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<document id="5B60106190AC0DBAD54336BD53FACACE" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub &amp; Caterino, Michael S" docDate="2024" docId="038C87BE170481794C8AFDC6FD5B7BA9" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Pseudacritus extinctus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop &amp; Caterino, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="13" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:title id="B9E3500E5FAFBAB2002311A7282BCB93">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="7E3AE694591127F1A8E1AA8A41B1ABA8">Simon-Pražák, Jan</mods:namePart>
<mods:affiliation id="542CFB6D870785EA2C5DCDA7B7968871">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="A40EB476E7AA3AB5B9AF8FE304DCF2CA" type="email">j.prazak@muzeumhk.cz</mods:nameIdentifier>
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<mods:namePart id="209850F15D698452CDDD21A2C426B923">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="5657D93EC92F95B9EF99D2D8380FAC3B">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="2F1847303176A0F7A161055553D4A3DF">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="F701F40D1E6F595B5BBB5FB2A82AD735">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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<mods:namePart id="07F742936297FE9D015CA6D1D55DA2FA">Fikáček, Martin</mods:namePart>
<mods:affiliation id="B7451EA61F9755EA8801C031F9725352">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="36986F5335C87790F6775B80C7810923">Prokop, Jakub</mods:namePart>
<mods:affiliation id="C83B05000F2EDAD6BF5CEEC041F92CFF">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="99B4A7BE9CEB46351CFE9C5818592C5B">Caterino, Michael S</mods:namePart>
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<mods:date id="C32318ACB7A3AE5AF13D28B5635780DD">2024</mods:date>
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<mods:title id="5ED9FDC1933549F31BB5E884F866D2B8">zlae 137</mods:title>
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<mods:number id="33684DCD99B9DF11EF4CBBA8BD80AF7E">2024-11-20</mods:number>
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<mods:number id="3CA9CC6CEA5C675BF570EE5692815AF2">202</mods:number>
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<treatment id="038C87BE170481794C8AFDC6FD5B7BA9" ID-DOI="http://doi.org/10.5281/zenodo.14516134" ID-Zenodo-Dep="14516134" LSID="urn:lsid:plazi:treatment:038C87BE170481794C8AFDC6FD5B7BA9" httpUri="http://treatment.plazi.org/id/038C87BE170481794C8AFDC6FD5B7BA9" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Abraeinae">
<subSubSection id="C33F65231704817A4C8AFDC6FA8A798F" box="[978,1322,614,641]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B9A36A81704817A4C8AFDC6FA8A798F" blockId="12.[978,1322,614,641]" box="[978,1322,614,641]" pageId="12" pageNumber="13">
<heading id="D0D281C41704817A4C8AFDC6FA8A798F" bold="true" box="[978,1322,614,641]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<emphasis id="B951EABA1704817A4C8AFDC6FA8A798F" bold="true" box="[978,1322,614,641]" pageId="12" pageNumber="13">
<taxonomicName id="4C254D2B1704817A4C8AFDC6FB6D798E" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[978,1229,614,641]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="extinctus" status="sp. nov.">
<emphasis id="B951EABA1704817A4C8AFDC6FB6D798E" bold="true" box="[978,1229,614,641]" italics="true" pageId="12" pageNumber="13">Pseudacritus extinctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11704817A4B8AFDC7FA8A798F" box="[1234,1322,615,641]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F65231704817A4B0FFD35FB0579A1" box="[1111,1189,661,688]" pageId="12" pageNumber="13" type="description">
<paragraph id="8B9A36A81704817A4B0FFD35FB0579A1" blockId="12.[1111,1189,661,688]" box="[1111,1189,661,688]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4B0FFD35FB0579A1" bold="true" box="[1111,1189,661,688]" pageId="12" pageNumber="13">
(
<figureCitation id="131E2A2D1704817A4B39FD36FB3A79BE" box="[1121,1178,662,688]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." figureDoi="http://doi.org/10.5281/zenodo.14516136" httpUri="https://zenodo.org/record/14516136/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33F65231704817A4C61FD76FB467842" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8B9A36A81704817A4C61FD76FB467842" blockId="12.[825,1475,726,844]" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4C61FD76FC3579E0" box="[825,917,726,750]" italics="true" pageId="12" pageNumber="13">Material:</emphasis>
<materialsCitation id="3B4D3CF51704817A4CC4FD76FB417842" collectionCode="MEBHK" location="Kachin" pageId="12" pageNumber="13" specimenCode="P1423/E" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A1704817A4CC4FD76FC4679E0" box="[924,998,726,750]" country="Myanmar" name="Kachin" pageId="12" pageNumber="13">Kachin</collectingRegion>
amber (
<emphasis id="B951EABA1704817A4B18FD77FBE979E0" box="[1088,1097,727,750]" italics="true" pageId="12" pageNumber="13">c</emphasis>
. 99 Mya). Amber piece dimensions: 16 × 5 ×
<quantity id="4CDD9B4D1704817A4CC2FD56FC757803" box="[922,981,758,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="2.0">2 mm</quantity>
. Amber clear with a large
<taxonomicName id="4C254D2B1704817A4BA2FD56FA297800" box="[1274,1417,758,782]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="order">Hymenoptera</taxonomicName>
fragment, a single specimen of
<taxonomicName id="4C254D2B1704817A4B3CFCB5FB4E7823" authorityName="Latreille" authorityYear="1802" box="[1124,1262,789,813]" class="Insecta" family="Trogossitidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
and large pieces of debris. Deposited at
<collectionCode id="ED34AE6D1704817A4B52FC95FBC67842" box="[1034,1126,821,844]" pageId="12" pageNumber="13">MEBHK</collectionCode>
(
<specimenCode id="DB839ED31704817A4B2CFC94FB797842" box="[1140,1241,820,844]" pageId="12" pageNumber="13">P1423/E</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F65231704817B4C61FCCEFE667DDD" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8B9A36A81704817B4C61FCCEFE667DDD" blockId="12.[825,1475,878,1967]" lastBlockId="13.[113,762,1629,1747]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B951EABA1704817A4C61FCCEFC6C788B" box="[825,972,878,901]" italics="true" pageId="12" pageNumber="13">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D1704817A4B30FCCEFB15788B" box="[1128,1205,878,902]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="mm" value="0.3">0.3 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D1704817A4CB5FC2DFBE878AA" box="[1005,1096,909,933]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="13" unit="mm" value="0.34">0.34 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D1704817A4CC9FC0CFC4C78CA" box="[913,1004,940,964]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="12" pageNumber="13" unit="mm" value="0.68">0.68 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D1704817A4B9DFC0CFAB378CA" box="[1221,1299,940,964]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="mm" value="0.4">0.4 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D1704817A4CD3FC6CFC4678ED" box="[907,998,972,996]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="13" unit="mm" value="0.48">0.48 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D1704817A4C61FC4BFC297F0C" box="[825,905,1003,1027]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="mm" value="0.7">0.7 mm</quantity>
.—
<emphasis id="B951EABA1704817A4CF0FC4BFC487F0D" bold="true" box="[936,1000,1003,1027]" italics="true" pageId="12" pageNumber="13">Body:</emphasis>
Round and dorsoventrally convex, brown in colour.—
<emphasis id="B951EABA1704817A4CC3FBAAFC7C7F2C" bold="true" box="[923,988,1034,1058]" italics="true" pageId="12" pageNumber="13">Head:</emphasis>
Without frontal stria. Frons smooth, slightly convex. Clypeus basally constricted, widening anteriorly. Labial as well as maxillary palpi exposed, the latter rather short, wide. Labrum with a transverse carinate median ridge, with a pair of prominent setae and several shorter setae. Terminal maxillary palpomere oval and pointed. Terminal labial palpomere short, pointed. Eyes large, convex. Antennal scape thickened, as long as antennomeres 27 combined. Pedicel approximately third scapes length, wider basally. Club large, oval, flattened, with two distinct annuli.—
<emphasis id="B951EABA1704817A4B7EFA84FBDA7E32" bold="true" box="[1062,1146,1316,1340]" italics="true" pageId="12" pageNumber="13">Thorax:</emphasis>
Pronotal disc almost smooth, with widely isolated small punctures and with a complete, punctuate antescutellar stria. Anterior corners acute. Marginal stria present. Lateral pronotal stria carinate, slightly expanding over pronotal margin medially and gradually distanced from pronotal margin basally. Scutellar shield small, triangular. Elytra dorsally with four weak costae. Elytral epipleuron concave, epipleural stria obscured. Elytral humeri with four short distinct costate striae. Prosternal keel short and wide, largely unobservable, laterally incised for the antennal passage. Round and relatively well-developed antennal cavities located at the anteriormost part of the prosternal hypomeron. Hypomeron excavate, creating a depression shaped to receive the profemur and protibia. When retracted the protibial would probably conceal the antennal club. Mesoventrite obscured by legs and fracturing. Metaventrite very finely punctuated, with a median depression, which may be an artefact. Postmesocoxal stria complete, recurved along mesocoxa. Lateral metaventral stria complete, continuous on to metepisternum. Metepisternum exposed.—
<emphasis id="B951EABA1704817A4A4FF8F8FAEE7C61" bold="true" box="[1303,1358,1880,1903]" italics="true" pageId="12" pageNumber="13">Legs:</emphasis>
All femora and tibiae thin, laterally flattened. Tarsal formula 5-5-5, ultimate tarsomeres as long as the preceding two.—
<emphasis id="B951EABA1704817A4A41F836FA267CA0" bold="true" box="[1305,1414,1942,1966]" italics="true" pageId="12" pageNumber="13">Abdomen:</emphasis>
First abdominal ventrite wide, with thin and complete postmetacoxal striae. Pygidium triangular, laterobasally with depressions, apex with a single short groove medially. Propygidium short and broad, with irregular depressions.
</paragraph>
</subSubSection>
<caption id="DF5A66201705817B4F28FA7DFF3D7D23" ID-DOI="http://doi.org/10.5281/zenodo.14516136" ID-Zenodo-Dep="14516136" httpUri="https://zenodo.org/record/14516136/files/figure.png" pageId="13" pageNumber="14" startId="13.[112,177,1501,1525]" targetBox="[114,1458,144,1473]" targetPageId="13" targetType="figure">
<paragraph id="8B9A36A81705817B4F28FA7DFF3D7D23" blockId="13.[112,1457,1501,1581]" pageId="13" pageNumber="14">
<emphasis id="B951EABA1705817B4F28FA7DFF687EFB" bold="true" box="[112,200,1501,1525]" pageId="13" pageNumber="14">Figure 9.</emphasis>
<taxonomicName id="4C254D2B1705817B4F95FA7DFE337EFB" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[205,403,1501,1525]" class="Insecta" family="Histeridae" genus="Pseudacritus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="extinctus" status="gen. et sp. nov">
<emphasis id="B951EABA1705817B4F95FA7DFE337EFB" box="[205,403,1501,1525]" italics="true" pageId="13" pageNumber="14">Pseudacritus extinctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11705817B4EC0FA7EFDBB7EFB" box="[408,539,1502,1526]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov</taxonomicNameLabel>
.. A, dorsal view. B, ventral view. C, lateroventral view. D, antenna. E, frontal view. F, pronotal base. Abbreviations: aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc.
</paragraph>
</caption>
<subSubSection id="C33F65231705817B4F29F955FAB27C5E" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8B9A36A81705817B4F29F955FAB27C5E" blockId="13.[113,763,1781,1961]" lastBlockId="13.[810,1460,1629,1872]" pageId="13" pageNumber="14">
<emphasis id="B951EABA1705817B4F29F955FEF17C02" box="[113,337,1781,1804]" italics="true" pageId="13" pageNumber="14">Taxonomic assignment:</emphasis>
This species can be tentatively placed in the
<taxonomicName id="4C254D2B1705817B4F29F8B4FF777C22" authorityName="W. S. MacLeay" authorityYear="1819" box="[113,215,1812,1836]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
subfamily, based on the absence of a distinct pronotal lobe, small body size of round shape, and strongly convex eyes (
<figureCitation id="131E2A2D1705817B4F24F8F3FF667C65" box="[124,198,1875,1899]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." figureDoi="http://doi.org/10.5281/zenodo.14516136" httpUri="https://zenodo.org/record/14516136/files/figure.png" pageId="13" pageNumber="14">Fig. 9A</figureCitation>
). This position was also supported by all phylogenetic analyses. The species bears characters strikingly similar to the extant
<taxonomicName id="4C254D2B1705817B4FEFF831FEA97CA7" authorityName="Wenzel" authorityYear="1944" box="[183,265,1937,1961]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
: position of the antennal cavity at the frontmost part of prosternal hypomeron (
<figureCitation id="131E2A2D1705817B4B34F9FDFB177D7B" box="[1132,1207,1629,1653]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." figureDoi="http://doi.org/10.5281/zenodo.14516136" httpUri="https://zenodo.org/record/14516136/files/figure.png" pageId="13" pageNumber="14">Fig. 9C</figureCitation>
) and the presence of an antescutellar stria on the pronotal disc (
<figureCitation id="131E2A2D1705817B4B95F9DDFA9A7D9A" box="[1229,1338,1661,1685]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." figureDoi="http://doi.org/10.5281/zenodo.14516136" httpUri="https://zenodo.org/record/14516136/files/figure.png" pageId="13" pageNumber="14">Fig. 9A, D</figureCitation>
). However, it lacks the key defining feature of
<taxonomicName id="4C254D2B1705817B4BC9F93CFB407DBA" authorityName="Wenzel" authorityYear="1944" box="[1169,1248,1692,1716]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritini">Acritini</taxonomicName>
, the four-segmented metatarsi. Moreover, the species exhibits a plesiomorphic annulate antennal club (
<figureCitation id="131E2A2D1705817B4CBEF97BFB8E7DFD" box="[998,1070,1755,1779]" captionStart="Figure 9" captionStartId="13.[112,177,1501,1525]" captionTargetBox="[114,1458,144,1473]" captionTargetId="figure-201@13.[114,1458,144,1473]" captionTargetPageId="13" captionText="Figure 9. Pseudacritus extinctus gen.et sp. nov.. A, dorsal view.B, ventral view. C, lateroventral view.D, antenna. E, frontal view. F, pronotal base. Abbreviations:aca, antennal cavity; acl, antennal club; as, antennal scape; ass, antescutellar stria; lab, labrum; mp, maxillary palpus; sd, scutellar disc." figureDoi="http://doi.org/10.5281/zenodo.14516136" httpUri="https://zenodo.org/record/14516136/files/figure.png" pageId="13" pageNumber="14">Fig. 9C</figureCitation>
), as opposed to most recent
<taxonomicName id="4C254D2B1705817B4A1FF97BFA0F7DFD" authorityName="W. S. MacLeay" authorityYear="1819" box="[1351,1455,1755,1779]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Abraeinae">Abraeinae</taxonomicName>
, except
<taxonomicName id="4C254D2B1705817B4C20F95AFBB87C1C" authorityName="Wenzel" authorityYear="1944" box="[888,1048,1786,1810]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
(
<bibRefCitation id="EFB44B591705817B4B77F95AFB1A7C1C" author="Wenzel RL" box="[1071,1210,1786,1810]" pageId="13" pageNumber="14" pagination="51 - 151" refId="ref19054" refString="Wenzel RL. On the classification of the histerid beetles. Fieldiana Zoology 1944; 28: 51 - 151. hups: // doi. org / 10.1093 / aesa / 37.3.375" type="journal article" year="1944">Wenzel 1944</bibRefCitation>
,
<bibRefCitation id="EFB44B591705817B4B94F95AFAEE7C1C" author="Mazur" box="[1228,1358,1786,1810]" pageId="13" pageNumber="14" pagination="27 - 32" refId="ref18506" refString="Mazur º. New species of histerid beetles (Coleoptera, Histeridae). Annales Zoologici 1997; 38: 27 - 32. hup: // rcin. org. pl" type="journal article" year="1997">Mazur 1997</bibRefCitation>
). Recent
<taxonomicName id="4C254D2B1705817B4C72F8B9FC6C7C3F" authorityName="Wenzel" authorityYear="1944" box="[810,972,1817,1841]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Acritomorphini">Acritomorphini</taxonomicName>
, however, have an oblong, elongated body shape. Hence, the placement within a tribe is uncertain.
</paragraph>
</subSubSection>
<subSubSection id="C33F6523170581784C72F8D3FA9A7CBA" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8B9A36A8170581784C72F8D3FA9A7CBA" blockId="13.[810,1459,1906,1961]" lastBlockId="14.[825,1475,1885,1972]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B951EABA1705817B4C72F8D3FC2A7C84" box="[810,906,1907,1930]" italics="true" pageId="13" pageNumber="14">Diagnosis:</emphasis>
Pronotum with antescutellar stria and small widely separated punctures.Elytra with four low costae and without an epipleural stria. Postmesocoxal stria complete, recurved along mesocoxa. Lateral metaventral stria complete, continuous onto metepisternum. The species can be distinguished from
<taxonomicName id="4C254D2B170681784EB6F83CFD0D7CBA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[494,685,1948,1972]" class="Insecta" family="Histeridae" genus="Palaeabraeus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="glabrus">
<emphasis id="B951EABA170681784EB6F83CFD0D7CBA" box="[494,685,1948,1972]" italics="true" pageId="14" pageNumber="15">Palaeabraeus glabrus</emphasis>
</taxonomicName>
(the only similar abraeine reported from Burmese ambers) by the presence of antescutellar stria, the anterior position of antennal cavities, presence of elytral costae, and absence of elytral epipleural stria.
</paragraph>
</subSubSection>
<caption id="DF5A6620170681784FD9F88BFC007C4D" ID-DOI="http://doi.org/10.5281/zenodo.14516138" ID-Zenodo-Dep="14516138" box="[129,928,1835,1859]" httpUri="https://zenodo.org/record/14516138/files/figure.png" pageId="14" pageNumber="15" startId="14.[129,194,1835,1859]" targetBox="[141,1461,143,1807]" targetPageId="14" targetType="figure">
<paragraph id="8B9A36A8170681784FD9F88BFC007C4D" blockId="14.[129,928,1835,1859]" box="[129,928,1835,1859]" pageId="14" pageNumber="15">
<emphasis id="B951EABA170681784FD9F88BFF457C4D" bold="true" box="[129,229,1835,1859]" pageId="14" pageNumber="15">Figure 10.</emphasis>
<taxonomicName id="4C254D2B170681784FB1F88BFE3F7C4D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[233,415,1835,1859]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="planatus" status="sp. nov.">
<emphasis id="B951EABA170681784FB1F88BFE3F7C4D" box="[233,415,1835,1859]" italics="true" pageId="14" pageNumber="15">Phasmister planatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170681784EFCF88BFE4A7C4C" box="[420,490,1835,1858]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(AC) and
<taxonomicName id="4C254D2B170681784D04F88BFCB07C4D" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[604,784,1835,1859]" class="Insecta" family="Histeridae" genus="Phasmister" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kraliceki" status="sp. nov.">
<emphasis id="B951EABA170681784D04F88BFCB07C4D" box="[604,784,1835,1859]" italics="true" pageId="14" pageNumber="15">Phasmister kraliceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C1170681784C4CF88BFCFB7C4C" box="[788,859,1835,1858]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(D, E).
</paragraph>
</caption>
<subSubSection id="C33F6523170781794F29FF30FD5B7BA9" box="[113,763,144,168]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8B9A36A8170781794F29FF30FD5B7BA9" blockId="15.[113,763,144,168]" box="[113,763,144,168]" pageId="15" pageNumber="16">
<emphasis id="B951EABA170781794F29FF30FF417BA6" box="[113,225,144,168]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
The specific name signifies the species extinct status.
</paragraph>
</subSubSection>
</treatment>
</document>