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<document id="1B0CF0CFB37F3B484512589CFDBEF2D7" ID-DOI="10.26107/RBZ-2021-0065" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174585" ID-ZooBank="66CD6BC0-B64A-4E01-8B21-82061D9DC342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665291273268" checkinUser="felipe" docAuthor="Mendoza, Jose Christopher E." docDate="2021" docId="038C3420FF82FFFEADCAFF4F08A4F817" docLanguage="en" docName="RafflesBZool.69.463480.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Exagorium fidelisi Naruse, Chuang &amp; Tangah 2015" docType="treatment" docVersion="5" lastPageNumber="465" masterDocId="FFB54C58FF80FFFCAD69FFF2095FFF99" masterDocTitle="Marine crabs new to Singapore, with a description of a new species of intertidal xanthid crab of the genus Macromedaeus Ward, 1942 (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="480" masterPageNumber="463" pageNumber="465" updateTime="1699380004941" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DE8B93AF3F81F9A21A0D5C892E5B68D8">Marine crabs new to Singapore, with a description of a new species of intertidal xanthid crab of the genus Macromedaeus Ward, 1942 (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="AF9DD2715F6501ADA0C2CC82AB0790D4">Mendoza, Jose Christopher E.</mods:namePart>
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<taxonomicName id="4C25FEB5FF82FFFEADCAFF4F0BB4FF4C" ID-CoL="3DJK5" authority="Naruse, Chung &amp; Tangah, 2015" authorityName="Naruse, Chuang &amp; Tangah" authorityYear="2015" box="[163,747,188,213]" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEADCAFF4F0BB4FF4C" bold="true" box="[163,747,188,213]" pageId="2" pageNumber="465">
<emphasis id="B9515924FF82FFFEADCAFF4F082DFF4D" bold="true" box="[163,370,188,213]" italics="true" pageId="2" pageNumber="465">Exagorium fidelisi</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEAC13FF4F0BB4FF4C" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[378,747,189,213]" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse, Chung &amp; Tangah, 2015</bibRefCitation>
</emphasis>
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(
<figureCitation id="131E99B3FF82FFFEACCFFF2F08B9FF6C" box="[422,486,221,245]" captionStart="Fig" captionStartId="1.[113,148,2007,2029]" captionTargetBox="[126,1438,189,1993]" captionTargetId="figure-8@1.[126,1438,189,1993]" captionTargetPageId="1" captionText="Fig. 1. Exagorium fidelisi Naruse, Chung &amp; Tangah, 2015 (ZRC 2019.0662), Sungei Buloh Wetland Reserve, Singapore. A, male, 8.5 × 6.0 mm; BG, male, 8.7 × 6.1 mm; H, I, ovig. female, 9.0 × 6.7 mm. A, H, habitus, dorsal view; B, cephalothorax, anterior view; C, male thoracic sternum and pleon; D, eyes, antennae, and third maxillipeds, anteroventral view; E, left chela, external view; F, left G1, sternal view; G, left G1, pleonal view; I, cephalothorax and pleon, ventral view. Scale bars: A, H = 5.0 mm; BE, I = 1.0 mm; F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7174587" httpUri="https://zenodo.org/record/7174587/files/figure.png" pageId="2" pageNumber="465">Fig. 1</figureCitation>
)
</paragraph>
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<subSubSection id="C33FD6BDFF82FFFEADE1FEEE08B2FED7" pageId="2" pageNumber="465" type="reference_group">
<paragraph id="8B9A8536FF82FFFEADE1FEEE08B2FED7" blockId="2.[136,775,283,334]" pageId="2" pageNumber="465">
<treatmentCitationGroup id="AB35A218FF82FFFEADE1FEEE08B2FED7" pageId="2" pageNumber="465">
<treatmentCitation id="0A84A327FF82FFFEADE1FEEE0B1EFEA8" LSID="urn:lsid:zoobank.org:act:F0E95BFE-0893-44E1-946B-E9A678C988DD" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[136,577,283,305]" httpUri="http://treatment.plazi.org/id/33462940FFA8FFC019F3FA01FE9E85F4" page="328" pageId="2" pageNumber="465" year="2015">
<taxonomicName id="4C25FEB5FF82FFFEADE1FEEE0B1EFEA8" ID-CoL="3DJK5" authority="Naruse et al., 2015: 328" authorityName="Naruse" authorityPageNumber="328" authorityYear="2015" box="[136,577,283,305]" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEADE1FEEE0860FEA9" box="[136,319,283,305]" italics="true" pageId="2" pageNumber="465">Exagorium fidelisi</emphasis>
<materialsCitation id="3B4D8F6BFF82FFFEAC20FEEE0B1EFEA8" box="[329,577,283,305]" country="Malaysia" location="Sabah" pageId="2" pageNumber="465" specimenCount="1" stateProvince="Sabah">
<bibRefCitation id="EFB4F8C7FF82FFFEAC20FEEE0B1EFEA8" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[329,577,283,305]" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse et al., 2015: 328</bibRefCitation>
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, figs. 1, 2, 3a, 4, 5 (
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locality:
<collectingRegion id="49E14BD4FF82FFFEAC57FECA0824FED7" box="[318,379,312,334]" country="Malaysia" name="Sabah" pageId="2" pageNumber="465">Sabah</collectingRegion>
,
<collectingCountry id="F332C5A6FF82FFFEACEFFECA08BDFED7" box="[390,482,312,334]" name="Malaysia" pageId="2" pageNumber="465">Malaysia</collectingCountry>
).
