243 lines
29 KiB
XML
243 lines
29 KiB
XML
<document id="5B2E89838DD568BF702C4FD16FE2A1AC" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub & Caterino, Michael S" docDate="2024" docId="038C87BE170781674C9CFE52FE227CBF" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Prodigister tricostatus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop & Caterino, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="16" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:titleInfo id="15778A305C9CDB8F6EC7CA7578794D5B">
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<mods:title id="E15DE019D59E214BCCF3EF3FB7DBB751">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:name id="C9FFD2DFEC2D1C42AD0301C0F2A6FFAD" type="personal">
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<mods:roleTerm id="53A18C7CD5DFEC1AF8668FC7C582F2D6">Author</mods:roleTerm>
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<mods:namePart id="F252757332206DFA24127A8D3212D105">Simon-Pražák, Jan</mods:namePart>
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<mods:affiliation id="458C974311B02CDFC523EB72F3735588">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic & Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="FE0838083B29654DA06430CF48F74D73" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
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<mods:roleTerm id="DAFD0615E0CE554DBD34855758949446">Author</mods:roleTerm>
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<mods:namePart id="9AEF88DEDE1925B01118CBBB7D99C113">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="0FF5F44216A9E56A838D087EAD33D966">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="8565275B74C1D2B6634B475F20A10105" type="personal">
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<mods:namePart id="0EFDAAED20638605559429E3CBE5981E">Lackner, Tomáš</mods:namePart>
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<mods:affiliation id="8F9DBEA2A4F6B205EC5153829BB38553">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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</mods:name>
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<mods:namePart id="B500C2D9916E39F5F70D1A1A8AFDC302">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="3A49AA99D7099C7948817FEBA26CCA2E">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan & Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="2E2F5106E2F88435D4A7510FCD8F8E8E">Prokop, Jakub</mods:namePart>
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<mods:affiliation id="F10E069D8745B1FAC08238B45868DF35">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic & Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="8034F0846BC6102C5C47575034A635C2">Caterino, Michael S</mods:namePart>
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<mods:title id="4920E56FF3874D61B2438753638C589F">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="656CB5741FC3148E6D903A95925D6162">
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<mods:date id="D858EA9FEC15ACF4715E9FC236442B72">2024</mods:date>
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<mods:title id="081A5EA2DAEF69DBD573ADF6009EFA3B">zlae 137</mods:title>
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<mods:number id="CE72ABCF7FADF3267205F8E4921E011E">2024-11-20</mods:number>
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<mods:detail id="35A332D07388958A231A6E6533611D82" type="volume">
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<mods:number id="181399C013B135F311D775E11D5ECE7D">202</mods:number>
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<mods:detail id="F1525CF378FCF6188EE7BD5C0CF18D50" type="issue">
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<mods:number id="17C8E7C6E743ABE62B40704712BCCE30">3</mods:number>
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<mods:start id="7ED8B619B94B397449C1B71C3BE84DC8">1</mods:start>
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<mods:end id="BC8E588BA4C15C4B42792EEA35E6362C">28</mods:end>
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<mods:url id="3EBF910B6B2E0B8CA9A46A9A2C47FE3E">https://doi.org/10.1093/zoolinnean/zlae137</mods:url>
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<mods:classification id="FA423EEF6E884FF6642C1BF8C29B7CEB">journal article</mods:classification>
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<mods:identifier id="17A782C1D765EA3EB4D0FC63CA9B39FE" type="DOI">10.1093/zoolinnean/zlae137</mods:identifier>
