211 lines
28 KiB
XML
211 lines
28 KiB
XML
<document id="AE2725642B136CA7F1E665378DD23500" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub & Caterino, Michael S" docDate="2024" docId="038C87BE171981654C86FDE6FD4C7C92" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Placatister cascus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop & Caterino, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="18" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:titleInfo id="A02FDB19244A2843DE72FE40095C6A3A">
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<mods:title id="EBC13E91188E57906285170B5E4E9B69">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:name id="5DCEBBBA3C511004AC5E1E457B6E34DC" type="personal">
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<mods:roleTerm id="B8F9C38242D256D760B4D10416F12D67">Author</mods:roleTerm>
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<mods:namePart id="71D37B4BCA6E1FC4296EBBEA7A7D6F21">Simon-Pražák, Jan</mods:namePart>
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<mods:affiliation id="F0AF49FCC8737AC65BAFBB567988DB72">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic & Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="EAA5B2266D59B1F8E6C798A507E2DC8D" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
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<mods:name id="770BDEDF0BAD61ABC4548957D798C02A" type="personal">
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<mods:roleTerm id="9EDA521802BE36294FD5871273B25276">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8A3FE75611F03771AD3C9EEDB24380A5">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="19B61C1013CCC203DDAFE88F025C79A0">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="347E4F083D720E71CBEAAF9C1414B743" type="personal">
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<mods:roleTerm id="F258E8371EF2ABDAD9E04F9531025C72">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D92BDB442789F5A6D40B13BB18D653E5">Lackner, Tomáš</mods:namePart>
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<mods:affiliation id="97AD8BF59FBD200A324D9F91F9F523F6">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name id="2510DCE438EDD80B22F812B0073507DA" type="personal">
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<mods:namePart id="724402D0DB62DF59B1FCC28CE960CB2C">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="C859F8639FABF911010BD0E12D080B93">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan & Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="57473D6D30CB5DFC964185B072A217DD">Prokop, Jakub</mods:namePart>
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<mods:affiliation id="FE05FE017F22D00741DE9FB646B3F11E">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic & Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="1C6B8CD80CDCE6BFAE4DEB0221F1F824">Caterino, Michael S</mods:namePart>
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<mods:title id="F6D0A36ACB6CDC63BBE385F024D0969F">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="768F5BEE6FF90348CDC91C4AA1AA045E">
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<mods:date id="4076076EC5E014F2E8761F7EF89F1097">2024</mods:date>
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<mods:title id="7556B3DACB9436D93B10529D4362FD34">zlae 137</mods:title>
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<mods:number id="A9F304B2D8601DF23C40488751FA1EB2">2024-11-20</mods:number>
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<mods:number id="DD09C2A3BC6081417A500D233C092438">202</mods:number>
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<mods:detail id="2DAB3303D9B1F2DB941FA05908C2CC14" type="issue">
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<mods:number id="8185A67D87C1BC0BCD6A2BF4449975EB">3</mods:number>
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<mods:start id="BE89F465C63157CC2E01AC02E0997B26">1</mods:start>
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<mods:url id="3854287D8E4216E4FE19563D06300BBA">https://doi.org/10.1093/zoolinnean/zlae137</mods:url>
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<mods:classification id="25C6C8AC0EB76898A9343D8D987BDD1C">journal article</mods:classification>
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<mods:identifier id="CB72B4899FB640B1938B770EB8D56D2D" type="DOI">10.1093/zoolinnean/zlae137</mods:identifier>
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<mods:identifier id="8AA881BD65F8E98AE780C24E9BCF4A54" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="E46E4F393655FF98A90F44AE41DB9B6D" type="Zenodo-Dep">14516114</mods:identifier>
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<treatment id="038C87BE171981654C86FDE6FD4C7C92" ID-DOI="http://doi.org/10.5281/zenodo.14516156" ID-Zenodo-Dep="14516156" LSID="urn:lsid:plazi:treatment:038C87BE171981654C86FDE6FD4C7C92" httpUri="http://treatment.plazi.org/id/038C87BE171981654C86FDE6FD4C7C92" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Onthophilinae">
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<subSubSection id="C33F6523171981674C86FDE6FB5F796F" box="[990,1279,582,609]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8B9A36A8171981674C86FDE6FB5F796F" blockId="17.[990,1279,582,609]" box="[990,1279,582,609]" pageId="17" pageNumber="18">
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<heading id="D0D281C4171981674C86FDE6FB5F796F" bold="true" box="[990,1279,582,609]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
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<emphasis id="B951EABA171981674C86FDE6FB5F796F" bold="true" box="[990,1279,582,609]" pageId="17" pageNumber="18">
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||
<taxonomicName id="4C254D2B171981674C86FDE6FB03796F" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[990,1187,582,609]" class="Insecta" family="Histeridae" genus="Placatister" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cascus" status="sp. nov.">
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<emphasis id="B951EABA171981674C86FDE6FB03796F" bold="true" box="[990,1187,582,609]" italics="true" pageId="17" pageNumber="18">Placatister cascus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26257C1171981674BF0FDE7FB5F796F" box="[1192,1279,583,609]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33F6523171981674B18FDD5FB3D7981" box="[1088,1181,629,656]" pageId="17" pageNumber="18" type="description">
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<paragraph id="8B9A36A8171981674B18FDD5FB3D7981" blockId="17.[1088,1181,629,656]" box="[1088,1181,629,656]" pageId="17" pageNumber="18">
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<emphasis id="B951EABA171981674B18FDD5FB3D7981" bold="true" box="[1088,1181,629,656]" pageId="17" pageNumber="18">
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||
(
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||
<figureCitation id="131E2A2D171981674B13FDD6FB32799E" box="[1099,1170,630,656]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." figureDoi="http://doi.org/10.5281/zenodo.14516146" httpUri="https://zenodo.org/record/14516146/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
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)
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</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33F6523171981674C72FD16FB477803" pageId="17" pageNumber="18" type="materials_examined">
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||
<paragraph id="8B9A36A8171981674C72FD16FB477803" blockId="17.[810,1460,694,781]" pageId="17" pageNumber="18">
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<emphasis id="B951EABA171981674C72FD16FC2679C0" box="[810,902,694,718]" italics="true" pageId="17" pageNumber="18">Material:</emphasis>
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<materialsCitation id="3B4D3CF5171981674CD5FD16FB437803" collectionCode="MEBHK" location="Kachin" pageId="17" pageNumber="18" specimenCode="P1425/E" specimenCount="1" stateProvince="Kachin">
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<collectingRegion id="49E1F84A171981674CD5FD16FC7779C0" box="[909,983,694,718]" country="Myanmar" name="Kachin" pageId="17" pageNumber="18">Kachin</collectingRegion>
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amber (
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<emphasis id="B951EABA171981674B69FD17FB9A79C0" box="[1073,1082,695,718]" italics="true" pageId="17" pageNumber="18">c</emphasis>
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. 99 Mya). Amber piece dimensions: 9 × 5 ×
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<quantity id="4CDD9B4D171981674C25FD76FC1879E3" box="[893,952,726,750]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
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. Amber clear with several insect fragments and air bubbles. Deposited at
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<collectionCode id="ED34AE6D171981674B54FD55FBC87802" box="[1036,1128,757,780]" pageId="17" pageNumber="18">MEBHK</collectionCode>
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(
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<specimenCode id="DB839ED3171981674B2EFD55FB7B7803" box="[1142,1243,757,781]" pageId="17" pageNumber="18">P1425/E</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6523171981654C72FC8FFF097876" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
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<paragraph id="8B9A36A8171981654C72FC8FFF097876" blockId="17.[809,1460,814,1966]" lastBlockId="19.[113,763,144,888]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<emphasis id="B951EABA171981674C72FC8FFC1C7848" box="[810,956,815,838]" italics="true" pageId="17" pageNumber="18">Measurements:</emphasis>
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Head width:
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<quantity id="4CDD9B4D171981674B0AFC8EFB0D7848" box="[1106,1197,814,838]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="17" pageNumber="18" unit="mm" value="0.36">0.36 mm</quantity>
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; width between anterior pronotal angles:
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<quantity id="4CDD9B4D171981674C86FCEDFB99786B" box="[990,1081,845,869]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="18" unit="mm" value="0.56">0.56 mm</quantity>
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; width between posterior pronotal angles:
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<quantity id="4CDD9B4D171981674C27FCCDFC7A788A" box="[895,986,877,901]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="17" pageNumber="18" unit="mm" value="0.92">0.92 mm</quantity>
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; pronotal length:
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<quantity id="4CDD9B4D171981674BF5FCCDFAA8788A" box="[1197,1288,877,901]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="18" unit="mm" value="0.58">0.58 mm</quantity>
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; sutural elytral length:
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<quantity id="4CDD9B4D171981674C26FC2CFC6B78AA" box="[894,971,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="mm" value="0.8">0.8 mm</quantity>
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; elytral width (in pair, across widest point):
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<quantity id="4CDD9B4D171981674C72FC0CFCDA78CD" box="[810,890,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
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.—
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<emphasis id="B951EABA171981674CC1FC0BFC7978CD" bold="true" box="[921,985,939,963]" italics="true" pageId="17" pageNumber="18">Body:</emphasis>
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Round, strongly dorsoventrally flattened (probably partially compressed). Cuticle regularly punctuate. Colour reddish brown to black.—
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<emphasis id="B951EABA171981674BCCFC4AFB757F0C" bold="true" box="[1172,1237,1002,1026]" italics="true" pageId="17" pageNumber="18">Head:</emphasis>
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Frons flat, wide and punctuate, strongly, squarely margined along inner edge of each eye, with a fine stria along their inner edges continuous with a complete, transverse frontal stria. Frontoclypeal suture not visible. Epistoma constricted, medially ridged, sides receding below broad frontal shelf. Labrum trapezoidal (width: length ratio 5: 1), emarginate anteriorly, with a pair of short setae at anterolateral corners, lacking punctuation. Mandibles wide and thin, basally with weakly dentate incisor edge, and a long, acute tip. Terminal maxillary palpomere wide and round; other ventral mouthparts damaged and/or missing. Antenna inserted under frontal shelf, strongly dorsoventrally flattened, possibly partially compressed. Scape short, widened, and curved to apex. Pedicel wide, subcylindrical, as long as antennomeres 3–5 combined. Antennomeres 3–7 similar in shape and size, short, wider than long (width: length ratio 3: 1). Antennomere 8 adjacent to the antennal club, 2.5× wider than antennomere 7. Antennomeres 9–11 form a wide, flat, and round antennal club with well-defined sutures.—
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<emphasis id="B951EABA171981674CCCF9BEFC497D38" bold="true" box="[916,1001,1566,1590]" italics="true" pageId="17" pageNumber="18">Thorax:</emphasis>
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Pronotal disc flat. Anterior margin widely emarginate, with fine marginal pronotal stria; not evident on lateral or basal margins. Basal margin broadly rounded, almost angularly projecting. Surface of pronotal disc densely punctuate, punctures separated by 1–2× their diameter. Scutellar shield not visible, obscured by bubbles. Elytra truncate, without dorsal striation, epipleural stria complete; elytral disc appears weakly depressed in humeral corners (possibly due to damage). Elytral punctuation with the same pattern as in pronotum. Prosternal lobe rather short, subtruncate anteriorly, set off from keel by pair of deep, setose foveae, weakly conjoined at their bases; prosternal keel flat, short, abruptly descending at front into two large approximate prosternal foveae, with fine longitudinal striae; finely punctate between; sides of prosternum widely incised for passage of antennal funicle, with deep channel reaching middle of base of prosternal keel; sides of prosternal keel appear concave. Mesoventrite wide and short, obscured by bubbles, probably finely punctuated. Meso-metaventral suture visible as a fine line. Metaventrite punctuate, with lateral metaventral stria directed posterolaterally to hind coxae. Other striation not visible. Metepisternum broadly exposed, rectangular.—
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<emphasis id="B951EABA171B81654D9AFECCFD5A7A8D" bold="true" box="[706,762,364,387]" italics="true" pageId="19" pageNumber="20">Legs:</emphasis>
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Profemur flatted, slightly widened. Protibia narrow and smooth, without dentation, few longer setae present along inner apical edge, without tibial spur. Tarsal groove present as an impressed line on the outer edge (along the entire protibial length). Tarsi 5-5-5; protarsomere 5 as long as protarsomeres 2–4 combined. Claws well developed, about half the length of protarsomere 5. Mesofemur swollen, mesotibia narrow. Three prominent setae can be seen in the median part of the left mesotibial outer margin. Mesotarsus identical to protarsus. Metafemur slender, slightly curved and swollen in the median part. Metatibia slender, with a couple stronger spines at outer apical corner; metatarsus identical to protarsus.—
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<emphasis id="B951EABA171B81654E1AFD43FE0F79F5" bold="true" box="[322,431,739,763]" italics="true" pageId="19" pageNumber="20">Abdomen:</emphasis>
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First visible abdominal sternite without visible striation. All visible abdominal sclerites finely punctuated. Propygidium very short, almost entirely covered by elytra. Pygidium weakly convex, about two-thirds as long as wide.
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</paragraph>
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</subSubSection>
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||
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|
||
<paragraph id="8B9A36A8171A81644FD9F88CFBD07C6E" blockId="18.[129,1441,1836,1888]" pageId="18" pageNumber="19">
|
||
<emphasis id="B951EABA171A81644FD9F88CFF457C4A" bold="true" box="[129,229,1836,1860]" pageId="18" pageNumber="19">Figure 12.</emphasis>
|
||
<taxonomicName id="4C254D2B171A81644FB1F88CFE237C4A" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[233,387,1836,1860]" class="Insecta" family="Histeridae" genus="Placatister" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cascus" status="gen. et sp. nov">
|
||
<emphasis id="B951EABA171A81644FB1F88CFE237C4A" box="[233,387,1836,1860]" italics="true" pageId="18" pageNumber="19">Placatister cascus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C1171A81644ED0F88DFDAC7C4A" box="[392,524,1837,1861]" pageId="18" pageNumber="19" rank="species">gen. et sp. nov</taxonomicNameLabel>
|
||
.. A, dorsal view. B, ventral view. C, lateral view. D, antennal club. E, mouthparts. F, frontal view. G, prosternal keel. H, antenna. Abbreviations: ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F6523171B81654F29FC3EFEF37D26" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="8B9A36A8171B81654F29FC3EFEF37D26" blockId="19.[113,763,925,1576]" pageId="19" pageNumber="20">
|
||
<emphasis id="B951EABA171B81654F29FC3EFEC178BB" box="[113,353,926,949]" italics="true" pageId="19" pageNumber="20">Taxonomic assignment:</emphasis>
|
||
The species displays several highly plesiomorphic characters, such as the absence of a prosternal lobe and antennal cavity, as well as in the shape of the antennal club with distinct segments (
|
||
<figureCitation id="131E2A2D171B81654EFCFC5BFE5E7F1D" box="[420,510,1019,1043]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." figureDoi="http://doi.org/10.5281/zenodo.14516146" httpUri="https://zenodo.org/record/14516146/files/figure.png" pageId="19" pageNumber="20">Fig. 12D</figureCitation>
|
||
). The flattened antenna with wide and short antennomeres 3–7 is especially unique among the entire
|
||
<taxonomicName id="4C254D2B171B81654E6BFB9AFE3E7F5C" box="[307,414,1082,1106]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
family. Prosternal structure, and particularly the prosternal foveae (
|
||
<figureCitation id="131E2A2D171B81654EB9FBF9FDFC7F7F" box="[481,604,1113,1137]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." figureDoi="http://doi.org/10.5281/zenodo.14516146" httpUri="https://zenodo.org/record/14516146/files/figure.png" pageId="19" pageNumber="20">Fig. 12B, G</figureCitation>
|
||
), are plausibly similar to those of some Saprininae (e.g.
|
||
<taxonomicName id="4C254D2B171B81654D4FFBD9FD2B7F9E" authorityName="Marseul" authorityYear="1862" box="[535,651,1145,1168]" class="Insecta" family="Histeridae" genus="Tomogenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171B81654D4FFBD9FD2B7F9E" box="[535,651,1145,1168]" italics="true" pageId="19" pageNumber="20">Tomogenius</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFB44B59171B81654DFFFBD8FF057FBE" author="Lackner T" pageId="19" pageNumber="20" pagination="1 - 254" refId="ref18143" refString="Lackner T. Review of the Palaearctic genera of ºaprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae 2010; 50: 1 - 254." type="journal article" year="2010">Lackner 2010</bibRefCitation>
|
||
). Nevertheless, the homology of these structures is highly unlikely. The presence (if true) of an antennal receptacle along the sides of the prosternal keel would also be similar to that of Saprininae. However, other characters, such as the distinctly sutured antennal club (
|
||
<figureCitation id="131E2A2D171B81654E1DFAB5FE3C7E23" box="[325,412,1301,1325]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." figureDoi="http://doi.org/10.5281/zenodo.14516146" httpUri="https://zenodo.org/record/14516146/files/figure.png" pageId="19" pageNumber="20">Fig. 12D</figureCitation>
|
||
), flattened body shape (
|
||
<figureCitation id="131E2A2D171B81654DCBFAB5FD497E23" box="[659,745,1301,1325]" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." figureDoi="http://doi.org/10.5281/zenodo.14516146" httpUri="https://zenodo.org/record/14516146/files/figure.png" pageId="19" pageNumber="20">Fig. 12C</figureCitation>
|
||
), and simple leg shape with slender tibiae without denticles (
|
||
<figureCitation id="131E2A2D171B81654D8CFA94FF3A7E62" captionStart="Figure 12" captionStartId="18.[129,194,1836,1860]" captionTargetBox="[175,1428,144,1808]" captionTargetId="figure-8@18.[175,1428,144,1808]" captionTargetPageId="18" captionText="Figure 12. Placatister cascus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateral view.D, antennal club. E, mouthparts. F, frontal view.G, prosternal keel.H, antenna.Abbreviations:ees, elytral epipleural stria; fs, frontal stria; pf, prosternal foveae." figureDoi="http://doi.org/10.5281/zenodo.14516146" httpUri="https://zenodo.org/record/14516146/files/figure.png" pageId="19" pageNumber="20">Fig. 12B</figureCitation>
|
||
), would be incompatible with this placement. The phylogenetic analyses did not resolve the position of this taxon, as it nested in various positions throughout the
|
||
<taxonomicName id="4C254D2B171B81654D75FA32FD387EA4" box="[557,664,1426,1450]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
tree (but outside the Saprininae and Histeromorphae clades). This species most likely represents a primitive stem lineage of
|
||
<taxonomicName id="4C254D2B171B81654DD2FA71FD567EE7" box="[650,758,1489,1513]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
, but a subfamily or tribal assignment is impossible based on the observable characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523171B81654F29F9EDFE477C4E" pageId="19" pageNumber="20" type="diagnosis">
|
||
<paragraph id="8B9A36A8171B81654F29F9EDFE477C4E" blockId="19.[113,762,1612,1856]" pageId="19" pageNumber="20">
|
||
<emphasis id="B951EABA171B81654F29F9EDFF777D6A" box="[113,215,1613,1636]" italics="true" pageId="19" pageNumber="20">Diagnosis:</emphasis>
|
||
Head with a complete frontal stria. Antennomeres short and wide, antennae dorsoventrally flattened. Prosternal process with a pair of setose foveae in apical part and with concave sides. Cuticle dorsally and ventrally punctuate. Legs narrow and smooth. The species may remotely resemble the flat
|
||
<taxonomicName id="4C254D2B171B81654DF3F96AFF577C0F" authorityName="Simon Prazak & Lackner" authorityYear="2023" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="complanatum">
|
||
<emphasis id="B951EABA171B81654DF3F96AFF577C0F" italics="true" pageId="19" pageNumber="20">Olexum complanatum</emphasis>
|
||
</taxonomicName>
|
||
or the round-bodied species of
|
||
<taxonomicName id="4C254D2B171B81654D19F949FD3B7C0F" authorityName="G. H. Horn" authorityYear="1873" box="[577,667,1769,1793]" class="Insecta" family="Histeridae" genus="Anapleus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171B81654D19F949FD3B7C0F" box="[577,667,1769,1793]" italics="true" pageId="19" pageNumber="20">Anapleus</emphasis>
|
||
</taxonomicName>
|
||
. It is distinguished by the wide and segmented antennal club and the wide and short segmented flagellum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523171B81654F29F8C4FD4C7C92" pageId="19" pageNumber="20" type="etymology">
|
||
<paragraph id="8B9A36A8171B81654F29F8C4FD4C7C92" blockId="19.[113,762,1892,1948]" pageId="19" pageNumber="20">
|
||
<emphasis id="B951EABA171B81654F29F8C4FF417C72" box="[113,225,1892,1916]" italics="true" pageId="19" pageNumber="20">Etymology:</emphasis>
|
||
This species represents a plesiomorphic set of characters, and the species’ name ‘cascus’ means ancient/primitive.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |