treatments-xml/data/03/8C/87/038C87C1D360EA3DFF1218CFFCD23765.xml

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<document id="C7F5B31A5EF2089B94152F2969F57133" ID-DOI="10.11646/zootaxa.5562.1.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595661" ID-ZooBank="14DDD852-231F-4150-AE26-550F867EF7EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1735931848199" checkinUser="plazi" docAuthor="Lee, Soo Bin, Nam, Gi Soo &amp; Park, Jong Kyun" docDate="2024" docId="038C87C1D360EA3DFF1218CFFCD23765" docLanguage="en" docName="zootaxa.5562.1.12.pdf" docOrigin="Zootaxa 5562 (1)" docSource="https://doi.org/10.11646/zootaxa.5562.1.12" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Necronectulus lazarus Lee, Nam &amp; Park, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="101" masterDocId="FFB5FFB9D366EA35FF851E28FFB23624" masterDocTitle="Cretaceous beetles from Jinju Formation: Trachypachidae (Coleoptera: Adephaga)" masterLastPageNumber="106" masterPageNumber="94" pageNumber="100" updateTime="1735933828614" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9CDBF6BC35E2B0CAAAFED2926A7CCB0E">Cretaceous beetles from Jinju Formation: Trachypachidae (Coleoptera: Adephaga)</mods:title>
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<mods:namePart id="2A956DA2D62E8EC0641F7C9F851AAA5B">Lee, Soo Bin</mods:namePart>
<mods:nameIdentifier id="8D075B6297D7C2BAC738B83973DD4AE5" type="ORCID">0000-0003-2112-3897</mods:nameIdentifier>
<mods:affiliation id="0DFBED922F90E62C5F086A40B89EEB4F">Korean Fossil Research Center, 50 - 2, Hangangro 2 Ga, YongsangGu, Seoul, Republic of Korea &amp; Department of Science Education, Gongju National University of Education, 27 Ungjinro, Gongjusi, Chungcheongnam-do 32553, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="A0113E59195F3FBE9FB57BA0B271B9B3" type="email">dinos20000@naver.com</mods:nameIdentifier>
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<mods:namePart id="2BBE659370F24A56DEECC313C10BAC7D">Nam, Gi Soo</mods:namePart>
<mods:nameIdentifier id="1BF0415E606127FCE36501E9FE96A4A3" type="ORCID">0000-0003-0570-8270</mods:nameIdentifier>
<mods:affiliation id="AD4A465529862D0D8207DA1FA70241FF">Department of Science Education, Gongju National University of Education, 27 Ungjinro, Gongjusi, Chungcheongnam-do 32553, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="2AB83726B88D16A25FD1CD8EFFE409B2" type="email">fossil@gjue.ac.kr</mods:nameIdentifier>
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<mods:namePart id="C6081AB8B34C266F83B18DC058B9F7D9">Park, Jong Kyun</mods:namePart>
<mods:nameIdentifier id="CD914215A2BA0190F35F965CD430ABD5" type="ORCID">0000-0002-4049-3546</mods:nameIdentifier>
<mods:affiliation id="6ECFAC08BAEC04C94D2A28634591F45E">College of Ecology and Environmental Science, Kyungpook National University, Sangju-si, Gyeongsangbuk-do 37224, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="F5254ACCEAEC242BE12384FBA970BDDA" type="email">entopark@knu.ac.kr</mods:nameIdentifier>
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<mods:date id="65F6D866BCF22D0BB4B1516A29C2CF6C">2024</mods:date>
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<subSubSection id="C33F655CD360EA33FF1218CFFDB93125" box="[151,523,1767,1794]" pageId="6" pageNumber="100" type="nomenclature">
<paragraph id="8B9A36D7D360EA33FF1218CFFDB93125" blockId="6.[151,523,1767,1794]" box="[151,523,1767,1794]" pageId="6" pageNumber="100">
<heading id="D0D281BBD360EA33FF1218CFFDB93125" bold="true" box="[151,523,1767,1794]" fontSize="11" level="1" pageId="6" pageNumber="100" reason="1">
<emphasis id="B951EAC5D360EA33FF1218CFFDB93125" bold="true" box="[151,523,1767,1794]" pageId="6" pageNumber="100">
<taxonomicName id="4C254D54D360EA33FF1218CFFE153126" authorityName="Lee &amp; Nam &amp; Park" authorityYear="2024" box="[151,423,1767,1794]" class="Insecta" family="Trachypachidae" genus="Necronectulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B951EAC5D360EA33FF1218CFFE153126" bold="true" box="[151,423,1767,1794]" italics="true" pageId="6" pageNumber="100">Necronectulus lazarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257BED360EA33FE2B18CFFDB93125" box="[430,523,1767,1793]" pageId="6" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C33F655CD360EA33FF121907FD8E316D" box="[151,572,1839,1865]" pageId="6" pageNumber="100" type="materials_examined">
<paragraph id="8B9A36D7D360EA33FF121907FD8E316D" blockId="6.[151,1436,1839,2009]" box="[151,572,1839,1865]" pageId="6" pageNumber="100">
<emphasis id="B951EAC5D360EA33FF121907FEB4316D" bold="true" box="[151,262,1839,1865]" pageId="6" pageNumber="100">Material.</emphasis>
<typeStatus id="549E8875D360EA33FE881907FEC7316D" box="[269,373,1839,1865]" pageId="6" pageNumber="100" type="holotype">Holotype</typeStatus>
KNUL-1114-110
</paragraph>
</subSubSection>
<subSubSection id="C33F655CD360EA33FF42197BFD8831FD" pageId="6" pageNumber="100" type="etymology">
<paragraph id="8B9A36D7D360EA33FF42197BFAEF31B5" blockId="6.[151,1436,1839,2009]" pageId="6" pageNumber="100">
<emphasis id="B951EAC5D360EA33FF42197BFEFC3149" bold="true" box="[199,334,1875,1901]" pageId="6" pageNumber="100">Etymology.</emphasis>
The species name
<taxonomicName id="4C254D54D360EA33FDA2197BFDCB3149" authorityName="Lee &amp; Nam &amp; Park" authorityYear="2024" box="[551,633,1875,1901]" class="Insecta" family="Trachypachidae" genus="Necronectulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="lazarus">
<emphasis id="B951EAC5D360EA33FDA2197BFDCB3149" box="[551,633,1875,1901]" italics="true" pageId="6" pageNumber="100">lazarus</emphasis>
</taxonomicName>
is derived from the term “Lazarus taxon”, which refers to a taxon that was initially considered to have disappeared from a particular period but later reappeared in the fossil record.
</paragraph>
<paragraph id="8B9A36D7D360EA33FF4219B3FD8831FD" blockId="6.[151,1436,1839,2009]" pageId="6" pageNumber="100">
<emphasis id="B951EAC5D360EA33FF4219B3FE6B3191" bold="true" box="[199,473,1947,1973]" pageId="6" pageNumber="100">Locality and horizon.</emphasis>
Goryeong-gun, Seongsan-myeon,
<collectingRegion id="49E1F835D360EA33FCFE19B3FBEB3191" box="[891,1113,1947,1973]" country="South Korea" name="Gyeongbuk" pageId="6" pageNumber="100">Gyeongsangbuk-do</collectingRegion>
,
<collectingCountry id="F3327647D360EA33FBE819B3FAE03192" box="[1133,1362,1947,1974]" name="South Korea" pageId="6" pageNumber="100">Republic of Korea</collectingCountry>
; Jinju Formation, Early Cretaceous, Albian.
</paragraph>
</subSubSection>
<caption id="DF5A665FD361EA32FF121BCDFCAD33D9" box="[151,799,1509,1533]" pageId="7" pageNumber="101" startId="7.[151,250,1509,1533]" targetType="figure">
<paragraph id="8B9A36D7D361EA32FF121BCDFCAD33D9" blockId="7.[151,799,1509,1533]" box="[151,799,1509,1533]" pageId="7" pageNumber="101">
<emphasis id="B951EAC5D361EA32FF121BCDFEA133D9" bold="true" box="[151,275,1509,1533]" pageId="7" pageNumber="101">FIGURE 5.</emphasis>
<taxonomicName id="4C254D54D361EA32FE9C1BCDFE4E33D9" authorityName="Lee &amp; Nam &amp; Park" authorityYear="2024" box="[281,508,1509,1533]" class="Insecta" family="Trachypachidae" genus="Necronectulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="101" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B951EAC5D361EA32FE9C1BCDFE4E33D9" box="[281,508,1509,1533]" italics="true" pageId="7" pageNumber="101">Necronectulus lazarus</emphasis>
</taxonomicName>
<emphasis id="B951EAC5D361EA32FD871BCDFDE133D9" bold="true" box="[514,595,1509,1533]" pageId="7" pageNumber="101">
<taxonomicNameLabel id="A26257BED361EA32FD871BCDFDE133D9" box="[514,595,1509,1533]" pageId="7" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(KNUL-1114-110).
</paragraph>
</caption>
<subSubSection id="C33F655CD361EA32FF421867FC4B304D" box="[199,1017,1615,1642]" pageId="7" pageNumber="101" type="diagnosis">
<paragraph id="8B9A36D7D361EA32FF421867FC4B304D" blockId="7.[151,1436,1615,2038]" box="[199,1017,1615,1642]" pageId="7" pageNumber="101">
<emphasis id="B951EAC5D361EA32FF421867FEF0304D" bold="true" box="[199,322,1615,1641]" pageId="7" pageNumber="101">Diagnosis.</emphasis>
Large scutellum; 9 longitudinal striae grooved on the elytron.
</paragraph>
</subSubSection>
<subSubSection id="C33F655CD361EA32FF42185BFB7A3189" pageId="7" pageNumber="101" type="description">
<paragraph id="8B9A36D7D361EA32FF42185BFB7A3189" blockId="7.[151,1436,1615,2038]" pageId="7" pageNumber="101">
<emphasis id="B951EAC5D361EA32FF42185BFEEB30A9" bold="true" box="[199,345,1651,1677]" pageId="7" pageNumber="101">Description.</emphasis>
Overall body oval shape, with greatest width at posterior part of pronotum extending to anterior margin of elytra. Total body length
<quantity id="4CDD9B32D361EA32FD9C18BFFDC23096" box="[537,624,1687,1714]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="101" unit="mm" value="7.5">7.5 mm</quantity>
, width
<quantity id="4CDD9B32D361EA32FD3B18BFFCA73096" box="[702,789,1687,1714]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="101" unit="mm" value="4.5">4.5 mm</quantity>
. Head wider than long, and extremely narrowed anteriorly; frons wider than long; width of frons and clypeus almost equal from posterior to anterior parts; labrum rounded anteriorly; maxillary palp with three segments; compound eyes large; antennae long, with 11 antennomeres; lateral margin of pronotum slightly rounded, with both anterior and posterior margins straight; anterior corners sharp; a longitudinal groove running medially from pronotum to frons; anterior margin of elytra straight, elytra formed with nine longitudinally grooved striae; epipleuron thick; scutellum large, triangular in shape; adephagus protruded and rounded posteriorly. Ventral part of body poorly preserved, but several traces evident; precoxae contiguous with each other, and profemur wider distally; metacoxal plate extending laterally to meet epipleuron.
</paragraph>
</subSubSection>
<subSubSection id="C33F655CD361EA3DFF42199FFCD23765" lastPageId="8" lastPageNumber="102" pageId="7" pageNumber="101" type="discussion">
<paragraph id="8B9A36D7D361EA3DFF42199FFCD23765" blockId="7.[151,1436,1615,2038]" lastBlockId="8.[151,1436,151,321]" lastPageId="8" lastPageNumber="102" pageId="7" pageNumber="101">
<emphasis id="B951EAC5D361EA32FF42199FFE8931F5" bold="true" box="[199,315,1975,2001]" pageId="7" pageNumber="101">Remarks.</emphasis>
Morphological features such as: head narrower than prothorax; Antennae inserted between the eyes and base of the mandible; Filiform shape antennae formed with 11 antennomeres; metacoxae extended laterally and meet with the lateral margins of body are features of the family
<taxonomicName id="4C254D54D36EEA3DFC101EBFFB603695" authority="(Ball 2000)" baseAuthorityName="Ball" baseAuthorityYear="2000" box="[917,1234,151,177]" class="Insecta" family="Trachypachidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="102" phylum="Arthropoda" rank="family">
Trachypachidae (
<bibRefCitation id="EFB44B26D36EEA3DFBD31EBFFB783695" author="Ball, G. E." box="[1110,1226,151,177]" pageId="8" pageNumber="102" pagination="144 - 146" refId="ref4107" refString="Ball, G. E. (2000) 9. Trachypachidae Thomson, C. G., 1857. In: Arnett, R. H. Jr. &amp; Thomas, M. C. (Eds). American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 144 - 146." type="book chapter" year="2000">Ball 2000</bibRefCitation>
)
</taxonomicName>
. The overall body shape of KNUL-1114-110 is oval, with the head being transverse and wider than long. The length of the head is greater than that of the pronotum. The pronotum tapers roundly anteriorly, with both the posterior margin of the pronotum and the anterior margin of the elytra being straight. The antennae are short and slender. These features are characteristic of the genus
<taxonomicName id="4C254D54D36EEA3DFE461F0FFCE93765" authority="(Ponomarenko 1977)" baseAuthorityName="Ponomarenko" baseAuthorityYear="1977" box="[451,859,295,321]" class="Insecta" family="Trachypachidae" genus="Necronectulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAC5D36EEA3DFE461F0FFDD63765" box="[451,612,295,321]" italics="true" pageId="8" pageNumber="102">Necronectulus</emphasis>
(
<bibRefCitation id="EFB44B26D36EEA3DFDF71F0FFCE13765" author="Ponomarenko, A. G." box="[626,851,295,321]" pageId="8" pageNumber="102" pagination="17 - 104" refId="ref7225" refString="Ponomarenko, A. G. (1977) Suborder Adephaga. In: Arnoldi L. V., Zherikhin V. V., Nikritin L. M. &amp; Ponomarenko, A. G. (Eds), Mezozoiskie zhestkokryiye. Nauka Publishers, Moscow, pp. 17 - 104. [in Russian]." type="book chapter" year="1977">Ponomarenko 1977</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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