" pageNumber="370">Description.</emphasis>
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Specimen U-shaped; off-white in alcohol, anterior and posterior ends greyish, length
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<quantity id="4CDD9B16BF6EFF92FA85FA787B88667C" box="[1346,1432,1524,1551]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="370" unit="mm" value="46.0">46 mm</quantity>
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; breadth in mid-body
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<quantity id="4CDD9B16BF6EFF92FE4FF99B7FDE6640" box="[392,462,1559,1587]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="370" unit="mm" value="5.0">5 mm</quantity>
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. Mouth and anus terminal, the latter with five anal teeth. Tube feet minute, retracted, scattered, more numerous ventrally with some indication of their arrangement in double rows anteriorly and posteriorly, with scattering in mid-body. Calcareous ring and associated structures absent. End plates present in tube feet but no ossicles were evident in body wall and tube feet.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6578BF6EFF92FF00F9247DB66720" pageId="9" pageNumber="370" type="discussion">
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<paragraph id="8B9A36F3BF6EFF92FF00F9247DB66720" blockId="9.[151,1437,1523,1875]" pageId="9" pageNumber="370">
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<emphasis id="B951EAE1BF6EFF92FF00F9247F2B66B1" bold="true" box="[199,315,1704,1730]" pageId="9" pageNumber="370">Remarks.</emphasis>
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It is suspected that because of the complete lack of body-wall ossicles and the presence of darker coloured anterior and posterior ends and tube feet end plates, the specimen might represent
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<taxonomicName id="4C254D70BF6EFF92FBABF9407B0B6695" authorityName="Norman" authorityYear="1869" box="[1132,1307,1740,1766]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="elegans">
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<emphasis id="B951EAE1BF6EFF92FBABF9407B0B6695" box="[1132,1307,1740,1766]" italics="true" pageId="9" pageNumber="370">Thyone elegans</emphasis>
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</taxonomicName>
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(?synonym
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<taxonomicName id="4C254D70BF6EFF92FF50F97D7F086779" authorityName="Herouard" authorityYear="1889" box="[151,280,1777,1802]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="roscovita">
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<emphasis id="B951EAE1BF6EFF92FF50F97D7F086779" box="[151,280,1777,1802]" italics="true" pageId="9" pageNumber="370">T. roscovita</emphasis>
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</taxonomicName>
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) but as the calcareous ring and associated structures are absent, a definite conclusion is not possible. The species could also be
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<taxonomicName id="4C254D70BF6EFF92FE49F8997C5C675D" authority="Heller" authorityName="Heller" authorityYear="1869" box="[398,588,1812,1838]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="inermis">
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<emphasis id="B951EAE1BF6EFF92FE49F8997FEE675D" box="[398,510,1813,1838]" italics="true" pageId="9" pageNumber="370">T. inermis</emphasis>
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Heller
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</taxonomicName>
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but
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<bibRefCitation id="EFB44B02BF6EFF92FDB8F8987D56675D" author="McKenzie, J. D." box="[639,838,1812,1838]" pageId="9" pageNumber="370" pagination="123 - 171" refId="ref11951" refString="McKenzie, J. D. (1991) The taxonomy and natural history of north European dendrochirote holothurians (Echinodermata). Journal of Natural History, 25 (1), 123 - 171. https: // doi. org / 10.1080 / 00222939100770091" type="journal article" year="1991">McKenzie (1991)</bibRefCitation>
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states that
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<taxonomicName id="4C254D70BF6EFF92FC00F8997A27675D" authorityName="Heller" authorityYear="1869" box="[967,1079,1813,1838]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="inermis">
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<emphasis id="B951EAE1BF6EFF92FC00F8997A27675D" box="[967,1079,1813,1838]" italics="true" pageId="9" pageNumber="370">T. inermis</emphasis>
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</taxonomicName>
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always has deposits in the anal region and is perhaps a clinal variation of
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<taxonomicName id="4C254D70BF6EFF92FDA9F8B57CE16721" authorityName="Herouard" authorityYear="1889" box="[622,753,1849,1874]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="roscovita">
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<emphasis id="B951EAE1BF6EFF92FDA9F8B57CE16721" box="[622,753,1849,1874]" italics="true" pageId="9" pageNumber="370">T. roscovita</emphasis>
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</taxonomicName>
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(?=
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<taxonomicName id="4C254D70BF6EFF92FCE2F8B57D866721" authorityName="Norman" authorityYear="1869" box="[805,918,1848,1874]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="370" phylum="Echinodermata" rank="species" species="elegans">
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<emphasis id="B951EAE1BF6EFF92FCE2F8B57D866721" box="[805,918,1848,1874]" italics="true" pageId="9" pageNumber="370">T. elegans</emphasis>
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</taxonomicName>
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|
).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |