339 lines
36 KiB
XML
339 lines
36 KiB
XML
<document id="A9003EA39ABC17520C1151A3EDBD0A4C" ID-DOI="http://dx.doi.org/10.3897/zookeys.34.268" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio & Munetoshi Maruyama" docDate="2010" docId="038951095C1A3D0466B6FF63ABB6FD92" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Pterorthochaetes tsurui Ballerio & Maruyama, 2010, sp. n." docType="treatment" docVersion="10" lastPageNumber="101" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageNumber="95" updateTime="1732486157858" updateUser="ExternalLinkService">
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<mods:mods id="F18FE53F128D6BE7C9BC155D77560754" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="33C9723A12D39CB1618AE30AD62B82D1">
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<mods:title id="94ADD09F85F4B3A23C18FD0892CCAE87">The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2AE5EB2284B6B2BD7FA417FD5084825B" type="personal">
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<mods:role id="71309A282E5C89A5F3C20876DA0E67AF">
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<mods:roleTerm id="247D0D3243B339F7883873DFBF3D15C2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="53B531407685FD1C543636FDE8F7A35E">Alberto Ballerio</mods:namePart>
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</mods:name>
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<mods:name id="579D613768FC1BA2975B7393F760437F" type="personal">
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<mods:role id="214FA04D6AEB6B43423F1DE334237BC4">
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<mods:roleTerm id="CE29BA061D0F1624D06A722B612CE4BF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B83EE65D44BA62F8C9F2F14FB247AD8F">Munetoshi Maruyama</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4A9BFBFE447BB307172B6356CE682D4B">text</mods:typeOfResource>
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<mods:relatedItem id="B8F5CDC8E0AED05D6D24E56ACBAFC622" type="host">
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<mods:titleInfo id="5A8A0BFD3113F3837DDF387AB74B3F4A">
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<mods:title id="D05AE4BDE2D09D98436B5CB760DE8415">Zookeys</mods:title>
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</mods:titleInfo>
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<mods:part id="4EEDAACF387B80A9B1690584CF0F67CB">
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<mods:date id="5C53FCC78816359C549B989AF950A715">2010</mods:date>
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<mods:detail id="1325420B511B6C14EC6F3A6C2127E12E" type="volume">
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<mods:number id="838FD64A66F0960ED6CEE1196B64AC11">34</mods:number>
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</mods:detail>
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<mods:extent id="1BCEE767167ECE393A7F866F281D385E" unit="page">
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<mods:start id="1998DCED9FAEDD3738F058CF8E6632BB">77</mods:start>
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<mods:end id="E6521B81D60BCF47143F39FA8F09B1D8">104</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="E82719FC3DD2811222C8F20FC7C10963">journal article</mods:classification>
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<mods:identifier id="C23D09168526E7979D9A3C498EBA4B93" type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
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</mods:mods>
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<treatment id="038951095C1A3D0466B6FF63ABB6FD92" ID-DOI="http://doi.org/10.5281/zenodo.6210813" ID-GBIF-Taxon="114923704" ID-Zenodo-Dep="6210813" LSID="urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0" httpUri="http://treatment.plazi.org/id/038951095C1A3D0466B6FF63ABB6FD92" lastPageId="24" lastPageNumber="101" pageId="18" pageNumber="95" scope_family="Hybosoridae" scope_order="Coleoptera" scope_superFamily="Scarabaeoidea">
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<subSubSection id="C33AB3945C1A3D0E66B6FF63A9EAFF17" box="[140,488,167,193]" pageId="18" pageNumber="95" type="nomenclature">
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<paragraph id="8B9FE01F5C1A3D0E66B6FF63A9EAFF17" box="[140,488,167,193]" pageId="18" pageNumber="95">
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<heading id="D0D757735C1A3D0E66B6FF63A9EAFF17" bold="true" box="[140,488,167,193]" fontSize="11" level="1" pageId="18" pageNumber="95" reason="6">
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<emphasis id="B9543C0D5C1A3D0E66B6FF63A9EAFF17" bold="true" box="[140,488,167,193]" pageId="18" pageNumber="95">
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<taxonomicName id="4C209B9C5C1A3D0E66B6FF63A9A2FF17" ID-CoL="4Q36G" LSID-ZBK="urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0 " box="[140,416,167,193]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
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<emphasis id="B9543C0D5C1A3D0E66B6FF63A9A2FF17" bold="true" box="[140,416,167,193]" italics="true" pageId="18" pageNumber="95">Pterorthochaetes tsurui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26781765C1A3D0E679DFF63A9EAFF17" box="[423,488,167,193]" pageId="18" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<paragraph id="8B9FE01F5C1A3D0E66B6FF0DA91BFED1" pageId="18" pageNumber="95">
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<subSubSection id="C33AB3945C1A3D0E66B6FF0DABBDFF32" box="[140,959,201,228]" pageId="18" pageNumber="95" type="description">urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0</subSubSection>
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<subSubSection id="C33AB3945C1A3D0E66B6FF29A91BFED1" box="[140,281,237,263]" pageId="18" pageNumber="95" type="description">
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(
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<figureCitation id="131BFC9A5C1A3D0E66AEFF29A8D4FED1" box="[148,214,237,263]" captionStart="Figure 3" captionStartId="18.[140,211,1590,1612]" captionTargetBox="[146,1100,502,1555]" captionTargetId="figure@18.[140,1108,487,1570]" captionTargetPageId="18" captionText="Figure 3. Ceratocanthinae of Ulu Gombak. A Pterorthochaetes insularis habitus dorsal B Pterorthochaetes sp. habitus dorsal C Pterorthochaetes tsurui sp. n. habitus dorsal D idem habitus lateral E Cyphopisthes sp. habitus dorsal." pageId="18" pageNumber="95">Fig. 3</figureCitation>
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C, D)
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</subSubSection>
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</paragraph>
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<subSubSection id="C33AB3945C1A3D0F66B6FEF0AA1BFA01" lastPageId="19" lastPageNumber="96" pageId="18" pageNumber="95" type="materials_examined">
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<paragraph id="8B9FE01F5C1A3D0E66B6FEF0ABDCFE98" box="[140,990,307,334]" pageId="18" pageNumber="95">
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<emphasis id="B9543C0D5C1A3D0E66B6FEF0A92FFE98" bold="true" box="[140,301,307,334]" pageId="18" pageNumber="95">Type locality:</emphasis>
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West Malaysia: Ulu Gombak (Selangor), 3°19
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<emphasis id="B9543C0D5C1A3D0E6501FEF0AB43FE98" box="[827,833,308,334]" italics="true" pageId="18" pageNumber="95">'</emphasis>
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N 101°45
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<emphasis id="B9543C0D5C1A3D0E6582FEF0ABBDFE98" box="[952,959,308,334]" italics="true" pageId="18" pageNumber="95">'</emphasis>
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E.
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</paragraph>
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<paragraph id="8B9FE01F5C1A3D0F6683FE93AA1BFA01" lastPageId="19" lastPageNumber="96" pageId="18" pageNumber="95">
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<emphasis id="B9543C0D5C1A3D0E6683FE93A9A3FEA7" bold="true" box="[185,417,343,369]" pageId="18" pageNumber="95">Material examined.</emphasis>
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<emphasis id="B9543C0D5C1A3D0E6790FE92AA0BFEA7" box="[426,521,342,369]" italics="true" pageId="18" pageNumber="95">Holotype</emphasis>
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, male: West Malaysia, Selangor, Ulu Gombak, 250 mt., 29.IV.2007, Leg. M. Maruyama /
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<taxonomicName id="4C209B9C5C1A3D0E6462FEBDAB52FE42" box="[600,848,377,404]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
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<emphasis id="B9543C0D5C1A3D0E6462FEBDAB52FE42" box="[600,848,377,404]" italics="true" pageId="18" pageNumber="95">Pterorthochaetes tsurui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26781765C1A3D0E6560FEBEAB98FE42" box="[858,922,378,404]" pageId="18" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
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det. Ballerio & Maruyama 2009 Holotypus (KUM). [distended specimen glued on a card, dissected, genitalia mounted in DMHF resin on a separate card, same pin].
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<emphasis id="B9543C0D5C1B3D0F654DFACAABD9FAF1" box="[887,987,1294,1319]" italics="true" pageId="19" pageNumber="96">Paratypes</emphasis>
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[one male and four females dissected]: 15.IV.2007, M. Maruyama (1 male); 26.IV.2007, M. Maruyama (1 female and 1 male); 28. IV. 2007, M. Maruyama (1 female); 29. IV. 2007, M. Maruyama (1 female); 30. IV. 2007, M. Maruyama (1 male and female); 3. V. 2007, M. Maruyama (1 female); 4. V. 2007, M. Maruyama (1 female). All from termite nest(s). (2 ABCB, 7 KUM)
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</paragraph>
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</subSubSection>
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<caption id="DF5FB0975C1A3D0E66B6F9F2A916F95A" pageId="18" pageNumber="95" targetBox="[146,1100,502,1555]" targetPageId="18">
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<paragraph id="8B9FE01F5C1A3D0E66B6F9F2A916F95A" pageId="18" pageNumber="95">
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<emphasis id="B9543C0D5C1A3D0E66B6F9F2A8ECF99A" bold="true" box="[140,238,1590,1612]" pageId="18" pageNumber="95">Figure 3.</emphasis>
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Ceratocanthinae of Ulu Gombak.
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<emphasis id="B9543C0D5C1A3D0E640FF9F2AA4AF99A" bold="true" box="[565,584,1590,1612]" pageId="18" pageNumber="95">A</emphasis>
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<taxonomicName id="4C209B9C5C1A3D0E6474F9F2AB2CF99A" box="[590,814,1590,1612]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="insularis">
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<emphasis id="B9543C0D5C1A3D0E6474F9F2AB2CF99A" box="[590,814,1590,1612]" italics="true" pageId="18" pageNumber="95">Pterorthochaetes insularis</emphasis>
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</taxonomicName>
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habitus dorsal
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<emphasis id="B9543C0D5C1A3D0E6586F9F2ABCFF99A" bold="true" box="[956,973,1590,1612]" pageId="18" pageNumber="95">B</emphasis>
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<emphasis id="B9543C0D5C1A3D0E65E9F9F2A8BDF9BA" italics="true" pageId="18" pageNumber="95">
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<taxonomicName id="4C209B9C5C1A3D0E65E9F9F2A8A7F9BA" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="genus">Pterorthochaetes</taxonomicName>
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sp.
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</emphasis>
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habitus dorsal
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<emphasis id="B9543C0D5C1A3D0E677CF992A95BF9BA" bold="true" box="[326,345,1622,1644]" pageId="18" pageNumber="95">C</emphasis>
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<taxonomicName id="4C209B9C5C1A3D0E6767F992AA26F9BA" box="[349,548,1622,1644]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
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<emphasis id="B9543C0D5C1A3D0E6767F992AA26F9BA" box="[349,548,1622,1644]" italics="true" pageId="18" pageNumber="95">Pterorthochaetes tsurui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26781765C1A3D0E6413F993AA58F9BA" box="[553,602,1623,1644]" pageId="18" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
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habitus dorsal
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<emphasis id="B9543C0D5C1A3D0E64D9F992AAF4F9BA" bold="true" box="[739,758,1622,1644]" pageId="18" pageNumber="95">D</emphasis>
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idem habitus lateral
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<emphasis id="B9543C0D5C1A3D0E658EF992ABC1F9BA" bold="true" box="[948,963,1622,1644]" pageId="18" pageNumber="95">E</emphasis>
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<emphasis id="B9543C0D5C1A3D0E65FDF992AC57F9BA" box="[967,1109,1622,1644]" italics="true" pageId="18" pageNumber="95">
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<taxonomicName id="4C209B9C5C1A3D0E65FDF992AC3CF9BA" box="[967,1086,1622,1644]" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
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sp.
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</emphasis>
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habitus dorsal.
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</paragraph>
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</caption>
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<caption id="DF5FB0975C1B3D0F66B6FB5AABC5FB02" pageId="19" pageNumber="96" targetBox="[156,1100,180,1155]" targetPageId="19">
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<paragraph id="8B9FE01F5C1B3D0F66B6FB5AABC5FB02" pageId="19" pageNumber="96">
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<emphasis id="B9543C0D5C1B3D0F66B6FB5AA8EFFB62" bold="true" box="[140,237,1182,1204]" pageId="19" pageNumber="96">Figure 4.</emphasis>
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Ceratocanthinae of Ulu Gombak.
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<emphasis id="B9543C0D5C1B3D0F640AFB5AAA41FB62" bold="true" box="[560,579,1182,1204]" pageId="19" pageNumber="96">A</emphasis>
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<emphasis id="B9543C0D5C1B3D0F6473FB5AAAFFFB62" box="[585,765,1182,1204]" italics="true" pageId="19" pageNumber="96">
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<taxonomicName id="4C209B9C5C1B3D0F6473FB5AAAD2FB62" box="[585,720,1182,1204]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
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sp. a
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</emphasis>
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habitus dorsal
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<emphasis id="B9543C0D5C1B3D0F65B3FB5AAB9BFB62" bold="true" box="[905,921,1182,1204]" pageId="19" pageNumber="96">B</emphasis>
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<emphasis id="B9543C0D5C1B3D0F65A5FB5AAC51FB62" box="[927,1107,1182,1204]" italics="true" pageId="19" pageNumber="96">
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<taxonomicName id="4C209B9C5C1B3D0F65A5FB5AAC24FB62" box="[927,1062,1182,1204]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
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sp. b
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</emphasis>
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habitus dorsal
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<emphasis id="B9543C0D5C1B3D0F6729FB7AA927FB02" bold="true" box="[275,293,1214,1236]" pageId="19" pageNumber="96">C</emphasis>
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<emphasis id="B9543C0D5C1B3D0F6711FB7AA9DFFB02" box="[299,477,1214,1236]" italics="true" pageId="19" pageNumber="96">
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<taxonomicName id="4C209B9C5C1B3D0F6711FB7AA9B0FB02" box="[299,434,1214,1236]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
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sp. c
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</emphasis>
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habitus dorsal
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<emphasis id="B9543C0D5C1B3D0F6450FB7AAA7FFB02" bold="true" box="[618,637,1214,1236]" pageId="19" pageNumber="96">D</emphasis>
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<taxonomicName id="4C209B9C5C1B3D0F64B9FB7AAB3AFB02" box="[643,824,1214,1236]" class="Insecta" family="Hybosoridae" genus="Ebbrittoniella" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="species" species="ignita">
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<emphasis id="B9543C0D5C1B3D0F64B9FB7AAB3AFB02" box="[643,824,1214,1236]" italics="true" pageId="19" pageNumber="96">Ebbrittoniella ignita</emphasis>
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</taxonomicName>
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habitus dorsal.
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</paragraph>
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</caption>
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<subSubSection id="C33AB3945C1B3D0B6683FA24AC11FF17" lastPageId="23" lastPageNumber="100" pageId="19" pageNumber="96" type="description">
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<paragraph id="8B9FE01F5C1B3D0F6683FA24A9FFF9C8" pageId="19" pageNumber="96">
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<emphasis id="B9543C0D5C1B3D0F6683FA24A94DFA2C" bold="true" box="[185,335,1504,1530]" pageId="19" pageNumber="96">Description.</emphasis>
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HL = 0.69 mm; HW = 1.44 mm; PL = 1.38 mm; PW = 2.33 mm; EL = 2.36 mm; EW = 2.27 mm.
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</paragraph>
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<paragraph id="8B9FE01F5C1B3D0F6683F9E2A910F9B2" pageId="19" pageNumber="96">
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Medium sized
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<taxonomicName id="4C209B9C5C1B3D0F6758F9E2AA12F997" box="[354,528,1574,1601]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">
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<emphasis id="B9543C0D5C1B3D0F6758F9E2AA12F997" box="[354,528,1574,1601]" italics="true" pageId="19" pageNumber="96">Pterorthochaetes</emphasis>
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</taxonomicName>
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, surface shiny, setose; “rolling up” coaptations perfect; volant.
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</paragraph>
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<paragraph id="8B9FE01F5C1B3D0F6683F9AAABDEF951" box="[185,988,1645,1671]" pageId="19" pageNumber="96">Dorsum brown, setation yellowish, sternum reddish, antennae reddish.</paragraph>
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<caption id="DF5FB0975C1C3D0866B6FB81AB7CFB4D" pageId="20" pageNumber="97" targetBox="[140,1107,165,1070]" targetPageId="20">
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<paragraph id="8B9FE01F5C1C3D0866B6FB81AB7CFB4D" pageId="20" pageNumber="97">
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<emphasis id="B9543C0D5C1C3D0866B6FB81A8EFFB8D" bold="true" box="[140,237,1093,1115]" pageId="20" pageNumber="97">Figure 5.</emphasis>
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SEM photographs of:
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<emphasis id="B9543C0D5C1C3D086786FB81A9CDFB8D" bold="true" box="[444,463,1093,1115]" pageId="20" pageNumber="97">A</emphasis>
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<emphasis id="B9543C0D5C1C3D0867EEFB81AA86FB8D" box="[468,644,1093,1115]" italics="true" pageId="20" pageNumber="97">
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<taxonomicName id="4C209B9C5C1C3D0867EEFB81AA3FFB8D" box="[468,573,1093,1115]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
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clypeale
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</emphasis>
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: male pronotum
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<emphasis id="B9543C0D5C1C3D08651EFB81AB36FB8D" bold="true" box="[804,820,1093,1115]" pageId="20" pageNumber="97">B</emphasis>
|
|
detail of male pronotum puncture
|
|
<emphasis id="B9543C0D5C1C3D086682FBA1A8C9FBAD" bold="true" box="[184,203,1125,1147]" pageId="20" pageNumber="97">C</emphasis>
|
|
female pronotum
|
|
<emphasis id="B9543C0D5C1C3D086746FBA1A992FBAD" bold="true" box="[380,400,1125,1147]" pageId="20" pageNumber="97">D</emphasis>
|
|
detail of female pronotum punctures
|
|
<emphasis id="B9543C0D5C1C3D0864C3FBA1AB0AFBAD" bold="true" box="[761,776,1125,1147]" pageId="20" pageNumber="97">E</emphasis>
|
|
and
|
|
<emphasis id="B9543C0D5C1C3D086503FBA1AB4AFBAD" bold="true" box="[825,840,1125,1147]" pageId="20" pageNumber="97">F</emphasis>
|
|
<taxonomicName id="4C209B9C5C1C3D086575FBA1AC1AFBAD" box="[847,1048,1125,1147]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="mirificum" status="sp.n.">
|
|
<emphasis id="B9543C0D5C1C3D086575FBA1AC1AFBAD" box="[847,1048,1125,1147]" italics="true" pageId="20" pageNumber="97">Madrasostes mirificum</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A26781765C1C3D08621AFBA2AC56FBAD" box="[1056,1108,1126,1147]" pageId="20" pageNumber="97">sp.n.</taxonomicNameLabel>
|
|
mouthparts: labium
|
|
<emphasis id="B9543C0D5C1C3D086771FB41A95CFB4D" bold="true" box="[331,350,1157,1179]" pageId="20" pageNumber="97">G</emphasis>
|
|
epipharynx
|
|
<emphasis id="B9543C0D5C1C3D0867E8FB41A9E4FB4D" bold="true" box="[466,486,1157,1179]" pageId="20" pageNumber="97">H</emphasis>
|
|
mandibles
|
|
<emphasis id="B9543C0D5C1C3D08646BFB41AA5BFB4D" bold="true" box="[593,601,1157,1179]" pageId="20" pageNumber="97">I</emphasis>
|
|
detail of galear brush
|
|
<emphasis id="B9543C0D5C1C3D08651CFB41AB2CFB4D" bold="true" box="[806,814,1157,1179]" pageId="20" pageNumber="97">J</emphasis>
|
|
maxilla.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8B9FE01F5C1C3D086683FB02AAA1FAB8" pageId="20" pageNumber="97">
|
|
<emphasis id="B9543C0D5C1C3D086683FB02A8F9FB37" box="[185,251,1222,1249]" italics="true" pageId="20" pageNumber="97">Head:</emphasis>
|
|
subrectangular, wider than long, fore margin irregularly sinuated, serrated, tip acute, interocular distance about 9 times the maximum width of dorsal ocular area, dorsal ocular area large, sculpure distally made of very coarse and deep wrinkles and proximally of impressed small dense horseshoe-shaped punctures, each one having a pore in the middle bearing an erect simple seta.
|
|
</paragraph>
|
|
<paragraph id="8B9FE01F5C1C3D086683FAB3AAFFF951" pageId="20" pageNumber="97">
|
|
<emphasis id="B9543C0D5C1C3D086683FAB3A92FFA47" box="[185,301,1399,1425]" italics="true" pageId="20" pageNumber="97">Pronotum:</emphasis>
|
|
wider than long (W/L ratio= 1.68), fore angles normally shaped, elytral lateral margins fringed with a row of long simple setae, spaced out by an interval about half of their length, the whole pronotal surface covered by medium sized round, shallow, ocellate punctures, each one having a pore in the middle bearing a long simple erect seta; puncturation relatively dense, the distance between punctures being inferior to their diameter, denser on disc and at sides, only at sides the ocellate punctures are replaced by horseshoe-shaped punctures with openings outwards. Pronotal pubescence relatively long, approximately as long as marginal setae.
|
|
</paragraph>
|
|
<caption id="DF5FB0975C1D3D0966B6FB72A9BBFADA" pageId="21" pageNumber="98" targetBox="[140,1107,165,1186]" targetPageId="21">
|
|
<paragraph id="8B9FE01F5C1D3D0966B6FB72A9BBFADA" pageId="21" pageNumber="98">
|
|
<emphasis id="B9543C0D5C1D3D0966B6FB72A8ECFB1A" bold="true" box="[140,238,1206,1228]" pageId="21" pageNumber="98">Figure 6.</emphasis>
|
|
Photos of alive specimens taken in habitat at Ulu Gombak (photos © M. Maruyama, 2008).
|
|
<emphasis id="B9543C0D5C1D3D0966B6FB12A89DFB3A" bold="true" box="[140,159,1238,1260]" pageId="21" pageNumber="98">A</emphasis>
|
|
<emphasis id="B9543C0D5C1D3D09669FFB12A974FB3A" box="[165,374,1238,1260]" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName id="4C209B9C5C1D3D09669FFB12A90CFB3A" box="[165,270,1238,1260]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
|
|
variolosum
|
|
</emphasis>
|
|
<emphasis id="B9543C0D5C1D3D096747FB12A98FFB3A" bold="true" box="[381,397,1238,1260]" pageId="21" pageNumber="98">B</emphasis>
|
|
<taxonomicName id="4C209B9C5C1D3D0967AEFB12AA3EFB3A" box="[404,572,1238,1260]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="agostii">
|
|
<emphasis id="B9543C0D5C1D3D0967AEFB12AA3EFB3A" box="[404,572,1238,1260]" italics="true" pageId="21" pageNumber="98">Madrasostes agostii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B9543C0D5C1D3D096479FB12AA57FB3A" bold="true" box="[579,597,1238,1260]" pageId="21" pageNumber="98">C</emphasis>
|
|
<emphasis id="B9543C0D5C1D3D096466FB12AB0DFB3A" box="[604,783,1238,1260]" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName id="4C209B9C5C1D3D096466FB12AAC7FB3A" box="[604,709,1238,1260]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
|
|
clypeale
|
|
</emphasis>
|
|
male
|
|
<emphasis id="B9543C0D5C1D3D096572FB12AB5EFB3A" bold="true" box="[840,860,1238,1260]" pageId="21" pageNumber="98">D</emphasis>
|
|
idem female
|
|
<emphasis id="B9543C0D5C1D3D0965E1FB12ABE9FB3A" bold="true" box="[987,1003,1238,1260]" pageId="21" pageNumber="98">E</emphasis>
|
|
<emphasis id="B9543C0D5C1D3D0965C8FB12A8E5FADA" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName id="4C209B9C5C1D3D0965C8FB12A8BDFADA" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. b
|
|
</emphasis>
|
|
<emphasis id="B9543C0D5C1D3D0966D7FB32A8F9FADA" bold="true" box="[237,251,1270,1292]" pageId="21" pageNumber="98">F</emphasis>
|
|
<emphasis id="B9543C0D5C1D3D09673BFB32A9B1FADA" box="[257,435,1270,1292]" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName id="4C209B9C5C1D3D09673BFB32A98AFADA" box="[257,392,1270,1292]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. c
|
|
</emphasis>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8B9FE01F5C1D3D096683FAF4AB3CFA9D" box="[185,830,1328,1355]" pageId="21" pageNumber="98">
|
|
<emphasis id="B9543C0D5C1D3D096683FAF4A92EFA9D" box="[185,300,1328,1355]" italics="true" pageId="21" pageNumber="98">Scutellum:</emphasis>
|
|
punctures horseshoe-shaped, thick and coarse.
|
|
</paragraph>
|
|
<paragraph id="8B9FE01F5C1D3D096683FA97AAC3FA01" pageId="21" pageNumber="98">Elytra: shape oval, longer than wide (W/L ratio= 0.96); elytral surface covered by medium sized shallow horseshoe-shaped punctures with opening directed backwards, spaced out by an interval inferior to their diameter. Each horseshoe-shaped puncture containing a pore bearing a long erect simple seta.</paragraph>
|
|
<paragraph id="8B9FE01F5C1D3D096683FA24A9E1F951" pageId="21" pageNumber="98">
|
|
<emphasis id="B9543C0D5C1D3D096683FA24A967FA2D" box="[185,357,1504,1531]" italics="true" pageId="21" pageNumber="98">Male genitalia:</emphasis>
|
|
parameres about as long as basal piece, slightly asymmetrical (
|
|
<figureCitation id="131BFC9A5C1D3D0966AFF9C7A8D1F9C8" box="[149,211,1539,1566]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="21" pageNumber="98">Fig.7</figureCitation>
|
|
A, B), apex distinctly bent, internal sac armed with longitudinal subrectangular sclerotization about as long as the basal piece with three short distal sclerites (
|
|
<figureCitation id="131BFC9A5C1D3D0965D1F9E2AC2EF997" box="[1003,1068,1574,1601]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="21" pageNumber="98">Fig. 7</figureCitation>
|
|
D), genital segment with manubrium long but strongly bent/twisted, so that in dorsal view it appears very short (
|
|
<figureCitation id="131BFC9A5C1D3D096745F9A9A9C3F951" box="[383,449,1645,1671]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="21" pageNumber="98">Fig. 7</figureCitation>
|
|
E).
|
|
</paragraph>
|
|
<caption id="DF5FB0975C1E3D0A66B6F9F2AB4FF95A" pageId="22" pageNumber="99" targetBox="[231,1090,199,1518]" targetPageId="22">
|
|
<paragraph id="8B9FE01F5C1E3D0A66B6F9F2AB4FF95A" pageId="22" pageNumber="99">
|
|
<emphasis id="B9543C0D5C1E3D0A66B6F9F2A8EDF99A" bold="true" box="[140,239,1590,1612]" pageId="22" pageNumber="99">Figure 7.</emphasis>
|
|
<taxonomicName id="4C209B9C5C1E3D0A66CFF9F2A9BCF99A" box="[245,446,1590,1612]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="99" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
|
|
<emphasis id="B9543C0D5C1E3D0A66CFF9F2A9BCF99A" box="[245,446,1590,1612]" italics="true" pageId="22" pageNumber="99">Pterorthochaetes tsurui</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A26781765C1E3D0A67FFF9F3A9F8F99A" box="[453,506,1591,1612]" pageId="22" pageNumber="99" rank="species">sp. n.</taxonomicNameLabel>
|
|
:
|
|
<emphasis id="B9543C0D5C1E3D0A643EF9F2AA15F99A" bold="true" box="[516,535,1590,1612]" pageId="22" pageNumber="99">A</emphasis>
|
|
parameres in lateral view (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1E3D0A65FCF9F2ABD4F99A" bold="true" box="[966,982,1590,1612]" pageId="22" pageNumber="99">B</emphasis>
|
|
parameres in dorsal view (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1E3D0A6797F992A9C2F9BA" bold="true" box="[429,448,1622,1644]" pageId="22" pageNumber="99">C</emphasis>
|
|
female bursal sclerites (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1E3D0A657CF992AB5BF9BA" bold="true" box="[838,857,1622,1644]" pageId="22" pageNumber="99">D</emphasis>
|
|
, sclerites of the internal sac of aedeagus (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1E3D0A678FF9B2A9C6F95A" bold="true" box="[437,452,1654,1676]" pageId="22" pageNumber="99">E</emphasis>
|
|
male genital segment (scale bar: 0,1 mm).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8B9FE01F5C1F3D0B6683FF62AC11FF17" box="[185,1043,166,193]" pageId="23" pageNumber="100">
|
|
<emphasis id="B9543C0D5C1F3D0B6683FF62A970FF17" box="[185,370,166,193]" italics="true" pageId="23" pageNumber="100">Female genitalia:</emphasis>
|
|
bursal sclerites weakly sclerotized and shaped as in
|
|
<figureCitation id="131BFC9A5C1F3D0B658DFF62ABFBFF17" box="[951,1017,166,193]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="23" pageNumber="100">Fig. 7</figureCitation>
|
|
C.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C33AB3945C1F3D046683FF0EAB2AFE98" lastPageId="24" lastPageNumber="101" pageId="23" pageNumber="100" type="diagnosis">
|
|
<paragraph id="8B9FE01F5C1F3D046683FF0EAB2AFE98" lastPageId="24" lastPageNumber="101" pageId="23" pageNumber="100">
|
|
<emphasis id="B9543C0D5C1F3D0B6683FF0EA966FF32" bold="true" box="[185,356,202,228]" pageId="23" pageNumber="100">Identification.</emphasis>
|
|
This species is characterized by the following combination of characters: a) pronotal puncturation made of ocellate, shallow, medium sized punctures, b) lateral margins of pronotum fringed by long, simple setae, setae relatively spaced out, c) elytral puncturation made of medium sized horseshoe-shaped punctures with opening backwards and bearing a long erect seta, d) shape of parameres (about as long as basal piece and with bent apex, e) shape of genital segment of males, with manubrium strongly twisted, f) shape of female bursal sclerites and, g) head subrectangular, with fore margin somewhat sinuated in a way very unusual for
|
|
<taxonomicName id="4C209B9C5C103D04652CFF29ABC1FED1" box="[790,963,237,263]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9543C0D5C103D04652CFF29ABC1FED1" box="[790,963,237,263]" italics="true" pageId="24" pageNumber="101">Pterorthochaetes</emphasis>
|
|
</taxonomicName>
|
|
. Perhaps the species closest to it could be
|
|
<taxonomicName id="4C209B9C5C103D0467F7FED4AAF9FEFD" box="[461,763,272,299]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="brevisetosus">
|
|
<emphasis id="B9543C0D5C103D0467F7FED4AAF9FEFD" box="[461,763,272,299]" italics="true" pageId="24" pageNumber="101">Pterorthochaetes brevisetosus</emphasis>
|
|
</taxonomicName>
|
|
, which, however, has pronotal and elytral puncturation much less dense and shorter setae.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="DF5FB0975C1F3D0B66B6F9D2ABF3F95A" pageId="23" pageNumber="100" targetBox="[145,1078,415,1513]" targetPageId="23">
|
|
<paragraph id="8B9FE01F5C1F3D0B66B6F9D2ABF3F95A" pageId="23" pageNumber="100">
|
|
<emphasis id="B9543C0D5C1F3D0B66B6F9D2A8F2F9FA" bold="true" box="[140,240,1558,1580]" pageId="23" pageNumber="100">Figure 8.</emphasis>
|
|
<taxonomicName id="4C209B9C5C1F3D0B66CDF9D2A9C2F9FA" box="[247,448,1558,1580]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="mirificum" status="sp. n.">
|
|
<emphasis id="B9543C0D5C1F3D0B66CDF9D2A9C2F9FA" box="[247,448,1558,1580]" italics="true" pageId="23" pageNumber="100">Madrasostes mirificum</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A26781765C1F3D0B67F2F9D3A9FCF9FA" box="[456,510,1559,1580]" pageId="23" pageNumber="100" rank="species">sp. n.</taxonomicNameLabel>
|
|
:
|
|
<emphasis id="B9543C0D5C1F3D0B6433F9D2AA1EF9FA" bold="true" box="[521,540,1558,1580]" pageId="23" pageNumber="100">A</emphasis>
|
|
male genital segment (scale bar: 0,2 mm)
|
|
<emphasis id="B9543C0D5C1F3D0B658AF9D2ABC3F9FA" bold="true" box="[944,961,1558,1580]" pageId="23" pageNumber="100">B</emphasis>
|
|
aedeagus (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1F3D0B6734F9F2A923F99A" bold="true" box="[270,289,1590,1612]" pageId="23" pageNumber="100">C</emphasis>
|
|
parameres in dorsal view (scale bar: 0,0 5 mm)
|
|
<emphasis id="B9543C0D5C1F3D0B64E3F9F2AAEFF99A" bold="true" box="[729,749,1590,1612]" pageId="23" pageNumber="100">D</emphasis>
|
|
<taxonomicName id="4C209B9C5C1F3D0B64C9F9F2AB9FF99A" box="[755,925,1590,1612]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="hasimi" status="sp. n.">
|
|
<emphasis id="B9543C0D5C1F3D0B64C9F9F2AB9FF99A" box="[755,925,1590,1612]" italics="true" pageId="23" pageNumber="100">Madrasostes hasimi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A26781765C1F3D0B659EF9F3ABDBF99A" box="[932,985,1591,1612]" pageId="23" pageNumber="100" rank="species">sp. n.</taxonomicNameLabel>
|
|
: male genital segment (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1F3D0B67A3F992A9ABF9BA" bold="true" box="[409,425,1622,1644]" pageId="23" pageNumber="100">E</emphasis>
|
|
parameres (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1F3D0B64F7F992AAD9F9BA" bold="true" box="[717,731,1622,1644]" pageId="23" pageNumber="100">F</emphasis>
|
|
sclerite of internal sac of aedeagus (scale bar: 0,1 mm)
|
|
<emphasis id="B9543C0D5C1F3D0B6737F9B2A922F95A" bold="true" box="[269,288,1654,1676]" pageId="23" pageNumber="100">G</emphasis>
|
|
<taxonomicName id="4C209B9C5C1F3D0B671CF9B2A9ECF95A" box="[294,494,1654,1676]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="depressum">
|
|
<emphasis id="B9543C0D5C1F3D0B671CF9B2A9ECF95A" box="[294,494,1654,1676]" italics="true" pageId="23" pageNumber="100">Madrasostes depressum</emphasis>
|
|
</taxonomicName>
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|
: sclerite of internal sac of aedeagus (scale bar: 0,1 mm).
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|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="C33AB3945C103D046683FE93A98AFE42" pageId="24" pageNumber="101" type="discussion">
|
|
<paragraph id="8B9FE01F5C103D046683FE93A98AFE42" pageId="24" pageNumber="101">
|
|
<emphasis id="B9543C0D5C103D046683FE93A939FEA7" bold="true" box="[185,315,343,369]" pageId="24" pageNumber="101">Variability.</emphasis>
|
|
The type series shows a strong uniformity as for size, shape, setation and microsculpturing.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C33AB3945C103D046683FE59A928FE28" pageId="24" pageNumber="101" type="etymology">
|
|
<paragraph id="8B9FE01F5C103D046683FE59A928FE28" pageId="24" pageNumber="101">
|
|
<emphasis id="B9543C0D5C103D046683FE59A942FE61" bold="true" box="[185,320,413,439]" pageId="24" pageNumber="101">Etymology.</emphasis>
|
|
Dedicated to Mr. Tomoyuki Tsuru, who found the huge termite nest from which the junior author collected more than 400 Ceratocanthinae, among which
|
|
<taxonomicName id="4C209B9C5C103D0466B6FE20A8E7FE28" box="[140,229,484,510]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" genus.bestMatchDistance="1276" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
|
|
<emphasis id="B9543C0D5C103D0466B6FE20A8E7FE28" box="[140,229,484,510]" italics="true" pageId="24" pageNumber="101">P. tsurui</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A26781765C103D0466D6FE20A928FE28" box="[236,298,484,510]" pageId="24" pageNumber="101" rank="species">sp. n.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C33AB3945C103D046683FDC3ABB6FD92" pageId="24" pageNumber="101" type="distribution">
|
|
<paragraph id="8B9FE01F5C103D046683FDC3ABB6FD92" pageId="24" pageNumber="101">
|
|
<emphasis id="B9543C0D5C103D046683FDC3A9E9FDF7" bold="true" box="[185,491,519,545]" pageId="24" pageNumber="101">Distribution and habitat.</emphasis>
|
|
This new species is know only from the type locality. For details on collecting circumstances see the introductory paragraphs.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |