413 lines
54 KiB
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413 lines
54 KiB
XML
<document id="409B5512DD00C4512528E8B4C797D6FB" ID-CLB-Dataset="217329" ID-DOI="10.37520/aemnp.2022.017" ID-GBIF-Dataset="795cf556-a4a9-477b-b55d-cb80ed8a0f2d" ID-ISSN="1804-6487" ID-Zenodo-Dep="7399702" ID-ZooBank="28D55112-98B1-49A5-B382-58B1B068570B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670240514228" checkinUser="felipe" docAuthor="Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas & Solodovnikov, Alexey" docDate="2022" docId="038987A0FFD24B30B0EFA5381011FEEA" docLanguage="en" docName="ActaEntMusNatPra.62.1.225-299.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.017" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Quedius unicolor Kiesenwetter 1847" docType="treatment" docVersion="8" lastPageNumber="266" masterDocId="FFB0FFD8FFFA4B19B016A005151AFF9F" masterDocTitle="Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)" masterLastPageNumber="299" masterPageNumber="225" pageNumber="265" updateTime="1699412962807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="3031B5C7237FAAB2382A9E860D4D3217">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="56013656B471FD8EE9652F7D2AFBF981">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="A5C7F9B89050C41E19AAC2AEF7DA0AB9">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark & Department of Bioscience, University of Aarhus, Ny Munkegade 116, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:namePart id="F87434D027840FD011C6134DFA5EFAD2">Brunke, Adam</mods:namePart>
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<mods:affiliation id="8B455E61E39D712F9D5C1ACB91E95587">Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, Ontario, K</mods:affiliation>
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<mods:namePart id="D54CDBDC56DA3383C6B508587D7DA9E9">Simonsen, Thomas</mods:namePart>
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<mods:affiliation id="13A40AE9BCCF29C0DD69CE8A63B77004">Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:namePart id="6CF6C1AF5939BC29FD870EC4DD7721A2">Solodovnikov, Alexey</mods:namePart>
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<mods:affiliation id="8A9930D1181FC854E2FDF60B678900D6">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Zoological Institute, Russian Academy of Science, Universitetskaja nab.</mods:affiliation>
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<mods:title id="C84FFE910AF871F5F29A8F77B1F5C3FF">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<treatment id="038987A0FFD24B30B0EFA5381011FEEA" ID-DOI="http://doi.org/10.5281/zenodo.7503577" ID-GBIF-Taxon="208533234" ID-Zenodo-Dep="7503577" LSID="urn:lsid:plazi:treatment:038987A0FFD24B30B0EFA5381011FEEA" httpUri="http://treatment.plazi.org/id/038987A0FFD24B30B0EFA5381011FEEA" lastPageId="41" lastPageNumber="266" pageId="40" pageNumber="265">
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<subSubSection id="C33A653DFFD24B31B0EFA538178FFACA" box="[249,661,1341,1365]" pageId="40" pageNumber="265" type="nomenclature">
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<paragraph id="8B9F36B6FFD24B31B0EFA538178FFACA" blockId="40.[249,661,1341,1365]" box="[249,661,1341,1365]" pageId="40" pageNumber="265">
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<heading id="D0D781DAFFD24B31B0EFA538178FFACA" box="[249,661,1341,1365]" centered="true" fontSize="10" level="2" pageId="40" pageNumber="265" reason="3">
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<taxonomicName id="4C204D35FFD24B31B0EFA538178FFACA" ID-CoL="78Q9W" authority="Kiesenwetter, 1847" authorityName="Kiesenwetter" authorityYear="1847" box="[249,661,1341,1365]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="265" phylum="Arthropoda" rank="species" species="unicolor">
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<emphasis id="B954EAA4FFD24B31B0EFA538178FFACA" bold="true" box="[249,661,1341,1365]" pageId="40" pageNumber="265">
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<emphasis id="B954EAA4FFD24B31B0EFA53814A3FACA" bold="true" box="[249,441,1341,1365]" italics="true" pageId="40" pageNumber="265">Quedius unicolor</emphasis>
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<bibRefCitation id="EFB14B47FFD24B31B1D6A53B178FFACA" author="KIESENWETTER H. & MARKEL F." box="[448,661,1342,1365]" pageId="40" pageNumber="265" pagination="73 - 87" refId="ref85769" refString="KIESENWETTER H. ඏඈඇ & MARKEL F. 1847: Eine entomologische Excursion im Riesengebirge im Juli 1846 (Fortsetzung). Entomologische Zeitung (Stettin) 8: 73 - 87." type="journal article" year="1847">Kiesenwetter, 1847</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C33A653DFFD24B31B146A5591724FAF0" box="[336,574,1372,1391]" pageId="40" pageNumber="265" type="description">
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<paragraph id="8B9F36B6FFD24B31B146A5591724FAF0" blockId="40.[151,761,1372,1452]" box="[336,574,1372,1391]" pageId="40" pageNumber="265">
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(
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<figureCitation id="131B2A33FFD24B31B140A5591496FAF0" box="[342,396,1372,1391]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="40" pageNumber="265">Figs 1</figureCitation>
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,
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<figureCitation id="131B2A33FFD24B31B181A55914A9FAF0" box="[407,435,1372,1391]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="40" pageNumber="265">2G</figureCitation>
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,
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<figureCitation id="131B2A33FFD24B31B1ADA55914DDFAF0" box="[443,455,1372,1391]" captionStart="Fig" captionStartId="12.[151,182,2008,2027]" captionTargetBox="[182,1406,199,1981]" captionTargetId="figure-13@12.[182,1407,189,1989]" captionTargetPageId="12" captionText="Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80." figureDoi="http://doi.org/10.5281/zenodo.7399710" httpUri="https://zenodo.org/record/7399710/files/figure.png" pageId="40" pageNumber="265">4</figureCitation>
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,
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<figureCitation id="131B2A33FFD24B31B1C7A55914F0FAF0" box="[465,490,1372,1391]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="40" pageNumber="265">7E</figureCitation>
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,
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<figureCitation id="131B2A33FFD24B31B1E5A5591703FAF0" box="[499,537,1372,1391]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="40" pageNumber="265">14A</figureCitation>
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,
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<figureCitation id="131B2A33FFD24B31B234A5591722FAF0" box="[546,568,1372,1391]" captionStart="Fig" captionStartId="60.[151,182,2032,2051]" captionTargetBox="[193,1394,189,2003]" captionTargetId="figure-13@60.[193,1394,189,2003]" captionTargetPageId="60" captionText="Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +." figureDoi="http://doi.org/10.5281/zenodo.7399750" httpUri="https://zenodo.org/record/7399750/files/figure.png" pageId="40" pageNumber="265">21</figureCitation>
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)
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</paragraph>
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<subSubSection id="C33A653DFFD24B31B081A57B17C3F8F9" pageId="40" pageNumber="265" type="reference_group">
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<paragraph id="8B9F36B6FFD24B31B081A57B14A3FA33" blockId="40.[151,761,1372,1452]" pageId="40" pageNumber="265">
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<taxonomicName id="4C204D35FFD24B31B081A57B14BAFA0E" ID-CoL="78Q9W" authority="Kiesenwetter" authorityName="Kiesenwetter" authorityYear="1847" box="[151,416,1406,1425]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="265" phylum="Arthropoda" rank="species" species="unicolor">
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<emphasis id="B954EAA4FFD24B31B081A57B1431FA0E" box="[151,299,1406,1425]" italics="true" pageId="40" pageNumber="265">Quedius unicolor</emphasis>
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Kiesenwetter
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</taxonomicName>
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in Kංൾඌൾඇඐൾඍඍൾඋ & Mඟඋĸൾඅ (1847: 75). [
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<typeStatus id="549B8814FFD24B31B0ACA59C15FDFA33" box="[186,231,1433,1452]" pageId="40" pageNumber="265">Type</typeStatus>
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locality: Riesengebirge]
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</paragraph>
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<paragraph id="8B9F36B6FFD24B31B081A5C617C3F8F9" blockId="40.[151,761,1475,1894]" pageId="40" pageNumber="265">
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<emphasis id="B954EAA4FFD24B31B081A5C61418FA49" bold="true" box="[151,258,1475,1494]" pageId="40" pageNumber="265">References.</emphasis>
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Kංൾඌൾඇඐൾඍඍൾඋ (1848): 53 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 825, (1857): 202, (1874): 199 (characters); Fൺංඋආൺංඋൾ & Lൺൻඈඎඅ-ൻජඇൾ (1856):536 (characters); Fൺඎඏൾඅ (1874):291 (characters);Mඎඅඌൺඇඍ & Rൾඒ (1876): 697 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Pඈඋඍൺ (1907):132 (characters);Rൾංඍඍൾඋ (1909):112 (characters); Gඋංൽൾඅඅං (1924): 80 (characters); Pඈඋඍൾඏංඇ (1929): 341 (characters); WඳඌඍHඈൿൿ (1938): fig. 24 (characters); Sආൾඍൺඇൺ (1955): 143, (1960a): 259, (1964): 168, (1966): 331, (1993): 50 (distribution); (1958): 363 (characters and biology); (1962a): 134 (characters);Cඈංൿൿൺංඍ (1961): 49,(1978):193 (characters);LඈHඌൾ (1964):211 (characters); Hൺඏൾඅĸൺ (1964):93 (distribution); Hඈඋංඈඇ (1965): 274 (distribution); Bඈඋൽඈඇං (1976a): 94 (characters); ZൾඋർHൾ (1977): 9 (distribution), TඬඍH (1984): 119 (characters); LඎർHඍ (1987):109 (distribution);Jൺඇගĸ (1992):91 (distribution);Hൾඋආൺඇ (2001): 50 (application to ICZN), Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Mൺඓඎඋ et al. (2007): 30 (distribution); MൺඍൾඅൾඌHĸඈ (2007): 184 (distribution); SർHൺඍඓ (2008): 388 (distribution); Zൺඇൾඍඍං (2015): 96 (distribution).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A653DFFD24B31B081A77811ABFCBE" pageId="40" pageNumber="265" type="materials_examined">
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<paragraph id="8B9F36B6FFD24B31B081A77811ABFCBE" blockId="40.[151,761,1917,2069]" lastBlockId="40.[826,1438,189,801]" pageId="40" pageNumber="265">
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<emphasis id="B954EAA4FFD24B31B081A77814A9F80F" bold="true" box="[151,435,1917,1936]" pageId="40" pageNumber="265">
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Material examined.
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<collectingCountry id="F3377626FFD24B31B15DA77814B4F80F" box="[331,430,1917,1936]" name="Austria" pageId="40" pageNumber="265">AUSTRIA</collectingCountry>
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:
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</emphasis>
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Golling, [47.59, 13.16], leg. Skalitzky (
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<specimenCount id="9D26FD3FFFD24B31B0B6A79215A6F835" box="[160,188,1937,1962]" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B0D6A79215E0F835" box="[192,250,1943,1962]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
); Kerschbaumeralm, [46.75, 12.76],
|
||
<date id="FF9E1076FFD24B31B23FA79217C8F835" box="[553,722,1943,1962]" pageId="40" pageNumber="265" value="1948-07-24" valueMax="1948-08-07" valueMin="1948-07-24">24.VII.-7.VIII.1948</date>
|
||
, leg. F. Schubert (
|
||
<specimenCount id="9D26FD3FFFD24B31B11FA7B71431F85B" box="[265,299,1963,1989]" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B126A7B71470F85A" box="[304,362,1970,1989]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
); Koralpe, [46.88, 14.99], leg. A. Otto (3 JJ
|
||
<collectionCode id="ED31AE73FFD24B31B081A7C815CBF87F" box="[151,209,1997,2016]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
); Lünersee, [47.05, 9.74],
|
||
<date id="FF9E1076FFD24B31B1A2A7C8174BF87F" box="[436,593,1997,2016]" pageId="40" pageNumber="265" value="1952-06-14" valueMax="1952-07-03" valueMin="1952-06-14">14.VI.-3.VII.1952</date>
|
||
, leg. F. Schubert (
|
||
<specimenCount id="9D26FD3FFFD24B31B2F8A7C815B2F865" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B0BBA7E215FDF865" box="[173,231,2023,2042]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
); Stuhleck, [47.56, 15.78], leg. Breit (
|
||
<specimenCount id="9D26FD3FFFD24B31B239A7E21755F865" box="[559,591,2017,2042]" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B242A7E21783F865" box="[596,665,2023,2042]" pageId="40" pageNumber="265">NHMD</collectionCode>
|
||
); Stuhleck,
|
||
<collectingRegion id="49E4F854FFD24B31B081A80715ECF78A" box="[151,246,2050,2069]" country="Austria" name="Steiermark" pageId="40" pageNumber="265">Steiermark</collectingRegion>
|
||
, [47.56, 15.78], leg.Wingelmüller (
|
||
<specimenCount id="9D26FD3FFFD24B31B237A8071727F78B" box="[545,573,2043,2069]" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B254A8071766F78A" box="[578,636,2050,2069]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
);
|
||
<collectingRegion id="49E4F854FFD24B31B29FA80717AFF78A" box="[649,693,2050,2069]" country="Austria" name="Tirol" pageId="40" pageNumber="265">Tirol</collectingRegion>
|
||
, [47.22, 11.51] (1
|
||
<collectionCode id="ED31AE73FFD24B31B39CA0C616A6FF49" box="[906,956,195,214]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
); Wechsel,[47.52, 15.91], leg. Scheerpeltz (
|
||
<specimenCount id="9D26FD3FFFD24B31B53EA0C61049FF49" box="[1320,1363,189,214]" pageId="40" pageNumber="265" type="female">3♀♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B54FA0C61089FF49" box="[1369,1427,195,214]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
); Wolayersee, [46.61, 12.86],
|
||
<date id="FF9E1076FFD24B31B424A0DB11A3FF6E" box="[1074,1209,222,241]" pageId="40" pageNumber="265" value="1949-08-03" valueMax="1949-08-11" valueMin="1949-08-03">3.-11.VIII.1949</date>
|
||
, leg. F. Schubert (2 JJ
|
||
<specimenCount id="9D26FD3FFFD24B31B584A0DB1654FE94" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B345A0FC1682FE93" box="[851,920,249,268]" pageId="40" pageNumber="265">NHMD</collectionCode>
|
||
); Zillertal nr. Mayrhofen, [47.16, 11.87],
|
||
<date id="FF9E1076FFD24B31B51CA0FC1068FE93" box="[1290,1394,249,268]" pageId="40" pageNumber="265" value="1914-08-04">4.VIII.1914</date>
|
||
, leg. H. Wagner (1
|
||
<collectionCode id="ED31AE73FFD24B31B3AFA11616F6FEB9" box="[953,1004,275,294]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
); Wolayersee, [46.61, 12.86], 3.11.
|
||
<date id="FF9E1076FFD24B31B50DA1161069FEB9" box="[1307,1395,275,294]" pageId="40" pageNumber="265" value="1949-08">VIII.1949</date>
|
||
, leg. F. Schubert (
|
||
<specimenCount id="9D26FD3FFFD24B31B3B8A12B16FAFEDE" box="[942,992,296,321]" pageId="40" pageNumber="265" type="female">2 ♀♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B3FEA12B1138FEDE" box="[1000,1058,302,321]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B425A12B11E7FEDE" bold="true" box="[1075,1277,302,321]" pageId="40" pageNumber="265">
|
||
<collectingCountry id="F3377626FFD24B31B425A12B11E3FEDE" box="[1075,1273,302,321]" name="Czech Republic" pageId="40" pageNumber="265">CZECH REPUBLIC</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Krkonoše [50.71, 15.66], leg. Obenberger (1 J
|
||
<specimenCount id="9D26FD3FFFD24B31B423A14C1179FEC4" box="[1077,1123,322,348]" pageId="40" pageNumber="265" type="female">2 ♀♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B47FA14C11B4FEC3" box="[1129,1198,329,348]" pageId="40" pageNumber="265">NHMD</collectionCode>
|
||
); Spindelmühle [Špindlerův Mlýn], [50.73, 15.61], leg. Skalitzky (1
|
||
<collectionCode id="ED31AE73FFD24B31B48AA16611FBFEE9" box="[1180,1249,355,374]" pageId="40" pageNumber="265">NHMD</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B4E7A166107DFEE9" bold="true" box="[1265,1383,355,374]" pageId="40" pageNumber="265">
|
||
<collectingCountry id="F3377626FFD24B31B4E7A1661079FEE9" box="[1265,1379,355,374]" name="Germany" pageId="40" pageNumber="265">GERMANY</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Harz, Brocken, [51.79, 10.61],
|
||
<date id="FF9E1076FFD24B31B40EA17B117DFE0E" box="[1048,1127,382,401]" pageId="40" pageNumber="265" value="1922-05-09">9.V.1922</date>
|
||
, leg. Uhmann (1 J
|
||
<collectionCode id="ED31AE73FFD24B31B50CA17B1057FE0E" box="[1306,1357,382,401]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
); Hercyn mont.[Harz Mountains], [51.74, 10.63],
|
||
<date id="FF9E1076FFD24B31B49BA19C11F5FE33" box="[1165,1263,409,428]" pageId="40" pageNumber="265" value="1914-06-14">14.VI.1914</date>
|
||
, leg. Beckenkamp (
|
||
<specimenCount id="9D26FD3FFFD24B31B584A19C1643FE59" pageId="40" pageNumber="265" type="female">3 ♀♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B376A1B61689FE59" box="[864,915,435,454]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
); Thüringer Wald Oberhof, [50.71, 10.73],
|
||
<date id="FF9E1076FFD24B31B516A1B61042FE59" box="[1280,1368,435,454]" pageId="40" pageNumber="265" value="1951-09-08">8.IX.1951</date>
|
||
, leg. W. Liebmann (
|
||
<specimenCount id="9D26FD3FFFD24B31B389A1CB16A6FE7E" box="[927,956,456,481]" pageId="40" pageNumber="265" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFD24B31B3D7A1CB16EEFE7E" box="[961,1012,462,481]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B417A1CB115DFE7E" bold="true" box="[1025,1095,462,481]" pageId="40" pageNumber="265">
|
||
<collectingCountry id="F3377626FFD24B31B417A1CB1159FE7E" box="[1025,1091,462,481]" name="Italy" pageId="40" pageNumber="265">ITALY</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Gressory [Gressoney] la Trinite, [45.87, 7.82],
|
||
<date id="FF9E1076FFD24B31B363A1EC16C5FE63" box="[885,991,489,508]" pageId="40" pageNumber="265" value="1901-07-15">15.VII.1901</date>
|
||
, leg. Künnemann (1
|
||
<collectionCode id="ED31AE73FFD24B31B48CA1EC11D7FE63" box="[1178,1229,489,508]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
); Malga Preghena Alta, Trento,
|
||
<geoCoordinate id="EE145071FFD24B31B36BA20616DFFD89" box="[893,965,515,534]" degrees="46.4148" direction="north" orientation="latitude" pageId="40" pageNumber="265" precision="5" value="46.4148">46.4148</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145071FFD24B31B3DAA2061109FD89" box="[972,1043,515,534]" degrees="10.8922" direction="east" orientation="longitude" pageId="40" pageNumber="265" precision="5" value="10.8922">10.8922</geoCoordinate>
|
||
,
|
||
<quantity id="4CD89B53FFD24B31B40CA2061146FD89" box="[1050,1116,515,534]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="40" pageNumber="265" unit="m" value="2100.0">2100 m</quantity>
|
||
, under stones along stream with moss,
|
||
<date id="FF9E1076FFD24B31B32AA21B16B1FDAE" box="[828,939,542,561]" pageId="40" pageNumber="265" value="2019-08-26">26.VIII.2019</date>
|
||
, leg.A.Zanetti (
|
||
<collectionCode id="ED31AE73FFD24B31B426A21B116FFDAE" box="[1072,1141,542,561]" pageId="40" pageNumber="265">NHMD</collectionCode>
|
||
);Valle Brembana, Strada di Porcile, [46.06, 9.73],
|
||
<quantity id="4CD89B53FFD24B31B3A2A23C1132FDD3" box="[948,1064,569,588]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="40" pageNumber="265" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800-2000 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFD24B31B439A23C1189FDD3" box="[1071,1171,569,588]" pageId="40" pageNumber="265" value="1961-06-26">26.VI.1961</date>
|
||
, leg. V. Rosa (1 J
|
||
<collectionCode id="ED31AE73FFD24B31B526A23C1070FDD3" box="[1328,1386,569,588]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="265">NMW</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B561A23C1665FDF9" bold="true" pageId="40" pageNumber="265">PO-LAND:</emphasis>
|
||
Silesia [Bad] Flinsberg [Świeradów-Zdrój], [50.92, 15.31], leg. Kraatz (1
|
||
<collectionCode id="ED31AE73FFD24B31B387A26B16DEFD1E" box="[913,964,622,641]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B3C4A26B1102FD1E" bold="true" box="[978,1048,622,641]" pageId="40" pageNumber="265">
|
||
<collectingCountry id="F3377626FFD24B31B3C4A26B110EFD1E" box="[978,1044,622,641]" name="Spain" pageId="40" pageNumber="265">SPAIN</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="49E4F854FFD24B31B40BA26B1168FD1E" box="[1053,1138,622,641]" country="Spain" name="Cantabria" pageId="40" pageNumber="265">Cantabria</collectingRegion>
|
||
, Alto Campoo, W Reinosa, [43.03, -4.37],
|
||
<date id="FF9E1076FFD24B31B36CA28C16CAFD03" box="[890,976,649,668]" pageId="40" pageNumber="265" value="1991-06-04">4.VI.1991</date>
|
||
<quantity id="4CD89B53FFD24B31B3C3A28C110BFD03" box="[981,1041,649,668]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="265" unit="m" value="2000.0">2000m</quantity>
|
||
leg. Zerche (1
|
||
<collectionCode id="ED31AE73FFD24B31B487A28C11DEFD03" box="[1169,1220,649,668]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B4C4A28C1052FD03" bold="true" box="[1234,1352,649,668]" pageId="40" pageNumber="265">
|
||
<collectingCountry id="F3377626FFD24B31B4C4A28C105EFD03" box="[1234,1348,649,668]" name="Slovakia" pageId="40" pageNumber="265">SLOVAKIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Ružomberok, [49.04, 19.23],
|
||
<date id="FF9E1076FFD24B31B3E6A2A6115AFD29" box="[1008,1088,675,694]" pageId="40" pageNumber="265" value="1952-07">VII.1952</date>
|
||
, leg. Přívora (1 J
|
||
<collectionCode id="ED31AE73FFD24B31B4C9A2A6103EFD29" box="[1247,1316,675,694]" pageId="40" pageNumber="265">NHMD</collectionCode>
|
||
); High Tatras, Mengušovská dolina, Vysoke Tatry, [49.16, 20.07],
|
||
<quantity id="4CD89B53FFD24B31B4E2A2BB102DFD4E" box="[1268,1335,702,721]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="40" pageNumber="265" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFD24B31B556A2BB1083FD4E" box="[1344,1433,702,721]" pageId="40" pageNumber="265" value="1991-05-17">17.V.1991</date>
|
||
, leg. J. Frisch (1 J
|
||
<collectionCode id="ED31AE73FFD24B31B3C8A2DC1105FD73" box="[990,1055,729,748]" pageId="40" pageNumber="265">ZMHB</collectionCode>
|
||
); Tatra, Kráľova hoľa, [48.88, 20.13],
|
||
<quantity id="4CD89B53FFD24B31B57CA2DC1698FC99" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="40" pageNumber="265" unit="m" value="1450.0" valueMax="1600.0" valueMin="1300.0">1300- 1600 m</quantity>
|
||
,
|
||
<date id="FF9E1076FFD24B31B39BA2F6111AFC99" box="[909,1024,755,774]" pageId="40" pageNumber="265" value="1981-08-12">12.VIII.1981</date>
|
||
, leg. Hieke (1 J
|
||
<collectionCode id="ED31AE73FFD24B31B4B7A2F611F9FC99" box="[1185,1251,755,774]" pageId="40" pageNumber="265">ZMHB</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFD24B31B4E2A2F61086FC99" bold="true" box="[1268,1436,755,774]" pageId="40" pageNumber="265">
|
||
<collectingCountry id="F3377626FFD24B31B4E2A2F61082FC99" box="[1268,1432,755,774]" name="Switzerland" pageId="40" pageNumber="265">SWITZERLAND</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Aarau, [47.38, 8.05], leg. Stierlin (1
|
||
<collectionCode id="ED31AE73FFD24B31B465A30B11BCFCBE" box="[1139,1190,782,801]" pageId="40" pageNumber="265">SDEI</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFD24B31B32AA33F1023F78E" pageId="40" pageNumber="265" type="description">
|
||
<paragraph id="8B9F36B6FFD24B31B32AA33F108DFB2E" blockId="40.[828,1437,820,2065]" pageId="40" pageNumber="265">
|
||
<emphasis id="B954EAA4FFD24B31B32AA33F16FBFCCE" bold="true" box="[828,993,826,849]" pageId="40" pageNumber="265">Redescription.</emphasis>
|
||
Measurements JJ (n = 5): HW = 1.47– 1.76 (1.58); HL = 1.31–1.44 (1.36); HL/HW 0.82–0.91 (0.86); PW = 1.96–2.33 (2.16); PL = 1.78–2.02 (1.91); PL/PW 0.86–0.92 (0.89); EW = 2.13–2.47 (2.32); EL = 2.18–2.47 (2.32); EL/EW 0.95–1.05 (1.00); EL/PL 1.15– 1.26 (1.22); PW/HW 1.47–1.65 (1.58); forebody length 5.36–5.93 (5.58).
|
||
<specimenCount id="9D26FD3FFFD24B31B417A3F6113DFB8D" box="[1025,1063,1011,1042]" pageId="40" pageNumber="265" type="female">♀♀</specimenCount>
|
||
(n = 5): HW = 1.58–1.67 (1.61); HL = 1.31–1.42 (1.37); HL/HW 0.79–0.89 (0.85); PW = 2.00–2.18 (2.08); PL = 1.82–1.89 (1.87); PL/PW 0.87–0.92 (0.90); EW = 2.22–2.40 (2.31); EL = 2.20–2.40 (2.31); EL/EW 0.96–1.01 (0.99); EL/PL 1.19–1.27 (1.23); PW/ HW 1.45–1.57 (1.52); forebody length 5.36–5.69 (5.52).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFD24B31B34CA4BF102DFB4E" blockId="40.[828,1437,820,2065]" box="[858,1335,1210,1233]" pageId="40" pageNumber="265">
|
||
Medium sized species; body black (
|
||
<figureCitation id="131B2A33FFD24B31B4CEA4BF1031FB4E" box="[1240,1323,1210,1233]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="40" pageNumber="265">Fig. 7E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFD24B31B34CA4DF114FFA0E" blockId="40.[828,1437,820,2065]" pageId="40" pageNumber="265">
|
||
Head black, distinctly transverse, with eyes rather small (EyL/TL = 1.59–2.00 (1.82)); microsculpture of fine mesh, especially on frons, appears dull; no interocular punctures between anterior frontal punctures (cf.
|
||
<figureCitation id="131B2A33FFD24B31B4CCA53F1030FACE" box="[1242,1322,1338,1361]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="40" pageNumber="265">Fig. 6F</figureCitation>
|
||
); antennae dark brown to black, all antennomeres slightly elongate; palpi dark brown to black.
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFD24B31B34CA59F1186F90E" blockId="40.[828,1437,820,2065]" pageId="40" pageNumber="265">Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of fine transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum punctured and pubescent; elytra black, uniformly pubescent, as long as wide, clearly longer than prontoum; legs dark, outer face of tibia and tarsi often paler.</paragraph>
|
||
<paragraph id="8B9F36B6FFD24B31B34CA69F16ECF94E" blockId="40.[828,1437,820,2065]" pageId="40" pageNumber="265">Abdomen black, tergites uniformly punctured, without clear iridescence.</paragraph>
|
||
<paragraph id="8B9F36B6FFD24B31B34CA6DC1023F78E" blockId="40.[828,1437,820,2065]" pageId="40" pageNumber="265">
|
||
<emphasis id="B954EAA4FFD24B31B34CA6DC1683F96E" bold="true" box="[858,921,1753,1777]" italics="true" pageId="40" pageNumber="265">Male.</emphasis>
|
||
Aedeagus (
|
||
<figureCitation id="131B2A33FFD24B31B401A6DF1161F96E" box="[1047,1147,1754,1777]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="40" pageNumber="265">Fig. 14A</figureCitation>
|
||
): paramere lanceolate with slight medial attenuation and extending into a slight expansion broadest below apex, apex extending into small blunt knob away from the median lobe, apex slightly asymmetric, reaching just beyond apex of median lobe, with sensory peg setae forming two rows fusing together towards apex; median lobe broad with gentle constriction to a point at apex, on parameral side with two small ridges continuing basad, positioned at level near basal level of peg setae band of the paramere; internal sac without C-sclerite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFD34B30B081A0BB15F6FC8A" pageId="41" pageNumber="266" type="diagnosis">
|
||
<paragraph id="8B9F36B6FFD34B30B081A0BB14C1FD2A" blockId="41.[151,761,190,2069]" pageId="41" pageNumber="266">
|
||
<emphasis id="B954EAA4FFD34B30B081A0BB148EFF4A" bold="true" box="[151,404,190,213]" pageId="41" pageNumber="266">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C204D35FFD34B30B18CA0BB174DFF4A" authorityName="Kiesenwetter" authorityYear="1847" box="[410,599,190,213]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B18CA0BB174DFF4A" box="[410,599,190,213]" italics="true" pageId="41" pageNumber="266">Quedius unicolor</emphasis>
|
||
</taxonomicName>
|
||
is very similar to other species with dark elytra from the
|
||
<emphasis id="B954EAA4FFD34B30B267A0DB1584FE8A" italics="true" pageId="41" pageNumber="266">molochinus-</emphasis>
|
||
group –
|
||
<taxonomicName id="4C204D35FFD34B30B113A0FB14ABFE8A" authorityName="Korge" authorityYear="1961" box="[261,433,254,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="subunicolor">
|
||
<emphasis id="B954EAA4FFD34B30B113A0FB14ABFE8A" box="[261,433,254,277]" italics="true" pageId="41" pageNumber="266">Q. subunicolor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C204D35FFD34B30B1D2A0FB1741FE8A" authorityName="Smetana" authorityYear="2003" box="[452,603,254,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="sundukovi">
|
||
<emphasis id="B954EAA4FFD34B30B1D2A0FB1741FE8A" box="[452,603,254,277]" italics="true" pageId="41" pageNumber="266">Q. sundukovi</emphasis>
|
||
</taxonomicName>
|
||
and the dark form of
|
||
<taxonomicName id="4C204D35FFD34B30B0E4A11B148BFEAA" box="[242,401,286,309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="molochinus">
|
||
<emphasis id="B954EAA4FFD34B30B0E4A11B148BFEAA" box="[242,401,286,309]" italics="true" pageId="41" pageNumber="266">Q. molochinus</emphasis>
|
||
</taxonomicName>
|
||
. It can be distinguished from
|
||
<taxonomicName id="4C204D35FFD34B30B2CBA11B1401FECA" authorityName="Korge" authorityYear="1961" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="subunicolor">
|
||
<emphasis id="B954EAA4FFD34B30B2CBA11B1401FECA" italics="true" pageId="41" pageNumber="266">Q. subunicolor</emphasis>
|
||
</taxonomicName>
|
||
by the dull frons due to meshed microsculpture (in
|
||
<taxonomicName id="4C204D35FFD34B30B109A15B14DEFEEA" authorityName="Korge" authorityYear="1961" box="[287,452,350,373]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="subunicolor">
|
||
<emphasis id="B954EAA4FFD34B30B109A15B14DEFEEA" box="[287,452,350,373]" italics="true" pageId="41" pageNumber="266">Q. subunicolor</emphasis>
|
||
</taxonomicName>
|
||
the microsculpture consists of fine transverse waves or only slightly meshed). Also, the genitalia and the distributions are clearly different in the two species. It usually differs from the dark form of
|
||
<taxonomicName id="4C204D35FFD34B30B0A3A1DB1449FE6A" box="[181,339,478,501]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="molochinus">
|
||
<emphasis id="B954EAA4FFD34B30B0A3A1DB1449FE6A" box="[181,339,478,501]" italics="true" pageId="41" pageNumber="266">Q. molochinus</emphasis>
|
||
</taxonomicName>
|
||
in the darker appendages and the finer microsculpture in addition to a clearly different aedagus, which is without a C-sclerite in
|
||
<taxonomicName id="4C204D35FFD34B30B1F2A21B1745FDAA" authorityName="Kiesenwetter" authorityYear="1847" box="[484,607,542,565]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B1F2A21B1745FDAA" box="[484,607,542,565]" italics="true" pageId="41" pageNumber="266">Q. unicolor</emphasis>
|
||
</taxonomicName>
|
||
, but with such in
|
||
<emphasis id="B954EAA4FFD34B30B0A4A23B170CFDCA" box="[178,534,574,597]" italics="true" pageId="41" pageNumber="266">
|
||
<taxonomicName id="4C204D35FFD34B30B0A4A23B144BFDCA" box="[178,337,574,597]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="molochinus">Q. molochinus</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C204D35FFD34B30B14CA23B170CFDCA" authorityName="Kiesenwetter" authorityYear="1847" box="[346,534,574,597]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">Quedius unicolor</taxonomicName>
|
||
</emphasis>
|
||
has non-overlapping distribution with
|
||
<taxonomicName id="4C204D35FFD34B30B140A25B173CFDEA" authorityName="Smetana" authorityYear="2003" box="[342,550,606,629]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="sundukovi">
|
||
<emphasis id="B954EAA4FFD34B30B140A25B173CFDEA" box="[342,550,606,629]" italics="true" pageId="41" pageNumber="266">Quedius sundukovi</emphasis>
|
||
</taxonomicName>
|
||
and can be clearly distinguished from the latter species by the presence of palisade fringe on tergite VII.
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFD34B30B0A3A2BB15F6FC8A" blockId="41.[151,761,190,2069]" pageId="41" pageNumber="266">
|
||
<taxonomicName id="4C204D35FFD34B30B0A3A2BB146BFD4A" authorityName="Kiesenwetter" authorityYear="1847" box="[181,369,702,725]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B0A3A2BB146BFD4A" box="[181,369,702,725]" italics="true" pageId="41" pageNumber="266">Quedius unicolor</emphasis>
|
||
</taxonomicName>
|
||
has historically been confused with
|
||
<taxonomicName id="4C204D35FFD34B30B081A2DB142DFD6A" authorityName="Korge" authorityYear="1961" box="[151,311,734,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="subunicolor">
|
||
<emphasis id="B954EAA4FFD34B30B081A2DB142DFD6A" box="[151,311,734,757]" italics="true" pageId="41" pageNumber="266">Q. subunicolor</emphasis>
|
||
</taxonomicName>
|
||
. For details see comments about the latter species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFD34B30B081A31B17E3F9EA" pageId="41" pageNumber="266" type="discussion">
|
||
<paragraph id="8B9F36B6FFD34B30B081A31B17E3F9EA" blockId="41.[151,761,190,2069]" pageId="41" pageNumber="266">
|
||
<emphasis id="B954EAA4FFD34B30B081A31B1403FCAA" bold="true" box="[151,281,798,821]" pageId="41" pageNumber="266">Comments.</emphasis>
|
||
<taxonomicName id="4C204D35FFD34B30B135A31B14F8FCAA" authorityName="Kiesenwetter" authorityYear="1847" box="[291,482,798,821]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B135A31B14F8FCAA" box="[291,482,798,821]" italics="true" pageId="41" pageNumber="266">Quedius unicolor</emphasis>
|
||
</taxonomicName>
|
||
was originally described from Riesengebirge [Krkonoše] (Kංൾඌൾඇඐൾඍඍൾඋ & Mඟඋĸൾඅ 1847), a mountain range at the border between
|
||
<collectingCountry id="F3377626FFD34B30B2BBA35B17EDFCEA" box="[685,759,862,885]" name="Poland" pageId="41" pageNumber="266">Poland</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3377626FFD34B30B0DCA37B149BFC0A" box="[202,385,894,917]" name="Czech Republic" pageId="41" pageNumber="266">Czech Republic</collectingCountry>
|
||
. It was later discovered in other mountainous area, including the Alps, the Carpathians, the Pyrenees, Picos de Europa, the Harz Mountains, and possibly extending into Sierra Nevada and even central
|
||
<collectingCountry id="F3377626FFD34B30B081A3FB15ECFB8A" box="[151,246,1022,1045]" name="Turkey" pageId="41" pageNumber="266">Anatolia</collectingCountry>
|
||
, although the latter records are not confirmed here (Cඈංൿൿൺංඍ 1961, AൻൺർංGංඅ et. al 2009;
|
||
<figureCitation id="131B2A33FFD34B30B261A41B17D3FBAA" box="[631,713,1054,1077]" captionStart="Fig" captionStartId="60.[151,182,2032,2051]" captionTargetBox="[193,1394,189,2003]" captionTargetId="figure-13@60.[193,1394,189,2003]" captionTargetPageId="60" captionText="Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +." figureDoi="http://doi.org/10.5281/zenodo.7399750" httpUri="https://zenodo.org/record/7399750/files/figure.png" pageId="41" pageNumber="266">Fig. 21</figureCitation>
|
||
). At these sites it seems to be restricted to a very specific habitat
|
||
<typeStatus id="549B8814FFD34B30B081A45B15D2FBEA" box="[151,200,1118,1141]" pageId="41" pageNumber="266">type</typeStatus>
|
||
, with most specimens being found under stones and in moss near mountain streams at higher elevations. COI barcodes of specimens from several sites, including the Austrian and Italian Alps, the Carpathians of
|
||
<collectingCountry id="F3377626FFD34B30B27CA4BB17D3FB4A" box="[618,713,1214,1237]" name="Slovakia" pageId="41" pageNumber="266">Slovakia</collectingCountry>
|
||
, and Picos de Europa in Spain displayed high genetic variability (
|
||
<figureCitation id="131B2A33FFD34B30B089A4FB15FAFA8A" box="[159,224,1278,1301]" captionStart="Fig" captionStartId="12.[151,182,2008,2027]" captionTargetBox="[182,1406,199,1981]" captionTargetId="figure-13@12.[182,1407,189,1989]" captionTargetPageId="12" captionText="Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80." figureDoi="http://doi.org/10.5281/zenodo.7399710" httpUri="https://zenodo.org/record/7399710/files/figure.png" pageId="41" pageNumber="266">Fig. 4</figureCitation>
|
||
), with the sites being grouped into three OTUs (the Carpathians, Picos de Europa, and the Alps) ranging from 3.5 % to 4.5 % divergence, with a 0.2% divergence between the Austrian and Italian Alps (
|
||
<tableCitation id="C6A2030DFFD34B30B1CFA55B1734FAEA" box="[473,558,1374,1397]" captionStart="Table 2" captionStartId="13.[151,201,190,209]" captionTargetPageId="13" captionText="Table 2. Genetic distance between clusters of CO1 barcodes.n = number of specimens barcoded. Number of OTUs found by species delimiter plugin in Geneious. Intraspecific distance within cluster. Interspecific distance is distance to nearest other cluster." httpUri="http://table.plazi.org/id/DF5F663EFFF74B14B081A0BB1713FEBE" pageId="41" pageNumber="266" tableUuid="DF5F663EFFF74B14B081A0BB1713FEBE">Table 2</tableCitation>
|
||
). With such a high genetic divergence in the barcode region, we carefully checked for morphological differences. However, we found no obvious differences corresponding to the molecular clusters. Since the wings seem to be fully developed in this species, these genetic differences may be attributed to isolation by distance, the very specific habitat requirements of the species and its close association to a now fragmented habitat, which would limit gene flow among populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFD34B30B081A67B1474F8AA" pageId="41" pageNumber="266" type="description">
|
||
<paragraph id="8B9F36B6FFD34B30B081A67B1474F8AA" blockId="41.[151,761,190,2069]" pageId="41" pageNumber="266">
|
||
<emphasis id="B954EAA4FFD34B30B081A67B140FF90A" bold="true" box="[151,277,1662,1685]" pageId="41" pageNumber="266">Bionomics.</emphasis>
|
||
<taxonomicName id="4C204D35FFD34B30B10CA67B14C8F90A" authorityName="Kiesenwetter" authorityYear="1847" box="[282,466,1662,1685]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B10CA67B14C8F90A" box="[282,466,1662,1685]" italics="true" pageId="41" pageNumber="266">Quedius unicolor</emphasis>
|
||
</taxonomicName>
|
||
seems to be very habitat selective and is only found in alpine sites in sphagnum moss and under rocks near bogs, swamps and creeks (
|
||
<figureCitation id="131B2A33FFD34B30B280A6BB17F6F94A" box="[662,748,1726,1749]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="41" pageNumber="266">Fig. 2G</figureCitation>
|
||
). It is found at altitudes between 900 and
|
||
<quantity id="4CD89B53FFD34B30B223A6DB179EF96A" box="[565,644,1758,1781]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="41" pageNumber="266" unit="m" value="2300.0">2300 m</quantity>
|
||
depending on the latitude of the locality, with higher elevations at more southern sites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFD34B30B081A73B1011FEEA" pageId="41" pageNumber="266" type="distribution">
|
||
<paragraph id="8B9F36B6FFD34B30B081A73B1011FEEA" blockId="41.[151,761,190,2069]" lastBlockId="41.[828,1437,190,373]" pageId="41" pageNumber="266">
|
||
<emphasis id="B954EAA4FFD34B30B081A73B143CF8CA" bold="true" box="[151,294,1854,1877]" pageId="41" pageNumber="266">Distribution.</emphasis>
|
||
<taxonomicName id="4C204D35FFD34B30B13DA73B14FAF8CA" authorityName="Kiesenwetter" authorityYear="1847" box="[299,480,1854,1877]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B13DA73B14FAF8CA" box="[299,480,1854,1877]" italics="true" pageId="41" pageNumber="266">Quedius unicolor</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the montane and subalpine zone of Central and southern Europe (
|
||
<figureCitation id="131B2A33FFD34B30B2DBA75B15AFF80A" captionStart="Fig" captionStartId="60.[151,182,2032,2051]" captionTargetBox="[193,1394,189,2003]" captionTargetId="figure-13@60.[193,1394,189,2003]" captionTargetPageId="60" captionText="Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +." figureDoi="http://doi.org/10.5281/zenodo.7399750" httpUri="https://zenodo.org/record/7399750/files/figure.png" pageId="41" pageNumber="266">Fig. 21</figureCitation>
|
||
). Specifically, it is known from sites in the Carpathians, the Alps, the Pyrenees, Picos de Europa, the Sudetes, the Harz and the Ore mountain ranges. Due to its very specific habitat requirement, it may be under-collected and may actually be present in mountains further south. Cඈංൿൿൺංඍ (1961) reported it from Sierra Nevada in Southern Spain, but this record may be in fact a misidentified
|
||
<taxonomicName id="4C204D35FFD34B30B511A0DB108FFF6A" authorityName="Bernhauer" authorityYear="1898" box="[1287,1429,222,245]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="hispanicus">
|
||
<emphasis id="B954EAA4FFD34B30B511A0DB108FFF6A" box="[1287,1429,222,245]" italics="true" pageId="41" pageNumber="266">Q. hispanicus</emphasis>
|
||
</taxonomicName>
|
||
. Another doubtful record of
|
||
<taxonomicName id="4C204D35FFD34B30B440A0FB11D7FE8A" authorityName="Kiesenwetter" authorityYear="1847" box="[1110,1229,254,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="266" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B954EAA4FFD34B30B440A0FB11D7FE8A" box="[1110,1229,254,277]" italics="true" pageId="41" pageNumber="266">Q. unicolor</emphasis>
|
||
</taxonomicName>
|
||
comes from Koçere Stream in the Kazdağları Mountain of the North-western
|
||
<collectingCountry id="F3377626FFD34B30B32AA13B1682FECA" box="[828,920,318,341]" name="Turkey" pageId="41" pageNumber="266">Anatolia</collectingCountry>
|
||
by AൻൺർංGංඅ et. al (2009), which is a location remote from the main range of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |