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<document id="8A4636BCDD84E207CC7778FB5D95675A" ID-DOI="10.11646/zootaxa.5575.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746442" ID-ZooBank="521D404D-EB9D-4110-989E-A8A273C10152" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737970514015" checkinUser="plazi" docAuthor="Génier, François, Gunter, Nicole L. &amp; Saxton, Natalie A." docDate="2025" docId="038987E8FFE66E08FF4615C0FDEAFD7D" docLanguage="en" docName="zootaxa.5575.1.3.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhytomus sodalis Saxton &amp; Genier 2025, new species" docType="treatment" docVersion="2" lastPageNumber="106" masterDocId="FFB0FF90FFF76E1BFFD11005FFC3FFB0" masterDocTitle="Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="87" pageNumber="104" updateTime="1737971416811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D0935E59EF3A721A73DCA2C400D0AFF8">Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="40143C7D89238D6000F978DE1042A025">Génier, François</mods:namePart>
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<mods:namePart id="D503B4DD84155E99A392B2CA6CB9CE57">Gunter, Nicole L.</mods:namePart>
<mods:affiliation id="B9F9D7DC05D80BDDE8E39EE12DA86C2B">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:namePart id="822C4033FFE700BFF612B96A1088E6F6">Saxton, Natalie A.</mods:namePart>
<mods:affiliation id="391F94C9B26CC0DD9171A9C305BA1689">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. &amp; Corresponding author</mods:affiliation>
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<taxonomicName id="4C204D7DFFE66E0AFF4615C0FD84FA6F" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,583,1476,1504]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="104" phylum="Arthropoda" rank="species" species="sodalis" status="sp. nov.">
<emphasis id="B954EAECFFE66E0AFF4615C0FEAFFA50" bold="true" box="[151,364,1477,1504]" italics="true" pageId="17" pageNumber="104">Rhytomus sodalis</emphasis>
Saxton &amp; Génier
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,
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<figureCitation id="131B2A7BFFE66E0AFF461608FF39F998" box="[151,250,1549,1576]" captionStart="FIGURE 7" captionStartId="10.[152,255,1714,1738]" captionTargetBox="[156,1425,184,1681]" captionTargetId="figure-22@10.[151,1435,179,1688]" captionTargetPageId="10" captionText="FIGURE 7. Habitus of Rhytomus species in mainland New Guinea. A, R. fervidus; B, R. heurni; C, R. howdeni; D, R. larseni; E, R. latidentatus; F, R. papuanus; G, R. setulosum; H, R. sodalis; I, R. vanus; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14746458" httpUri="https://zenodo.org/record/14746458/files/figure.png" pageId="17" pageNumber="104">Figs. 7H</figureCitation>
,
<figureCitation id="131B2A7BFFE66E0AFED71608FEE9F997" box="[262,298,1549,1575]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="17" pageNumber="104">8H</figureCitation>
,
<figureCitation id="131B2A7BFFE66E0AFEE71608FEA4F997" box="[310,359,1549,1575]" captionStart="FIGURE 10" captionStartId="14.[152,255,1979,2003]" captionTargetBox="[155,1431,490,1949]" captionTargetId="figure-166@14.[151,1435,486,1953]" captionTargetPageId="14" captionText="FIGURE 10. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral. A, R. latidentatus; B, R. papuanus; C, R. sodalis; D, R. vanus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746466" httpUri="https://zenodo.org/record/14746466/files/figure.png" pageId="17" pageNumber="104">10C</figureCitation>
,
<figureCitation id="131B2A7BFFE66E0AFEA21608FE62F997" box="[371,417,1549,1575]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="17" pageNumber="104">12C</figureCitation>
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<emphasis id="B954EAECFFE66E0AFF461650FE06F9DF" bold="true" box="[151,453,1621,1647]" pageId="17" pageNumber="104">Name-bearing type data.</emphasis>
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(CMNC): ||
<collectingCountry id="F337766EFFE66E0AFD3D1653FBC4F9DF" box="[748,1031,1621,1648]" name="Papua New Guinea" pageId="17" pageNumber="104">PAPUA NEW GUINEA</collectingCountry>
: Western | Prov., P’nvang, CI RAP Survey | Camp 1.
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|
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.
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. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049583 || [QR code label]; ||
<typeStatus id="549B885CFFE66E0AFCDF1698FC61F907" box="[782,930,1693,1719]" pageId="17" pageNumber="104" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D26FD77FFE66E0AFC78169BFC7DF908" box="[937,958,1694,1720]" count="1" pageId="17" pageNumber="104" type="male">♂</specimenCount>
|
<taxonomicName id="4C204D7DFFE66E0AFC021698FB82F907" box="[979,1089,1693,1719]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFE66E0AFC021698FB82F907" box="[979,1089,1693,1719]" italics="true" pageId="17" pageNumber="104">Rhytomus</emphasis>
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|
<taxonomicName id="4C204D7DFFE66E0AFB841698FB67F907" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1109,1188,1693,1719]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="104" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="B954EAECFFE66E0AFB841698FB67F907" box="[1109,1188,1693,1719]" italics="true" pageId="17" pageNumber="104">sodalis</emphasis>
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| Des. N. Saxton &amp; F. Génier, 2024 || [red card].
</paragraph>
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<emphasis id="B954EAECFFE66E0AFF1616E0FE6FF94F" bold="true" box="[199,428,1765,1791]" pageId="17" pageNumber="104">Material examined</emphasis>
(
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).
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<collectingCountry id="F337766EFFE66E0AFD5B16E0FC6FF94F" box="[650,940,1765,1791]" name="Papua New Guinea" pageId="17" pageNumber="104">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
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, CI RAP
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1, P’nvang,
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05°29’
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141°32’
<collectingRegion id="49E4F81CFFE66E0AFDA6170CFD48F893" box="[631,651,1801,1827]" country="Papua New Guinea" name="E" pageId="17" pageNumber="104">E</collectingRegion>
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,
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<collectingDate id="EFDAE9D6FFE66E0AFD44170CFD20F893" box="[661,739,1801,1827]" pageId="17" pageNumber="104" value="2013-05">v.2013</collectingDate>
</date>
, T.
<collectorName id="26D55328FFE66E0AFCD6170CFC97F893" box="[775,852,1801,1827]" pageId="17" pageNumber="104">Larsen</collectorName>
—
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,
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(
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),
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,
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(
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),
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,
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<specimenCount id="9D26FD77FFE66E0AFE60172BFE05F8F8" box="[433,454,1838,1864]" count="1" pageId="17" pageNumber="104" type="female">♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFE1F1728FDE8F8F7" box="[462,555,1837,1863]" pageId="17" pageNumber="104" type="allotype">allotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="3B483CA3FFE66E0AFDE61728FBECF803" collectingDate="2013-05" collectionCode="CMNC, MNHN, NMPC, USNM, QM" collectorName="T. Larsen" country="Papua New Guinea" elevation="800" latitude="-5.5333333" location="Survey Camp" longLatPrecision="1305" longitude="141.55" municipality="Survey Camp" pageId="17" pageNumber="104" specimenCount="85" specimenCount-female="21" specimenCount-male="64" stateProvince="S" typeStatus="paratype">
<specimenCount id="9D26FD77FFE66E0AFDE61728FD44F8F8" box="[567,647,1837,1864]" count="23" pageId="17" pageNumber="104" type="male">23 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFD451728FD21F8F8" box="[660,738,1837,1864]" count="12" pageId="17" pageNumber="104" type="female">12 ♀♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFD3A172BFC95F8F8" box="[747,854,1838,1864]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFCB71728FC7DF8F7" box="[870,958,1837,1863]" pageId="17" pageNumber="104">CMNC</collectionCode>
),
<specimenCount id="9D26FD77FFE66E0AFC031728FBD7F8F8" box="[978,1044,1837,1864]" count="2" pageId="17" pageNumber="104" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFBF01728FB8FF8F8" box="[1057,1100,1837,1864]" count="1" pageId="17" pageNumber="104" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFB84172BFB03F8F8" box="[1109,1216,1838,1864]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFB011728FAE8F8F7" box="[1232,1323,1837,1863]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="104">MNHN</collectionCode>
),
<specimenCount id="9D26FD77FFE66E0AFAEE1728FA42F8F8" box="[1343,1409,1837,1864]" count="2" pageId="17" pageNumber="104" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFA5F1728FF6FF8DC" count="1" pageId="17" pageNumber="104" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFF621757FEDDF8DC" box="[179,286,1874,1900]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFEFC1754FE41F8DB" box="[301,386,1873,1899]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="17" pageNumber="104">NMPC</collectionCode>
),
<specimenCount id="9D26FD77FFE66E0AFE471754FE01F8DC" box="[406,450,1873,1900]" count="1" pageId="17" pageNumber="104" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFE1F1754FE3BF8DC" box="[462,504,1873,1900]" count="1" pageId="17" pageNumber="104" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFE2E1757FDA9F8DC" box="[511,618,1874,1900]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFDA81754FD13F8DB" box="[633,720,1873,1899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="104" type="Museum">USNM</collectionCode>
); CI RAP
<collectingMunicipality id="6BFBAC84FFE66E0AFC921754FC1FF8DB" box="[835,988,1873,1899]" pageId="17" pageNumber="104">Survey Camp</collectingMunicipality>
2, P’nvang,
<quantity id="4CD89B1BFFE66E0AFBBA1754FB77F8DC" box="[1131,1204,1873,1900]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="104" unit="m" value="800.0">
<elevation id="000DD1CDFFE66E0AFBBA1754FB77F8DC" box="[1131,1204,1873,1900]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="104" unit="m" value="800.0">800 m</elevation>
</quantity>
,
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05°30’
<collectingRegion id="49E4F81CFFE66E0AFADF1754FADDF8DC" box="[1294,1310,1873,1900]" country="Papua New Guinea" name="S" pageId="17" pageNumber="104">S</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="EE145039FFE66E0AFAF81754FA54F8DB" box="[1321,1431,1873,1899]" degrees="141" direction="east" minutes="34" orientation="longitude" pageId="17" pageNumber="104" precision="925" value="141.56667">
141°34’
<collectingRegion id="49E4F81CFFE66E0AFA521754FA54F8DB" box="[1411,1431,1873,1899]" country="Papua New Guinea" name="E" pageId="17" pageNumber="104">E</collectingRegion>
</geoCoordinate>
,
<date id="FF9E103EFFE66E0AFF461770FF26F83F" box="[151,229,1909,1935]" pageId="17" pageNumber="104" value="2013-05">
<collectingDate id="EFDAE9D6FFE66E0AFF461770FF26F83F" box="[151,229,1909,1935]" pageId="17" pageNumber="104" value="2013-05">v.2013</collectingDate>
</date>
,
<collectorName id="26D55328FFE66E0AFF3F1773FE94F820" box="[238,343,1909,1936]" pageId="17" pageNumber="104">T. Larsen</collectorName>
—
<specimenCount id="9D26FD77FFE66E0AFEA51770FE7DF820" box="[372,446,1909,1936]" count="15" pageId="17" pageNumber="104" type="male">15 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFE161770FDC7F820" box="[455,516,1909,1936]" count="3" pageId="17" pageNumber="104" type="female">3 ♀♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFDD81773FDB7F820" box="[521,628,1910,1936]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFD501770FD19F83F" box="[641,730,1909,1935]" pageId="17" pageNumber="104">CMNC</collectionCode>
); CI RAP
<location id="8EFF6025FFE66E0AFC971770FC1EF83F" LSID="urn:lsid:plazi:treatment:038987E8FFE66E08FF4615C0FDEAFD7D:8EFF6025FFE66E0AFC971770FC1EF83F" box="[838,989,1909,1935]" country="Papua New Guinea" latitude="-5.5333333" longLatPrecision="1305" longitude="141.55" municipality="Survey Camp" name="Survey Camp" pageId="17" pageNumber="104" stateProvince="S">Survey Camp</location>
3, P’nvang,
<quantity id="4CD89B1BFFE66E0AFBB41770FB7BF820" box="[1125,1208,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="17" pageNumber="104" unit="m" value="1110.0">
<elevation id="000DD1CDFFE66E0AFBB41770FB7BF820" box="[1125,1208,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="17" pageNumber="104" unit="m" value="1110.0">1110 m</elevation>
</quantity>
,
<geoCoordinate id="EE145039FFE66E0AFB131770FAE3F820" box="[1218,1312,1909,1936]" degrees="05" direction="south" minutes="32" orientation="latitude" pageId="17" pageNumber="104" precision="925" value="-5.5333333">
05°32’
<collectingRegion id="49E4F81CFFE66E0AFAC11770FAE3F820" box="[1296,1312,1909,1936]" country="Papua New Guinea" name="S" pageId="17" pageNumber="104">S</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="EE145039FFE66E0AFAFB1770FA54F83F" box="[1322,1431,1909,1935]" degrees="141" direction="east" minutes="33" orientation="longitude" pageId="17" pageNumber="104" precision="925" value="141.55">
141°33’
<collectingRegion id="49E4F81CFFE66E0AFA521770FA54F83F" box="[1411,1431,1909,1935]" country="Papua New Guinea" name="E" pageId="17" pageNumber="104">E</collectingRegion>
</geoCoordinate>
,
<date id="FF9E103EFFE66E0AFF46179CFF26F803" box="[151,229,1945,1971]" pageId="17" pageNumber="104" value="2013-05">
<collectingDate id="EFDAE9D6FFE66E0AFF46179CFF26F803" box="[151,229,1945,1971]" pageId="17" pageNumber="104" value="2013-05">v.2013</collectingDate>
</date>
,
<collectorName id="26D55328FFE66E0AFF21179FFE98F803" box="[240,347,1945,1972]" pageId="17" pageNumber="104">T. Larsen</collectorName>
—
<specimenCount id="9D26FD77FFE66E0AFEA9179CFE07F804" box="[376,452,1945,1972]" count="19" pageId="17" pageNumber="104" type="male">19 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFE01179CFE39F804" box="[464,506,1945,1972]" count="1" pageId="17" pageNumber="104" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFDD0179FFDAFF804" box="[513,620,1946,1972]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFDAA179CFD10F803" box="[635,723,1945,1971]" pageId="17" pageNumber="104">CMNC</collectionCode>
),
<specimenCount id="9D26FD77FFE66E0AFD37179CFCE4F804" box="[742,807,1945,1972]" count="2" pageId="17" pageNumber="104" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE66E0AFCE2179CFCB1F804" box="[819,882,1945,1972]" count="2" pageId="17" pageNumber="104" type="female">2 ♀♀</specimenCount>
<typeStatus id="549B885CFFE66E0AFCA8179FFC27F804" box="[889,996,1946,1972]" pageId="17" pageNumber="104" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE66E0AFC25179CFBE4F803" box="[1012,1063,1945,1971]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="17" pageNumber="104" type="Museum">QM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5F6676FFE56E09FF491652FCF2F96B" ID-DOI="http://doi.org/10.5281/zenodo.14746472" ID-Zenodo-Dep="14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="18" pageNumber="105" startId="18.[152,255,1623,1647]" targetBox="[151,1435,179,1597]" targetPageId="18" targetType="figure">
<paragraph id="8B9F36FEFFE56E09FF491652FCF2F96B" blockId="18.[151,1437,1623,1755]" pageId="18" pageNumber="105">
<emphasis id="B954EAECFFE56E09FF491652FEEBF9DF" bold="true" box="[152,296,1623,1647]" pageId="18" pageNumber="105">FIGURE 12.</emphasis>
Characters used for species identification. A, elytral surface with microsculpture (
<taxonomicName id="4C204D7DFFE56E09FB5D165DFAE0F9DF" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1164,1315,1623,1647]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="105" phylum="Arthropoda" rank="species" species="latidentatus">
<emphasis id="B954EAECFFE56E09FB5D165DFAE0F9DF" box="[1164,1315,1623,1647]" italics="true" pageId="18" pageNumber="105">R. latidentatus</emphasis>
</taxonomicName>
); B, elytral surface glossy (
<taxonomicName id="4C204D7DFFE56E09FEEC1679FE6CF923" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[317,431,1659,1683]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="105" phylum="Arthropoda" rank="species" species="fervidus">
<emphasis id="B954EAECFFE56E09FEEC1679FE6CF923" box="[317,431,1659,1683]" italics="true" pageId="18" pageNumber="105">R. fervidus</emphasis>
</taxonomicName>
); C, male metatibial apicointernal tooth in line with posterior tibial edge (
<taxonomicName id="4C204D7DFFE56E09FB191679FAECF923" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1224,1327,1659,1683]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="105" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="B954EAECFFE56E09FB191679FAECF923" box="[1224,1327,1659,1683]" italics="true" pageId="18" pageNumber="105">R. sodalis</emphasis>
</taxonomicName>
); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (
<taxonomicName id="4C204D7DFFE56E09FC8616A5FC78F907" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[855,955,1695,1719]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="105" phylum="Arthropoda" rank="species" species="larseni">
<emphasis id="B954EAECFFE56E09FC8616A5FC78F907" box="[855,955,1695,1719]" italics="true" pageId="18" pageNumber="105">R. larseni</emphasis>
</taxonomicName>
); E, lateral view of strongly convex elytra (
<taxonomicName id="4C204D7DFFE56E09FA5716A5FF37F96B" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Balthasar" baseAuthorityYear="1965" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="105" phylum="Arthropoda" rank="species" species="setulosus">
<emphasis id="B954EAECFFE56E09FA5716A5FF37F96B" italics="true" pageId="18" pageNumber="105">R. setulosus</emphasis>
</taxonomicName>
); F, lateral view of moderately convex elytra (
<taxonomicName id="4C204D7DFFE56E09FD1A16C1FCE0F96B" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[715,803,1732,1755]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="105" phylum="Arthropoda" rank="species" species="vanus">
<emphasis id="B954EAECFFE56E09FD1A16C1FCE0F96B" box="[715,803,1732,1755]" italics="true" pageId="18" pageNumber="105">R. vanus</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<subSubSection id="C33A6575FFE56E08FF161728FC4CFE45" lastPageId="19" lastPageNumber="106" pageId="18" pageNumber="105" type="description">
<paragraph id="8B9F36FEFFE56E08FF161728FC4CFE45" blockId="18.[151,1437,1837,2043]" lastBlockId="19.[151,1437,151,717]" lastPageId="19" lastPageNumber="106" pageId="18" pageNumber="105">
<emphasis id="B954EAECFFE56E09FF161728FDDBF8F7" bold="true" box="[199,536,1837,1863]" pageId="18" pageNumber="105">Description. Measurements.</emphasis>
Body length 3.0–
<quantity id="4CD89B1BFFE56E09FD361728FC82F8F8" box="[743,833,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="18" pageNumber="105" unit="mm" value="4.6">4.6 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFE56E09FB821728FB1CF8F8" box="[1107,1247,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.7" metricValueMin="2.1" pageId="18" pageNumber="105" unit="mm" value="2.4" valueMax="2.7" valueMin="2.1">2.1–2.7 mm</quantity>
.
<typeStatus id="549B885CFFE56E09FB3A1728FA9BF8F7" box="[1259,1368,1837,1863]" pageId="18" pageNumber="105" type="holotype">Holotype</typeStatus>
: body length
<quantity id="4CD89B1BFFE56E09FF351754FEF8F8DC" box="[228,315,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="18" pageNumber="105" unit="mm" value="3.8">3.8 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFE56E09FD921754FD5FF8DC" box="[579,668,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="105" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="B954EAECFFE56E09FD771754FD29F8DB" bold="true" box="[678,746,1873,1899]" pageId="18" pageNumber="105">Body.</emphasis>
Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third.
<emphasis id="B954EAECFFE56E09FDF21770FDA9F83F" bold="true" box="[547,618,1909,1935]" pageId="18" pageNumber="105">Head.</emphasis>
Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.2 times eye width.
<emphasis id="B954EAECFFE56E09FDCF17B8FD5DF867" bold="true" box="[542,670,1981,2007]" pageId="18" pageNumber="105">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression.
<emphasis id="B954EAECFFE46E08FE7A1092FE3DFF01" bold="true" box="[427,510,151,177]" pageId="19" pageNumber="106">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit wide and deep, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures in some individuals.
<emphasis id="B954EAECFFE46E08FC031122FBD2FEF1" bold="true" box="[978,1041,295,321]" pageId="19" pageNumber="106">Legs.</emphasis>
Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge (
<figureCitation id="131B2A7BFFE46E08FECC116AFE47FE39" box="[285,388,367,393]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="19" pageNumber="106">Fig. 12C</figureCitation>
).
<emphasis id="B954EAECFFE46E08FE49116BFDD1FE38" bold="true" box="[408,530,366,392]" pageId="19" pageNumber="106">Pygidium.</emphasis>
Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove with coarse transverse microsculptures (
<figureCitation id="131B2A7BFFE46E08FC321196FBF8FE1D" box="[995,1083,403,429]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="19" pageNumber="106">Fig. 8H</figureCitation>
), pygidium of female depressed basally.
<emphasis id="B954EAECFFE46E08FF2011B3FEA8FE60" bold="true" box="[241,363,438,464]" pageId="19" pageNumber="106">Aedeagus.</emphasis>
Left paramere apex truncated apically, distinctly angular externally (
<figureCitation id="131B2A7BFFE46E08FB8C11B2FB02FE61" box="[1117,1217,439,465]" captionStart="FIGURE 10" captionStartId="14.[152,255,1979,2003]" captionTargetBox="[155,1431,490,1949]" captionTargetId="figure-166@14.[151,1435,486,1953]" captionTargetPageId="14" captionText="FIGURE 10. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral. A, R. latidentatus; B, R. papuanus; C, R. sodalis; D, R. vanus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746466" httpUri="https://zenodo.org/record/14746466/files/figure.png" pageId="19" pageNumber="106">Fig. 10C</figureCitation>
).
<emphasis id="B954EAECFFE46E08FB0211B2FA5FFE60" bold="true" box="[1235,1436,438,465]" pageId="19" pageNumber="106">Female genitalia.</emphasis>
Spermatheca with two apodemes basally and one apodeme apically.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE46E08FF1611FAFBB7FD8D" pageId="19" pageNumber="106" type="etymology">
<paragraph id="8B9F36FEFFE46E08FF1611FAFBB7FD8D" blockId="19.[151,1437,151,717]" pageId="19" pageNumber="106">
<emphasis id="B954EAECFFE46E08FF1611FAFE8DFDA9" bold="true" box="[199,334,511,537]" pageId="19" pageNumber="106">Etymology.</emphasis>
The specific epithet of this species is a masculine noun in apposition
<taxonomicName id="4C204D7DFFE46E08FB8C11FAFB79FDA9" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1117,1210,511,537]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="106" phylum="Arthropoda" rank="species" species="sodalis">“sŏdālis</taxonomicName>
” (companion, mate, fellow) relating to the fact that it was collected along with
<taxonomicName id="4C204D7DFFE46E08FCF61226FC57FD8D" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[807,916,547,573]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="106" phylum="Arthropoda" rank="species" species="larseni">
<emphasis id="B954EAECFFE46E08FCF61226FC57FD8D" box="[807,916,547,573]" italics="true" pageId="19" pageNumber="106">R. larseni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C204D7DFFE46E08FC1B1226FBAEFD8D" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[970,1133,547,573]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="106" phylum="Arthropoda" rank="species" species="latidentatus">
<emphasis id="B954EAECFFE46E08FC1B1226FBAEFD8D" box="[970,1133,547,573]" italics="true" pageId="19" pageNumber="106">R. latidentatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE46E08FF161242FDEAFD7D" pageId="19" pageNumber="106" type="discussion">
<paragraph id="8B9F36FEFFE46E08FF161242FDEAFD7D" blockId="19.[151,1437,151,717]" pageId="19" pageNumber="106">
<emphasis id="B954EAECFFE46E08FF161242FEF8FDD1" bold="true" box="[199,315,583,609]" pageId="19" pageNumber="106">Remarks.</emphasis>
<specimenCount id="9D26FD77FFE46E08FE951242FE04FDD1" box="[324,455,583,609]" count="1" pageId="19" pageNumber="106" type="female">One female</specimenCount>
specimen (excluded from
<typeStatus id="549B885CFFE46E08FD2B1242FCE9FDD1" box="[762,810,583,609]" pageId="19" pageNumber="106">type</typeStatus>
series) was discovered from the same collecting event (i.e., Camp
<quantity id="4CD89B1BFFE46E08FECF126EFE45FD35" box="[286,390,618,645]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="19" pageNumber="106" unit="m" value="1575.0">1, 575 m</quantity>
) that had coarser pronotal punctation and a barely indicated meso-metaventrite suture. This specimen likely represents a fourth species. However, because no male specimens could be found at the time of this study, we refrain from describing it.
</paragraph>
</subSubSection>
</treatment>
</document>
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