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<mods:identifier id="8E6DA2C3A4027A344C0882F5E8B63A24" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="E6B374CF5EFB6C39A142912D2E585AE4" type="Zenodo-Dep">14516114</mods:identifier>
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<mods:identifier id="A831CDDB4B0758E62D1EABDA67C4CFB0" type="ZooBank">9059AA2-5086-46AD-85C6-DBDA56CA72E0</mods:identifier>
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<treatment id="038C87BE170781674C9CFE52FE227CBF" ID-DOI="http://doi.org/10.5281/zenodo.14516148" ID-Zenodo-Dep="14516148" LSID="urn:lsid:plazi:treatment:038C87BE170781674C9CFE52FE227CBF" httpUri="http://treatment.plazi.org/id/038C87BE170781674C9CFE52FE227CBF" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Onthophilinae">
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<subSubSection id="C33F6523170781794C9CFE52FAB97902" box="[964,1305,498,525]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8B9A36A8170781794C9CFE52FAB97902" blockId="15.[964,1305,498,525]" box="[964,1305,498,525]" pageId="15" pageNumber="16">
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<heading id="D0D281C4170781794C9CFE52FAB97902" bold="true" box="[964,1305,498,525]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
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<emphasis id="B951EABA170781794C9CFE52FAB97902" bold="true" box="[964,1305,498,525]" pageId="15" pageNumber="16">
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<taxonomicName id="4C254D2B170781794C9CFE52FB1D7902" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[964,1213,498,525]" class="Insecta" family="Histeridae" genus="Prodigister" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
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<emphasis id="B951EABA170781794C9CFE52FB1D7902" bold="true" box="[964,1213,498,525]" italics="true" pageId="15" pageNumber="16">Prodigister tricostatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26257C1170781794B9AFE52FAB97902" box="[1218,1305,498,524]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33F6523170781794B18FD81FB3D7935" box="[1088,1181,545,571]" pageId="15" pageNumber="16" type="description">
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<paragraph id="8B9A36A8170781794B18FD81FB3D7935" blockId="15.[1088,1181,545,571]" box="[1088,1181,545,571]" pageId="15" pageNumber="16">
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<emphasis id="B951EABA170781794B18FD81FB3D7935" bold="true" box="[1088,1181,545,571]" pageId="15" pageNumber="16">
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||
(
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||
<figureCitation id="131E2A2D170781794B13FD81FB327935" box="[1099,1170,545,571]" captionStart="Figure 11" captionStartId="16.[129,194,1836,1860]" captionTargetBox="[213,1389,144,1808]" captionTargetId="figure-8@16.[213,1389,144,1808]" captionTargetPageId="16" captionText="Figure 11. Prodigister tricostatus sp. nov.. A, dorsal view.B, lateroventral view.C, lateral view. D, elytra.E, frontal view.F, mouthparts. Abbreviations:C1–C3, costae 1–3; ds, dorsal setae; gal, galea; lab, labrum; man, mandible; mp, maxillary palpus; mt, mesotibia; pt, protibia." figureDoi="http://doi.org/10.5281/zenodo.14516140" httpUri="https://zenodo.org/record/14516140/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
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)
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</emphasis>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33F6523170781794C72FDC2FAF979B7" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="8B9A36A8170781794C72FDC2FAF979B7" blockId="15.[810,1460,610,697]" pageId="15" pageNumber="16">
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<emphasis id="B951EABA170781794C72FDC2FC267974" box="[810,902,610,634]" italics="true" pageId="15" pageNumber="16">Material:</emphasis>
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<materialsCitation id="3B4D3CF5170781794CD5FDC2FAF579B7" collectionCode="HUM" location="Kachin" pageId="15" pageNumber="16" specimenCode="SEHU-0000121220" specimenCount="1" stateProvince="Kachin">
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<collectingRegion id="49E1F84A170781794CD5FDC2FC777974" box="[909,983,610,634]" country="Myanmar" name="Kachin" pageId="15" pageNumber="16">Kachin</collectingRegion>
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amber (
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<emphasis id="B951EABA170781794B69FDC3FB9A7974" box="[1073,1082,611,634]" italics="true" pageId="15" pageNumber="16">c</emphasis>
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. 99 Mya). Amber piece dimensions: 15 × 8 ×
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<quantity id="4CDD9B4D170781794CD2FD21FC667997" box="[906,966,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="2.0">2 mm</quantity>
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. Amber clear with several insect fragments and air bubbles. Deposited at
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<collectionCode id="ED34AE6D170781794B74FD01FBCB79B6" box="[1068,1131,673,696]" pageId="15" pageNumber="16">HUM</collectionCode>
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(
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<specimenCode id="DB839ED3170781794B2FFD01FAEE79B7" box="[1143,1358,673,697]" pageId="15" pageNumber="16">SEHU-0000121220</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6523170781674C72FD7BFD9F7973" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
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<paragraph id="8B9A36A8170781674C72FD7BFD9F7973" blockId="15.[810,1460,730,1976]" lastBlockId="17.[113,763,144,638]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<emphasis id="B951EABA170781794C72FD7BFC0179FC" box="[810,929,731,754]" italics="true" pageId="15" pageNumber="16">Dimensions.</emphasis>
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Head width:
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<quantity id="4CDD9B4D170781794B12FD7AFB3879FF" box="[1098,1176,730,754]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="mm" value="0.4">0.4 mm</quantity>
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; width between anterior pronotal angles:
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<quantity id="4CDD9B4D170781794C86FD59FB99781F" box="[990,1081,761,785]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="16" unit="mm" value="0.42">0.42 mm</quantity>
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; width between posterior pronotal angles:
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<quantity id="4CDD9B4D170781794CDCFCB9FC72783E" box="[900,978,793,817]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="mm" value="0.7">0.7 mm</quantity>
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; pronotal length:
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<quantity id="4CDD9B4D170781794BEAFCB8FB5F783E" box="[1202,1279,792,816]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="mm" value="0.5">0.5 mm</quantity>
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; sutural elytral length:
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<quantity id="4CDD9B4D170781794C24FC98FC777841" box="[892,983,824,848]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="16" unit="mm" value="0.95">0.95 mm</quantity>
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; elytral width (in pair, across widest point):
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<quantity id="4CDD9B4D170781794C72FCF7FC277861" box="[810,903,855,879]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="16" unit="mm" value="0.75">0.75 mm</quantity>
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.—
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<emphasis id="B951EABA170781794CFEFCF7FC467861" bold="true" box="[934,998,855,879]" italics="true" pageId="15" pageNumber="16">Body:</emphasis>
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Elongate, subparallel-sided, almost rectangular in shape. Pronotum irregularly sculptured. Elytra with several very prominent costae. Dorsal cuticle with fine setation. Thorax and abdomen strongly compressed (deformed) laterally.—
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<emphasis id="B951EABA170781794CAEFC74FB9778E2" bold="true" box="[1014,1079,980,1004]" italics="true" pageId="15" pageNumber="16">Head:</emphasis>
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Frons and clypeus with sparse setae. Frontoclypeal suture absent. Frontal stria complete, weakly carinate. Labrum long (width: length ratio is 3: 2), plurisetose and punctuate. Mandibles with several prominent setae on outer margin and a blunt subapical tooth, prostheca well developed. Terminal maxillary palpomere elongate and pointed, as long as palpomeres 2–3 combined. Scape straight and parallel-sided with several setae apically. Pedicel short and bulbous, as long as antennomeres 3–4 combined, wider basally than distally. Antennomere 3 short (about half of antennomere 4 length). Antennomeres 3–8 becoming wider apically. Antennal club round and flattened, setose, without annuli.—
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<emphasis id="B951EABA170781794BAAFA8DFAE77E4B" bold="true" box="[1266,1351,1325,1349]" italics="true" pageId="15" pageNumber="16">Thorax:</emphasis>
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Pronotum with irregular sculpturing, mostly convex medially, with a pair of large depressions posterolaterally and one mediobasally, elevations on either side aligned with longitudinal elytral ridges; pronotal sides margined, sinuate, widening posterad, narrowed anterad; marginal pronotal striae not evident. Basal pronotal margin rounded. Marginal prosternal stria complete. Pronotum sparsely covered with fine, short setae. Scutellar shield not visible. Elytra with fine punctuation and vestiture (setae 5× as long as in pronotum). Dorsal elytral striation not visible. Epipleural stria complete. Outer elytral margin carinate and weakly elevated. Medioanterior part strongly depressed. Sutural area strongly elevated forming a keel that is sharp on anterior two-thirds, thence gradually diminishing posterad; each elytron with strong lateral longitudinal costa, rather sharp in anterior half of elytra, becoming lower and broader apically; elytral apices obscured by cracks and evidently distorted. Elytral epipleural stria complete. Ventral body surface strongly compressed, creating a (probably artificial) keel. Prosternal lobe short and broad, punctuate. The presence and character of antennal cavity cannot be determined due to the ventral deformation. Prosternal keel without apparent striation or punctuation. Mesoventrite strongly compressed ventrally, striation not visible. Metaventrite finely punctuate, lateral metaventral stria visible on anterior third.—
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<emphasis id="B951EABA171981674F9DFF4EFF5C7A0B" bold="true" box="[197,252,238,261]" italics="true" pageId="17" pageNumber="18">Legs:</emphasis>
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Profemur very wide, flattened. Protibia strongly dilated with denticles along its outer margin. Protarsal groove present. Protibial spur not developed. Terminal protarsomere as long as protarsomeres 1–4 combined. Mesofemur flattened, broad basally and becoming narrower apically. Mesotibia approximately half of the mesofemoral length, slightly dilated with thick setae along the outer margin, diminishing in size basally. Terminal mesotarsomere as long as mesotarsomeres 2–4 combined. Metafemur broad. Metatibia narrow with only two to three thick setae apically. Metatarsus identical to mesotarsus.—
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<emphasis id="B951EABA171981674F29FD87FF7E7931" bold="true" box="[113,222,551,575]" italics="true" pageId="17" pageNumber="18">Abdomen:</emphasis>
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First visible abdominal sternite long and wide, finely punctuate. Rest of the abdomen hardly observable due to cracks and contamination. Pygidium weakly costate.
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</paragraph>
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</subSubSection>
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<paragraph id="8B9A36A8171881664FD9F88CFA087C6E" blockId="16.[129,1448,1836,1888]" pageId="16" pageNumber="17">
|
||
<emphasis id="B951EABA171881664FD9F88CFF457C4A" bold="true" box="[129,229,1836,1860]" pageId="16" pageNumber="17">Figure 11.</emphasis>
|
||
<taxonomicName id="4C254D2B171881664FB1F88CFE0B7C4A" authorityName="Jan & ºimon-Pražák & Yamamoto & Lackner & Fikáček & Prokop & Michael & Caterino" authorityYear="2024" box="[233,427,1836,1860]" class="Insecta" family="Histeridae" genus="Prodigister" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tricostatus" status="sp. nov.">
|
||
<emphasis id="B951EABA171881664FB1F88CFE0B7C4A" box="[233,427,1836,1860]" italics="true" pageId="16" pageNumber="17">Prodigister tricostatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26257C1171881664EE8F88CFE517C4D" box="[432,497,1836,1859]" pageId="16" pageNumber="17" rank="species">sp. nov</taxonomicNameLabel>
|
||
.. A, dorsal view. B, lateroventral view. C, lateral view. D, elytra. E, frontal view. F, mouthparts. Abbreviations: C1–C3, costae 1–3; ds, dorsal setae; gal, galea; lab, labrum; man, mandible; mp, maxillary palpus; mt, mesotibia; pt, protibia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33F6523171981674F29FD00FF7F7D2A" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="8B9A36A8171981674F29FD00FF7F7D2A" blockId="17.[113,763,671,1572]" pageId="17" pageNumber="18">
|
||
<emphasis id="B951EABA171981674F29FD00FEFF79B9" box="[113,351,672,695]" italics="true" pageId="17" pageNumber="18">Taxonomic assignment:</emphasis>
|
||
Few characters that would serve as diagnostic at higher levels are clearly visible in this specimen. Furthermore, it is considerably deformed by lateral compression, especially ventrally, and sternal characters are severely altered. The strongly convex pronotum and costate elytra, being reasonably bilaterally symmetrical, show no signs of being artefacts of deformation, and distinguish this taxon from any known histerids, extinct or extant. These are not similar to costae in onthophiline genera, and this genus could not be placed there. The antennal club is not easily visible, but it is sufficiently clear that it lacks distinct annuli, as are seen in a number of other Cretaceous ‘basal grade’ histeroids. The strongly expanded protibiae are a remarkably common feature among Cretaceous taxa, seen in
|
||
<taxonomicName id="4C254D2B171981674E41FB96FEC87F40" authorityName="Simon Prazak & Lackner" authorityYear="2023" box="[281,360,1078,1102]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674E41FB96FEC87F40" box="[281,360,1078,1102]" italics="true" pageId="17" pageNumber="18">Olexum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D2B171981674E27FB96FE687F40" authorityName="Caterino" authorityYear="2021" box="[383,456,1078,1102]" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674E27FB96FE687F40" box="[383,456,1078,1102]" italics="true" pageId="17" pageNumber="18">Yethiha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D2B171981674E87FB97FD9A7F40" authorityName="Caterino" authorityYear="2021" box="[479,570,1079,1102]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674E87FB97FD9A7F40" box="[479,570,1079,1102]" italics="true" pageId="17" pageNumber="18">Druantia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D2B171981674D09FB96FD1C7F40" authorityName="Simon Prazak & Lackner" authorityYear="2023" box="[593,700,1078,1102]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674D09FB96FD1C7F40" box="[593,700,1078,1102]" italics="true" pageId="17" pageNumber="18">Platycretus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C254D2B171981674F29FBF6FF567F63" box="[113,246,1110,1133]" class="Insecta" family="Histeridae" genus="Promyrmister" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674F29FBF6FF567F63" box="[113,246,1110,1133]" italics="true" pageId="17" pageNumber="18">Promyrmister</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB44B59171981674E5EFBF6FE0C7F60" author="Zhou YL & Slipinski A & Ren D" box="[262,428,1110,1134]" pageId="17" pageNumber="18" refId="ref19248" refString="Zhou YL, Slipinski A, Ren D et al. A Mesozoic clown beetle myrmecophile (Coleoptera: Histeridae). eLife 2019; 8: e 44985. hups: // doi. org / 10.7554 / eLife. 44985" type="journal volume" year="2019">
|
||
Zhou
|
||
<emphasis id="B951EABA171981674E1CFBF6FED37F60" box="[324,371,1110,1134]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB44B59171981674EEFFBF6FDEA7F60" author="Caterino" box="[439,586,1110,1134]" pageId="17" pageNumber="18" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino 2021</bibRefCitation>
|
||
, Simon-Pražák
|
||
<emphasis id="B951EABA171981674DB1FBF6FF2A7F83" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
2023b). Of these, the species bears some characters linking it to
|
||
<taxonomicName id="4C254D2B171981674FFFFB34FEB27FA2" authorityName="Simon Prazak & Lackner" authorityYear="2023" box="[167,274,1172,1196]" class="Insecta" family="Histeridae" genus="Platycretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674FFFFB34FEB27FA2" box="[167,274,1172,1196]" italics="true" pageId="17" pageNumber="18">Platycretus</emphasis>
|
||
</taxonomicName>
|
||
: very wide and flat profemur, and mesofemur longer than mesotibia (Simon-Pražák
|
||
<emphasis id="B951EABA171981674D5BFB14FD967FC2" box="[515,566,1204,1228]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
2023b). However, the plurisetose labrum, as well as fused antennal club, would suggest a closer relationship to
|
||
<taxonomicName id="4C254D2B171981674EEFFB52FDF97E04" authorityName="Reitter" authorityYear="1909" box="[439,601,1266,1290]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Dendrophilinae">Dendrophilinae</taxonomicName>
|
||
taxa—
|
||
<taxonomicName id="4C254D2B171981674DFEFB52FD557E04" authorityName="Simon Prazak & Lackner" authorityYear="2023" box="[678,757,1266,1290]" class="Insecta" family="Histeridae" genus="Olexum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674DFEFB52FD557E04" box="[678,757,1266,1290]" italics="true" pageId="17" pageNumber="18">Olexum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C254D2B171981674F29FAB2FF1A7E24" authorityName="Caterino" authorityYear="2021" box="[113,186,1298,1322]" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674F29FAB2FF1A7E24" box="[113,186,1298,1322]" italics="true" pageId="17" pageNumber="18">Yethiha</emphasis>
|
||
</taxonomicName>
|
||
, or
|
||
<taxonomicName id="4C254D2B171981674F87FAB2FE9A7E27" authorityName="Caterino" authorityYear="2021" box="[223,314,1298,1321]" class="Insecta" family="Histeridae" genus="Druantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B951EABA171981674F87FAB2FE9A7E27" box="[223,314,1298,1321]" italics="true" pageId="17" pageNumber="18">Druantia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB44B59171981674E11FAB2FE7C7E24" author="Caterino" box="[329,476,1298,1322]" pageId="17" pageNumber="18" pagination="211 - 21" refId="ref17393" refString="Caterino Mº. New fossil histerid species from Cretaceous Burmese amber (Coleoptera: Histeridae). Ne Coleopterists Bulletin 2021; 75: 211 - 21." type="journal article" year="2021">Caterino 2021</bibRefCitation>
|
||
, Simon-Pražák
|
||
<emphasis id="B951EABA171981674D20FAB2FD067E24" box="[632,678,1298,1322]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
2023b). The position of this genus could not be sufficiently resolved via phylogenetic analyses either, as it nested in various positions in
|
||
<taxonomicName id="4C254D2B171981674F29FAD0FF7C7E86" box="[113,220,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
outside the
|
||
<taxonomicName id="4C254D2B171981674E38FAD0FE4F7E86" authorityName="Zhou, Yu-Lingxi, Caterino, Dong Ren & Ślipiński" authorityYear="2020" box="[352,495,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Antigracilinae">Antigracilinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C254D2B171981674D7DFAD0FD2D7E86" authorityName="W. S. MacLeay" authorityYear="1819" box="[549,653,1392,1416]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
|
||
clades. As some of the key characters for higher taxonomic assignment are not visible due to deformation, we leave this genus unplaced to a higher taxon for now, and hope that better preserved specimens materialize to reveal more phylogenetically significant characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523171981674F29F9E6FEC47C56" pageId="17" pageNumber="18" type="diagnosis">
|
||
<paragraph id="8B9A36A8171981674F29F9E6FEC47C56" blockId="17.[113,762,1606,1880]" pageId="17" pageNumber="18">
|
||
<emphasis id="B951EABA171981674F29F9E6FF777D53" box="[113,215,1606,1629]" italics="true" pageId="17" pageNumber="18">Diagnosis:</emphasis>
|
||
Prolonged, parallel-sided body shape with several elytral dorsal costae. Dorsal cuticle setose. Labrum long and multisetose. Head without a frontoclypeal suture, frontal stria complete. Antennal club fused, without annuli. Basal pronotal margin widely rounded. Wide and flat profemur. Protibia strongly dilated with denticles. Mesotibia narrow and spinose, metatibia narrow and almost smooth. The species is unlike any known recent or extinct
|
||
<taxonomicName id="4C254D2B171981674E26F881FE497C37" box="[382,489,1825,1849]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Histeridae</taxonomicName>
|
||
species and is easily, unequivocally identifiable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33F6523171981674F29F8DAFE227CBF" pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph id="8B9A36A8171981674F29F8DAFE227CBF" blockId="17.[113,762,1914,1969]" pageId="17" pageNumber="18">
|
||
<emphasis id="B951EABA171981674F29F8DAFF417C9C" box="[113,225,1914,1938]" italics="true" pageId="17" pageNumber="18">Etymology:</emphasis>
|
||
The species name ‘tricostatus’ refers to the three dorsal costae on the elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |