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<document id="95C841EBFB4D7A049C672E8E0F953D00" ID-CLB-Dataset="306634" ID-DOI="10.1093/zoolinnean/zlae137" ID-GBIF-Dataset="b9ccfb40-9194-4228-abd2-acaf275f5749" ID-ISSN="0024-4082" ID-Zenodo-Dep="14516114" ID-ZooBank="9059AA2-5086-46AD-85C6-DBDA56CA72E0" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033959621" checkinUser="plazi" docAuthor="Simon-Pražák, Jan, Yamamoto, Shûhei, Lackner, Tomáš, Fikáček, Martin, Prokop, Jakub &amp; Caterino, Michael S" docDate="2024" docId="038C87BE1701817F4E4AFE09FC6078E5" docLanguage="en" docName="zlae137.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 137) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae137" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Pantostictus hirsutus Simon-Pražák, Yamamoto, Lackner, Fikáček, Prokop &amp; Caterino, 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="10" masterDocId="FFB5FFC6170881764F58FFA0FFA07B0E" masterDocTitle="A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="10" updateTime="1734537818702" updateUser="ExternalLinkService">
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<mods:title id="CD8CED9551DA8B085632049E445F8830">A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="35490DD0D1AB74915CDB479908F84829">Simon-Pražák, Jan</mods:namePart>
<mods:affiliation id="D72C7611E5A8DC12F545E3BE1113EBBD">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Králové, Eliščino nábřeží 465, 500 03 Hradec Králové 3, Czech Republic</mods:affiliation>
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<mods:namePart id="2C5798637597A6BD04AAA36585E5AD6B">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="DEEEE661148A84EA29293C08D742EDA7">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="100E26EBFF4C3914561D3B0A83E16136">Lackner, Tomáš</mods:namePart>
<mods:affiliation id="4B27420F62577D94A4EC8208644C06BC">Department of Environmental Systems Science, ETH Zürich, Weinbergstrasse 56, 8092 Zürich, Switzerland</mods:affiliation>
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<mods:namePart id="4AF41DA378D6C41134FA2B80BF7CE4FB">Fikáček, Martin</mods:namePart>
<mods:affiliation id="E29447407F3ED3A46516D3B296EE1F35">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusová 1740 CZ- 19300 Praha 9, Czech Republic</mods:affiliation>
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<mods:namePart id="41253260F82A6A42ABB006AC03FCEB1E">Prokop, Jakub</mods:namePart>
<mods:affiliation id="45C0E4DD3F004BD6CD720D3CD10759E2">Charles University, Faculty of Science, Department of Zoology, Viničná 7, 12800 Prague 2, Czech Republic &amp; Department of Plant and Environmental Sciences, Clemson University, Clemson, SC 29634, United States</mods:affiliation>
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<mods:namePart id="5285F80F605E0F2B896FADB0CC8DBC6A">Caterino, Michael S</mods:namePart>
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<mods:date id="A64AA9681221481172CF1A1A1382048F">2024</mods:date>
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<mods:number id="3633DBBD5565F1B2C5E8564D9A8BB5B7">2024-11-20</mods:number>
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<treatment id="038C87BE1701817F4E4AFE09FC6078E5" ID-DOI="http://doi.org/10.5281/zenodo.14424550" ID-Zenodo-Dep="14424550" LSID="urn:lsid:plazi:treatment:038C87BE1701817F4E4AFE09FC6078E5" httpUri="http://treatment.plazi.org/id/038C87BE1701817F4E4AFE09FC6078E5" lastPageNumber="10" pageId="9" pageNumber="10" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Abraeinae" scope_tribe="Pantostictini">
<subSubSection id="C33F65231701817F4E4AFE09FDF97ACA" box="[274,601,425,452]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9A36A81701817F4E4AFE09FDF97ACA" blockId="9.[274,601,425,452]" box="[274,601,425,452]" pageId="9" pageNumber="10">
<heading id="D0D281C41701817F4E4AFE09FDF97ACA" bold="true" box="[274,601,425,452]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B951EABA1701817F4E4AFE09FDF97ACA" bold="true" box="[274,601,425,452]" pageId="9" pageNumber="10">
<taxonomicName id="4C254D2B1701817F4E4AFE09FE5D7ACA" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[274,509,425,452]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
<emphasis id="B951EABA1701817F4E4AFE09FE5D7ACA" bold="true" box="[274,509,425,452]" italics="true" pageId="9" pageNumber="10">Pantostictus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257C11701817F4D5AFE0AFDF97ACA" box="[514,601,426,452]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C33F65231701817F4ED7FE7BFC6078E5" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9A36A81701817F4ED7FE7BFE7D7AFB" blockId="9.[399,477,475,502]" box="[399,477,475,502]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4ED7FE7BFE7D7AFB" bold="true" box="[399,477,475,502]" pageId="9" pageNumber="10">
(
<figureCitation id="131E2A2D1701817F4EC1FE7BFE727AF8" box="[409,466,475,502]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516130" httpUri="https://zenodo.org/record/14516130/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A36A81701817F4F29FDBCFDCD797D" blockId="9.[113,763,540,627]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29FDBCFF6D793A" box="[113,205,540,564]" italics="true" pageId="9" pageNumber="10">Material:</emphasis>
<materialsCitation id="3B4D3CF51701817F4F8BFDBCFDC9797D" collectionCode="HUM" location="Kachin" pageId="9" pageNumber="10" specimenCode="SEHU-121212" specimenCount="1" stateProvince="Kachin">
<collectingRegion id="49E1F84A1701817F4F8BFDBCFEBD793A" box="[211,285,540,564]" country="Myanmar" name="Kachin" pageId="9" pageNumber="10">Kachin</collectingRegion>
amber (c
<emphasis id="B951EABA1701817F4E26FDBDFE27793A" box="[382,391,541,564]" italics="true" pageId="9" pageNumber="10">c</emphasis>
. 99 Mya). Amber piece dimensions: 12 × 7 ×
<quantity id="4CDD9B4D1701817F4F8AFD9BFEAD795D" box="[210,269,571,595]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1.0">1 mm</quantity>
. Amber clear with several insect fragments and air bubbles. Deposited at
<collectionCode id="ED34AE6D1701817F4E2BFDFBFE12797C" box="[371,434,603,626]" pageId="9" pageNumber="10">HUM</collectionCode>
(
<specimenCode id="DB839ED31701817F4EE7FDFBFDC1797D" box="[447,609,603,627]" pageId="9" pageNumber="10">SEHU-121212</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A36A81701817F4F29FD35FD127D98" blockId="9.[113,763,660,1686]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29FD35FEA479A2" box="[113,260,661,684]" italics="true" pageId="9" pageNumber="10">Measurements:</emphasis>
Head width:
<quantity id="4CDD9B4D1701817F4EC1FD34FE5479A5" box="[409,500,660,684]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="0.25">0.25 mm</quantity>
; width between anterior pronotal angles:
<quantity id="4CDD9B4D1701817F4E7DFD13FE2079C5" box="[293,384,691,715]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="0.35">0.35 mm</quantity>
; width between posterior pronotal angles:
<quantity id="4CDD9B4D1701817F4F9FFD72FE8279E4" box="[199,290,722,746]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="10" unit="mm" value="0.55">0.55 mm</quantity>
; pronotal length:
<quantity id="4CDD9B4D1701817F4EADFD72FDEF79E4" box="[501,591,722,746]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="0.35">0.35 mm</quantity>
; sutural elytral length:
<quantity id="4CDD9B4D1701817F4F9BFD52FEBE7807" box="[195,286,754,778]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="0.35">0.35 mm</quantity>
; elytral width (in pair, across widest point):
<quantity id="4CDD9B4D1701817F4F29FCB1FF6E7827" box="[113,206,785,809]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="mm" value="0.65">0.65 mm</quantity>
.—
<emphasis id="B951EABA1701817F4FB5FCB1FE8D7827" bold="true" box="[237,301,785,809]" italics="true" pageId="9" pageNumber="10">Body:</emphasis>
Oval with a densely and coarsely punctuated cuticle. Dorsal surface with sparse short setation.—
<emphasis id="B951EABA1701817F4D21FC90FD1A7846" bold="true" box="[633,698,816,840]" italics="true" pageId="9" pageNumber="10">Head:</emphasis>
Frons and clypeus flat, coarsely punctuate, without striae. Labrum short and wide, rounded apically, plurisetose. Left mandible appearing bidentate. Antennal club flat and triangular (most likely deformed).—
<emphasis id="B951EABA1701817F4E19FC0EFE3578C8" bold="true" box="[321,405,942,966]" italics="true" pageId="9" pageNumber="10">Thorax:</emphasis>
Pronotal disc coarsely punctuate, broadly rounded basally, anterior angles pointed. Lateral margin finely crenulate. Scutellar shield hidden. Elytra expanded laterally in basal third, surface of elytral disc coarsely punctuate and finely setose, without striae. Epipleura punctate, but not striate. Prosternum short and wide. Prosternal process widely rectangular anteriorly, coarsely punctuated with crenulate margin and lateral fissures for antennal passage. Antennal clubs rest in shallow cavities located in anterior prosternal angles. Prosternal lobe not distinct. Meso- and metaventrite coarsely punctuated. Metaventrite with recurved postcoxal striae.—
<emphasis id="B951EABA1701817F4F29FAA7FF097E10" bold="true" box="[113,169,1287,1310]" italics="true" pageId="9" pageNumber="10">Legs:</emphasis>
Profemur flattened, with a groove for protibial reception. Protibia slightly expanding apically, outer margin rounded in apical half, bearing few short spines; protibial apex with a small tibial spur. Protarsomeres short, terminal as long as 34 combined. Middle leg similar to front leg. Metafemur swollen with setae dorsally, metatibia slightly expanded apically, with several small setae along outer margin increasing in size apically and with a tibial spur. Tarsomeres short, terminal tarsomere as long as tarsomeres 34 combined; tarsomeres 14 bearing single pair of apicoventral spines.—
<emphasis id="B951EABA1701817F4E25F980FE4A7D36" bold="true" box="[381,490,1568,1592]" italics="true" pageId="9" pageNumber="10">Abdomen:</emphasis>
First abdominal ventrite punctuated with the same coarseness as metaventrite. Other visible abdominal sclerites (incl. pygidium) with finer punctuation. Propygidium short and wide, largely concealed by elytra.
</paragraph>
<paragraph id="8B9A36A81701817F4F29F961FBB87AEF" blockId="9.[113,763,1728,1972]" lastBlockId="9.[810,1463,144,481]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4F29F961FEF77DD6" box="[113,343,1729,1752]" italics="true" pageId="9" pageNumber="10">Taxonomic assignment:</emphasis>
We assign this new species to the genus
<taxonomicName id="4C254D2B1701817F4F29F940FF457DF9" authorityName="Poinar and Brown" authorityYear="2009" box="[113,229,1760,1783]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B951EABA1701817F4F29F940FF457DF9" box="[113,229,1760,1783]" italics="true" pageId="9" pageNumber="10">Pantostictus</emphasis>
</taxonomicName>
(in the monotypic Pantosticini;
<bibRefCitation id="EFB44B591701817F4D19F940FD4F7DF6" author="Zhou Y &amp; Ren D" box="[577,751,1760,1784]" pageId="9" pageNumber="10" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
Zhou
<emphasis id="B951EABA1701817F4DDAF940FD147DF6" box="[642,692,1760,1784]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2020
</bibRefCitation>
) based on the ventrally flat and oval body shape, broad and rectangular apex of prosternal process, and the coarse punctuation. The genus was further characterized by
<bibRefCitation id="EFB44B591701817F4D6AF89EFF1C7C7B" author="Poinar G &amp; Brown AE" pageId="9" pageNumber="10" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown (2009)</bibRefCitation>
as having an acute, triangular-shaped antennal club, and by lacking postmesocoxal and ventral abdominal striae. Despite appearing somewhat similar in the sole specimen of this new species, we believe that the odd antennal club shape is a postmortem deformity caused by collapse of a poorly sclerotized structure, as its exact shape varies considerably among available specimens (compare, e.g. Poinar and Browns figs 2 vs. 3). Further, although the tribal separation of
<taxonomicName id="4C254D2B1701817F4A6AFEADFA137A2B" box="[1330,1459,269,293]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Pantostictini">Pantostictini</taxonomicName>
from
<taxonomicName id="4C254D2B1701817F4C3EFE8CFC7A7A4A" authorityName="Portevin" authorityYear="1929" box="[870,986,300,324]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Plegaderini">Plegaderini</taxonomicName>
is well justified on several characters (
<bibRefCitation id="EFB44B591701817F4A22FE8CFC357A6A" author="Zhou Y &amp; Ren D" pageId="9" pageNumber="10" pagination="521 - 39" refId="ref19290" refString="Zhou Y, Caterino Mº, Ren D et al. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 2020; 36: 521 - 39. hups: // doi. org / 10.1111 / cla. 12418" type="journal article" year="2020">
Zhou
<emphasis id="B951EABA1701817F4C72FEECFCFB7A6A" box="[810,859,332,356]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2020
</bibRefCitation>
), the alleged absence of postcoxal striae (on the lateral metaventrite and on abdominal ventrite 1) should be disregarded. Their absence is difficult to assess in the specimens of
<taxonomicName id="4C254D2B1701817F4AFAFE2BFC3C7ACC" authorityName="Poinar and Brown" authorityYear="2009" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4AFAFE2BFC3C7ACC" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
, and the postmesocoxal striae, at least, are visible in
<taxonomicName id="4C254D2B1701817F4C72FE6AFC2D7AEF" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[810,909,457,481]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4C72FE6AFC2D7AEF" box="[810,909,457,481]" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
(see
<figureCitation id="131E2A2D1701817F4C9AFE69FBA87AEE" box="[962,1032,457,481]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516130" httpUri="https://zenodo.org/record/14516130/files/figure.png" pageId="9" pageNumber="10">Fig. 7F</figureCitation>
).
</paragraph>
<paragraph id="8B9A36A81701817F4C72FDA2FCCF789C" blockId="9.[810,1463,514,914]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72FDA2FBA07914" box="[810,1024,514,538]" italics="true" pageId="9" pageNumber="10">Differential diagnosis:</emphasis>
The species appears to be closely related to
<taxonomicName id="4C254D2B1701817F4C1EFD82FA967934" authority="Poinar and Brown, 2009" authorityName="Poinar and Brown" authorityYear="2009" box="[838,1334,546,570]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C1EFD82FB937934" box="[838,1075,546,570]" italics="true" pageId="9" pageNumber="10">Pantostictus burmanicus</emphasis>
<bibRefCitation id="EFB44B591701817F4B61FD82FA967934" author="Poinar G &amp; Brown AE" box="[1081,1334,546,570]" pageId="9" pageNumber="10" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown, 2009</bibRefCitation>
</taxonomicName>
. It is similar in overall appearance, but there are significant differential characters. The elytral punctuation is coarser in
<taxonomicName id="4C254D2B1701817F4BACFDC1FADE7976" authorityName="Poinar and Brown" authorityYear="2009" box="[1268,1406,608,632]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4BACFDC1FADE7976" box="[1268,1406,608,632]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
. The lateral pronotal margins are finely crenulate in
<taxonomicName id="4C254D2B1701817F4A4CFD20FADA7996" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1300,1402,640,664]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4A4CFD20FADA7996" box="[1300,1402,640,664]" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A2D1701817F4AD5FD20FCE679B8" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516130" httpUri="https://zenodo.org/record/14516130/files/figure.png" pageId="9" pageNumber="10">Fig. 7E</figureCitation>
), whereas in
<taxonomicName id="4C254D2B1701817F4C95FD00FBF579B9" authorityName="Poinar and Brown" authorityYear="2009" box="[973,1109,671,695]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C95FD00FBF579B9" box="[973,1109,671,695]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
they are smooth. In
<taxonomicName id="4C254D2B1701817F4A7CFD00FA2779B9" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" box="[1316,1415,671,695]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4A7CFD00FA2779B9" box="[1316,1415,671,695]" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
, the pronotum is wider basally and lateral margins are simply rounded and parallel-sided in the posterior half (
<figureCitation id="131E2A2D1701817F4BEEFD7DFB5E79F8" box="[1206,1278,733,758]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516130" httpUri="https://zenodo.org/record/14516130/files/figure.png" pageId="9" pageNumber="10">Fig. 7A</figureCitation>
), while the lateral pronotal margins of
<taxonomicName id="4C254D2B1701817F4B59FD5EFB2C781B" authorityName="Poinar and Brown" authorityYear="2009" box="[1025,1164,765,789]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4B59FD5EFB2C781B" box="[1025,1164,765,789]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
are sinuate. Furthermore,
<taxonomicName id="4C254D2B1701817F4AFAFD5EFCD7783A" authorityName="Jan &amp; ºimon-Pražák &amp; Yamamoto &amp; Lackner &amp; Fikáček &amp; Prokop &amp; Michael &amp; Caterino" authorityYear="2024" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B951EABA1701817F4AFAFD5EFCD7783A" italics="true" pageId="9" pageNumber="10">P. hirsutus</emphasis>
</taxonomicName>
displays a setose dorsal cuticle (
<figureCitation id="131E2A2D1701817F4B9BFCBCFAAE783A" box="[1219,1294,796,820]" captionStart="Figure 7" captionStartId="10.[129,194,1484,1508]" captionTargetBox="[129,1473,144,1456]" captionTargetId="figure-220@10.[129,1473,144,1456]" captionTargetPageId="10" captionText="Figure 7. Pantostictus hirsutus sp. nov.. A, dorsal view. B, ventral view.C, lateral view. D, elytral setation, E, dorsal view. F, metaventrite, abdomen.Abbreviations:ac, antennal club; as, antennal scape; ls, labral setae; pl, prosternal lobe; pmss, postmesocoxal stria." figureDoi="http://doi.org/10.5281/zenodo.14516130" httpUri="https://zenodo.org/record/14516130/files/figure.png" pageId="9" pageNumber="10">Fig. 7D</figureCitation>
). Setation presence in
<taxonomicName id="4C254D2B1701817F4C26FC9CFBA9785D" authorityName="Poinar and Brown" authorityYear="2009" box="[894,1033,827,851]" class="Insecta" family="Histeridae" genus="Pantostictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="burmanicus">
<emphasis id="B951EABA1701817F4C26FC9CFBA9785D" box="[894,1033,827,851]" italics="true" pageId="9" pageNumber="10">P. burmanicus</emphasis>
</taxonomicName>
cannot be determined based on the original description and published photographs (
<bibRefCitation id="EFB44B591701817F4BA4FCFBFCFE789C" author="Poinar G &amp; Brown AE" pageId="9" pageNumber="10" pagination="38 - 46" refId="ref18750" refString="Poinar G, Brown AE. Pantostictus burmanicus, a new genus and species of Cretaceous beetles (Coleoptera: Hydrophiloidea: Histeridae) in Burmese amber. Proceedings of the Entomological Society of Washington 2009; 111: 38 - 46. hups: // doi. org / 10.4289 / 0013 - 8797 - 111.1.38" type="journal article" year="2009">Poinar and Brown 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A36A81701817F4C72FC13FC6078E5" blockId="9.[810,1459,947,1003]" pageId="9" pageNumber="10">
<emphasis id="B951EABA1701817F4C72FC13FC3978C5" box="[810,921,947,971]" italics="true" pageId="9" pageNumber="10">Etymology:</emphasis>
The name of this species was chosen to point out its hirsute cuticle.
</paragraph>
</subSubSection>
</treatment>
</document>