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<mods:title id="31DC93B898CE5431ED741114E151E2F5">New or rare Madagascar tiger beetles- 24. Pogonostoma (Microgeniatum) signifemorale sp. nov. with revised key to species of the subgenus Microgeniatum Rivalier (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="C947F9EBF8C777E8BECFF3BDEE250A8C">Moravec, Jiří</mods:namePart>
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<mods:affiliation id="0BBC81109ABAEB4D20D126AFF1C769E8">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. &amp; jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410</mods:affiliation>
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<mods:namePart id="66B62C0BBDE75F19CB6C77C7E1A2A2E0">Trýzna, Miloš</mods:namePart>
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<mods:affiliation id="A66FDBD7381736F7DEACF95CE9F17E45">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. &amp; anthribidae @ gmail. com; https: // orcid. org / 0000 - 0002 - 0426 - 3348</mods:affiliation>
<mods:nameIdentifier id="7FB5312DB18D903D10A0CC6EE6936D82" type="email">anthribidae@gmail.com</mods:nameIdentifier>
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<mods:date id="0B2BDF92230F446E96C36AF9E9F6C56C">2021</mods:date>
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<paragraph id="8B9F36ADFFD75824FF5DFC50FD54DCC4" blockId="2.[151,646,906,933]" box="[151,646,906,933]" pageId="2" pageNumber="526">
<heading id="D0D781C1FFD75824FF5DFC50FD54DCC4" bold="true" box="[151,646,906,933]" fontSize="11" level="1" pageId="2" pageNumber="526" reason="1">
<emphasis id="B954EABFFFD75824FF5DFC50FD54DCC4" bold="true" box="[151,646,906,933]" pageId="2" pageNumber="526">
Subgenus
<taxonomicName id="4C204D2EFFD75824FEDDFC51FD54DCC4" ID-CoL="8MGG8" authority="Rivalier, 1970" authorityName="Rivalier" authorityYear="1970" box="[279,646,906,933]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FEDDFC51FE00DCC5" bold="true" box="[279,466,907,933]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
<bibRefCitation id="EFB14B5CFFD75824FE10FC50FD54DCC4" author="Rivalier, E." box="[474,646,906,932]" pageId="2" pageNumber="526" pagination="269 - 338" refId="ref6843" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de Franc e, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<paragraph id="8B9F36ADFFD75824FF5DFC0AFD2DDC87" blockId="2.[151,1030,975,1061]" box="[151,767,975,999]" pageId="2" pageNumber="526">
<treatmentCitationGroup id="AB301183FFD75824FF5DFC0AFD2DDC87" box="[151,767,975,999]" pageId="2" pageNumber="526">
<treatmentCitation id="0A8110BCFFD75824FF5DFC0AFD4DDC87" author="Rivalier, E." box="[151,671,975,999]" page="306" pageId="2" pageNumber="526" year="1970">
<taxonomicName id="4C204D2EFFD75824FF5DFC0AFD4DDC87" ID-CoL="8MGG8" authority="Rivalier, 1970: 306" authorityName="Rivalier" authorityPageNumber="306" authorityYear="1970" box="[151,671,975,999]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FF5DFC0AFEF0DC87" box="[151,290,976,999]" italics="true" pageId="2" pageNumber="526">Pogonostoma</emphasis>
(
<emphasis id="B954EABFFFD75824FEFBFC0AFE1EDC87" box="[305,460,976,999]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
)
<bibRefCitation id="EFB14B5CFFD75824FE11FC15FD4DDC87" author="Rivalier, E." box="[475,671,975,999]" pageId="2" pageNumber="526" pagination="269 - 338" refId="ref6843" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de Franc e, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970: 306</bibRefCitation>
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(partim).
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</paragraph>
<paragraph id="8B9F36ADFFD75824FF5DFC35FD62DB66" blockId="2.[151,1030,975,1061]" box="[151,688,1006,1030]" pageId="2" pageNumber="526">
<treatmentCitationGroup id="AB301183FFD75824FF5DFC35FD62DB66" box="[151,688,1006,1030]" pageId="2" pageNumber="526">
<taxonomicName id="4C204D2EFFD75824FF5DFC35FE0ADB66" ID-CoL="8MGG8" authority="" box="[151,472,1006,1030]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FF5DFC35FEF0DB66" box="[151,290,1007,1030]" italics="true" pageId="2" pageNumber="526">Pogonostoma</emphasis>
(
<emphasis id="B954EABFFFD75824FEFBFC34FE1EDB65" box="[305,460,1006,1029]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="0A8110BCFFD75824FE28FC34FD7EDB66" author="Moravec, J." box="[482,684,1006,1030]" page="447" pageId="2" pageNumber="526" year="2007">
<bibRefCitation id="EFB14B5CFFD75824FE28FC34FD7EDB66" author="Moravec, J." box="[482,684,1006,1030]" pageId="2" pageNumber="526" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007: 447</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C33A6526FFD75824FF5DFBD7FCBFD995" pageId="2" pageNumber="526" type="type_taxon">
<paragraph id="8B9F36ADFFD75824FF5DFBD7FBD4DB44" blockId="2.[151,1030,975,1061]" box="[151,1030,1036,1061]" pageId="2" pageNumber="526">
<emphasis id="B954EABFFFD75824FF5DFBD7FECCDB44" bold="true" box="[151,286,1036,1061]" pageId="2" pageNumber="526">
<typeStatus id="549B880FFFD75824FF5DFBD7FF1EDB45" box="[151,204,1037,1061]" pageId="2" pageNumber="526">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C204D2EFFD75824FEE0FBD7FD2CDB45" ID-CoL="4KW2V" authority="W. Horn, 1934" authorityName="W. Horn" authorityYear="1934" box="[298,766,1036,1061]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="pseudominimum">
<emphasis id="B954EABFFFD75824FEE0FBD7FDB0DB44" box="[298,610,1036,1060]" italics="true" pageId="2" pageNumber="526">Pogonostoma pseudominimum</emphasis>
W.
<bibRefCitation id="EFB14B5CFFD75824FD40FBD7FD2CDB45" author="Horn, W." box="[650,766,1037,1061]" pageId="2" pageNumber="526" pagination="7 - 31" refId="ref5494" refString="Horn, W. (1934) Catalogue Bibliographique et synonymique des Cicindelides de Madagascar. In: Horn, W. &amp; Olsoufieff, G., Les Cicindelides de Madagascar (premiere partie). Memories de l'Academie Malgache, 20, 7 - 31 + 3 pls." type="journal article" year="1934">Horn, 1934</bibRefCitation>
</taxonomicName>
(by original designation).
</paragraph>
<paragraph id="8B9F36ADFFD75824FF5DFB88FDF3D97D" blockId="2.[151,1437,1106,2034]" pageId="2" pageNumber="526">
<emphasis id="B954EABFFFD75824FF5DFB88FE29DB0C" bold="true" box="[151,507,1106,1132]" pageId="2" pageNumber="526">Revised differential diagnosis</emphasis>
. Because of their extremely small to small,
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long, black bodies, and 4setose labrum, the four known species of the subgenus
<taxonomicName id="4C204D2EFFD75824FCE0FBADFC04DBF0" ID-CoL="8MGG8" box="[810,982,1143,1168]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FCE0FBADFC04DBF0" box="[810,982,1143,1168]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
</taxonomicName>
may resemble some species of the subgenus
<taxonomicName id="4C204D2EFFD75824FF2AFB41FDE4DBD5" ID-CoL="8MMD6" authority="Jeannel, 1946" authorityName="Jeannel" authorityYear="1946" box="[224,566,1179,1205]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microstenocera">
<emphasis id="B954EABFFFD75824FF2AFB41FE42DBD4" box="[224,400,1179,1204]" italics="true" pageId="2" pageNumber="526">Microstenocera</emphasis>
<bibRefCitation id="EFB14B5CFFD75824FE52FB41FDE4DBD5" author="Jeannel, R." box="[408,566,1179,1205]" pageId="2" pageNumber="526" refId="ref5545" refString="Jeannel, R. (1946) Coleopteres Carabiques de la region Malgache (premiere partie). Faune de l'empire francais. VI. Libraire Larosse, Paris, 372 pp." type="book" year="1946">Jeannel, 1946</bibRefCitation>
</taxonomicName>
. However, in contrast to the irregular elytral setal vesture in
<taxonomicName id="4C204D2EFFD75824FB2FFB41FA47DBD4" ID-CoL="8MMD6" authorityName="Jeannel" authorityYear="1946" box="[1253,1429,1179,1204]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microstenocera">
<emphasis id="B954EABFFFD75824FB2FFB41FA47DBD4" box="[1253,1429,1179,1204]" italics="true" pageId="2" pageNumber="526">Microstenocera</emphasis>
</taxonomicName>
, the species of
<taxonomicName id="4C204D2EFFD75824FEF1FB65FE35DBB8" ID-CoL="8MGG8" box="[315,487,1215,1240]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FEF1FB65FE35DBB8" box="[315,487,1215,1240]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
</taxonomicName>
are characterized by their regular elytral setal vesture, consisting of ornamental se- tae, which are confined to punctures—each ornamental seta (microtrichia) arising from anterior margin of corre- sponding puncture (as in the nominotypical subgenus, which, however, houses species with much larger body with multisetose labrum). Elytra shiny black, notably coarsely punctate throughout, with smooth, glabrous intervals. Pronotal disc sculptured by coarse, or very coarse, transverse-wavy, or partly anastomosing ridges. Head temples moderately long, approximately 2 times shorter than eyes. Galea entirely black (or with slightly brownish apex of terminal galeomere). Labial palpi black, or partly or entirely testaceous. Antennae entirely black, or black-brown, or (in one species) predominantly ivory-testaceous. Legs with apical half of mesofemora, or also metafemora, more or less conspicuously subclavate-dilated, entirely black, except for one species with markedly bicoloured femora and tibiae. Aedeagus of various shapes.
</paragraph>
<paragraph id="8B9F36ADFFD75824FF0DF9FCFCBFD995" blockId="2.[151,1437,1106,2034]" pageId="2" pageNumber="526">
<emphasis id="B954EABFFFD75824FF0DF9FCFE6BD921" bold="true" box="[199,441,1574,1601]" pageId="2" pageNumber="526">Nomenclatorial note</emphasis>
. Because of their regular elytral setal vesture, species of the subgenus
<taxonomicName id="4C204D2EFFD75824FB08F9FDFABCD920" ID-CoL="8MGG8" box="[1218,1390,1575,1600]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FB08F9FDFABCD920" box="[1218,1390,1575,1600]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
</taxonomicName>
had been treated by
<bibRefCitation id="EFB14B5CFFD75824FE87F991FDC7D905" author="Olsoufieff, G." box="[333,533,1611,1637]" pageId="2" pageNumber="526" pagination="31 - 73" refId="ref6697" refString="Olsoufieff, G. (1934) Essai de Revision systematique et Biologie des Cicindelides de Madagascar. In: Horn, W. &amp; Olsoufieff, G., Les Cicindelides de Madagascar (deuxieme partie). Memoires de l'Academie Malgache, 20, pp. 31 - 73." type="journal article" year="1934">Olsoufieff (1934)</bibRefCitation>
,
<bibRefCitation id="EFB14B5CFFD75824FDE8F991FD1BD906" author="Jeannel, R." box="[546,713,1611,1638]" pageId="2" pageNumber="526" refId="ref5545" refString="Jeannel, R. (1946) Coleopteres Carabiques de la region Malgache (premiere partie). Faune de l'empire francais. VI. Libraire Larosse, Paris, 372 pp." type="book" year="1946">Jeannel (1946)</bibRefCitation>
and
<bibRefCitation id="EFB14B5CFFD75824FCC9F991FC5DD905" author="Horn, W." box="[771,911,1611,1637]" pageId="2" pageNumber="526" pagination="7 - 31" refId="ref5494" refString="Horn, W. (1934) Catalogue Bibliographique et synonymique des Cicindelides de Madagascar. In: Horn, W. &amp; Olsoufieff, G., Les Cicindelides de Madagascar (premiere partie). Memories de l'Academie Malgache, 20, 7 - 31 + 3 pls." type="journal article" year="1934">Horn (1934)</bibRefCitation>
in the nominotypical subgenus
<taxonomicName id="4C204D2EFFD75824FB31F991FA47D904" ID-CoL="8KYDW" authorityName="Klug" authorityYear="1835" box="[1275,1429,1611,1636]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B954EABFFFD75824FB31F991FA47D904" box="[1275,1429,1611,1636]" italics="true" pageId="2" pageNumber="526">Pogonostoma</emphasis>
</taxonomicName>
. When
<bibRefCitation id="EFB14B5CFFD75824FF2BF9B5FE5FD9E9" author="Rivalier, E." box="[225,397,1647,1673]" pageId="2" pageNumber="526" pagination="269 - 338" refId="ref6843" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de Franc e, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier (1970)</bibRefCitation>
established the subgenus
<taxonomicName id="4C204D2EFFD75824FD7AF9B5FC8ED9E8" ID-CoL="8MGG8" box="[688,860,1647,1672]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FD7AF9B5FC8ED9E8" box="[688,860,1647,1672]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
</taxonomicName>
for the three above-mentioned species, he included also
<taxonomicName id="4C204D2EFFD75824FF06F949FD57D9CD" ID-CoL="4KW2K" authority="W. Horn, 1934" authorityName="W. Horn" authorityYear="1934" box="[204,645,1683,1709]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B954EABFFFD75824FF06F949FE04D9CD" box="[204,470,1683,1709]" italics="true" pageId="2" pageNumber="526">Pogonostoma gladiator</emphasis>
W.
<bibRefCitation id="EFB14B5CFFD75824FDCFF949FD57D9CD" author="Horn, W." box="[517,645,1683,1709]" pageId="2" pageNumber="526" pagination="7 - 31" refId="ref5494" refString="Horn, W. (1934) Catalogue Bibliographique et synonymique des Cicindelides de Madagascar. In: Horn, W. &amp; Olsoufieff, G., Les Cicindelides de Madagascar (premiere partie). Memories de l'Academie Malgache, 20, 7 - 31 + 3 pls." type="journal article" year="1934">Horn, 1934</bibRefCitation>
</taxonomicName>
there. Due to extremely distinctive diagnostic characters of
<emphasis id="B954EABFFFD75824FAFBF949FF1DD9B0" italics="true" pageId="2" pageNumber="526">Pogonos- toma</emphasis>
<taxonomicName id="4C204D2EFFD75824FF1FF96DFDD5D9B1" ID-CoL="4KW2K" authorityName="W. Horn" authorityYear="1934" box="[213,519,1719,1745]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="species" species="gladiator" subGenus="Hornogeniatum">
(
<emphasis id="B954EABFFFD75824FF14F96DFE5DD9B0" box="[222,399,1719,1744]" italics="true" pageId="2" pageNumber="526">Hornogeniatum</emphasis>
)
<emphasis id="B954EABFFFD75824FE55F96DFDD5D9B1" box="[415,519,1719,1745]" italics="true" pageId="2" pageNumber="526">gladiator</emphasis>
</taxonomicName>
(unique within the whole genus), the species was placed later (
<bibRefCitation id="EFB14B5CFFD75824FB74F96DFAB1D9B1" author="Moravec, J." box="[1214,1379,1719,1745]" pageId="2" pageNumber="526" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
) into a separate, monobasic subgenus
<taxonomicName id="4C204D2EFFD75824FDCEF901FCB8D995" authority="Moravec, 2007" authorityName="Moravec" authorityYear="2007" box="[516,874,1755,1781]" class="Insecta" family="Carabidae" genus="Hornogeniatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B954EABFFFD75824FDCEF901FD67D994" box="[516,693,1755,1780]" italics="true" pageId="2" pageNumber="526">Hornogeniatum</emphasis>
<bibRefCitation id="EFB14B5CFFD75824FD76F901FCB8D995" author="Moravec, J." box="[700,874,1755,1781]" pageId="2" pageNumber="526" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec, 2007</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6526FFD75825FF0DF924FA87DECD" lastPageId="3" lastPageNumber="527" pageId="2" pageNumber="526" type="distribution">
<paragraph id="8B9F36ADFFD75825FF0DF924FA87DECD" blockId="2.[151,1437,1106,2034]" lastBlockId="3.[151,1437,151,429]" lastPageId="3" lastPageNumber="527" pageId="2" pageNumber="526">
<emphasis id="B954EABFFFD75824FF0DF924FE8ED878" bold="true" box="[199,348,1790,1816]" pageId="2" pageNumber="526">Distribution</emphasis>
(
<figureCitation id="131B2A28FFD75824FEBAF925FE25D879" box="[368,503,1791,1817]" captionStart="FIGURES 1114" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[233,1354,193,1908]" captionTargetId="figure-29@7.[218,1369,181,1920]" captionTargetPageId="7" captionText="FIGURES 1114. Distribution of Pogonostoma (Microgeniatum) in Madagascar. 11—P. (M.) pseudominimum W. Horn; 12— P. (M.) infimum Rivalier; 13—P. (M.) flavopalpale Jeannel; 14—P. (M.) signifemorale sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5778849" httpUri="https://zenodo.org/record/5778849/files/figure.png" pageId="2" pageNumber="526">Figs 1114</figureCitation>
). Species of the subgenus are very rare, absent in most collections (including BMNH—see
<bibRefCitation id="EFB14B5CFFD75824FEF9F8F9FD97D85D" author="Moravec, J. &amp; Gillett, P. D. T." box="[307,581,1827,1853]" pageId="2" pageNumber="526" pagination="131 - 154" refId="ref6003" refString="Moravec, J. &amp; Gillett, P. D. T. (2009) Annotated list of specimens of Madagascan Cicindelidae deposited in The Natural History Museum London. Folia Heyrovskyana, Series A, 16, 131 - 154." type="journal article" year="2009">Moravec &amp; Gillett 2009</bibRefCitation>
). Behaviour of adults is similar to most species of the silvicolous and strictly arboreal genus; developmental stages unknown. Somewhat more common is only
<emphasis id="B954EABFFFD75824FBE0F89DFBE9D800" box="[1066,1083,1863,1888]" italics="true" pageId="2" pageNumber="526">P</emphasis>
. (
<taxonomicName id="4C204D2EFFD75824FB98F89DFAD2D800" box="[1106,1280,1863,1888]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD75824FB98F89DFAD2D800" box="[1106,1280,1863,1888]" italics="true" pageId="2" pageNumber="526">Microgeniatum</emphasis>
</taxonomicName>
)
<emphasis id="B954EABFFFD75824FADAF89DFF1CD8E4" italics="true" pageId="2" pageNumber="526">pseudomini- mum</emphasis>
W.
<bibRefCitation id="EFB14B5CFFD75824FF30F8B1FEA9D8E5" author="Horn, W." box="[250,379,1899,1925]" pageId="2" pageNumber="526" pagination="7 - 31" refId="ref5494" refString="Horn, W. (1934) Catalogue Bibliographique et synonymique des Cicindelides de Madagascar. In: Horn, W. &amp; Olsoufieff, G., Les Cicindelides de Madagascar (premiere partie). Memories de l'Academie Malgache, 20, 7 - 31 + 3 pls." type="journal article" year="1934">Horn, 1934</bibRefCitation>
(type species of the subgenus), which is known from eastern
<collectingCountry id="F337763DFFD75824FBF9F8B1FB6FD8E5" box="[1075,1213,1899,1925]" name="Madagascar" pageId="2" pageNumber="526">Madagascar</collectingCountry>
(Antongil Bay, dis- trict of Maroantsetra), penetrating through the forest of the Mananara Nord National Park (Province of
<collectingRegion id="49E4F84FFFD75824FAD0F855FA47D8CA" box="[1306,1429,1935,1962]" country="Madagascar" name="Toamasina" pageId="2" pageNumber="526">Toamasina</collectingRegion>
) to central area of Central Highlands, through Antsianaka near Lac Alaotra and the forest of Didy (both in the district of Ambatondrazaka) up to the forest of Analamazaotra-Perinet (type locality) east of Moramanga. The
<typeStatus id="549B880FFFD75824FAD6F80DFAACD891" box="[1308,1406,2007,2033]" pageId="2" pageNumber="526" type="syntype">syntypes</typeStatus>
(
<specimenCount id="9D26FD24FFD75825FA5AF80DFF7CDFD1" lastPageId="3" lastPageNumber="527" pageId="2" pageNumber="526" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD24FFD65825FF70FF4DFF36DFD1" box="[186,228,151,177]" pageId="3" pageNumber="527" type="female">1 ♀</specimenCount>
,
<typeStatus id="549B880FFFD65825FF38FF4DFE8BDFD1" box="[242,345,151,177]" pageId="3" pageNumber="527" type="lectotype">lectotype</typeStatus>
not designated) of
<taxonomicName id="4C204D2EFFD65825FDFAFF4DFC01DFD1" authorityName="W. Horn" authorityYear="1934" box="[560,979,151,177]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="pseudominimum" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD65825FDFAFF4DFD93DFD0" box="[560,577,151,176]" italics="true" pageId="3" pageNumber="527">P</emphasis>
. (
<emphasis id="B954EABFFFD65825FD93FF4DFCD5DFD0" box="[601,775,151,176]" italics="true" pageId="3" pageNumber="527">Microgeniatum</emphasis>
)
<emphasis id="B954EABFFFD65825FCD3FF4DFC01DFD1" box="[793,979,151,177]" italics="true" pageId="3" pageNumber="527">pseudominimum</emphasis>
</taxonomicName>
are deposited in SDEI; other specimens examined are in MNHN, CCJM, CJVB and JWCW (
<bibRefCitation id="EFB14B5CFFD65825FD3BFF61FC4ADFB5" author="Moravec, J." box="[753,920,187,213]" pageId="3" pageNumber="527" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
and recent personal data). The following two species have not been confirmed recently.
<taxonomicName id="4C204D2EFFD65825FDBFFF05FB5ADF99" authority="Jeannel, 1946" authorityName="Jeannel" authorityYear="1946" box="[629,1160,223,249]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="flavopalpale" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD65825FDBFFF05FD54DF98" box="[629,646,223,248]" italics="true" pageId="3" pageNumber="527">P</emphasis>
. (
<emphasis id="B954EABFFFD65825FD57FF05FC9BDF98" box="[669,841,223,248]" italics="true" pageId="3" pageNumber="527">Microgeniatum</emphasis>
)
<emphasis id="B954EABFFFD65825FC90FF05FC34DF99" box="[858,998,223,249]" italics="true" pageId="3" pageNumber="527">flavopalpale</emphasis>
<bibRefCitation id="EFB14B5CFFD65825FC27FF05FB5ADF99" author="Jeannel, R." box="[1005,1160,223,249]" pageId="3" pageNumber="527" refId="ref5545" refString="Jeannel, R. (1946) Coleopteres Carabiques de la region Malgache (premiere partie). Faune de l'empire francais. VI. Libraire Larosse, Paris, 372 pp." type="book" year="1946">Jeannel, 1946</bibRefCitation>
</taxonomicName>
is known only from
<specimenCount id="9D26FD24FFD65825FABEFF05FF1FDE7D" pageId="3" pageNumber="527" type="male">one male</specimenCount>
and
<specimenCount id="9D26FD24FFD65825FECEFED9FE52DE7D" box="[260,384,259,285]" pageId="3" pageNumber="527" type="female">one female</specimenCount>
(MNHN), caught in the forest of Piton Saint-Louis, north of Tolagnaro (= Fort Dauphin), 121 years ago. Only the male
<typeStatus id="549B880FFFD65825FE7CFEFDFDCADE21" box="[438,536,295,321]" pageId="3" pageNumber="527" type="holotype">holotype</typeStatus>
(MNHN) of
<taxonomicName id="4C204D2EFFD65825FD64FEFDFB47DE21" authority="Rivalier, 1970" authorityName="Rivalier" authorityYear="1970" box="[686,1173,295,321]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="infimum" subGenus="Microgeniatum">
<emphasis id="B954EABFFFD65825FD64FEFDFD6DDE20" box="[686,703,295,320]" italics="true" pageId="3" pageNumber="527">P</emphasis>
. (
<emphasis id="B954EABFFFD65825FD1CFEFDFC50DE20" box="[726,898,295,320]" italics="true" pageId="3" pageNumber="527">Microgeniatum</emphasis>
)
<emphasis id="B954EABFFFD65825FC59FEFDFC3DDE21" box="[915,1007,295,321]" italics="true" pageId="3" pageNumber="527">infimum</emphasis>
<bibRefCitation id="EFB14B5CFFD65825FC3CFEFDFB47DE21" author="Rivalier, E." box="[1014,1173,295,321]" pageId="3" pageNumber="527" pagination="269 - 338" refId="ref6843" refString="Rivalier, E. (1970) Le genre Pogonostoma (Col. Cicindelidae). Revision avec description d'especes nouvelles. Annales de la Societe entomologique de Franc e, New Series, 6, 269 - 338." type="journal article" year="1970">Rivalier, 1970</bibRefCitation>
</taxonomicName>
exists; it was caught in the deciduous forest of Analamerana,
<quantity id="4CD89B48FFD65825FD82FE90FD43DE05" box="[584,657,330,357]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="527" unit="km" value="50.0">50 km</quantity>
south-east of
<collectingRegion id="49E4F84FFFD65825FCF8FE91FC6BDE05" box="[818,953,331,357]" country="Madagascar" name="Antsiranana" pageId="3" pageNumber="527">Antsiranana</collectingRegion>
(= Diego Suarez) in northern
<collectingCountry id="F337763DFFD65825FADFFE91FA4EDE05" box="[1301,1436,331,357]" name="Madagascar" pageId="3" pageNumber="527">Madagascar</collectingCountry>
(in
<date id="FF9E106DFFD65825FF74FEB5FE8BDEE9" box="[190,345,366,393]" pageId="3" pageNumber="527" value="1959-01">January 1959</date>
). Finally,
<taxonomicName id="4C204D2EFFD65825FE00FEB5FC99DEE9" authority="Moravec &amp; Trýzna, 2021" authorityName="Moravec &amp; Trýzna" authorityYear="2021" box="[458,843,367,393]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="signifemorale" status="sp. nov." subGenus="Microgeniatum">
<emphasis id="B954EABFFFD65825FE00FEB5FE09DEE8" box="[458,475,367,392]" italics="true" pageId="3" pageNumber="527">P</emphasis>
. (
<emphasis id="B954EABFFFD65825FE3EFEB5FD72DEE8" box="[500,672,367,392]" italics="true" pageId="3" pageNumber="527">Microgeniatum</emphasis>
)
<emphasis id="B954EABFFFD65825FD7BFEB5FC99DEE9" box="[689,843,367,393]" italics="true" pageId="3" pageNumber="527">signifemorale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757C4FFD65825FC99FEB5FC7DDEE9" box="[851,943,367,393]" pageId="3" pageNumber="527" rank="species">
<emphasis id="B954EABFFFD65825FC99FEB5FC7ADEE9" bold="true" box="[851,936,367,393]" pageId="3" pageNumber="527">sp. nov</emphasis>
.
</taxonomicNameLabel>
is known only from the single male caught recently in the dense, dry deciduous forest of the Ankarafantsika National Park in northwestern
<collectingCountry id="F337763DFFD65825FB0DFE49FA82DECD" box="[1223,1360,403,429]" name="Madagascar" pageId="3" pageNumber="527">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>