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<document id="1EAD35CD0F38474907AA0ADAFFF26408" ID-DOI="10.11646/zootaxa.5575.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746442" ID-ZooBank="521D404D-EB9D-4110-989E-A8A273C10152" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737970514015" checkinUser="plazi" docAuthor="Génier, François, Gunter, Nicole L. &amp; Saxton, Natalie A." docDate="2025" docId="038987E8FFF86E0BFF4616DCFD52FBA0" docLanguage="en" docName="zootaxa.5575.1.3.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhytomus latidentatus Saxton &amp; Genier 2025, new species" docType="treatment" docVersion="2" lastPageNumber="103" masterDocId="FFB0FF90FFF76E1BFFD11005FFC3FFB0" masterDocTitle="Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="87" pageNumber="102" updateTime="1737971416811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="292639D3462885FD6A6CCBF39C3D1CA8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C9F6C9B06290549EA27CDF451F97F817">Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="633CE8E4F6B4844C1ADC364A260465FF" type="personal">
<mods:role id="CDF17DAB8EBC3459F5E7C270558089DC">
<mods:roleTerm id="87DFE4B5502419A0C554F8CC5AB3A806">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5AAEA94F4FE9F84C613B2AFB7A6E6B7F">Génier, François</mods:namePart>
<mods:nameIdentifier id="621FD31F6588EDF4635C46AF6496F05C" type="ORCID">0000-0002-5399-8412</mods:nameIdentifier>
<mods:nameIdentifier id="C713C4C48221E5A08BDB024DB8600892" type="email">enier@nature.ca</mods:nameIdentifier>
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<mods:namePart id="F3B578537FE3B4F5A23B16E0AEBBF272">Gunter, Nicole L.</mods:namePart>
<mods:affiliation id="189D6295E8B93136431F7064A6280AB0">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
</mods:name>
<mods:name id="3493E79752535A80675686246B1BC2E4" type="personal">
<mods:role id="3CE94B0E9E11CC6CD52FD92BA6E32A2A">
<mods:roleTerm id="AF865E45D0B677124D09FF05F551CE3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B43E2A017BCAF383FFD0EB51E34875F8">Saxton, Natalie A.</mods:namePart>
<mods:affiliation id="4BC64124AF42B6C66320D8834FF50F14">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. &amp; Corresponding author</mods:affiliation>
</mods:name>
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<mods:title id="1DB6E8D845EAD4AED92D352626F34390">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DF78E44E492A56890469E9CC11A63460">
<mods:date id="328F5E91006542CA616220DAE1B9BE0D">2025</mods:date>
<mods:detail id="F3D475CBCF1A4358FDC3C35345664C56" type="pubDate">
<mods:number id="02D89D1373CBBB331E3A971F440C3937">2025-01-22</mods:number>
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<mods:number id="4BF42AD86C58D1CB95719E023A9773AD">5575</mods:number>
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<mods:url id="03CF360E7CDCAE20A884EB79F85A2744">https://doi.org/10.11646/zootaxa.5575.1.3</mods:url>
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<treatment id="038987E8FFF86E0BFF4616DCFD52FBA0" LSID="urn:lsid:plazi:treatment:038987E8FFF86E0BFF4616DCFD52FBA0" httpUri="http://treatment.plazi.org/id/038987E8FFF86E0BFF4616DCFD52FBA0" lastPageId="16" lastPageNumber="103" pageId="15" pageNumber="102">
<subSubSection id="C33A6575FFF86E14FF4616DCFC0EF8A7" pageId="15" pageNumber="102" type="nomenclature">
<paragraph id="8B9F36FEFFF86E14FF4616DCFC0EF8A7" blockId="15.[151,973,1752,1852]" pageId="15" pageNumber="102">
<heading id="D0D78192FFF86E14FF4616DCFCDDF943" bold="true" box="[151,798,1752,1780]" fontSize="11" level="1" pageId="15" pageNumber="102" reason="1">
<emphasis id="B954EAECFFF86E14FF4616DCFCDDF943" bold="true" box="[151,798,1752,1780]" pageId="15" pageNumber="102">
<taxonomicName id="4C204D7DFFF86E14FF4616DCFD42F943" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,641,1752,1780]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="102" phylum="Arthropoda" rank="species" species="latidentatus" status="sp. nov.">
<emphasis id="B954EAECFFF86E14FF4616DCFE64F944" bold="true" box="[151,423,1753,1780]" italics="true" pageId="15" pageNumber="102">Rhytomus latidentatus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C33A6575FFF86E14FF461724FE2CF85F" pageId="15" pageNumber="102" type="description">
<paragraph id="8B9F36FEFFF86E14FF461724FE3AF88B" blockId="15.[151,973,1752,1852]" box="[151,505,1825,1852]" pageId="15" pageNumber="102">
<figureCitation id="131B2A7BFFF86E14FF461724FEE5F88B" box="[151,294,1825,1851]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="15" pageNumber="102">Figs. 4C, 4E</figureCitation>
,
<figureCitation id="131B2A7BFFF86E14FEE01727FE91F88C" box="[305,338,1826,1852]" captionStart="FIGURE 7" captionStartId="10.[152,255,1714,1738]" captionTargetBox="[156,1425,184,1681]" captionTargetId="figure-22@10.[151,1435,179,1688]" captionTargetPageId="10" captionText="FIGURE 7. Habitus of Rhytomus species in mainland New Guinea. A, R. fervidus; B, R. heurni; C, R. howdeni; D, R. larseni; E, R. latidentatus; F, R. papuanus; G, R. setulosum; H, R. sodalis; I, R. vanus; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14746458" httpUri="https://zenodo.org/record/14746458/files/figure.png" pageId="15" pageNumber="102">7E</figureCitation>
,
<figureCitation id="131B2A7BFFF86E14FE8F1724FE43F88B" box="[350,384,1825,1851]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="15" pageNumber="102">8E</figureCitation>
,
<figureCitation id="131B2A7BFFF86E14FE5A1724FE7DF88B" box="[395,446,1825,1851]" captionStart="FIGURE 10" captionStartId="14.[152,255,1979,2003]" captionTargetBox="[155,1431,490,1949]" captionTargetId="figure-166@14.[151,1435,486,1953]" captionTargetPageId="14" captionText="FIGURE 10. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral. A, R. latidentatus; B, R. papuanus; C, R. sodalis; D, R. vanus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746466" httpUri="https://zenodo.org/record/14746466/files/figure.png" pageId="15" pageNumber="102">10A</figureCitation>
,
<figureCitation id="131B2A7BFFF86E14FE1B1724FE3AF88B" box="[458,505,1825,1851]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="15" pageNumber="102">12A</figureCitation>
</paragraph>
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<emphasis id="B954EAECFFF86E14FF46176CFE06F833" bold="true" box="[151,453,1897,1923]" pageId="15" pageNumber="102">Name-bearing type data.</emphasis>
<typeStatus id="549B885CFFF86E14FE1E176CFDF4F833" box="[463,567,1897,1923]" pageId="15" pageNumber="102" type="holotype">Holotype</typeStatus>
<specimenCount id="9D26FD77FFF86E14FD90176FFD95F834" box="[577,598,1898,1924]" count="1" pageId="15" pageNumber="102" type="male">♂</specimenCount>
(CMNC). ||
<collectingCountry id="F337766EFFF86E14FD3D176CFBC5F833" box="[748,1030,1897,1924]" name="Papua New Guinea" pageId="15" pageNumber="102">PAPUA NEW GUINEA</collectingCountry>
: Western | Prov., P’nvang, CI RAP Survey | Camp 1.
<geoCoordinate id="EE145039FFF86E14FE8E1788FE06F818" box="[351,453,1933,1960]" degrees="05" direction="south" minutes="29.5" orientation="latitude" pageId="15" pageNumber="102" precision="92" value="-5.491667">05º29.5S</geoCoordinate>
<geoCoordinate id="EE145039FFF86E14FE1A1788FD83F817" box="[459,576,1933,1959]" degrees="141" direction="east" minutes="32.6" orientation="longitude" pageId="15" pageNumber="102" precision="92" value="141.54333">141º32.6E</geoCoordinate>
|
<specimenCount id="9D26FD77FFF86E14FD821788FD62F818" box="[595,673,1933,1960]" count="573" pageId="15" pageNumber="102" type="generic">
<quantity id="4CD89B1BFFF86E14FD821788FD5EF818" box="[595,669,1933,1960]" metricMagnitude="2" metricUnit="m" metricValue="5.7299999999999995" pageId="15" pageNumber="102" unit="m" value="573.0">573 m</quantity>
.
</specimenCount>
<date id="FF9E103EFFF86E14FD76178BFCDDF817" box="[679,798,1933,1960]" pageId="15" pageNumber="102" value="2013-05">May 2013</date>
. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049540 || [QR code label]; ||
<typeStatus id="549B885CFFF86E14FCC117B4FC67F87B" box="[784,932,1969,1995]" pageId="15" pageNumber="102" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D26FD77FFF86E14FC7D17B7FC02F87C" box="[940,961,1970,1996]" count="1" pageId="15" pageNumber="102" type="male">♂</specimenCount>
|
<taxonomicName id="4C204D7DFFF86E14FC0417B4FB80F87B" box="[981,1091,1969,1995]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFF86E14FC0417B4FB80F87B" box="[981,1091,1969,1995]" italics="true" pageId="15" pageNumber="102">Rhytomus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C204D7DFFF86E14FB8817B4FB1EF87B" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1113,1245,1969,1995]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="102" phylum="Arthropoda" rank="species" species="latidentatus">
<emphasis id="B954EAECFFF86E14FB8817B4FB1EF87B" box="[1113,1245,1969,1995]" italics="true" pageId="15" pageNumber="102">latidentatus</emphasis>
</taxonomicName>
| Des. N. Saxton &amp; F. Génier, 2024 || [red card].
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE76E0BFF161093FD5CFEF0" pageId="16" pageNumber="103" type="materials_examined">
<paragraph id="8B9F36FEFFE76E0BFF161093FD5CFEF0" blockId="16.[151,1437,150,1041]" pageId="16" pageNumber="103">
<emphasis id="B954EAECFFE76E0BFF161093FC7DFF00" bold="true" box="[199,958,150,180]" pageId="16" pageNumber="103">
Material examined (
<specimenCount id="9D26FD77FFE76E0BFE121093FDD5FF04" box="[451,534,150,180]" count="22" pageId="16" pageNumber="103" type="male">22 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFDF81093FDBFFF04" box="[553,636,150,180]" count="12" pageId="16" pageNumber="103" type="female">12 ♀♀</specimenCount>
).
<collectingCountry id="F337766EFFE76E0BFD491093FC7DFF00" box="[664,958,150,176]" name="Papua New Guinea" pageId="16" pageNumber="103">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
<materialsCitation id="3B483CA3FFE76E0BFC031093FD1DFF48" collectingDate="1976-08-12" collectingDateMax="1976-08-14" collectingDateMin="1976-08-12" collectionCode="CMNC, MNHN" collectorName="O. Kukal" country="Papua New Guinea" county="Kokoda Trak" elevation="700" latitude="-9.191667" location="Menari" longLatPrecision="21" longitude="147.625" municipality="Kauai River" pageId="16" pageNumber="103" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Central" typeStatus="paratype">
<collectingRegion id="49E4F81CFFE76E0BFC031093FB2EFF00" box="[978,1261,150,176]" country="Papua New Guinea" name="Central" pageId="16" pageNumber="103">CENTRAL PROVINCE</collectingRegion>
,
<collectingCounty id="62FE4E72FFE76E0BFB2C1093FA5BFF00" box="[1277,1432,150,176]" pageId="16" pageNumber="103">Kokoda Trak</collectingCounty>
,
<collectingMunicipality id="6BFBAC84FFE76E0BFF4610BFFEE7FF64" box="[151,292,186,212]" pageId="16" pageNumber="103">Kauai River</collectingMunicipality>
,
<location id="8EFF6025FFE76E0BFEE210BFFE44FF64" LSID="urn:lsid:plazi:treatment:038987E8FFF86E0BFF4616DCFD52FBA0:8EFF6025FFE76E0BFEE210BFFE44FF64" box="[307,391,186,212]" country="Papua New Guinea" county="Kokoda Trak" latitude="-9.191667" longLatPrecision="21" longitude="147.625" municipality="Kauai River" name="Menari" pageId="16" pageNumber="103" stateProvince="Central">Menari</location>
,
<quantity id="4CD89B1BFFE76E0BFE4410BFFE22FF65" box="[405,481,186,213]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="103" unit="m" value="700.0">
<elevation id="000DD1CDFFE76E0BFE4410BFFE22FF65" box="[405,481,186,213]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="103" unit="m" value="700.0">700 m</elevation>
</quantity>
, [
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09°11’30’’
<collectingRegion id="49E4F81CFFE76E0BFDA210BFFD40FF64" box="[627,643,186,212]" country="Papua New Guinea" name="S" pageId="16" pageNumber="103">S</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="EE145039FFE76E0BFD4310BFFCEDFF64" box="[658,814,186,212]" degrees="147" direction="east" minutes="37" orientation="longitude" pageId="16" pageNumber="103" precision="15" seconds="30" value="147.625">147°37’30’’E</geoCoordinate>
],
<date id="FF9E103EFFE76E0BFC9410BFFC3AFF64" box="[837,1017,186,212]" pageId="16" pageNumber="103" value="1976-08-12" valueMax="1976-08-14" valueMin="1976-08-12">
<collectingDate id="EFDAE9D6FFE76E0BFC9410BFFC3AFF64" box="[837,1017,186,212]" pageId="16" pageNumber="103" value="1976-08-12" valueMax="1976-08-14" valueMin="1976-08-12">12–14.viii.1976</collectingDate>
</date>
, rainforest, dung,
<collectorName id="26D55328FFE76E0BFB1C10BFFAFAFF64" box="[1229,1337,186,212]" pageId="16" pageNumber="103">O. Kukal</collectorName>
—
<specimenCount id="9D26FD77FFE76E0BFA8710BFFA54FF65" box="[1366,1431,186,213]" count="3" pageId="16" pageNumber="103" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFF4610DBFF00FF49" box="[151,195,222,249]" count="1" pageId="16" pageNumber="103" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFF1D10DAFEF4FF49" box="[204,311,223,249]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFE9910DBFE63FF48" box="[328,416,222,248]" pageId="16" pageNumber="103">CMNC</collectionCode>
),
<specimenCount id="9D26FD77FFE76E0BFE6410DBFE36FF49" box="[437,501,222,249]" count="2" pageId="16" pageNumber="103" type="male">2 ♂♂</specimenCount>
<typeStatus id="549B885CFFE76E0BFE2F10DAFDAAFF49" box="[510,617,223,249]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFDAB10DBFD15FF48" box="[634,726,222,248]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="103">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CA3FFE76E0BFD3A10DBFBB1FEAC" collectingDate="2013-05" collectionCode="ANIC" collectorName="T. Larsen" country="Papua New Guinea" elevation="575" latitude="-5.483333" location="Survey Camp" longLatPrecision="1305" longitude="141.53334" pageId="16" pageNumber="103" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Western" typeStatus="paratype">
<collectingRegion id="49E4F81CFFE76E0BFD3A10DBFBC4FF48" box="[747,1031,222,248]" country="Papua New Guinea" name="Western" pageId="16" pageNumber="103">WESTERN PROVINCE</collectingRegion>
, CI RAP
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1, P’nvang,
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<elevation id="000DD1CDFFE76E0BFF461107FF23FEAD" box="[151,224,258,285]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="16" pageNumber="103" unit="m" value="575.0">575 m</elevation>
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,
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05°29’
<collectingRegion id="49E4F81CFFE76E0BFEEA1107FE88FEAD" box="[315,331,258,285]" country="Papua New Guinea" name="S" pageId="16" pageNumber="103">S</collectingRegion>
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,
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141°32’
<collectingRegion id="49E4F81CFFE76E0BFE601107FE06FEAC" box="[433,453,258,284]" country="Papua New Guinea" name="E" pageId="16" pageNumber="103">E</collectingRegion>
</geoCoordinate>
,
<date id="FF9E103EFFE76E0BFE001107FDDDFEAC" box="[465,542,258,284]" pageId="16" pageNumber="103" value="2013-05">
<collectingDate id="EFDAE9D6FFE76E0BFE001107FDDDFEAC" box="[465,542,258,284]" pageId="16" pageNumber="103" value="2013-05">v.2013</collectingDate>
</date>
,
<collectorName id="26D55328FFE76E0BFDF81106FD54FEAC" box="[553,663,258,285]" pageId="16" pageNumber="103">T. Larsen</collectorName>
—
<specimenCount id="9D26FD77FFE76E0BFD651107FD18FEAD" box="[692,731,258,285]" count="1" pageId="16" pageNumber="103" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFD361107FCD1FEAD" box="[743,786,258,285]" count="1" pageId="16" pageNumber="103" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFCC81106FC47FEAD" box="[793,900,259,285]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFC421107FC1EFEAC" box="[915,989,258,284]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="16" pageNumber="103">ANIC</collectionCode>
),
<specimenCount id="9D26FD77FFE76E0BFC201106FBC5FEAD" box="[1009,1030,259,285]" count="1" pageId="16" pageNumber="103" type="male">♂</specimenCount>
<typeStatus id="549B885CFFE76E0BFBDC1107FBB1FEAC" box="[1037,1138,258,284]" pageId="16" pageNumber="103" type="holotype">holotype</typeStatus>
</materialsCitation>
,
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<specimenCount id="9D26FD77FFE76E0BFBAC1106FB51FEAD" box="[1149,1170,259,285]" count="1" pageId="16" pageNumber="103" type="female">♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFB481107FB35FEAC" box="[1177,1270,258,284]" pageId="16" pageNumber="103" type="allotype">allotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="3B483CA3FFE76E0BFAD31107FD58FEF0" collectionCode="CMNC, QM" country="Papua New Guinea" pageId="16" pageNumber="103" specimenCount="24" specimenCount-female="9" specimenCount-male="15" typeStatus="paratype">
<specimenCount id="9D26FD77FFE76E0BFAD31107FA92FEAD" box="[1282,1361,258,285]" count="14" pageId="16" pageNumber="103" type="male">14 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFA8C1107FA5FFEAD" box="[1373,1436,258,285]" count="8" pageId="16" pageNumber="103" type="female">8 ♀♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFF461122FEC1FEF1" box="[151,258,295,321]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFEC01123FEAAFEF0" box="[273,361,294,320]" pageId="16" pageNumber="103">CMNC</collectionCode>
),
<specimenCount id="9D26FD77FFE76E0BFEAC1123FE6AFEF1" box="[381,425,294,321]" count="1" pageId="16" pageNumber="103" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFE641123FE1CFEF1" box="[437,479,294,321]" count="1" pageId="16" pageNumber="103" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFE371122FD92FEF1" box="[486,593,295,321]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFDB11123FD50FEF0" box="[608,659,294,320]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="16" pageNumber="103" type="Museum">QM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE76E0BFF16114FFBA1FC30" pageId="16" pageNumber="103" type="description">
<paragraph id="8B9F36FEFFE76E0BFF16114FFBA1FC30" blockId="16.[151,1437,150,1041]" pageId="16" pageNumber="103">
<emphasis id="B954EAECFFE76E0BFF16114FFDD4FED4" bold="true" box="[199,535,330,356]" pageId="16" pageNumber="103">Description. Measurements.</emphasis>
Body length
<quantity id="4CD89B1BFFE76E0BFD60114FFCF8FED5" box="[689,827,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="6.3" metricValueMin="4.5" pageId="16" pageNumber="103" unit="mm" value="5.4" valueMax="6.3" valueMin="4.5">4.5–6.3 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFE76E0BFB99114FFB10FED5" box="[1096,1235,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="4.2" metricValueMin="3.1" pageId="16" pageNumber="103" unit="mm" value="3.65" valueMax="4.2" valueMin="3.1">3.1–4.2 mm</quantity>
.
<typeStatus id="549B885CFFE76E0BFB0F114FFA88FED4" box="[1246,1355,330,356]" pageId="16" pageNumber="103" type="holotype">Holotype</typeStatus>
: length
<quantity id="4CD89B1BFFE76E0BFF46116BFF2CFE39" box="[151,239,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="103" unit="mm" value="5.5">5.5 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFE76E0BFE26116BFD93FE39" box="[503,592,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="16" pageNumber="103" unit="mm" value="3.9">3.9 mm</quantity>
.
<emphasis id="B954EAECFFE76E0BFD8B116BFD5DFE38" bold="true" box="[602,670,366,392]" pageId="16" pageNumber="103">Body.</emphasis>
Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex.
<emphasis id="B954EAECFFE76E0BFEB91197FE6CFE1C" bold="true" box="[360,431,402,428]" pageId="16" pageNumber="103">Head.</emphasis>
Clypeal edge lateral to clypeal teeth lacking emargination, clypeal teeth wide, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.1 times eye width.
<emphasis id="B954EAECFFE76E0BFDE711DFFD75FE44" bold="true" box="[566,694,474,500]" pageId="16" pageNumber="103">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.6, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline with fine longitudinal depression.
<emphasis id="B954EAECFFE76E0BFE7A1227FE3DFD8C" bold="true" box="[427,510,546,572]" pageId="16" pageNumber="103">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures with distinct microsculptures (
<figureCitation id="131B2A7BFFE76E0BFE8A126FFE06FD34" box="[347,453,618,644]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="16" pageNumber="103">Fig. 12A</figureCitation>
), basal elytral pit absent, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae with the odd unaligned punctures on interstriae 3 and 5 and scattered unaligned punctures on interstria 4.
<emphasis id="B954EAECFFE76E0BFB0412B7FAD7FD7C" bold="true" box="[1237,1300,690,716]" pageId="16" pageNumber="103">Legs.</emphasis>
Profemoral anterior surface flat, male metatibial apicointernal teeth approximately in line with posterior tibial edge.
<emphasis id="B954EAECFFE76E0BFAF312D3FA5FFD40" bold="true" box="[1314,1436,726,752]" pageId="16" pageNumber="103">Pygidium.</emphasis>
Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (
<figureCitation id="131B2A7BFFE76E0BFB9012FFFB55FCA4" box="[1089,1174,762,788]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="16" pageNumber="103">Fig. 8E</figureCitation>
), surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex.
<emphasis id="B954EAECFFE76E0BFDE81347FD70FCEC" bold="true" box="[569,691,834,860]" pageId="16" pageNumber="103">Aedeagus.</emphasis>
Left paramere apex rounded in a dorsoventral plane (
<figureCitation id="131B2A7BFFE76E0BFAF41347FA53FCEC" box="[1317,1424,834,860]" captionStart="FIGURE 10" captionStartId="14.[152,255,1979,2003]" captionTargetBox="[155,1431,490,1949]" captionTargetId="figure-166@14.[151,1435,486,1953]" captionTargetPageId="14" captionText="FIGURE 10. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral. A, R. latidentatus; B, R. papuanus; C, R. sodalis; D, R. vanus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746466" httpUri="https://zenodo.org/record/14746466/files/figure.png" pageId="16" pageNumber="103">Fig. 10A</figureCitation>
).
<emphasis id="B954EAECFFE76E0BFF461363FEA0FC30" bold="true" box="[151,355,870,896]" pageId="16" pageNumber="103">Female genitalia.</emphasis>
Spermatheca with two apodemes basally and one apodeme apically.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE76E0BFF16138FFAA8FC78" pageId="16" pageNumber="103" type="etymology">
<paragraph id="8B9F36FEFFE76E0BFF16138FFAA8FC78" blockId="16.[151,1437,150,1041]" pageId="16" pageNumber="103">
<emphasis id="B954EAECFFE76E0BFF16138FFE8DFC14" bold="true" box="[199,334,906,932]" pageId="16" pageNumber="103">Etymology.</emphasis>
The specific epithet of this species is a combination of the Latin adjective “
<emphasis id="B954EAECFFE76E0BFB63138FFB2BFC14" box="[1202,1256,906,932]" italics="true" pageId="16" pageNumber="103">latus</emphasis>
” meaning wide and the adjective “
<emphasis id="B954EAECFFE76E0BFEB913ABFE04FC78" box="[360,455,942,968]" italics="true" pageId="16" pageNumber="103">dentatus</emphasis>
” meaning dentate in reference to this species’ characteristically wide clypeal teeth.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE76E0BFF1613D7FD52FBA0" pageId="16" pageNumber="103" type="discussion">
<paragraph id="8B9F36FEFFE76E0BFF1613D7FD52FBA0" blockId="16.[151,1437,150,1041]" pageId="16" pageNumber="103">
<emphasis id="B954EAECFFE76E0BFF1613D7FEF8FC5C" bold="true" box="[199,315,978,1004]" pageId="16" pageNumber="103">Remarks.</emphasis>
<bibRefCitation id="EFB14B0FFFE76E0BFE9313D7FE24FC5D" author="Paulian, R." box="[322,487,978,1005]" pageId="16" pageNumber="103" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian (1985)</bibRefCitation>
studied
<specimenCount id="9D26FD77FFE76E0BFD9413D7FD33FC5C" box="[581,752,978,1004]" count="4" pageId="16" pageNumber="103" type="generic">four specimens</specimenCount>
of this species which he identified as “
<emphasis id="B954EAECFFE76E0BFB7213D7FA43FC5C" box="[1187,1408,978,1004]" italics="true" pageId="16" pageNumber="103">
<taxonomicName id="4C204D7DFFE76E0BFB7213D7FAB6FC5C" authorityName="Paulian" authorityYear="1937" box="[1187,1397,978,1004]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="103" phylum="Arthropoda" rank="species" species="heurni">Tesserodon heurni</taxonomicName>
”
</emphasis>
in his work on the Canthonines of New
<collectingCountry id="F337766EFFE76E0BFDE813F3FD4DFBA0" box="[569,654,1014,1040]" name="Guinea-Bissau" pageId="16" pageNumber="103">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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