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</paragraph>
</subSubSection>
<subSubSection id="C33FD6BDFF82FFFEADE1FE840A58FE76" pageId="2" pageNumber="465" type="materials_examined">
<paragraph id="8B9A8536FF82FFFEADE1FE840A58FE76" blockId="2.[136,775,374,495]" pageId="2" pageNumber="465">
<emphasis id="B9515924FF82FFFEADE1FE840838FE17" bold="true" box="[136,359,374,398]" pageId="2" pageNumber="465">Material examined.</emphasis>
<materialsCitation id="3B4D8F6BFF82FFFEAC07FE850A5BFE76" collectingDate="2017-02-09" collectionCode="ZRC" collectorName="Tan H. H." country="Singapore" location="Sungei Buloh Wetland Reserve" pageId="2" pageNumber="465" specimenCode="ZRC 2019.0662" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D234EBFFF82FFFEAC07FE85089BFE17" box="[366,452,374,399]" pageId="2" pageNumber="465" type="male">2 males</specimenCount>
, 8.5 × 6.0 mm, 8.7 ×
<quantity id="4CDD28D3FF82FFFEAFC7FE840A5CFE17" box="[686,771,374,398]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="465" unit="mm" value="6.1">6.1 mm</quantity>
, 1 ovig. female, 9.0×
<quantity id="4CDD28D3FF82FFFEAC02FE6408E1FE37" box="[363,446,406,430]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="465" unit="mm" value="6.7">6.7 mm</quantity>
(
<specimenCode id="DB832D4DFF82FFFEACA4FE650BDEFE37" box="[461,641,406,431]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="465">ZRC 2019.0662</specimenCode>
),
<collectingCountry id="F332C5A6FF82FFFEAFFDFE640A5CFE37" box="[660,771,406,430]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
,
<location id="8EFAD3EDFF82FFFEADE1FE44088BFE57" LSID="urn:lsid:plazi:treatment:038C3420FF82FFFEADCAFF4F08A4F817:8EFAD3EDFF82FFFEADE1FE44088BFE57" box="[136,468,438,462]" country="Singapore" name="Sungei Buloh Wetland Reserve" pageId="2" pageNumber="465">Sungei Buloh Wetland Reserve</location>
, back mangrove, in soft mud, under wooden plank, coll.
<collectorName id="26D0E0E0FF82FFFEACCBFE250B53FE77" box="[418,524,471,494]" pageId="2" pageNumber="465">Tan H. H.</collectorName>
et al.,
<date id="FF9BA3F6FF82FFFEAF38FE250A5BFE76" box="[593,772,470,495]" pageId="2" pageNumber="465" value="2017-02-09">
<collectingDate id="EFDF5A1EFF82FFFEAF38FE250A5BFE76" box="[593,772,470,495]" pageId="2" pageNumber="465" value="2017-02-09">9 February 2017</collectingDate>
</date>
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.
</paragraph>
</subSubSection>
<subSubSection id="C33FD6BDFF82FFFEADE1FDE508A4F817" pageId="2" pageNumber="465" type="discussion">
<paragraph id="8B9A8536FF82FFFEADE1FDE50BBCFCF7" blockId="2.[136,775,534,879]" pageId="2" pageNumber="465">
<emphasis id="B9515924FF82FFFEADE1FDE509ABFDB6" bold="true" box="[136,244,535,559]" pageId="2" pageNumber="465">Remarks.</emphasis>
The camptandriid genus
<taxonomicName id="4C25FEB5FF82FFFEAC9DFDE50862FDD7" authority="Naruse, Chung &amp; Tangah, 2015" authorityName="Naruse, Chung &amp; Tangah" authorityYear="2015" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="B9515924FF82FFFEAC9DFDE50B37FDB7" box="[500,616,535,558]" italics="true" pageId="2" pageNumber="465">Exagorium</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEAF04FDE50862FDD7" author="Naruse T &amp; Chung AYC &amp; Tangah J" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse, Chung &amp; Tangah, 2015</bibRefCitation>
</taxonomicName>
, is currently represented only by the
<typeStatus id="549E3B94FF82FFFEAFB0FDC50A58FDD6" box="[729,775,567,591]" pageId="2" pageNumber="465">type</typeStatus>
species,
<taxonomicName id="4C25FEB5FF82FFFEAD8DFDA50BE5FDF7" authority="Naruse, Chung &amp; Tanagah, 2015" authorityName="Naruse, Chung &amp; Tanagah" authorityYear="2015" box="[228,698,598,623]" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEAD8DFDA50815FDF7" box="[228,330,598,622]" italics="true" pageId="2" pageNumber="465">E. fidelisi</emphasis>
Naruse, Chung &amp; Tanagah, 2015
</taxonomicName>
, which was discovered in the estuarine habitats of Tundon Buhangin, Lower Kinabatangan-Segama Wetlands, in
<collectingRegion id="49E14BD4FF82FFFEAF3BFD640BC9FD37" box="[594,662,662,686]" country="Malaysia" name="Sabah" pageId="2" pageNumber="465">Sabah</collectingRegion>
,
<collectingCountry id="F332C5A6FF82FFFEAFF6FD640A5CFD37" box="[671,771,662,686]" name="Malaysia" pageId="2" pageNumber="465">Malaysia</collectingCountry>
, near the northeastern tip of Borneo. It is distinguishable from other mangrove camptandriid crabs by its hexagonal carapace with cristate anterolateral margins, pale ivory colouration, male pleon which completely conceals the G1, thin pleonal somite 1 which partially covers the P5 coxa, and slender, recurved, terminally bifurcate G1 (
<bibRefCitation id="EFB4F8C7FF82FFFEAF6AFCA50B87FCF7" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[515,728,854,879]" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse et al., 2015</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A8536FF82FFFEADE1FC6408A4F817" blockId="2.[136,776,918,1934]" pageId="2" pageNumber="465">
The present material (
<figureCitation id="131E99B3FF82FFFEACE1FC640893FC36" box="[392,460,918,943]" captionStart="Fig" captionStartId="1.[113,148,2007,2029]" captionTargetBox="[126,1438,189,1993]" captionTargetId="figure-8@1.[126,1438,189,1993]" captionTargetPageId="1" captionText="Fig. 1. Exagorium fidelisi Naruse, Chung &amp; Tangah, 2015 (ZRC 2019.0662), Sungei Buloh Wetland Reserve, Singapore. A, male, 8.5 × 6.0 mm; BG, male, 8.7 × 6.1 mm; H, I, ovig. female, 9.0 × 6.7 mm. A, H, habitus, dorsal view; B, cephalothorax, anterior view; C, male thoracic sternum and pleon; D, eyes, antennae, and third maxillipeds, anteroventral view; E, left chela, external view; F, left G1, sternal view; G, left G1, pleonal view; I, cephalothorax and pleon, ventral view. Scale bars: A, H = 5.0 mm; BE, I = 1.0 mm; F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7174587" httpUri="https://zenodo.org/record/7174587/files/figure.png" pageId="2" pageNumber="465">Fig. 1</figureCitation>
) agrees well with the
<typeStatus id="549E3B94FF82FFFEAFBEFC650A59FC36" box="[727,774,919,943]" pageId="2" pageNumber="465">type</typeStatus>
series (cf.
<bibRefCitation id="EFB4F8C7FF82FFFEAC6DFC4508BBFC57" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[260,484,950,974]" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse et al., 2015</bibRefCitation>
: figs. 1, 2, 3a, 4, 5), and there is no doubt that they are conspecific. The habitat of
<taxonomicName id="4C25FEB5FF82FFFEADE1FC0509B1FB97" authorityName="Naruse, Chung &amp; Tanagah" authorityYear="2015" box="[136,238,1014,1038]" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEADE1FC0509B1FB97" box="[136,238,1014,1038]" italics="true" pageId="2" pageNumber="465">E. fidelisi</emphasis>
</taxonomicName>
in
<collectingCountry id="F332C5A6FF82FFFEAC7AFC040820FB97" box="[275,383,1014,1038]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
is slightly different from that of the
<typeStatus id="549E3B94FF82FFFEADE1FBE509E9FBB6" box="[136,182,1047,1071]" pageId="2" pageNumber="465">type</typeStatus>
locality. The present material was collected from the back mangrove zone dominated by
<taxonomicName id="4C25FEB5FF82FFFEAF48FBC50B96FBD7" box="[545,713,1079,1102]" class="Magnoliopsida" family="Rhizophoraceae" genus="Bruguiera" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="465" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9515924FF82FFFEAF48FBC50BCDFBD7" box="[545,658,1079,1102]" italics="true" pageId="2" pageNumber="465">Bruguiera</emphasis>
spp.
</taxonomicName>
(Tan H. H., pers. comm.), while the
<typeStatus id="549E3B94FF82FFFEACB7FBA50B53FBF6" box="[478,524,1111,1135]" pageId="2" pageNumber="465">type</typeStatus>
material was collected from a muddy riverbank dominated by nipa palms. The collecting method is also different—the present material was picked by hand from beneath a wooden plank that had been laid on top of the soft mud (Tan H. H., pers. comm.), whereas the
<typeStatus id="549E3B94FF82FFFEAC7FFB05081BFA96" box="[278,324,1271,1295]" pageId="2" pageNumber="465">type</typeStatus>
material was collected from submerged mud burrows by yabby pump. The
<specimenCount id="9D234EBFFF82FFFEAF73FAE40B94FAB7" box="[538,715,1302,1326]" pageId="2" pageNumber="465" type="generic">three specimens</specimenCount>
(
<specimenCount id="9D234EBFFF82FFFEAFB4FAE50999FAD7" pageId="2" pageNumber="465" type="male">two males</specimenCount>
and one ovigerous female) were collected with other camptandriid crabs,
<taxonomicName id="4C25FEB5FF82FFFEAC1AFAA40998FA16" authority="(Tweedie, 1937)" baseAuthorityName="Tweedie" baseAuthorityYear="1937" class="Malacostraca" family="Camptandriidae" genus="Ilyogynnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="microcheirum">
<emphasis id="B9515924FF82FFFEAC1AFAA40BD3FAF7" box="[371,652,1366,1390]" italics="true" pageId="2" pageNumber="465">Ilyogynnis microcheirum</emphasis>
(
<bibRefCitation id="EFB4F8C7FF82FFFEAFF6FAA4099FFA16" author="Tweedie MWF" pageId="2" pageNumber="465" pagination="140 - 170" refId="ref13421" refString="Tweedie MWF (1937) On the crabs of the family Ocypodidae in the collection of the Raffles Museum. Bulletin of the Raffles Museum, 13: 140 - 170." type="journal article" year="1937">Tweedie, 1937</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C25FEB5FF82FFFEAD90FA840B97FA17" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[249,712,1398,1422]" class="Malacostraca" family="Camptandriidae" genus="Paracleistostoma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="depressum">
<emphasis id="B9515924FF82FFFEAD90FA840B74FA17" box="[249,555,1398,1422]" italics="true" pageId="2" pageNumber="465">Paracleistostoma depressum</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEAF58FA850B97FA17" author="De Man JG" box="[561,712,1398,1422]" pageId="2" pageNumber="465" pagination="677 - 708" refId="ref10836" refString="De Man JG (1895 - 1898) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4): 485 - 609, 9 (1): 75 - 218, 9 (2): 339 - 386, 9 (4): 459 - 514, 9 (6): 725 - 790, pls. 12 - 14, 10 (6): 677 - 708, pls. 28 - 38." type="journal article" year="1895">De Man, 1895</bibRefCitation>
</taxonomicName>
(both of which were more abundant). The presence of
<taxonomicName id="4C25FEB5FF82FFFEAFE1FA650BB3FA37" authorityName="Naruse, Chung &amp; Tanagah" authorityYear="2015" box="[648,748,1430,1454]" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEAFE1FA650BB3FA37" box="[648,748,1430,1454]" italics="true" pageId="2" pageNumber="465">E. fidelisi</emphasis>
</taxonomicName>
in
<collectingCountry id="F332C5A6FF82FFFEADE1FA4409ABFA57" box="[136,244,1462,1486]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
is perhaps not surprising considering its relative proximity to the
<typeStatus id="549E3B94FF82FFFEAC55FA250835FA76" box="[316,362,1495,1519]" pageId="2" pageNumber="465">type</typeStatus>
locality in
<collectingRegion id="49E14BD4FF82FFFEAC8CFA240B76FA77" box="[485,553,1494,1518]" country="Malaysia" name="Sabah" pageId="2" pageNumber="465">Sabah</collectingRegion>
, eastern
<collectingCountry id="F332C5A6FF82FFFEAFEEFA240BB3FA77" box="[647,748,1494,1518]" name="Malaysia" pageId="2" pageNumber="465">Malaysia</collectingCountry>
. It is, however, important to note that this species has eluded previous surveys of the mangroves of
<collectingCountry id="F332C5A6FF82FFFEAF42F9E40BC5F9B7" box="[555,666,1558,1582]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
, and does not appear in published accounts of Singapores ocypodoid crab fauna (e.g.,
<bibRefCitation id="EFB4F8C7FF82FFFEAC25F9A408AAF9F6" author="Tweedie MWF" box="[332,501,1622,1647]" pageId="2" pageNumber="465" pagination="140 - 170" refId="ref13421" refString="Tweedie MWF (1937) On the crabs of the family Ocypodidae in the collection of the Raffles Museum. Bulletin of the Raffles Museum, 13: 140 - 170." type="journal article" year="1937">Tweedie, 1937</bibRefCitation>
;
<bibRefCitation id="EFB4F8C7FF82FFFEAF6DF9A50B99F9F6" author="Tan CGS &amp; Ng PKL" box="[516,710,1623,1647]" pageId="2" pageNumber="465" pagination="75 - 84" refId="ref13342" refString="Tan CGS &amp; Ng PKL (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285: 75 - 84." type="journal article" year="1994">Tan &amp; Ng, 1994</bibRefCitation>
). The probable reason is that while the present habitat in which
<taxonomicName id="4C25FEB5FF82FFFEAF99F9850990F937" authorityName="Naruse, Chung &amp; Tanagah" authorityYear="2015" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEAF99F9850990F937" italics="true" pageId="2" pageNumber="465">E. fidelisi</emphasis>
</taxonomicName>
was found, the Sungei Buloh Wetland Reserve, has been extensively surveyed, the survey methods used then did not specifically target deeply burrowing infauna (contrasted to the use of a yabby pump by
<bibRefCitation id="EFB4F8C7FF82FFFEAC86F9050B97F897" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[495,712,1782,1806]" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse et al., 2015</bibRefCitation>
). The present specimens, therefore, were found serendipitously. This is the first published record of
<taxonomicName id="4C25FEB5FF82FFFEAC90F8C50B02F8D7" authorityName="Naruse, Chung &amp; Tanagah" authorityYear="2015" box="[505,605,1846,1870]" class="Malacostraca" family="Camptandriidae" genus="Exagorium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="fidelisi">
<emphasis id="B9515924FF82FFFEAC90F8C50B02F8D7" box="[505,605,1846,1870]" italics="true" pageId="2" pageNumber="465">E. fidelisi</emphasis>
</taxonomicName>
from
<collectingCountry id="F332C5A6FF82FFFEAFF2F8C40A58F8D7" box="[667,775,1846,1870]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
and only the second time it has been recorded since it was described by
<bibRefCitation id="EFB4F8C7FF82FFFEAC73F88508A8F817" author="Naruse T &amp; Chung AYC &amp; Tangah J" box="[282,503,1910,1934]" pageId="2" pageNumber="465" pagination="327 - 333" refId="ref12305" refString="Naruse T, Chung AYC &amp; Tangah J (2015) Description of a new genus and a new species of the family Camptandriidae Stimpson, 1858 (Crustacea: Decapoda: Brachyura) from Lower Kinabatangan-Segama Wetlands, Sabah, Malaysia. Raffles Bulletin of Zoology, 63: 327 - 333." type="journal article" year="2015">Naruse et al. (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>